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<document id="7080C3CAFFC8BA8CAAEC8C0B8823F868" ID-CLB-Dataset="46294" ID-DOI="10.11646/zootaxa.3774.2.2" ID-GBIF-Dataset="8c347efa-46ad-406f-ab1b-493686aab077" ID-ISSN="1175-5326" ID-Zenodo-Dep="225828" ID-ZooBank="BAAD0752-E3B9-4C7A-AC0B-AA7E7E35F402" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460392384907" checkinUser="plazi" docAuthor="Mironov, Sergey V., Ehrnsberger, Rainer &amp; Dabert, Jacek" docDate="2014" docId="8912CE787A55092BCE83F8DEFBE78A00" docLanguage="en" docName="zt03774p151.pdf" docOrigin="Zootaxa 3774 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Protolichus placentis Mironov, Ehrnsberger &amp; Dabert, 2014, sp. n." docType="treatment" docVersion="8" lastPageNumber="139" masterDocId="752BB6007A530923CE14FFB6FF8B8D28" masterDocTitle="New species of the feather mite genus Protolichus Trouessart, 1884 (Astigmata, Pterolichidae) from lories and lorikeets (Aves: Psittaciformes)" masterLastPageNumber="151" masterPageNumber="131" pageNumber="137" updateTime="1698298364341" updateUser="ExternalLinkService">
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<mods:title id="404267A91447BF3A478E0757A75E203A">New species of the feather mite genus Protolichus Trouessart, 1884 (Astigmata, Pterolichidae) from lories and lorikeets (Aves: Psittaciformes)</mods:title>
</mods:titleInfo>
<mods:name id="1580DE1802CDC8F29D470CA9BC848983" type="personal">
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<mods:namePart id="606218C3329B2AF732D1D3C45595B2CB">Mironov, Sergey V.</mods:namePart>
</mods:name>
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<mods:role id="5263FCBD0EF99D29616FDA754EDE24B6">
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<mods:namePart id="E75C8C9416BED3A0EB07E735ECF2A133">Ehrnsberger, Rainer</mods:namePart>
</mods:name>
<mods:name id="418F5EC1543BEFAF2A5D030DCE20F1C2" type="personal">
<mods:role id="1413AD988DB62C83B29ED4B6CD99EE52">
<mods:roleTerm id="18DFAEAE41D4901A05A041A39D96E356">Author</mods:roleTerm>
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<mods:namePart id="9D4890565DEB64262DBC951198C1291B">Dabert, Jacek</mods:namePart>
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<mods:typeOfResource id="2985C335C5090C70E7D5312A9576024F">text</mods:typeOfResource>
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<mods:title id="34AB4B3C847BCD37D5128F81BC7B6CE2">Zootaxa</mods:title>
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<mods:part id="B80DDC7CF6C54A8EACFCBF208A4707B7">
<mods:date id="4C389A9F1B72422308489A6914B9E3E3">2014</mods:date>
<mods:detail id="47589938828202249396BFB344581FAB" type="volume">
<mods:number id="760E2B0ACC0C0119E301CB1FD28ECBA4">3774</mods:number>
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<mods:identifier id="DFBD53A10EFFB0CEC80241C025FF3539" type="DOI">10.11646/zootaxa.3774.2.2</mods:identifier>
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<treatment id="8912CE787A55092BCE83F8DEFBE78A00" ID-DOI="http://doi.org/10.5281/zenodo.6129364" ID-GBIF-Taxon="119417126" ID-Zenodo-Dep="6129364" LSID="urn:lsid:plazi:treatment:8912CE787A55092BCE83F8DEFBE78A00" httpUri="http://treatment.plazi.org/id/8912CE787A55092BCE83F8DEFBE78A00" lastPageId="8" lastPageNumber="139" pageId="6" pageNumber="137">
<subSubSection id="49A12CE57A550925CE83F8DEFEE88A8D" pageId="6" pageNumber="137" type="nomenclature">
<paragraph id="01047F6E7A550925CE83F8DEFE548AAB" blockId="6.[151,479,1896,1957]" box="[151,479,1896,1923]" pageId="6" pageNumber="137">
<heading id="5A4CC8027A550925CE83F8DEFE548AAB" bold="true" box="[151,479,1896,1923]" fontSize="11" level="1" pageId="6" pageNumber="137" reason="1">
<emphasis id="33CFA37C7A550925CE83F8DEFE548AAB" bold="true" box="[151,479,1896,1923]" pageId="6" pageNumber="137">
<taxonomicName id="C6BB04ED7A550925CE83F8DEFE1E8AAA" box="[151,405,1896,1922]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="placentis" status="sp. nov.">
<emphasis id="33CFA37C7A550925CE83F8DEFE1E8AAA" bold="true" box="[151,405,1896,1922]" italics="true" pageId="6" pageNumber="137">Protolichus placentis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28FC1E077A550925CF89F8DFFE548AAB" box="[413,479,1897,1923]" pageId="6" pageNumber="137" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="01047F6E7A550925CE83F83AFEE88A8D" blockId="6.[151,479,1896,1957]" box="[151,355,1932,1957]" pageId="6" pageNumber="137">
(
<figureCitation id="998063EB7A550925CE8BF83AFF668A8C" box="[159,237,1932,1957]" captionStart="FIGURE 4" captionStartId="6.[151,250,1764,1786]" captionTargetBox="[158,1429,200,1737]" captionTargetId="figure@6.[151,1436,193,1742]" captionTargetPageId="6" captionText="FIGURE 4. Dorsal view of male opisthosoma of Protolichus ornatus sp. n. (A, B) and Protolichus placentis sp. n. (C, D). A, C—heteromorph males, B, D—homeomorph males." httpUri="https://zenodo.org/record/225832/files/figure.png" pageId="6" pageNumber="137">Figs. 4</figureCitation>
C, D, 5, 6)
</paragraph>
</subSubSection>
<subSubSection id="49A12CE57A550924CE83F862FCAE8DD0" lastPageId="7" lastPageNumber="138" pageId="6" pageNumber="137" type="materials_examined">
<paragraph id="01047F6E7A550924CE83F862FEDC8DFD" blockId="6.[151,1436,2004,2029]" lastBlockId="7.[151,1436,151,248]" lastPageId="7" lastPageNumber="138" pageId="6" pageNumber="137">
<emphasis id="33CFA37C7A550925CE83F862FEC18AC5" bold="true" box="[151,330,2004,2029]" pageId="6" pageNumber="137">
<typeStatus id="DE00C1CC7A550925CE83F862FF598AC5" box="[151,210,2004,2029]" pageId="6" pageNumber="137">Type</typeStatus>
material.
</emphasis>
Heteromorph male
<typeStatus id="DE00C1CC7A550925CC2EF862FD148AC5" box="[570,671,2004,2029]" pageId="6" pageNumber="137" type="holotype">holotype</typeStatus>
, 2 heteromorph male and 1 homeomorph male
<typeStatus id="DE00C1CC7A550925CACEF863FACE8AC5" box="[1242,1349,2005,2029]" pageId="6" pageNumber="137" type="paratype">paratypes</typeStatus>
(AMU 01752) from
<taxonomicName id="C6BB04ED7A540924CF38FF21FC898D98" authority="Temminck, 1835" authorityName="Temminck" authorityYear="1835" box="[300,770,151,176]" class="Aves" family="Psittacidae" genus="Charmosyna" kingdom="Animalia" order="Psittaciformes" pageId="7" pageNumber="138" phylum="Chordata" rank="species" species="placentis">
<emphasis id="33CFA37C7A540924CF38FF21FDA38D98" box="[300,552,151,176]" italics="true" pageId="7" pageNumber="138">Charmosyna placentis</emphasis>
(Temminck, 1835)
</taxonomicName>
,
<collectingCountry id="79AC3FFE7A540924CD1BFF21FCF48D98" box="[783,895,151,176]" name="Indonesia" pageId="7" pageNumber="138">Indonesia</collectingCountry>
, Obi Archipelago,
<date id="750559AE7A540924CA41FF2EFAB88D98" box="[1109,1331,151,176]" pageId="7" pageNumber="138" value="1853-09-19">19 September 1853</date>
, no other data (
<collectionCode id="67AAE7AB7A540924CEC0FF0AFE9E8DFC" box="[212,277,188,212]" httpUri="http://grbio.org/cool/9tx7-8fc2" name="Uebersee-Museum" pageId="7" pageNumber="138">UMB</collectionCode>
257).
</paragraph>
<paragraph id="01047F6E7A540924CED3FF69FCAE8DD0" blockId="7.[151,1436,151,248]" box="[199,805,223,248]" pageId="7" pageNumber="138">
<emphasis id="33CFA37C7A540924CED3FF69FE078DD0" bold="true" box="[199,396,223,248]" pageId="7" pageNumber="138">
<typeStatus id="DE00C1CC7A540924CED3FF69FE898DD0" box="[199,258,223,248]" pageId="7" pageNumber="138">Type</typeStatus>
depository.
</emphasis>
<typeStatus id="DE00C1CC7A540924CF87FF69FE708DD0" box="[403,507,223,248]" pageId="7" pageNumber="138" type="holotype">Holotype</typeStatus>
and all paratypes—AMU.
</paragraph>
</subSubSection>
<caption id="55C42FE67A540924CE83F9C8FBEA8BBC" box="[151,1121,1662,1684]" httpUri="https://zenodo.org/record/225833/files/figure.png" pageId="7" pageNumber="138" targetBox="[159,1432,307,1637]" targetPageId="7">
<paragraph id="01047F6E7A540924CE83F9C8FBEA8BBC" blockId="7.[151,1121,1662,1684]" box="[151,1121,1662,1684]" pageId="7" pageNumber="138">
<emphasis id="33CFA37C7A540924CE83F9C8FE878BBC" bold="true" box="[151,268,1662,1684]" pageId="7" pageNumber="138">FIGURE 5</emphasis>
.
<taxonomicName id="C6BB04ED7A540924CF0DF9C8FE648BBC" box="[281,495,1662,1684]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="placentis" status="sp. nov.">
<emphasis id="33CFA37C7A540924CF0DF9C8FE648BBC" box="[281,495,1662,1684]" italics="true" pageId="7" pageNumber="138">Protolichus placentis</emphasis>
</taxonomicName>
<emphasis id="33CFA37C7A540924CFE2F9C9FDA48BBC" bold="true" box="[502,559,1663,1684]" pageId="7" pageNumber="138">
<taxonomicNameLabel id="28FC1E077A540924CFE2F9C9FDA48BBC" box="[502,559,1663,1684]" pageId="7" pageNumber="138" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, heteromorph male. A—dorsal view, B— ventral view.
</paragraph>
</caption>
<subSubSection id="49A12CE57A54092BCED3F90AFD598A2D" lastPageId="8" lastPageNumber="139" pageId="7" pageNumber="138" type="description">
<paragraph id="01047F6E7A54092BCED3F90AFDB38CDC" blockId="7.[151,1436,1724,2037]" lastBlockId="8.[151,1437,151,1832]" lastPageId="8" lastPageNumber="139" pageId="7" pageNumber="138">
<emphasis id="33CFA37C7A540924CED3F90AFED28BFD" bold="true" box="[199,345,1724,1749]" pageId="7" pageNumber="138">Description.</emphasis>
Heteromorph male (
<typeStatus id="DE00C1CC7A540924CC5CF90AFD258BFD" box="[584,686,1724,1749]" pageId="7" pageNumber="138" type="holotype">holotype</typeStatus>
, range for 2
<typeStatus id="DE00C1CC7A540924CD50F90BFC3B8BFD" box="[836,944,1725,1749]" pageId="7" pageNumber="138" type="paratype">paratypes</typeStatus>
in parentheses). Idiosoma, length × width, 568 (570593) × 283 (282298). Subcapitulum with posterior margin convex, postero-lateral parts with oblique curved folds, area around bases of setae
<emphasis id="33CFA37C7A540924CC4CF8B2FD078A35" box="[600,652,1796,1821]" italics="true" pageId="7" pageNumber="138">subc</emphasis>
not outlined (
<figureCitation id="998063EB7A540924CD33F8B2FCE08A35" box="[807,875,1796,1821]" captionStart="FIGURE 6" captionStartId="9.[151,250,1905,1927]" captionTargetBox="[177,1421,204,1867]" captionTargetId="figure@9.[153,1438,193,1868]" captionTargetPageId="9" captionText="FIGURE 6. Protolichus placentis sp. n., details of male. A—subcapitulum of heteromorph male, ventral view, B—leg I of heteromorph male, dorsal view, C—leg I of heteromorph male, ventral view, D—leg II of heteromorph male, dorsal view, E leg II of heteromorph male, ventral view, F—tarsus IV of heteromorph male, dorsal view, G—subcapitulum of homeomorph male, ventral view, H—leg I of homeomorph male, dorsal view, I—leg II of homeomorph male, dorsal view." httpUri="https://zenodo.org/record/225834/files/figure.png" pageId="7" pageNumber="138">Fig. 6</figureCitation>
A). Prodorsal shield with posterior margin slightly convex, length 164 (150165), surface monotonously punctate except transverse band at level of scapular setae. Setae
<emphasis id="33CFA37C7A540924CEF5F8FBFF7F8A4C" box="[225,244,1869,1892]" italics="true" pageId="7" pageNumber="138">si</emphasis>
spiculiform, 37 (3540) long. Distance between scapular setae:
<emphasis id="33CFA37C7A540924CDE8F8FBFBBF8A4C" box="[1020,1076,1869,1892]" italics="true" pageId="7" pageNumber="138">se:se</emphasis>
67 (6267),
<emphasis id="33CFA37C7A540924CAC3F8FBFA8D8A4C" box="[1239,1286,1869,1892]" italics="true" pageId="7" pageNumber="138">si:si</emphasis>
32 (3034). Hysterosoma 410 (420432) long. Length of hysteronotal shield: 405 (410415), greatest width 250 (220250), anterior margin straight, surface monotonously punctate. Opisthosomal lobes slightly divergent, acute apically, postero-lateral margin oblique, dorsal surface of lobes with small transverse crests posterior to base
<emphasis id="33CFA37C7A540924CB13F801FAAF8AF8" box="[1287,1316,1975,2000]" italics="true" pageId="7" pageNumber="138">h3</emphasis>
(
<figureCitation id="998063EB7A540924CB20F801FA088AF8" box="[1332,1411,1975,2000]" captionStart="FIGURE 4" captionStartId="6.[151,250,1764,1786]" captionTargetBox="[158,1429,200,1737]" captionTargetId="figure@6.[151,1436,193,1742]" captionTargetPageId="6" captionText="FIGURE 4. Dorsal view of male opisthosoma of Protolichus ornatus sp. n. (A, B) and Protolichus placentis sp. n. (C, D). A, C—heteromorph males, B, D—homeomorph males." httpUri="https://zenodo.org/record/225832/files/figure.png" pageId="7" pageNumber="138">Figs. 4</figureCitation>
C, 5A). Terminal cleft as a inverted rounded V, 70 (6872) long, 30 (3034) in width at level of setae
<emphasis id="33CFA37C7A540924CB05F86AFAA08ADC" box="[1297,1323,2012,2036]" italics="true" pageId="7" pageNumber="138">e1</emphasis>
. Setae
<emphasis id="33CFA37C7A540924CB95F86AFA178ADC" box="[1409,1436,2012,2036]" italics="true" pageId="7" pageNumber="138">c2</emphasis>
spiculiform, 53 (lost in
<typeStatus id="DE00C1CC7A5B092BCFAEFF2EFDAC8D98" box="[442,551,152,176]" pageId="8" pageNumber="139" type="paratype">paratypes</typeStatus>
) long; setae
<emphasis id="33CFA37C7A5B092BCCDDFF21FD6E8D98" box="[713,741,151,176]" italics="true" pageId="8" pageNumber="139">d2</emphasis>
filiform, 28 (2632) long; setae
<emphasis id="33CFA37C7A5B092BCA68FF2EFB1C8D98" box="[1148,1175,152,176]" italics="true" pageId="8" pageNumber="139">e2</emphasis>
thick, represented by macrosetae without filiform apices, 145 (145150) long, extending slightly beyond lobar apices. Setae
<emphasis id="33CFA37C7A5B092BCB08FF0AFABC8DFC" box="[1308,1335,188,212]" italics="true" pageId="8" pageNumber="139">e1</emphasis>
filiform, situated on median margins of terminal cleft, approximately at level of macrosetae
<emphasis id="33CFA37C7A5B092BCA5FFF69FBEC8DD0" box="[1099,1127,223,248]" italics="true" pageId="8" pageNumber="139">h2</emphasis>
. Setae
<emphasis id="33CFA37C7A5B092BCAAFFF56FB598DD0" box="[1211,1234,224,248]" italics="true" pageId="8" pageNumber="139">f2</emphasis>
spatuliform, with oblique bidentate apex (
<figureCitation id="998063EB7A5B092BCFB6FEB2FE6D8C34" box="[418,486,260,285]" captionStart="FIGURE 4" captionStartId="6.[151,250,1764,1786]" captionTargetBox="[158,1429,200,1737]" captionTargetId="figure@6.[151,1436,193,1742]" captionTargetPageId="6" captionText="FIGURE 4. Dorsal view of male opisthosoma of Protolichus ornatus sp. n. (A, B) and Protolichus placentis sp. n. (C, D). A, C—heteromorph males, B, D—homeomorph males." httpUri="https://zenodo.org/record/225832/files/figure.png" pageId="8" pageNumber="139">Fig. 4</figureCitation>
C). Distance between dorsal setae:
<emphasis id="33CFA37C7A5B092BCD7FFEB2FC218C35" box="[875,938,260,285]" italics="true" pageId="8" pageNumber="139">c2:d2</emphasis>
144 (135144),
<emphasis id="33CFA37C7A5B092BCA77FEB2FB288C35" box="[1123,1187,260,285]" italics="true" pageId="8" pageNumber="139">d2:e2</emphasis>
150 (155162),
<emphasis id="33CFA37C7A5B092BCB48FEB2FA178C35" box="[1372,1436,260,285]" italics="true" pageId="8" pageNumber="139">e2:h3</emphasis>
80 (7880),
<emphasis id="33CFA37C7A5B092BCF32FE9EFEEF8C68" box="[294,356,296,320]" italics="true" pageId="8" pageNumber="139">e2:e2</emphasis>
120 (118123),
<emphasis id="33CFA37C7A5B092BCC33FE91FDE38C68" box="[551,616,295,320]" italics="true" pageId="8" pageNumber="139">h2:h2</emphasis>
80 (8086),
<emphasis id="33CFA37C7A5B092BCD15FE91FCC98C68" box="[769,834,295,320]" italics="true" pageId="8" pageNumber="139">h3:h3</emphasis>
73 (8085),
<emphasis id="33CFA37C7A5B092BCDC8FE9EFBBF8C68" box="[988,1076,296,320]" italics="true" pageId="8" pageNumber="139">ps1:ps1</emphasis>
50 (5560),
<emphasis id="33CFA37C7A5B092BCAD8FE91FA808C68" box="[1228,1291,295,320]" italics="true" pageId="8" pageNumber="139">e2:h2</emphasis>
57 (5458),
<emphasis id="33CFA37C7A5B092BCE83FEFAFF538C4D" box="[151,216,332,357]" italics="true" pageId="8" pageNumber="139">h2:h3</emphasis>
26 (2226),
<emphasis id="33CFA37C7A5B092BCF6EFEFAFE4C8C4D" box="[378,455,332,357]" italics="true" pageId="8" pageNumber="139">ps1:h3</emphasis>
15 (1416). Genital apparatus 28 (28310) × 19 (1519), its base at midlevel of trochanters IV. Paragenital apodemes with anterior ends free from inner tips of epimerites IIa, but connected with epimerites IIIa, middle parts of these apodemes connected by wide but poorly sclerotized transverse bridge (
<figureCitation id="998063EB7A5B092BCB7AFE22FF2D8CF8" captionStart="FIGURE 5" captionStartId="7.[151,250,1662,1684]" captionTargetBox="[159,1432,307,1637]" captionTargetId="figure@7.[151,1436,301,1641]" captionTargetPageId="7" captionText="FIGURE 5. Protolichus placentis sp. n., heteromorph male. A—dorsal view, B— ventral view." httpUri="https://zenodo.org/record/225833/files/figure.png" pageId="8" pageNumber="139">Fig. 5</figureCitation>
B). Distance between ventral setae:
<emphasis id="33CFA37C7A5B092BCC2BFE01FD0A8CF8" box="[575,641,439,464]" italics="true" pageId="8" pageNumber="139">3b:3a</emphasis>
10 (912),
<emphasis id="33CFA37C7A5B092BCD1CFE0EFCB78CF8" box="[776,828,440,464]" italics="true" pageId="8" pageNumber="139">3a:g</emphasis>
27 (2628),
<emphasis id="33CFA37C7A5B092BCDC5FE0EFB8E8CF8" box="[977,1029,440,464]" italics="true" pageId="8" pageNumber="139">g:4a</emphasis>
64 (6470),
<emphasis id="33CFA37C7A5B092BCA8EFE0EFB6D8CF8" box="[1178,1254,440,464]" italics="true" pageId="8" pageNumber="139">4a:ps3</emphasis>
125 (117125). Diameter of anal suckers 20 (1920).
</paragraph>
<paragraph id="01047F6E7A5B092BCED3FDB6FE8A8E10" blockId="8.[151,1437,151,1832]" pageId="8" pageNumber="139">
Legs II approximately 1.5 times longer than legs I. Tarsus I with crest-like ventral extension stretching along whole segment, proximal end of this extension widely rounded (
<figureCitation id="998063EB7A5B092BCD7AFD92FC368F15" box="[878,957,548,573]" captionStart="FIGURE 6" captionStartId="9.[151,250,1905,1927]" captionTargetBox="[177,1421,204,1867]" captionTargetId="figure@9.[153,1438,193,1868]" captionTargetPageId="9" captionText="FIGURE 6. Protolichus placentis sp. n., details of male. A—subcapitulum of heteromorph male, ventral view, B—leg I of heteromorph male, dorsal view, C—leg I of heteromorph male, ventral view, D—leg II of heteromorph male, dorsal view, E leg II of heteromorph male, ventral view, F—tarsus IV of heteromorph male, dorsal view, G—subcapitulum of homeomorph male, ventral view, H—leg I of homeomorph male, dorsal view, I—leg II of homeomorph male, dorsal view." httpUri="https://zenodo.org/record/225834/files/figure.png" pageId="8" pageNumber="139">Figs. 6</figureCitation>
B, C). Tibia and genu I without apophyses but with some ventral verrucae. Femora I with longitudinal ventral crest and with ventral spine in distal part. Tarsus II with large ventral tubercle bearing seta
<emphasis id="33CFA37C7A5B092BCC6DFDDBFD198FAC" box="[633,658,621,644]" italics="true" pageId="8" pageNumber="139">ra</emphasis>
and
<emphasis id="33CFA37C7A5B092BCCD8FDDBFD668FAC" box="[716,749,621,644]" italics="true" pageId="8" pageNumber="139">wa</emphasis>
(
<figureCitation id="998063EB7A5B092BCCEAFDDAFCC68FAD" box="[766,845,620,645]" captionStart="FIGURE 6" captionStartId="9.[151,250,1905,1927]" captionTargetBox="[177,1421,204,1867]" captionTargetId="figure@9.[153,1438,193,1868]" captionTargetPageId="9" captionText="FIGURE 6. Protolichus placentis sp. n., details of male. A—subcapitulum of heteromorph male, ventral view, B—leg I of heteromorph male, dorsal view, C—leg I of heteromorph male, ventral view, D—leg II of heteromorph male, dorsal view, E leg II of heteromorph male, ventral view, F—tarsus IV of heteromorph male, dorsal view, G—subcapitulum of homeomorph male, ventral view, H—leg I of homeomorph male, dorsal view, I—leg II of homeomorph male, dorsal view." httpUri="https://zenodo.org/record/225834/files/figure.png" pageId="8" pageNumber="139">Figs. 6</figureCitation>
D, E). Tibiae II with long and high dorsal crest and with verrucous thumb like ventral process. Solenidia
<emphasis id="33CFA37C7A5B092BCCFFFD27FD718F80" box="[747,762,657,680]" italics="true" pageId="8" pageNumber="139">φ</emphasis>
of tibiae I, II verrucous. Seta
<emphasis id="33CFA37C7A5B092BCA5DFD26FBE18F8F" box="[1097,1130,656,679]" italics="true" pageId="8" pageNumber="139">cG</emphasis>
of genu II flattened, bladelike, 70 (7075) long, approximately 1.41.5 times longer than segment; anterior part of genu II with short dorsal crest, ventral surface verrucous. Genu II with verrucous ventral side posterior to setae
<emphasis id="33CFA37C7A5B092BCA4FFD6EFB0B8FC7" box="[1115,1152,728,751]" italics="true" pageId="8" pageNumber="139">mG</emphasis>
. Femur II with small and acute dorsal spine and with longitudinal ventral crest. Setae
<emphasis id="33CFA37C7A5B092BCD51FD4AFCE48E3C" box="[837,879,764,789]" italics="true" pageId="8" pageNumber="139">d, e</emphasis>
of tarsus IV spiculiform, with distinct basal ring (
<figureCitation id="998063EB7A5B092BCE8BFCA9FF688E10" box="[159,227,799,824]" captionStart="FIGURE 6" captionStartId="9.[151,250,1905,1927]" captionTargetBox="[177,1421,204,1867]" captionTargetId="figure@9.[153,1438,193,1868]" captionTargetPageId="9" captionText="FIGURE 6. Protolichus placentis sp. n., details of male. A—subcapitulum of heteromorph male, ventral view, B—leg I of heteromorph male, dorsal view, C—leg I of heteromorph male, ventral view, D—leg II of heteromorph male, dorsal view, E leg II of heteromorph male, ventral view, F—tarsus IV of heteromorph male, dorsal view, G—subcapitulum of homeomorph male, ventral view, H—leg I of homeomorph male, dorsal view, I—leg II of homeomorph male, dorsal view." httpUri="https://zenodo.org/record/225834/files/figure.png" pageId="8" pageNumber="139">Fig. 6</figureCitation>
F).
</paragraph>
<paragraph id="01047F6E7A5B092BCED3FCF2FBC18EC5" blockId="8.[151,1437,151,1832]" pageId="8" pageNumber="139">
Homeomorph male (1
<typeStatus id="DE00C1CC7A5B092BCFC7FCF2FDB28E74" box="[467,569,836,860]" pageId="8" pageNumber="139" type="paratype">paratype</typeStatus>
). Subcapitulum with posterior margin convex, with short oblique folds in postero-lateral angles (
<figureCitation id="998063EB7A5B092BCF83FCD1FE508EA8" box="[407,475,871,896]" captionStart="FIGURE 6" captionStartId="9.[151,250,1905,1927]" captionTargetBox="[177,1421,204,1867]" captionTargetId="figure@9.[153,1438,193,1868]" captionTargetPageId="9" captionText="FIGURE 6. Protolichus placentis sp. n., details of male. A—subcapitulum of heteromorph male, ventral view, B—leg I of heteromorph male, dorsal view, C—leg I of heteromorph male, ventral view, D—leg II of heteromorph male, dorsal view, E leg II of heteromorph male, ventral view, F—tarsus IV of heteromorph male, dorsal view, G—subcapitulum of homeomorph male, ventral view, H—leg I of homeomorph male, dorsal view, I—leg II of homeomorph male, dorsal view." httpUri="https://zenodo.org/record/225834/files/figure.png" pageId="8" pageNumber="139">Fig. 6</figureCitation>
G). Terminal cleft between opisthosomal lobes very narrow, margins bearing setae
<emphasis id="33CFA37C7A5B092BCB95FCDEFA178EA8" box="[1409,1436,872,896]" italics="true" pageId="8" pageNumber="139">e1</emphasis>
nearly touching; dorsal surface of lobes posterior to base
<emphasis id="33CFA37C7A5B092BCD27FC3AFCC58E8D" box="[819,846,908,933]" italics="true" pageId="8" pageNumber="139">h3</emphasis>
without small transverse crests (
<figureCitation id="998063EB7A5B092BCAD8FC3AFA988E8C" box="[1228,1299,908,933]" captionStart="FIGURE 4" captionStartId="6.[151,250,1764,1786]" captionTargetBox="[158,1429,200,1737]" captionTargetId="figure@6.[151,1436,193,1742]" captionTargetPageId="6" captionText="FIGURE 4. Dorsal view of male opisthosoma of Protolichus ornatus sp. n. (A, B) and Protolichus placentis sp. n. (C, D). A, C—heteromorph males, B, D—homeomorph males." httpUri="https://zenodo.org/record/225832/files/figure.png" pageId="8" pageNumber="139">Fig. 4</figureCitation>
D). Setae
<emphasis id="33CFA37C7A5B092BCB92FC3AFA108E8C" box="[1414,1435,908,932]" italics="true" pageId="8" pageNumber="139">f2</emphasis>
narrowly spatuliform, with bidentate apices. Anterior ends of paragenital apodemes free from epimerites IIa and IIIa; transverse bridge between middle parts of paragenital apodemes not developed.
</paragraph>
<paragraph id="01047F6E7A5B092BCED3FC4EFE8F8955" blockId="8.[151,1437,151,1832]" pageId="8" pageNumber="139">
Legs II about 1.2 times longer than legs I. Tarsi I, II with blunt-angular ventral margin. Tibiae I with small apico-ventral spine in anterior part (
<figureCitation id="998063EB7A5B092BCC59FBAAFD29891D" box="[589,674,1052,1077]" captionStart="FIGURE 6" captionStartId="9.[151,250,1905,1927]" captionTargetBox="[177,1421,204,1867]" captionTargetId="figure@9.[153,1438,193,1868]" captionTargetPageId="9" captionText="FIGURE 6. Protolichus placentis sp. n., details of male. A—subcapitulum of heteromorph male, ventral view, B—leg I of heteromorph male, dorsal view, C—leg I of heteromorph male, ventral view, D—leg II of heteromorph male, dorsal view, E leg II of heteromorph male, ventral view, F—tarsus IV of heteromorph male, dorsal view, G—subcapitulum of homeomorph male, ventral view, H—leg I of homeomorph male, dorsal view, I—leg II of homeomorph male, dorsal view." httpUri="https://zenodo.org/record/225834/files/figure.png" pageId="8" pageNumber="139">Figs. 6</figureCitation>
H, I). Seta
<emphasis id="33CFA37C7A5B092BCD26FBABFCD8891C" box="[818,851,1053,1076]" italics="true" pageId="8" pageNumber="139">cG</emphasis>
of genu II thickened, with narrow membranous enlargement in basal part, 50 long, approximately 1.3 times longer than segment. Other segments of legs I, II not modified.
</paragraph>
<paragraph id="01047F6E7A5B092BCED3FB3EFB72887C" blockId="8.[151,1437,151,1832]" pageId="8" pageNumber="139">
Measurements: Idiosoma, length × width, 440 × 254. Prodorsal shield: length 140. Setae
<emphasis id="33CFA37C7A5B092BCAAEFB3FFB468988" box="[1210,1229,1161,1184]" italics="true" pageId="8" pageNumber="139">si</emphasis>
30 long. Distance between scapular setae:
<emphasis id="33CFA37C7A5B092BCFB2FB1BFE5589EC" box="[422,478,1197,1220]" italics="true" pageId="8" pageNumber="139">se:se</emphasis>
66,
<emphasis id="33CFA37C7A5B092BCC1AFB1BFDB689EC" box="[526,573,1197,1220]" italics="true" pageId="8" pageNumber="139">si:si</emphasis>
35. Hysterosoma 360 long. Hysteronotal shield: greatest length 345, greatest width 222, length of terminal cleft 58, width at level of setae
<emphasis id="33CFA37C7A5B092BCD2BFB66FCEE89C0" box="[831,869,1232,1256]" italics="true" pageId="8" pageNumber="139">ps1</emphasis>
9. Lateral setae
<emphasis id="33CFA37C7A5B092BCA08FB66FBBC89C0" box="[1052,1079,1232,1256]" italics="true" pageId="8" pageNumber="139">c2</emphasis>
45 long; setae
<emphasis id="33CFA37C7A5B092BCACBFB79FB7089C0" box="[1247,1275,1231,1256]" italics="true" pageId="8" pageNumber="139">d2</emphasis>
12 long; setae
<emphasis id="33CFA37C7A5B092BCE83FB42FF3A8824" box="[151,177,1268,1292]" italics="true" pageId="8" pageNumber="139">e2</emphasis>
thick, represented by macrosetae 103 long. Distances between dorsal setae:
<emphasis id="33CFA37C7A5B092BCA33FB42FBEC8825" box="[1063,1127,1268,1293]" italics="true" pageId="8" pageNumber="139">c2:d2</emphasis>
125,
<emphasis id="33CFA37C7A5B092BCABFFB42FB608825" box="[1195,1259,1268,1293]" italics="true" pageId="8" pageNumber="139">d2:e2</emphasis>
130,
<emphasis id="33CFA37C7A5B092BCB3BFB42FAE48825" box="[1327,1391,1268,1293]" italics="true" pageId="8" pageNumber="139">e2:h3</emphasis>
90,
<emphasis id="33CFA37C7A5B092BCE83FAAEFF5E8818" box="[151,213,1304,1328]" italics="true" pageId="8" pageNumber="139">
e2:
<geoCoordinate id="648F19A97A5B092BCEAFFAAEFF5E8818" box="[187,213,1304,1328]" direction="east" orientation="longitude" pageId="8" pageNumber="139" precision="925" value="1.9">e1</geoCoordinate>
</emphasis>
54,
<emphasis id="33CFA37C7A5B092BCF1EFAAEFEC38818" box="[266,328,1304,1328]" italics="true" pageId="8" pageNumber="139">e2:e2</emphasis>
110,
<emphasis id="33CFA37C7A5B092BCF9DFAA1FE408818" box="[393,459,1303,1328]" italics="true" pageId="8" pageNumber="139">h2:h2</emphasis>
54,
<emphasis id="33CFA37C7A5B092BCFEBFAA1FDCA8818" box="[511,577,1303,1328]" italics="true" pageId="8" pageNumber="139">h3:h3</emphasis>
42,
<emphasis id="33CFA37C7A5B092BCC61FAAEFD468818" box="[629,717,1304,1328]" italics="true" pageId="8" pageNumber="139">ps1:ps1</emphasis>
27,
<emphasis id="33CFA37C7A5B092BCD15FAA1FCCA8818" box="[769,833,1303,1328]" italics="true" pageId="8" pageNumber="139">e2:h2</emphasis>
50,
<emphasis id="33CFA37C7A5B092BCD61FAA1FC3C8818" box="[885,951,1303,1328]" italics="true" pageId="8" pageNumber="139">h2:h3</emphasis>
21,
<emphasis id="33CFA37C7A5B092BCDFFFAA1FBB38818" box="[1003,1080,1303,1328]" italics="true" pageId="8" pageNumber="139">ps1:h3</emphasis>
14. Genital apparatus 27 × 20. Distance between ventral setae:
<emphasis id="33CFA37C7A5B092BCFEBFA8AFDCA887D" box="[511,577,1340,1365]" italics="true" pageId="8" pageNumber="139">3b:3a</emphasis>
7,
<emphasis id="33CFA37C7A5B092BCC70FA8AFD1C887C" box="[612,663,1340,1364]" italics="true" pageId="8" pageNumber="139">3a:g</emphasis>
26,
<emphasis id="33CFA37C7A5B092BCCDCFA8AFD77887C" box="[712,764,1340,1364]" italics="true" pageId="8" pageNumber="139">g:4a</emphasis>
58,
<emphasis id="33CFA37C7A5B092BCD38FA8AFCF2887C" box="[812,889,1340,1364]" italics="true" pageId="8" pageNumber="139">4a:ps3</emphasis>
108. Diameter of anal suckers 18.
</paragraph>
<paragraph id="01047F6E7A5B092BCED3FAE9FE1B8850" blockId="8.[151,1437,151,1832]" box="[199,400,1375,1400]" pageId="8" pageNumber="139">Female unknown.</paragraph>
<paragraph id="01047F6E7A5B092BCED3FA32FD598A2D" blockId="8.[151,1437,151,1832]" pageId="8" pageNumber="139">
<emphasis id="33CFA37C7A5B092BCED3FA32FE5B88B5" bold="true" box="[199,464,1412,1437]" pageId="8" pageNumber="139">Differential diagnosis.</emphasis>
The new species belongs to the
<emphasis id="33CFA37C7A5B092BCD59FA33FC2288B4" box="[845,937,1413,1436]" italics="true" pageId="8" pageNumber="139">crassior</emphasis>
group and is most similar to
<taxonomicName id="C6BB04ED7A5B092BCB17FA33FED988E8" authority="Trouessart, 1884" authorityName="Trouessart" authorityYear="1884" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="casuarinus">
<emphasis id="33CFA37C7A5B092BCB17FA33FA1788B4" box="[1283,1436,1413,1436]" italics="true" pageId="8" pageNumber="139">P. casuarinus</emphasis>
<bibRefCitation id="652A029F7A5B092BCE83FA1EFED988E8" author="Trouessart" box="[151,338,1448,1472]" pageId="8" pageNumber="139" refString="Trouessart, E. L. (1884) Les Sarcoptides plumicoles. Journal de Micrographie, 8, 527 - 532, 572 - 579." type="journal article" year="1884">Trouessart, 1884</bibRefCitation>
</taxonomicName>
by having a blade-shaped seta
<emphasis id="33CFA37C7A5B092BCCB8FA1EFD468897" box="[684,717,1448,1471]" italics="true" pageId="8" pageNumber="139">cG</emphasis>
II, a dorsal crest on genu II, and a long ventral process on tibia II in heteromorph males. The new species differs from
<taxonomicName id="C6BB04ED7A5B092BCCF1FA7BFCF088CC" box="[741,891,1485,1508]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="casuarinus">
<emphasis id="33CFA37C7A5B092BCCF1FA7BFCF088CC" box="[741,891,1485,1508]" italics="true" pageId="8" pageNumber="139">P. casuarinus</emphasis>
</taxonomicName>
by the following features. In both male forms of
<taxonomicName id="C6BB04ED7A5B092BCE83FA47FE908B20" box="[151,283,1519,1544]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="placentis">
<emphasis id="33CFA37C7A5B092BCE83FA47FE908B20" box="[151,283,1519,1544]" italics="true" pageId="8" pageNumber="139">P. placentis</emphasis>
</taxonomicName>
, setae
<emphasis id="33CFA37C7A5B092BCF64FA46FE018B20" box="[368,394,1520,1544]" italics="true" pageId="8" pageNumber="139">e2</emphasis>
are relatively long and definitely extend beyond the lobar apices; in heteromorph males, femur I has an acute ventral spine in the distal part of the segment and a ventral longitudinal crest strongly enlarged proximally, and the ventral extension of tibia II is thumb-like, rounded apically, with a verrucous ventral side; in homeomorph males, the subcapitulum has oblique folds, slightly smaller than in heteromorph males. In both male forms of
<taxonomicName id="C6BB04ED7A5B092BCEE8F937FE1A8BB0" box="[252,401,1665,1688]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="casuarinus">
<emphasis id="33CFA37C7A5B092BCEE8F937FE1A8BB0" box="[252,401,1665,1688]" italics="true" pageId="8" pageNumber="139">P. casuarinus</emphasis>
</taxonomicName>
, setae
<emphasis id="33CFA37C7A5B092BCFC9F936FE7C8BB0" box="[477,503,1664,1688]" italics="true" pageId="8" pageNumber="139">e2</emphasis>
are shorter and do not extend to the level of the lobar apices; in heteromorph males, femur I does not have a ventral spine and the ventral longitudinal crest is narrow, and the ventral extension of tibia II is claw-shaped, acute apically and smooth; in homeomorph males, the subcapitulum lacks oblique folds and bears wavy transverse striae on the ventral surface.
</paragraph>
</subSubSection>
<subSubSection id="49A12CE57A5B092BCED3F8B9FBE78A00" box="[199,1132,1807,1832]" pageId="8" pageNumber="139" type="etymology">
<paragraph id="01047F6E7A5B092BCED3F8B9FBE78A00" blockId="8.[151,1437,151,1832]" box="[199,1132,1807,1832]" pageId="8" pageNumber="139">
<emphasis id="33CFA37C7A5B092BCED3F8B9FEC58A00" bold="true" box="[199,334,1807,1832]" pageId="8" pageNumber="139">Etymology.</emphasis>
The specific epithet was taken from the specific name of the
<typeStatus id="DE00C1CC7A5B092BCDEAF8A6FBA58A00" box="[1022,1070,1808,1832]" pageId="8" pageNumber="139">type</typeStatus>
host.
</paragraph>
</subSubSection>
</treatment>
</document>