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<document ID-CLB-Dataset="7610" ID-DOI="http://dx.doi.org/10.3897/zookeys.1026.61554" ID-GBIF-Dataset="0a2d723c-d292-4f0c-9d00-6cdd176c49a6" ID-PMC="PMC8016817" ID-Pensoft-Pub="1313-2970-1026-45" ID-Pensoft-UUID="2EFE20EFF6705E34BFA20CF497608C4D" ID-PubMed="33850418" ID-ZooBank="A3212DAACEF24A1F93CC1823940C0108" ModsDocID="1313-2970-1026-45" checkinTime="1616697129133" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Surbakti, Suriani, Sumoked, Bob &amp; Balke, Michael" docDate="2021" docId="7427E30DF6B85974B2937820FFBC61A3" docLanguage="en" docName="ZooKeys 1026: 45-67" docOrigin="ZooKeys 1026" docPubDate="2021-03-25" docSource="http://dx.doi.org/10.3897/zookeys.1026.61554" docTitle="Exocelina apistefti Shaverdo, Surbakti &amp; Balke 2021, sp. nov." docType="treatment" docUuid="7862C435-10FA-4ED3-B73E-BACCE3D10EBB" docUuidSource="ZooBank" docVersion="7" id="2EFE20EFF6705E34BFA20CF497608C4D" lastPageNumber="45" masterDocId="2EFE20EFF6705E34BFA20CF497608C4D" masterDocTitle="Seven new species of the Exocelina ekari group from New Guinea central and coastal mountains (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="67" masterPageNumber="45" pageNumber="45" updateTime="1732748175969" updateUser="ExternalLinkService">
<mods:mods id="84FFC45AF67BEB091145C370AA35993D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="47CC8BB63F6B7F401785033755D580A3">
<mods:title id="8913EF5D0514E9585D8B912A250388CB">Seven new species of the Exocelina ekari group from New Guinea central and coastal mountains (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
</mods:titleInfo>
<mods:name id="1BCDE23C898346F5233855CDD2A11CC5" type="personal">
<mods:role id="DC8E2F49908FC35932B4F48B35D0AB51">
<mods:roleTerm id="0054EBE3C6FD3F86FDD358B12983529E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="21009C05074F25F2FCCF0C9C3F5457BB">Shaverdo, Helena</mods:namePart>
<mods:affiliation id="F8D241D7F7F7676DA4DC634473726708">Naturhistorisches Museum Wien, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
<mods:nameIdentifier id="BAB0796874E7F812CD73977F31D04812" type="email">shaverdo@mail.ru</mods:nameIdentifier>
</mods:name>
<mods:name id="390F5095B4D76460A35C47C21EF6C498" type="personal">
<mods:role id="99D2C26D20AE3FCF58C9202D4115FFBA">
<mods:roleTerm id="29769C9961D63F7F9A23182058425F81">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FC431C914282CCDB6F4563B56EF37842">Surbakti, Suriani</mods:namePart>
<mods:affiliation id="707CC1B29927BD4C994B1CDBEE301E92">Department of Biology, Universitas Cendrawasih, Waena, Papua, Indonesia</mods:affiliation>
</mods:name>
<mods:name id="50BEA86B9EA2A54AAD34B2E1CB06E1C7" type="personal">
<mods:role id="49F6421DBCD2EE7C240062BEB52AB640">
<mods:roleTerm id="A4AA647519B371A95902555988F083EC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B5847A40E123D6A1F78F849EA8ECA2CB">Sumoked, Bob</mods:namePart>
<mods:affiliation id="4554170D95EC6574A94B33344DEF0C66">Walian 2, Tomohon Selatan, N Sulawesi 95439, Indonesia</mods:affiliation>
</mods:name>
<mods:name id="44F24E2680C7459AEEB396044E312E72" type="personal">
<mods:role id="7DC0F4A4AA5D60D2EBAC4A0056DBD81C">
<mods:roleTerm id="93AA65ADAB8A40E89A2D53CE1543B0BB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="99F37727056348AE4559B240578903D8">Balke, Michael</mods:namePart>
<mods:nameIdentifier id="B01F0C64A0BB99CA6798FFC9BE6D73D4" type="ORCID">https://orcid.org/0000-0002-3773-6586</mods:nameIdentifier>
<mods:affiliation id="98C81B988B39D226067B3A56348A5195">SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247 Munich, Germany &amp; GeoBioCenter, Ludwig-Maximilians-University, Munich, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource id="00420A1F062D8D346F8C8C340150ED06">text</mods:typeOfResource>
<mods:relatedItem id="3541FADD81B33E44C94BDBEBF85C2F55" type="host">
<mods:titleInfo id="69F909422CE58A9943F9FC42F8223535">
<mods:title id="306372C15A18C312D5F590C4A4F16C92">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="C89D5FBA14697E8157CE1E890FCBEE7F">
<mods:date id="0494542BEA219F0484424C0FC2A36B45">2021</mods:date>
<mods:detail id="F8744FAE1DAC3E04D8E106697DE903E8" type="pubDate">
<mods:number id="6B7C4B8A252113AF93FE1D1FCD89D97B">2021-03-25</mods:number>
</mods:detail>
<mods:detail id="013D14E79EACF007151DE9657DB5F01C" type="volume">
<mods:number id="5F6A7870A64AC4DCB528C0A4E22D407E">1026</mods:number>
</mods:detail>
<mods:extent id="28B053E3C7A89157C3F0456DD4D07844" unit="page">
<mods:start id="813F69679D1A38E4B8E6254D4AAEF69E">45</mods:start>
<mods:end id="F60BBFA58AFBF098CD02752360F107F2">67</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="0BDAF0F7ECE97771243CA8E9CD3E407A">
<mods:url id="6FA70B3478A6BF03652A1712AF791B1B">http://dx.doi.org/10.3897/zookeys.1026.61554</mods:url>
</mods:location>
<mods:classification id="BE0988E090A2037C1D8312F2DD06AEEF">journal article</mods:classification>
<mods:identifier id="1B847BCFBE6D92FC9AB0ECD0DFEA632F" type="DOI">http://dx.doi.org/10.3897/zookeys.1026.61554</mods:identifier>
<mods:identifier id="640338859395ADBAEDAB91920E720691" type="Pensoft-Pub">1313-2970-1026-45</mods:identifier>
<mods:identifier id="8BE6B0D5BDAAF97BA9C9172452419436" type="ZooBank">A3212DAACEF24A1F93CC1823940C0108</mods:identifier>
<mods:identifier id="E95B33953078FC428E3C3E196FDE3DBC" type="Pensoft-UUID">2EFE20EFF6705E34BFA20CF497608C4D</mods:identifier>
</mods:mods>
<treatment id="7427E30DF6B85974B2937820FFBC61A3" ID-GBIF-Taxon="180680995" LSID="urn:lsid:zoobank.org:act:7862C435-10FA-4ED3-B73E-BACCE3D10EBB" httpUri="http://treatment.plazi.org/id/7427E30DF6B85974B2937820FFBC61A3" lastPageNumber="45" pageId="0" pageNumber="45" scope_family="Dytiscidae" scope_order="Coleoptera" scope_subFamily="Copelatinae">
<subSubSection id="DF55D6774FC555DEBD72F83B32B7B6AA" pageId="0" pageNumber="45" type="nomenclature">
<paragraph id="889D3AF3E8C19368544EBE17C704BED3" pageId="0" pageNumber="45">
<taxonomicName id="119B1B31DD19C92B14977D04B1FFBE4D" ID-CoL="8LZZX" LSID="http://zoobank.org/7862C435-10FA-4ED3-B73E-BACCE3D10EBB" authority="Shaverdo, Surbakti &amp; Balke" authorityName="Shaverdo, Surbakti &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina apistefti" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="apistefti" status="sp. nov.">Exocelina apistefti Shaverdo, Surbakti &amp; Balke</taxonomicName>
<taxonomicNameLabel id="C801BCAF45C2B6137892EA045EC5AF24" pageId="0" pageNumber="45">sp. nov.</taxonomicNameLabel>
<figureCitation id="EA56DF1C270F40195429860E53D37FFD" captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus and colouration of holotype 1 Exocelina foja sp. nov. 2 E. waaf sp. nov. 3 E. riberai sp. nov. 4 E. apistefti sp. nov." figureDoi="10.3897/zookeys.1026.61554.figures1-4" httpUri="https://binary.pensoft.net/fig/524445" pageId="0" pageNumber="45">Figures 4</figureCitation>
<figureCitation id="AD36063C3E4D10A8A1973666C9BACE72" captionStart="Figures 7, 8" captionStartId="F3" captionText="Figures 7, 8. 7 Exocelina riberai sp. nov., holotype 8 E. apistefti sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures7-8" httpUri="https://binary.pensoft.net/fig/524447" pageId="0" pageNumber="45">, 8</figureCitation>
<figureCitation id="42FB1A4B0115F83293A153BE991F72B8" captionStart="Figure 15" captionStartId="F7" captionText="Figure 15. Map of the western part of New Guinea showing the species distribution." figureDoi="10.3897/zookeys.1026.61554.figure15" httpUri="https://binary.pensoft.net/fig/524451" pageId="0" pageNumber="45">, 15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="3CA5921523C8F4AD59789A37C7D414A6" pageId="0" pageNumber="45" type="reference_group">
<paragraph id="58DA50D02D288971E4B2062A5991A475" pageId="0" pageNumber="45">
<taxonomicName id="6D801B81ED5AB5800068637268F9BF30" authority="Shaverdo &amp; Surbakti &amp; Sumoked &amp; Balke, 2021" authorityName="Shaverdo &amp; Surbakti &amp; Sumoked &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina brahminensis" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="brahminensis" status="sp. nov.">Exocelina nr. brahminensis</taxonomicName>
#7287:
<bibRefCitation id="5F998A7A17774FEC1727C8E1104F5BA1" DOI="https://doi.org/10.1186/s12862-021-01764-2" author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="45" refId="B10" refString="Toussaint, EFA, White, LT, Shaverdo, H, Lam, A, Surbakti, S, Panjaitan, R, Sumoked, B, von Rintelen, T, Sagata, K, Balke, M, 2021. [in press] New Guinean orogenic dynamics and biota evolution revealed using a custom geospatial analyses pipeline. BMS Evolutionary Biology. https://doi.org/10.1186/s12862-021-01764-2" title="[in press] New Guinean orogenic dynamics and biota evolution revealed using a custom geospatial analyses pipeline. BMS Evolutionary Biology." url="https://doi.org/10.1186/s12862-021-01764-2" year="2021">Toussaint et al. 2021</bibRefCitation>
: figs 3-6.
</paragraph>
</subSubSection>
<subSubSection id="7892E271F081E9D6E8C3EBEEEBF611E8" pageId="0" pageNumber="45" type="type locality">
<paragraph id="3C98F48E28D530809ECDC6B57C09FCA5" pageId="0" pageNumber="45">
<typeStatus id="4F70DCC4F228026DC661FF47C6083B81">Type</typeStatus>
locality.
</paragraph>
<paragraph id="B1030F9A79EED8667815AC6AAA704C35" pageId="0" pageNumber="45">
<materialsCitation id="FE426F92F7EF26426E72C8EF2EFE9251" ID-GBIF-Occurrence="3062138321" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Indonesia" elevation="1700" latitude="-2.5718334" location="Foja Mts" longLatPrecision="1" longitude="138.71724" municipality="Sarmi Regency" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
<collectingCountry id="38E7023C98948113E27FAB2C3F6B4068" name="Indonesia">Indonesia</collectingCountry>
:
<collectingRegion id="3284BEEC70F68E77DB289E776D3371C7" country="Indonesia" name="Papua Barat">Papua Province</collectingRegion>
,
<collectingMunicipality id="DA30402EE337B222BB79798CB6534273">Sarmi Regency</collectingMunicipality>
,
<location id="262B8B0DC67DE37EBE707C5FE03779D0" LSID="urn:lsid:plazi:treatment:7427E30DF6B85974B2937820FFBC61A3:262B8B0DC67DE37EBE707C5FE03779D0" country="Indonesia" latitude="-2.5718334" longLatPrecision="1" longitude="138.71724" municipality="Sarmi Regency" name="Foja Mts" stateProvince="Papua Barat">Foja Mts</location>
,
<geoCoordinate id="D5D1BB1FA9D609D9AA5A8A59F99D74DE" degrees="02" direction="south" minutes="34" orientation="latitude" precision="1" seconds="18.6" value="-2.5718334">02°34'18.6&quot;S</geoCoordinate>
,
<geoCoordinate id="D9121490A340E55EDB37EE8FD149294D" degrees="138" direction="east" minutes="43" orientation="longitude" precision="1" seconds="02.1" value="138.71724">138°43'02.1&quot;E</geoCoordinate>
,
<elevation id="F4AA835AF9436EA683F50FBCF8188B30" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<quantity id="B381D8EB4E386D4655DB433C07755943" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700 m</quantity>
a.s.l.
</elevation>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E6F62D8E389824DA4BB2AD1DE908CAFD" pageId="0" pageNumber="45" type="type material">
<paragraph id="42B8F6F4E76DE91AEC6F7B5AD8045622" pageId="0" pageNumber="45">
<typeStatus id="3ABA9CD7246B3B61BA960AE4252D596B">Type</typeStatus>
material.
</paragraph>
<paragraph id="39D2F21D7EAB8F9ED16F0FB4A24359D2" pageId="0" pageNumber="45">
<emphasis id="821125DA435D0047CE8E29136CFF1428" bold="true" pageId="0" pageNumber="45">
<emphasis id="22E852EFBB50D64210C22113BDA9286B" italics="true" pageId="0" pageNumber="45">
<typeStatus id="88C6B7EAA9A9264DA837046B1B25464A">Holotype</typeStatus>
</emphasis>
:
</emphasis>
<materialsCitation id="47867F5BEE043F65BDD9945B64B34F4E" ID-GBIF-Occurrence="3062138339" collectingDate="2016-05-23" collectingDateMax="2016-06-03" collectingDateMin="2016-05-23" collectionCode="MZB" country="Indonesia" elevation="1700" latitude="-2.571839" location="Foja Mountains" longLatPrecision="1" longitude="138.71725" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
male &quot;
<collectingCountry id="C75C50DAEE102A9D3E8706F7A1752FE7" name="Indonesia">Indonesia</collectingCountry>
:
<collectingRegion id="FFBEA3EDA2582510C69B7C4976AD5CD2" country="Indonesia" name="Papua Barat">Papua</collectingRegion>
,
<location id="8D62B64180210C7DCA41E6DD99853E8D" LSID="urn:lsid:plazi:treatment:7427E30DF6B85974B2937820FFBC61A3:8D62B64180210C7DCA41E6DD99853E8D" country="Indonesia" latitude="-2.571839" longLatPrecision="1" longitude="138.71725" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
, bog camp,
<quantity id="7A96BD7BEDE9DB08A02519C00FFCAEDA" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<elevation id="B00F3A1B12C62E86FF28718C40F3CFD7" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700m</elevation>
</quantity>
,
<date id="DB7AED12F8A1AAA1760972015AD04F01" value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">
<collectingDate id="B2ED5C54491D863606BF3D22BF2DBE7C" value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">23.v.-3.vi.2016</collectingDate>
</date>
,
<geoCoordinate id="CEE34F6135BD04AAAA5740BEF8F7101F" degrees="2.571839" direction="south" orientation="latitude" precision="1" value="-2.571839">-2.571839</geoCoordinate>
<geoCoordinate id="3ED5E64872B4F2BCE3FDAE88E8732DCE" degrees="138.717250" direction="east" orientation="longitude" precision="1" value="138.71725">138.717250</geoCoordinate>
, Sumoked (Pap058)&quot;,
<normalizedToken id="B2649E3ACF738F18226BF47083BDD33A" originalValue="“7287”">&quot;7287&quot;</normalizedToken>
[green text] (
<collectionCode id="4AC2C32D1395CE18A56856F0FA95A212" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="E0F48CA9980AC39C43BE72F08DC2A551" pageId="0" pageNumber="45">
<materialsCitation id="129A96D5252C277841DECA31F7137C16" ID-GBIF-Occurrence="3062138334" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="KSP" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<emphasis id="C293224BCFD6ADCBC54F25BB61D0B29A" bold="true" pageId="0" pageNumber="45">
<emphasis id="04DF6BF762F0D507E18BE2AC0538AAE8" italics="true" pageId="0" pageNumber="45">
<typeStatus id="EA362384BB99EAE0897F28AD368E59C1">Paratypes</typeStatus>
</emphasis>
:
</emphasis>
<specimenCount id="9020DB7A39A6D2611F0C39576B2AF6DD" type="female">2 females</specimenCount>
with the same label as the
<typeStatus id="DFD4D4296DA32AB0A10302EA5C2EE536">holotype</typeStatus>
(
<collectionCode id="6BA4BDDBCD5849779A481C058A78F6E2" country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="4C13C0470C0251B7AB9BCF8E425B3023" ID-GBIF-Occurrence="3062138320" collectingDate="2016-05-23" collectingDateMax="2016-06-03" collectingDateMin="2016-05-23" collectionCode="MZB, KSP" country="Indonesia" elevation="1600" latitude="-2.561006" location="Foja Mountains" longLatPrecision="1" longitude="138.71149" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount id="970860D0E360EDA291C532F53CA486A5" type="male">1 male</specimenCount>
,
<specimenCount id="CAE28784798224A43C1CE832CD4EA663" type="female">1 female</specimenCount>
&quot;
<collectingCountry id="04A18AD5105919C82FD9FF1B07671311" name="Indonesia">Indonesia</collectingCountry>
:
<collectingRegion id="D1E107137B07C5279D200DA70CEA1893" country="Indonesia" name="Papua Barat">Papua</collectingRegion>
,
<location id="C5F1BEF221719927C587AB1FC529DB04" LSID="urn:lsid:plazi:treatment:7427E30DF6B85974B2937820FFBC61A3:C5F1BEF221719927C587AB1FC529DB04" country="Indonesia" latitude="-2.561006" longLatPrecision="1" longitude="138.71149" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
, river camp,
<quantity id="B29C76450E9D8A30EF630286E5D2FC3F" metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">
<elevation id="C769120F529E67089EF84607EF5D70C3" metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">1600m</elevation>
</quantity>
,
<date id="0ECD25B816F8963A76A5280EBD9AED82" value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">
<collectingDate id="D85A7C0245DC50408C67AEBDF5B917FC" value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">23.v.-3.vi.2016</collectingDate>
</date>
,
<geoCoordinate id="1EF7061223994E13DE085111B0D9C5C5" degrees="2.561006" direction="south" orientation="latitude" precision="1" value="-2.561006">-2.561006</geoCoordinate>
<geoCoordinate id="14F847D11D8DE5E779E4ADAF457E6E45" degrees="138.711487" direction="east" orientation="longitude" precision="1" value="138.71149">138.711487</geoCoordinate>
, Sumoked (Pap059)&quot; (
<collectionCode id="86BD7A50D42D6898656FDC800114ED66" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
,
<collectionCode id="9836D7202FF61DF2EE01DEB64ECAA7E5" country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8CE3D95C6D3E1AF8371425CB4163BC8" pageId="0" pageNumber="45" type="description">
<paragraph id="2C4D0AD992862C7FC594A2279BA98130" pageId="0" pageNumber="45">Description.</paragraph>
<paragraph id="9A670A4B124670F24D53990ADE364B6E" pageId="0" pageNumber="45">
<emphasis id="628D1BDF33ECDE058ED7FB372BA99352" bold="true" pageId="0" pageNumber="45">
<emphasis id="416503D38E88ACD4C2C12F253BC9503E" italics="true" pageId="0" pageNumber="45">Body size and form</emphasis>
:
</emphasis>
Beetle small: TL-H
<quantity id="CF0E6333D83CF73A9E82D44E31D67D1B" metricMagnitude="-3" metricUnit="m" metricValue="3.575" metricValueMax="3.7" metricValueMin="3.45" unit="mm" value="3.575" valueMax="3.7" valueMin="3.45">3.45-3.7 mm</quantity>
, TL
<quantity id="043591E5DD7DE773F32EC5A282D6264B" metricMagnitude="-3" metricUnit="m" metricValue="3.925" metricValueMax="4.05" metricValueMin="3.8" unit="mm" value="3.925" valueMax="4.05" valueMin="3.8">3.8-4.05 mm</quantity>
, MW
<quantity id="4B4E5BC24D0200D9C58BE7742CB95C26" metricMagnitude="-3" metricUnit="m" metricValue="1.925" metricValueMax="2.0" metricValueMin="1.85" unit="mm" value="1.925" valueMax="2.0" valueMin="1.85">1.85-2 mm</quantity>
(
<typeStatus id="EEEA4702FBA9718B24CA93EB6F3CA082">holotype</typeStatus>
: TL-H
<quantity id="11CCC2A79230F50DCFADB05C0A2C7775" metricMagnitude="-3" metricUnit="m" metricValue="3.5" unit="mm" value="3.5">3.5 mm</quantity>
, TL
<quantity id="4968D9C00327D44EFF6D4C3AAE1B76E3" metricMagnitude="-3" metricUnit="m" metricValue="3.9" unit="mm" value="3.9">3.9 mm</quantity>
,
<collectionCode id="D5331BAD61821AB2484466B4D7684413" country="Netherlands" name="Museum Wasmann">MW</collectionCode>
<quantity id="629D690EAAE89D3D532D7BCB143D5BF6" metricMagnitude="-3" metricUnit="m" metricValue="1.9" unit="mm" value="1.9">1.9 mm</quantity>
), with oblong-oval habitus (Fig.
<figureCitation id="4CD7736A98F2C722155281E622262148" captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus and colouration of holotype 1 Exocelina foja sp. nov. 2 E. waaf sp. nov. 3 E. riberai sp. nov. 4 E. apistefti sp. nov." figureDoi="10.3897/zookeys.1026.61554.figures1-4" httpUri="https://binary.pensoft.net/fig/524445" pageId="0" pageNumber="45">4</figureCitation>
).
</paragraph>
<paragraph id="CB6E47C2A7521D8F3181DC2835089BD9" pageId="0" pageNumber="45">
<emphasis id="716F4FC75F480160858C0D734F229959" bold="true" pageId="0" pageNumber="45">
<emphasis id="86275D56E48B9B291AA14A73AA0AB6C8" italics="true" pageId="0" pageNumber="45">Colouration</emphasis>
:
</emphasis>
Dorsally piceous, with paler lateral sides of pronotum (Fig.
<figureCitation id="745101079A2EC4BD9D9D8A508DC1FA5C" captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus and colouration of holotype 1 Exocelina foja sp. nov. 2 E. waaf sp. nov. 3 E. riberai sp. nov. 4 E. apistefti sp. nov." figureDoi="10.3897/zookeys.1026.61554.figures1-4" httpUri="https://binary.pensoft.net/fig/524445" pageId="0" pageNumber="45">4</figureCitation>
). Head piceous, with slightly paler, dark brown, anterior margin; pronotum piceous, slightly paler towards lateral sides, lateral sides brown to dark brown, yellowish to reddish brown at anterior angles; elytra uniformly piceous; head appendages and proximal part of legs yellowish brown, legs distally brown.
</paragraph>
<paragraph id="10981346F6E1222D430D265D97B93C7E" pageId="0" pageNumber="45">
<emphasis id="719118A194D952CB84E87EB26A3682BE" bold="true" pageId="0" pageNumber="45">
<emphasis id="8CC3A27C583F786C54AC6D722AF0F4A3" italics="true" pageId="0" pageNumber="45">Surface sculpture</emphasis>
:
</emphasis>
Shiny dorsally, with fine microreticulation and almost invisible punctation on elytra. Head with dense and coarse punctation (spaces between punctures 0-3 times size of punctures), distinctly finer and sparser anteriorly and posteriorly; diameter of punctures equal to diameter of cells of microreticulation. Pronotum with distinctly finer and sparser punctation than on head. Elytra with very sparse and fine punctation, almost invisible. Elytra with weakly impressed microreticulation; pronotum and especially head with stronger microreticulation. Metaventrite and metacoxa distinctly but weakly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal ventrites with weak microreticulation, strioles, and fine sparse punctation, coarser and denser on two last abdominal ventrites.
</paragraph>
<paragraph id="EC7A088A4141205E40D801C85651735D" pageId="0" pageNumber="45">
<emphasis id="BA923AC64132CED738F652EC12BC26C6" bold="true" pageId="0" pageNumber="45">
<emphasis id="0C77F6BF34530ACD0F9A3AAD5DAF833E" italics="true" pageId="0" pageNumber="45">Structures</emphasis>
:
</emphasis>
Pronotum without lateral bead. Base of prosternum and neck of prosternal process with ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively narrow, slightly convex medially, with distinct bead and few setae. Abdominal ventrite 6 broadly rounded apically.
</paragraph>
<paragraph id="D4F85E7CF429CD70961C6E5B22BB654C" pageId="0" pageNumber="45">
<emphasis id="20BCCA465084A5CF31E8BC0B8A92E10A" bold="true" pageId="0" pageNumber="45">
<emphasis id="4A63B21B9F99B64A17AD298796AD4CC8" italics="true" pageId="0" pageNumber="45">Male</emphasis>
:
</emphasis>
Antenna simple. Pro- and mesotarsomeres 1-3 not dilated, narrow. Protarsomere 4 cylindrical, narrow, with medium-sized, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior row of 13 and posterior row of six short setae (Fig.
<figureCitation id="A3B6B6A07FCDEEE5ECCB51766C58DF71" captionStart="Figures 7, 8" captionStartId="F3" captionText="Figures 7, 8. 7 Exocelina riberai sp. nov., holotype 8 E. apistefti sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures7-8" httpUri="https://binary.pensoft.net/fig/524447" pageId="0" pageNumber="45">8A</figureCitation>
). Median lobe with distinctly discontinuous outline; in lateral view, almost straight, with apex broad, curved downwards, and pointed at tip; in ventral view, with distinct submedian constriction, distal part narrower than proximal one, apex deeply and narrowly concave (Fig.
<figureCitation id="A96824CE58118C9E6BBCA27F8815F49A" captionStart="Figures 7, 8" captionStartId="F3" captionText="Figures 7, 8. 7 Exocelina riberai sp. nov., holotype 8 E. apistefti sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures7-8" httpUri="https://binary.pensoft.net/fig/524447" pageId="0" pageNumber="45">8B, C</figureCitation>
). Paramere with strong notch on dorsal side, with median notch tip sharply pointed, and subdistal part large and elongate; subdistal setae long and dense, of two kind: more numerous upper ones thin and lower setae shorter, thicker and flattened; proximal setae hair-like, distinctly more inconspicuous than subdistal ones (Fig.
<figureCitation id="3740CAB11E04F356CC671377709FA94C" captionStart="Figures 7, 8" captionStartId="F3" captionText="Figures 7, 8. 7 Exocelina riberai sp. nov., holotype 8 E. apistefti sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures7-8" httpUri="https://binary.pensoft.net/fig/524447" pageId="0" pageNumber="45">8D</figureCitation>
). Abdominal ventrite 6 broadly rounded, with 9-11 lateral striae on each side.
</paragraph>
<caption id="5291481BF4A335CA70CFD74B00019AA7" doi="10.3897/zookeys.1026.61554.figures7-8" httpUri="https://binary.pensoft.net/fig/524447" pageId="0" pageNumber="45" start="Figures 7, 8" startId="F3">
<paragraph id="5ABD96F5EB1F6CF51D9076E167140DE5" pageId="0" pageNumber="45">
<emphasis id="016C63948C298367B6D52D538756EDDB" bold="true" pageId="0" pageNumber="45">Figures 7, 8.</emphasis>
<emphasis id="9B02C80EE8E7E0696DD34F598F7D1B8F" bold="true" pageId="0" pageNumber="45">7</emphasis>
<taxonomicName id="2082728ABD5D10A106DC7158178119C0" authorityName="Shaverdo, Surbakti &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina riberai" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="riberai">
<emphasis id="9A5271C29C234CECD05332311229005D" italics="true" pageId="0" pageNumber="45">Exocelina riberai</emphasis>
</taxonomicName>
sp. nov.,
<typeStatus id="1EDEE3924C3394542494A7587FA272E7">holotype</typeStatus>
<emphasis id="5721339B22FFBEEAAB7A2FC171928FEE" bold="true" pageId="0" pageNumber="45">8</emphasis>
<taxonomicName id="7AFE21D19174A0AC8DD050B9E3D29650" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. apistefti" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="apistefti">
<emphasis id="E4D0E5F1BC8859397BB60F87DDB53B08" italics="true" pageId="0" pageNumber="45">E. apistefti</emphasis>
</taxonomicName>
sp. nov.,
<typeStatus id="3958F01F433AD8AA18BFC3A67C8C8D35">holotype</typeStatus>
<emphasis id="5B22292EB0A2E54489655346E986630C" bold="true" pageId="0" pageNumber="45">A</emphasis>
right protarsomeres 4 and
<quantity id="24FE2DBEAC5B8539B71C6E19D5A6A801" metricMagnitude="-1" metricUnit="m" metricValue="1.27" unit="in" value="5.0">5 in</quantity>
ventral view
<emphasis id="4927FE9D073C6C9125503437616F2752" bold="true" pageId="0" pageNumber="45">B</emphasis>
median lobe in ventral view
<emphasis id="7B50D26616BAA2C02C2A589A982EC2B7" bold="true" pageId="0" pageNumber="45">C</emphasis>
median lobe in lateral view
<emphasis id="D7F05AA1AE569DE513A29E5470B2EB5B" bold="true" pageId="0" pageNumber="45">D</emphasis>
right paramere in external view.
</paragraph>
</caption>
<paragraph id="E90EE51BF0F4FE9123E62352A6EBFE7C" pageId="0" pageNumber="45">
<emphasis id="6EBB9E574B9E00DA4A683D4DFF371717" bold="true" pageId="0" pageNumber="45">
<emphasis id="3AB2A46FA8D11400A00645DA371A61D4" italics="true" pageId="0" pageNumber="45">Female</emphasis>
:
</emphasis>
Pro- and mesotarsi not modified. Abdominal ventrite 6 without striae.
</paragraph>
</subSubSection>
<subSubSection id="2863F0892716FC4D54A80619C1B8F389" pageId="0" pageNumber="45" type="affinities">
<paragraph id="94AE0CC04379804706DCEC8368E7F8FD" pageId="0" pageNumber="45">Affinities.</paragraph>
<paragraph id="10E89FAB1ACB075B2A183EAE81C28FBB" pageId="0" pageNumber="45">
The species evidently belongs to the
<taxonomicName id="1C9BF5CA5E7F0482E9FDDDB26A3221A7" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. ekari" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="ekari">
<emphasis id="2CF811FD1EC75D7D02962800821AF914" italics="true" pageId="0" pageNumber="45">E. ekari</emphasis>
</taxonomicName>
group due to the discontinuous outline of its median lobe. The species is very similar to
<taxonomicName id="1BF81E815948DE1357CE07D5035376E5" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. brahminensis" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="brahminensis">
<emphasis id="6581B315A754331EC5D50FF6453DA85D" italics="true" pageId="0" pageNumber="45">E. brahminensis</emphasis>
</taxonomicName>
<bibRefCitation id="5EFB3D2E6BF85D116A01B3964C012F8C" DOI="https://doi.org/10.3897/zookeys.250.3715" author="Shaverdo, HV" journalOrPublisher="ZooKeys" pageId="0" pageNumber="45" pagination="1 - 76" refId="B3" refString="Shaverdo, HV, Surbakti, S, Hendrich, LM, Balke, M, 2012. Introduction of the Exocelina ekari -group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 250: 1 - 76, DOI: https://doi.org/10.3897/zookeys.250.3715" title="Introduction of the Exocelina ekari - group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)." url="https://doi.org/10.3897/zookeys.250.3715" volume="250" year="2012">
Shaverdo et al.,
<quantity id="287E21BF5CB28731EC63011CECD47ABD" metricMagnitude="1" metricUnit="m" metricValue="5.11048" unit="in" value="2012.0">2012 in</quantity>
</bibRefCitation>
general appearance and structure of male genitalia, especially in the sharply pointed median notch tip of paramere, but differs from it in lager size (TL-H
<quantity id="57525692489879C153A87747478EAB5E" metricMagnitude="-3" metricUnit="m" metricValue="3.2249999999999996" metricValueMax="3.3" metricValueMin="3.15" unit="mm" value="3.2249999999999996" valueMax="3.3" valueMin="3.15">3.15-3.3 mm</quantity>
for
<taxonomicName id="8459582327D5FD422540776C888A53DD" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. brahminensis" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="brahminensis">
<emphasis id="5615FBF414F15E0B63C10482334BFA8D" italics="true" pageId="0" pageNumber="45">E. brahminensis</emphasis>
</taxonomicName>
), darker dorsal colouration, shape of the median lobe, and paramere setation.
</paragraph>
</subSubSection>
<subSubSection id="451925782F5EF2EE2A667E91ED082B96" pageId="0" pageNumber="45" type="distribution">
<paragraph id="DBE84DEDDA9021DFAECA9333166CC25E" pageId="0" pageNumber="45">Distribution.</paragraph>
<paragraph id="097AB29731CFD0BCABB552CB7F5F6FBB" pageId="0" pageNumber="45">
<materialsCitation id="354A45A802D3914E8DA65E1ACBC43DB2" ID-GBIF-Occurrence="3062138322" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Indonesia" location="Foja Mountains" municipality="This" specimenCount="1" stateProvince="Papua Barat">
<collectingCountry id="72DED4841B221BA29087BC343AECE5CB" name="Indonesia">Indonesia</collectingCountry>
:
<collectingRegion id="09D0103FB1371E2FE2AC5BD087FDBFDA" country="Indonesia" name="Papua Barat">Papua Province</collectingRegion>
.
<collectingMunicipality id="4A5EA226B1EB541DC3CDD288C7B65B1D">This</collectingMunicipality>
species is known only from
<location id="A185B40B3578D71336FE84E7FDA4F5D5" LSID="urn:lsid:plazi:treatment:7427E30DF6B85974B2937820FFBC61A3:A185B40B3578D71336FE84E7FDA4F5D5" country="Indonesia" municipality="This" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
, from and near the
<typeStatus id="50B5B0EA3702FEF4F588923121D7BD4B">type</typeStatus>
locality (Fig.
<figureCitation id="6174C47E3EAA7170659C00B521C01C0D" captionStart="Figure 15" captionStartId="F7" captionText="Figure 15. Map of the western part of New Guinea showing the species distribution." figureDoi="10.3897/zookeys.1026.61554.figure15" httpUri="https://binary.pensoft.net/fig/524451" pageId="0" pageNumber="45">15</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C10F1E5A077E15B5D5A9663AB8F74749" pageId="0" pageNumber="45" type="habitat">
<paragraph id="F344B2C50D9897B30317CD707E16D88F" pageId="0" pageNumber="45">Habitat.</paragraph>
<paragraph id="BC19F1E5F4485F156D906E5A8443EA4E" pageId="0" pageNumber="45">The specimens were collected from small, shallow forest creeks.</paragraph>
</subSubSection>
<subSubSection id="BBFE9BDECB014F2CF49EB0416F8F2D24" pageId="0" pageNumber="45" type="etymology">
<paragraph id="243178CDD6ECBFC4744618A186C7E4D2" pageId="0" pageNumber="45">Etymology.</paragraph>
<paragraph id="858EDA2022931FB2634BE024EF0BD3CD" pageId="0" pageNumber="45">
The species name
<taxonomicName id="D3B65CE587C7FE511B1865212EAB7760" authorityName="Shaverdo, Surbakti &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina apistefti" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="apistefti">
<emphasis id="1B549C898C76054D1BC7ED79361DA360" italics="true" pageId="0" pageNumber="45">Exocelina apistefti</emphasis>
</taxonomicName>
derives from Greek
<emphasis id="B0FBB8F2C7B504902B72AE414F78D5A9" italics="true" pageId="0" pageNumber="45">
<normalizedToken id="49F243F5BA768841915851ACF1A1F503" originalValue="απίστευτος">απίστευτος</normalizedToken>
</emphasis>
(feminine
<emphasis id="8AE31141B1146D5D66882844A27503C3" italics="true" pageId="0" pageNumber="45">
<normalizedToken id="8D10956C2A6C007720D975BC8A94C93F" originalValue="απίστευτη">απίστευτη</normalizedToken>
</emphasis>
) meaning unbelievable. The name is an adjective in the nominative singular.
</paragraph>
</subSubSection>
</treatment>
</document>
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