165 lines
17 KiB
XML
165 lines
17 KiB
XML
<document id="67E4A147DBD91C3898CCD063D71819CF" ID-DOI="10.3897/zookeys.1223.139346" ID-ZooBank="451749E9-009E-43E9-A6F1-892035BDF1B0" ID-publisher-id="139346" URI-arpha="6EA2BAFE-82AB-5570-95B8-EE1F5FA60D46" URI-zoobank="http://zoobank.org/451749E9-009E-43E9-A6F1-892035BDF1B0" article-type="research-article" checkinTime="1736349736103" checkinUser="pensoft" docAuthor="Wesener, Thomas, Akkari, Nesrine & Golovatch, Sergei I." docDate="2025" docId="4F9FCBD8F2765C5BB032EDB13FA1CEE1" docLanguage="en" docName="ZooKeys 1223: 185-220" docOrigin="ZooKeys 1223" docSource="https://zookeys.pensoft.net/article/139346/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Dalodesmus orator Hoffman 1974" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="220" masterDocId="6EA2BAFE82AB557095B8EE1F5FA60D46" masterDocTitle="Revision of the millipede family Dalodesmidae in Madagascar, with descriptions of two new Malagasy species of Dalodesmus Cook, 1896 (Diplopoda, Polydesmida)" masterLastPageNumber="220" masterPageNumber="185" pageNumber="185" updateTime="1736350173991" updateUser="ExternalLinkService">
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<mods:titleInfo id="194CE86170F8779DA747F8DD41BD4484">
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<mods:title id="0105D0C283F61DBC2C1AA7FF1434B5EF">Revision of the millipede family Dalodesmidae in Madagascar, with descriptions of two new Malagasy species of Dalodesmus Cook, 1896 (Diplopoda, Polydesmida)</mods:title>
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<mods:roleTerm id="A683A125AE4F92477DABD08CA08844EF">Author</mods:roleTerm>
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<mods:namePart id="3CC288E68DE62680701D61FE841326E5">Wesener, Thomas</mods:namePart>
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<mods:nameIdentifier id="38118E3108E7B3614E2F79E2F4619410" type="ORCID">0000-0002-2028-3541</mods:nameIdentifier>
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<mods:affiliation id="F27773EFA50BEB28BB41E343625E6867">Zoological Research Museum Alexander Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, D- 53113 Bonn, Germany</mods:affiliation>
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<mods:name id="B092D01E64A001E10C802AFE81E873D5" type="personal">
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<mods:roleTerm id="8906E41C3118E037FA179B27865012D1">Author</mods:roleTerm>
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<mods:namePart id="06A699B00449FE13E4BD4721241E17DA">Akkari, Nesrine</mods:namePart>
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<mods:nameIdentifier id="ADF84993522BAA6D12546169A1C390F8" type="ORCID">0000-0001-5019-4833</mods:nameIdentifier>
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<mods:affiliation id="1411D3D8CD67F20AF8E87BC470923280">Third Zoological Department, Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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<mods:namePart id="A6458918389DBB594A5E8534095F588C">Golovatch, Sergei I.</mods:namePart>
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<mods:affiliation id="55D4F1CFA2C7A2919691C0BC04F80182">Institute of Ecology and Evolution, Russian Academy of Sciences, Leninsky prospekt 33, Moscow 119071, Russia</mods:affiliation>
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<mods:typeOfResource id="9E8508D3BF2C9DAB06459C84EBCBC052">text</mods:typeOfResource>
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<mods:title id="86986E83C5A866BDF19ED78E38712ED4">ZooKeys</mods:title>
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<mods:part id="759513883F11A8E682C635CC6A980CF4">
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<mods:date id="AC60AACAE03E910C5B4FDFDAE3A5DE5D">2025</mods:date>
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<mods:number id="ECFA4B4FC29CE8A60FC33D15D58882D3">2025-01-08</mods:number>
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<mods:number id="2372D296E485D835457A8657F1125081">1223</mods:number>
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<mods:start id="EEA2D8BAC689405D4353EEBA84D6CE14">185</mods:start>
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<mods:identifier id="06C9EC4D15462F9CAD9EF284B27B1BDE" type="DOI">10.3897/zookeys.1223.139346</mods:identifier>
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<mods:identifier id="16565652006D842D2E89F9B44FB2EFF8" type="ZooBank">451749E9-009E-43E9-A6F1-892035BDF1B0</mods:identifier>
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<treatment id="4F9FCBD8F2765C5BB032EDB13FA1CEE1" ID-DOI="http://doi.org/10.5281/zenodo.14616987" ID-Zenodo-Dep="14616987" ID-arpha="4F9FCBD8-F276-5C5B-B032-EDB13FA1CEE1" LSID="urn:lsid:plazi:treatment:4F9FCBD8F2765C5BB032EDB13FA1CEE1" httpUri="http://treatment.plazi.org/id/4F9FCBD8F2765C5BB032EDB13FA1CEE1">
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<subSubSection id="AA64F46A960008799B27B539378A9DD0" type="nomenclature">
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<paragraph id="407527DA04D55EDDB86D23718E7E96F7">
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<taxonomicName id="2CC43020B8F5AEFE5584DBD7B5467027" ID-CoL="6CBHG" authority="Hoffman, 1974" authorityName="Hoffman" authorityYear="1974" class="Diplopoda" family="Dalodesmidae" genus="Dalodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="orator">
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<emphasis id="FE1225184478E361C5A852CA6866CDA1" italics="true">Dalodesmus orator</emphasis>
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<bibRefCitation id="211F18874179F1F919EDFCB326978C23" author="Hoffman RL" issue="2" journalOrPublisher="Wasmann Journal of Biology" pagination="221-246" refId="B14" refString="Hoffman RL (1974) Short studies on dalodesmid millipeds from South Africa and Madagascar. Wasmann Journal of Biology 32 (2): 221 - 246." title="Short studies on dalodesmid millipeds from South Africa and Madagascar." volume="32" year="1974">Hoffman, 1974</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C01164C36F115C215A48F95828D52E91" type="description">
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<paragraph id="17092146E75B2E5A0EB7A3BBE5745765">
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<figureCitation id="3EA4913FD1505E23924C6DEE996F66FA" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Dalodesmus orator Hoffman, 1974, ♂ holotype, modified from Hoffman 1974 A midbody tergite, dorsal view B gonopod, posterior view C acropodite, mesal view D gonopod, lateral view. Abbreviations: lb = lateral branch; mb = mesal branch; pfe = postfemorite; sl = solenomere branch. Not to scale." figureDoi="10.3897/zookeys.1223.139346.figure10" httpUri="https://binary.pensoft.net/fig/1215834">Fig. 10</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="BA69BD3DD9888E82A42AD3CF0B3739B2" type="reference_group">
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<paragraph id="2502A6F554B048B68B19BDDCF3B8650D">
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<treatmentCitationGroup id="095EE4E8C635AE9849375C64A4AF0906">
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<treatmentCitation id="7BB926C5048640822DB8A6030BA5D0D7" author="Hoffman RL" page="225" year="1974">
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<taxonomicName id="E4363CACF47725FCED213C7371502D3B" ID-CoL="6CBHG" authority="Hoffman, 1974: 225" authorityName="Hoffman" authorityPageNumber="225" authorityYear="1974" class="Diplopoda" family="Dalodesmidae" genus="Dalodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="orator">
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<emphasis id="EEBFE0B0CF6678BE113904680CDE0106" italics="true">Dalodesmus orator</emphasis>
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<comment id="A43E87F7456D4948B4587ECD51F1F8EE">
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<bibRefCitation id="73FE7A99E67A4A8D46BF5F04A5E70DE3" author="Hoffman RL" issue="2" journalOrPublisher="Wasmann Journal of Biology" pagination="221-246" refId="B14" refString="Hoffman RL (1974) Short studies on dalodesmid millipeds from South Africa and Madagascar. Wasmann Journal of Biology 32 (2): 221 - 246." title="Short studies on dalodesmid millipeds from South Africa and Madagascar." volume="32" year="1974">Hoffman, 1974: 225</bibRefCitation>
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, figs 1–9 (D);
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<treatmentCitation id="9D78DA5C80BC26B84FCE9FE46F502EF1" author="Golovatch SI & Hoffman RL" page="162" year="1989">
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<bibRefCitation id="BC2F7E499CAC9B8F9A2BCB4B758C757A" DOI="10.1080/03946975.1989.10539436" author="Golovatch SI & Hoffman RL" firstAuthor="Golovatch" journalOrPublisher="Tropical Zoology" page="162" pagination="159-164" refId="B12" refString="Golovatch SI, Hoffman RL (1989) Identity of Polydesmus hamatus Brandt 1841, a Malagasy milliped (Diplopoda Polydesmida Dalodesmidae). Tropical Zoology 2: 159–164. https://doi.org/10.1080/03946975.1989.10539436" title="Identity of Polydesmus hamatus Brandt 1841, a Malagasy milliped (Diplopoda Polydesmida Dalodesmidae)." volume="2" year="1989">Golovatch and Hoffman 1989: 162</bibRefCitation>
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</treatmentCitation>
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(L);
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<treatmentCitation id="4FDCC0C6654FB7BDABAE30DFF09EC1C5" author="Enghoff H" page="623" year="2003">
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<bibRefCitation id="AA5669181D86B8B3516C8E9822922C21" author="Enghoff H" editor="Goodman SM & Benstead JP" firstAuthor="Enghoff" journalOrPublisher="University of Chicago Press, Chicago and London" page="623" pagination="617-627" refId="B10" refString="Enghoff H (2003) Diplopoda, millipedes. In: Goodman SM, Benstead JP (Eds) The Natural History of Madagascar. University of Chicago Press, Chicago and London, 617–627." title="Diplopoda, millipedes." volumeTitle="The Natural History of Madagascar." year="2003">Enghoff 2003: 623</bibRefCitation>
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</treatmentCitation>
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(L);
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<treatmentCitation id="468FAA7B5BD987CF043E10CF73B095A5" author="Wesener T & Enghoff H" page="926" year="2022">
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<bibRefCitation id="698D4E2379342A57CAB3A6D8397B87EE" DOI="10.2307/j.ctv2ks6tbb.118" author="Wesener T & Enghoff H" editor="Fisher BL & Goodman SM" firstAuthor="Wesener" journalOrPublisher="Princeton University Press USA" page="926" pagination="918-933" refId="B22" refString="Wesener T, Enghoff H (2022) Diplopoda, millipedes, Menavetraka, Ankodabitra, Ankodiavitry, Marotanana, Sakolavitsy. In: Fisher BL, Goodman SM (Eds) The New Natural History of Madagascar. Princeton University Press USA, 918–933. https://doi.org/10.2307/j.ctv2ks6tbb.118" title="Diplopoda, millipedes, Menavetraka, Ankodabitra, Ankodiavitry, Marotanana, Sakolavitsy." volumeTitle="The New Natural History of Madagascar." year="2022">Wesener and Enghoff 2022: 926</bibRefCitation>
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</treatmentCitation>
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(L).
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</comment>
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</taxonomicName>
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</treatmentCitation>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="SECID0E6JBG" type="remarks">
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<paragraph id="7C7AC45355A976E923A065B8757DCA22">
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<heading id="71F57D70EDE5ABA64C22830F28CD1B30" reason="title">Remark.</heading>
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</paragraph>
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<paragraph id="7E01EDB9F84267BF21A16E17914FF2EC">
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This species was described from a
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<specimenCount id="48359AAFB2E59D6631CDE5D33BDF1F8B" count="1" type="male">♂</specimenCount>
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<typeStatus id="FEC8075E0332FC8785D9997E73F1C741" type="holotype">holotype</typeStatus>
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,
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<specimenCount id="FE8207B7DA117D25A730E7D5D1DCD099" count="2" type="male">two ♂</specimenCount>
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and
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<specimenCount id="857D8F4A0E39CEE7CDA9A317A8F1F93B" count="2" type="female">two ♀</specimenCount>
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<typeStatus id="BB4FAF3C3F7001F3672CC01523AD5A8B" type="paratype">paratypes</typeStatus>
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, all coming from Ambohimitombo, central
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<collectingCountry id="4EEC452C7C33FA09574D9CA574CF20CF" name="Madagascar">Madagascar</collectingCountry>
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(
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<bibRefCitation id="8CE9023579FD541B1C1A9776FC74662C" author="Hoffman" firstAuthor="Hoffman" issue="2" journalOrPublisher="Wasmann Journal of Biology" pagination="221-246" refId="B14" refString="Hoffman RL (1974) Short studies on dalodesmid millipeds from South Africa and Madagascar. Wasmann Journal of Biology 32 (2): 221–246." title="Short studies on dalodesmid millipeds from South Africa and Madagascar." volume="32" year="1974">Hoffman 1974</bibRefCitation>
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), in the collection of the Natural History Museum in London,
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<collectingCountry id="DD021A4097A19D98CFDF378B14AE0B41" name="United Kingdom">United Kingdom</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="SECID0EJKBG" type="description">
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<paragraph id="C2B05E31768C144FBF9BCA069277A106">
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<heading id="CBA8809040493DBAE32C754905DC76C4" reason="title">Brief description.</heading>
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</paragraph>
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<paragraph id="4D2AD22B3236127F25C67328975220FC">
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(After
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<bibRefCitation id="8C979D5A6A7D0DFDF263B59A9FF1237D" author="Hoffman" firstAuthor="Hoffman" issue="2" journalOrPublisher="Wasmann Journal of Biology" pagination="221-246" refId="B14" refString="Hoffman RL (1974) Short studies on dalodesmid millipeds from South Africa and Madagascar. Wasmann Journal of Biology 32 (2): 221–246." title="Short studies on dalodesmid millipeds from South Africa and Madagascar." volume="32" year="1974">Hoffman 1974</bibRefCitation>
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.)
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<specimenCount id="D1E4D64E0D6E378A055EF6BDB16E8CBC" count="1" type="male">♂</specimenCount>
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<typeStatus id="67281D0DC21DE807D963AF3E5D041BD1" type="holotype">holotype</typeStatus>
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~
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<quantity id="E2C38F45CD27502B914922C9250F60BA" metricMagnitude="-2" metricUnit="m" metricValue="2.65" unit="mm" value="26.5">26.5 mm</quantity>
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long and up to 2.3 and
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<quantity id="F73085FE3B92048E8BC5DE383EC8676F" metricMagnitude="-3" metricUnit="m" metricValue="5.3" unit="mm" value="5.3">5.3 mm</quantity>
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wide on pro- and metazona, respectively;
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<specimenCount id="C4BD887FD9DEA471AC0D6BEC402558B0" count="1" type="female">♀</specimenCount>
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up to
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<quantity id="28F90729D5E2834BF31D65EA68D0D6CF" metricMagnitude="-3" metricUnit="m" metricValue="4.5" unit="mm" value="4.5">4.5 mm</quantity>
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wide. Colouration light reddish brown with yellow paratergal apices and legs. Paraterga largely clearly upturned above dorsum and directed more laterad than caudad, at ca 45 °, tips elongate and sharpened caudally; dorsal surface between paraterga mostly vaguely areate (Fig.
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<figureCitation id="F583842CF0B20F406F9AC0986AD7CB17" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Dalodesmus orator Hoffman, 1974, ♂ holotype, modified from Hoffman 1974 A midbody tergite, dorsal view B gonopod, posterior view C acropodite, mesal view D gonopod, lateral view. Abbreviations: lb = lateral branch; mb = mesal branch; pfe = postfemorite; sl = solenomere branch. Not to scale." figureDoi="10.3897/zookeys.1223.139346.figure10" httpUri="https://binary.pensoft.net/fig/1215834">10 A</figureCitation>
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).
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</paragraph>
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<caption id="77D7669980EC3076C8AC7277303AC770" ID-DOI="10.3897/zookeys.1223.139346.figure10" ID-arpha="0CA33CAD-EA2A-505C-AD59-8F88DC9223D9" httpUri="https://binary.pensoft.net/fig/1215834" startId="F10">
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<paragraph id="3FE38AE627F4E2713A8DD80394C6A5F3">
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<label id="2D929BBA32958BFEAB3EC7B1DDC28780">Figure 10.</label>
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</paragraph>
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<paragraph id="20D3B3F123141CF743146521F82BA1C1">
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<taxonomicName id="DC00CD26CDC88AF9026BF126AB6EFA40" authority="Hoffman, 1974" authorityName="Hoffman" authorityYear="1974" class="Diplopoda" family="Dalodesmidae" genus="Dalodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="orator">
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<emphasis id="031F71696BF1B3A826ADB2172A340268" italics="true">Dalodesmus orator</emphasis>
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<bibRefCitation id="7B408535BF6F27EB264B9D26DBA609B5" author="Hoffman RL" issue="2" journalOrPublisher="Wasmann Journal of Biology" pagination="221-246" refId="B14" refString="Hoffman RL (1974) Short studies on dalodesmid millipeds from South Africa and Madagascar. Wasmann Journal of Biology 32 (2): 221 - 246." title="Short studies on dalodesmid millipeds from South Africa and Madagascar." volume="32" year="1974">Hoffman, 1974</bibRefCitation>
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||
</taxonomicName>
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||
, ♂ holotype, modified from
|
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<bibRefCitation id="D4E301BD566B8104450FBAF57D7DDD95" author="Hoffman" firstAuthor="Hoffman" issue="2" journalOrPublisher="Wasmann Journal of Biology" pagination="221-246" refId="B14" refString="Hoffman RL (1974) Short studies on dalodesmid millipeds from South Africa and Madagascar. Wasmann Journal of Biology 32 (2): 221–246." title="Short studies on dalodesmid millipeds from South Africa and Madagascar." volume="32" year="1974">Hoffman 1974</bibRefCitation>
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||
<emphasis id="A4B2BDB78970DFE998FC338A3B84454A" bold="true">A</emphasis>
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midbody tergite, dorsal view
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<emphasis id="3D28A04A1800A52A0EEA5C68298D13C5" bold="true">B</emphasis>
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gonopod, posterior view
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<emphasis id="59CB1A3000059FB89C6D2E8D64BE420C" bold="true">C</emphasis>
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acropodite, mesal view
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<emphasis id="E010DEBCFF5AC87AA02C609279F6B5B8" bold="true">D</emphasis>
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gonopod, lateral view. Abbreviations: lb = lateral branch; mb = mesal branch; pfe = postfemorite; sl = solenomere branch. Not to scale.
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</paragraph>
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</caption>
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<paragraph id="E41ED996890EBBE74F27340472B37209">
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Gonopods (Fig.
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<figureCitation id="D719800215EDB7F2FE6F361546971477" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Dalodesmus orator Hoffman, 1974, ♂ holotype, modified from Hoffman 1974 A midbody tergite, dorsal view B gonopod, posterior view C acropodite, mesal view D gonopod, lateral view. Abbreviations: lb = lateral branch; mb = mesal branch; pfe = postfemorite; sl = solenomere branch. Not to scale." figureDoi="10.3897/zookeys.1223.139346.figure10" httpUri="https://binary.pensoft.net/fig/1215834">10 B – D</figureCitation>
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) showing bare femorites (fe), coupled with rather complex acropodites: an unusually short and small solenomere (sl) directed ventrad, lying between the highest, suberect, distally unequally bifid mesal branch (mb) and a much shorter and strongly folded lateral branch (lb) directed ventrad.
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</paragraph>
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</subSubSection>
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</treatment>
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||
</document> |