180 lines
20 KiB
XML
180 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.308.5559" ID-Pensoft-Pub="1313-2970-308-1" ID-Pensoft-UUID="FFA2FFFEFFE74632FFB7FFC7D553CE26" ID-Zenodo-Dep="577662" ID-ZooBank="EEC31850AAAB4081B05AB80A2D944658" ModsDocID="1313-2970-308-1" checkinTime="1451247225725" checkinUser="pensoft" docAuthor="Rodrigues, Taissa & Kellner, Alexander Wilhelm Armin" docDate="2013" docId="4F5C803DA15C35276FFDD064D375CB70" docLanguage="en" docName="ZooKeys 308: 1-112" docOrigin="ZooKeys 308" docPubDate="2013-06-12" docSource="http://dx.doi.org/10.3897/zookeys.308.5559" docTitle="Ornithocheirus dentatus Seeley 1870, (nomen dubium)" docType="treatment" docVersion="5" id="FFA2FFFEFFE74632FFB7FFC7D553CE26" lastPageNumber="44" masterDocId="FFA2FFFEFFE74632FFB7FFC7D553CE26" masterDocTitle="Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England" masterLastPageNumber="112" masterPageNumber="1" pageNumber="43" updateTime="1634706563434" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rodrigues, Taissa</mods:namePart>
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<mods:affiliation>Department of Biology, Agrarian Sciences Center, Universidade Federal do Espirito Santo. Alto Universitario s / n, Caixa Postal 16, Guararema, CEP 29500 - 000, Alegre, ES, Brazil</mods:affiliation>
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<mods:nameIdentifier type="email">taissa.rodrigues@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kellner, Alexander Wilhelm Armin</mods:namePart>
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<mods:affiliation>Laboratory of Systematics and Taphonomy of Fossil Vertebrates, Department of Geology and Paleontology, Museu Nacional / Universidade Federal do Rio de Janeiro. Quinta da Boa Vista s / n, Sao Cristovao, CEP 20940 - 040, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2013-06-12</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>308</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>112</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.308.5559</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.308.5559</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-308-1</mods:identifier>
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<mods:identifier type="ZooBank">EEC31850AAAB4081B05AB80A2D944658</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFA2FFFEFFE74632FFB7FFC7D553CE26</mods:identifier>
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<mods:identifier type="Zenodo-Dep">577662</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152045000" LSID="urn:lsid:plazi:treatment:4F5C803DA15C35276FFDD064D375CB70" httpUri="http://treatment.plazi.org/id/4F5C803DA15C35276FFDD064D375CB70" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
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<subSubSection pageId="42" pageNumber="43" type="nomenclature">
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<paragraph pageId="42" pageNumber="43">
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<taxonomicName LSID="4F5C803D-A15C-3527-6FFD-D064D375CB70" authority="Seeley, 1870" authorityName="Seeley" authorityYear="1870" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Ornithocheirus dentatus" order="Pterosauria" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="dentatus" status="(nomen dubium)">Ornithocheirus dentatus Seeley, 1870</taxonomicName>
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<taxonomicNameLabel pageId="42" pageNumber="43">(nomen dubium)</taxonomicNameLabel>
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<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. A-D Ornithocheirus crassidens, holotype CAMSM B 54499 (Albian, Cambridge Greensand), anterior fragment of the rostrum. A right lateral view B respective line drawing C ventral view D respective line drawing. E-H Ornithocheirus dentatus, holotype CAMSM B 54544 (Albian, Cambridge Greensand), anterior part of the rostrum E right lateral view F respective line drawing G ventral view H respective line drawing. Abbreviations :: m - maxillae, pm - premaxillae, prid - palatal ridge. Arrows and numbers indicate alveoli or teeth and their respective position. Scale bar = 10 mm." httpUri="https://binary.pensoft.net/fig/22085" pageId="42" pageNumber="43">Fig. 17E-H</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="42" pageNumber="43" type="reference_group">
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<paragraph pageId="42" pageNumber="43">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Ornithocheiridae" genus="Ptenodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="dentatus" status="(nomen dubium)">
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<normalizedToken originalValue="“Ptenodactylus”">"Ptenodactylus"</normalizedToken>
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dentatus
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</taxonomicName>
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Seeley:
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B77" refString="Seeley, HG, 1869. Index to the fossil remains of Aves, Ornithosauria, and Reptilia, from the Secondary System of Strata arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell and Co., Cambridge, xxiii + 143 pp." title="Index to the fossil remains of Aves, Ornithosauria, and Reptilia, from the Secondary System of Strata arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell and Co., Cambridge, xxiii + 143 pp." year="1869">Seeley 1869</bibRefCitation>
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: p. xvi [disclaimed]
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</paragraph>
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<paragraph pageId="42" pageNumber="43">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="dentatus" status="(nomen dubium)">Ornithocheirus dentatus</taxonomicName>
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Seeley:
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley 1870</bibRefCitation>
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: p. 119
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</paragraph>
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<paragraph pageId="42" pageNumber="43">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="dentatus" status="(nomen dubium)">Ornithocheirus dentatus</taxonomicName>
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Seeley:
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<bibRefCitation author="Hooley, RW" journalOrPublisher="Annals and Magazine of Natural History" pageId="66" pageNumber="67" pagination="529 - 557" publicationUrl="10.1080/00222931408693521" refId="B26" refString="Hooley, RW, 1914. On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge. Annals and Magazine of Natural History 78: 529 - 557, 10.1080/00222931408693521" title="On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge." url="10.1080/00222931408693521" volume="78" year="1914">Hooley 1914</bibRefCitation>
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: p. 535
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</paragraph>
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<paragraph pageId="42" pageNumber="43">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Anhanguera" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="cuvieri" status="(nomen dubium)">Anhanguera cuvieri</taxonomicName>
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(Bowerbank):
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<bibRefCitation author="Unwin, DM" journalOrPublisher="Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe" pageId="69" pageNumber="70" pagination="189 - 221" publicationUrl="10.1002/mmng.20010040112" refId="B85" refString="Unwin, DM, 2001. An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England. Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe 4: 189 - 221, 10.1002/mmng.20010040112" title="An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England." url="10.1002/mmng.20010040112" volume="4" year="2001">Unwin 2001</bibRefCitation>
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: table 1
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</paragraph>
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</subSubSection>
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<subSubSection pageId="42" pageNumber="43" type="holotype">
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<paragraph pageId="42" pageNumber="43">Holotype.</paragraph>
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<paragraph pageId="42" pageNumber="43">
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CAMSM B 54544, anterior portion of the rostrum (
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<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. A-D Ornithocheirus crassidens, holotype CAMSM B 54499 (Albian, Cambridge Greensand), anterior fragment of the rostrum. A right lateral view B respective line drawing C ventral view D respective line drawing. E-H Ornithocheirus dentatus, holotype CAMSM B 54544 (Albian, Cambridge Greensand), anterior part of the rostrum E right lateral view F respective line drawing G ventral view H respective line drawing. Abbreviations :: m - maxillae, pm - premaxillae, prid - palatal ridge. Arrows and numbers indicate alveoli or teeth and their respective position. Scale bar = 10 mm." httpUri="https://binary.pensoft.net/fig/22085" pageId="42" pageNumber="43">Fig. 17E-H</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="43" pageNumber="44" type="type locality">
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<paragraph pageId="43" pageNumber="44">
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<pageBreakToken pageId="43" pageNumber="44" start="start">Type</pageBreakToken>
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locality.
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</paragraph>
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<paragraph pageId="43" pageNumber="44">Cambridge, Cambridgeshire, England.</paragraph>
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</subSubSection>
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<subSubSection pageId="43" pageNumber="44" type="type horizon">
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<paragraph pageId="43" pageNumber="44">Type horizon.</paragraph>
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<paragraph pageId="43" pageNumber="44">Cambridge Greensand (Cenomanian; fossils Albian in age).</paragraph>
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</subSubSection>
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<subSubSection pageId="43" pageNumber="44" type="description">
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<paragraph pageId="43" pageNumber="44">Description.</paragraph>
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<paragraph pageId="43" pageNumber="44">
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley (1870)</bibRefCitation>
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described
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="dentatus">
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<emphasis italics="true" pageId="43" pageNumber="44">Ornithocheirus dentatus</emphasis>
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</taxonomicName>
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based on a fragmentary anterior portion of an upper jaw. Interestingly, CAMSM B 54544 lacks a fully developed alveolus on the right side (possibly the third), but differing from
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Owen" baseAuthorityYear="1859" class="Reptilia" genus="Camposipterus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="sedgwickii">
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<emphasis italics="true" pageId="43" pageNumber="44">Camposipterus(?) sedgwickii</emphasis>
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</taxonomicName>
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and
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<taxonomicName class="Reptilia" family="Anhangueridae" genus="Anhanguera" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="blittersdorffi">
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<emphasis italics="true" pageId="43" pageNumber="44">Anhanguera blittersdorffi</emphasis>
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</taxonomicName>
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(Pz-DBAV UERJ 40) because its margins are slightly marked. In the other specimens, the margins are completely absent.
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</paragraph>
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<paragraph pageId="43" pageNumber="44">
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley (1870)</bibRefCitation>
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compared and distinguished
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="dentatus">
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<emphasis italics="true" pageId="43" pageNumber="44">Ornithocheirus dentatus</emphasis>
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</taxonomicName>
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from
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<taxonomicName authorityName="Seeley" authorityYear="1869" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="43" pageNumber="44" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="43" pageNumber="44">Ornithocheirus</emphasis>
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</taxonomicName>
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[=
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<taxonomicName authority="(?)" class="Reptilia" genus="Camposipterus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="43" pageNumber="44" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="43" pageNumber="44">Camposipterus(?)</emphasis>
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</taxonomicName>
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]
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<emphasis italics="true" pageId="43" pageNumber="44">sedgwickii</emphasis>
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based on a wider palate and smaller teeth. It is hard to understand what he meant by wider palate, since CAMSM B 54544 is quite smaller individual in comparison to the holotype of
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Owen" baseAuthorityYear="1859" class="Reptilia" genus="Camposipterus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="sedgwickii">
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<emphasis italics="true" pageId="43" pageNumber="44">Camposipterus(?) sedgwickii</emphasis>
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</taxonomicName>
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, but both species can be further distinguished by the lack of an anterior expansion and of a dorsally curved palate in
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="dentatus">
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<emphasis italics="true" pageId="43" pageNumber="44">Ornithocheirus dentatus</emphasis>
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</taxonomicName>
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, besides a lower rostrum in the latter.
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</paragraph>
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<paragraph pageId="43" pageNumber="44">
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<bibRefCitation author="Unwin, DM" journalOrPublisher="Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe" pageId="69" pageNumber="70" pagination="189 - 221" publicationUrl="10.1002/mmng.20010040112" refId="B85" refString="Unwin, DM, 2001. An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England. Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe 4: 189 - 221, 10.1002/mmng.20010040112" title="An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England." url="10.1002/mmng.20010040112" volume="4" year="2001">Unwin (2001)</bibRefCitation>
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synonymized
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="dentatus">
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<emphasis italics="true" pageId="43" pageNumber="44">Ornithocheirus dentatus</emphasis>
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</taxonomicName>
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with
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<taxonomicName class="Reptilia" family="Anhangueridae" genus="Anhanguera" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="43" pageNumber="44" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="43" pageNumber="44">Anhanguera</emphasis>
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</taxonomicName>
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[=
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" genus="Cimoliopterus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="43" pageNumber="44" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="43" pageNumber="44">Cimoliopterus</emphasis>
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</taxonomicName>
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]
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<emphasis italics="true" pageId="43" pageNumber="44">cuvieri</emphasis>
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. However,
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley (1870)</bibRefCitation>
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had already noticed that both species can be differentiated by the presence of smaller alveoli, which are placed more closely together. It can also be excluded from the
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Lonchodraconidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="43" pageNumber="44" phylum="Chordata" rank="family">Lonchodraconidae</taxonomicName>
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as it does not possess a deep palatal ridge or alveoli placed on elevations. Due to fragmentary state of CAMSM B 54544, we regard
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="dentatus">
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<emphasis italics="true" pageId="43" pageNumber="44">Ornithocheirus dentatus</emphasis>
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</taxonomicName>
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as a
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<emphasis italics="true" pageId="43" pageNumber="44">nomen dubium</emphasis>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |