155 lines
18 KiB
XML
155 lines
18 KiB
XML
<document id="3A8C64A86E3DF1316915C2C64FBD9BAF" ID-DOI="10.11646/zootaxa.4039.3.3" ID-GBIF-Dataset="0ad500ca-d193-4331-90af-70a6018a5cfc" ID-ISSN="1175-5326" ID-Zenodo-Dep="238528" ID-ZooBank="87A108DC-6967-4CAD-8F6C-DAF7ADD622CB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461331934834" checkinUser="plazi" docAuthor="Irmler, Ulrich" docDate="2015" docId="07009178FF8ABB20FF4886413A052BED" docLanguage="en" docName="zt04039p444.pdf" docOrigin="Zootaxa 4039 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thoracophorus zicsii Irmler, 2015, n. sp." docType="treatment" docVersion="7" lastPageNumber="436" masterDocId="FB39E900FF8FBB25FFDF840938402C7E" masterDocTitle="Four new species of the genus Thoracophorus Motschulsky, 1857 for the Neotropical region (Coleoptera: Staphylinidae: Osoriinae)" masterLastPageNumber="444" masterPageNumber="431" pageNumber="436" updateTime="1698618579127" updateUser="plazi">
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<mods:title id="63A2E90A7040B3275DF484136EC730C3">Four new species of the genus Thoracophorus Motschulsky, 1857 for the Neotropical region (Coleoptera: Staphylinidae: Osoriinae)</mods:title>
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<mods:namePart id="FC338D4DD3649D608FE6C6BCF2748A73">Irmler, Ulrich</mods:namePart>
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<mods:title id="BB933BF2435AFADCE1A2341C6C2F6F9B">Zootaxa</mods:title>
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<mods:part id="7ED6753DECF7EB3151CA294C880CC4BE">
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<mods:date id="F4A0C840BE035D12D4A2B5B6DF5749A3">2015</mods:date>
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<mods:number id="89440988755B5C408C6A422684CF78A1">4039</mods:number>
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<mods:identifier id="08433FFEC143477D1D533F821F6E758F" type="DOI">10.11646/zootaxa.4039.3.3</mods:identifier>
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<treatment id="07009178FF8ABB20FF4886413A052BED" ID-DOI="http://doi.org/10.5281/zenodo.6107932" ID-GBIF-Taxon="119644041" ID-Zenodo-Dep="6107932" LSID="urn:lsid:plazi:treatment:07009178FF8ABB20FF4886413A052BED" httpUri="http://treatment.plazi.org/id/07009178FF8ABB20FF4886413A052BED" lastPageNumber="436" pageId="5" pageNumber="436">
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<subSubSection id="C7B373E5FF8ABB20FF48864139082EFD" pageId="5" pageNumber="436" type="nomenclature">
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<paragraph id="8F16206EFF8ABB20FF488641399F2E1C" blockId="5.[151,479,584,643]" box="[151,479,584,610]" pageId="5" pageNumber="436">
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<heading id="D45E9702FF8ABB20FF488641399F2E1C" bold="true" box="[151,479,584,610]" fontSize="11" level="1" pageId="5" pageNumber="436" reason="1">
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<emphasis id="BDDDFC7CFF8ABB20FF488641399F2E1C" bold="true" box="[151,479,584,610]" pageId="5" pageNumber="436">
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<taxonomicName id="48A95BEDFF8ABB20FF48864139D52E1C" ID-CoL="7BYBL" box="[151,405,584,610]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="436" phylum="Arthropoda" rank="species" species="zicsii" status="sp. nov.">
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<emphasis id="BDDDFC7CFF8ABB20FF48864139D52E1C" bold="true" box="[151,405,584,610]" italics="true" pageId="5" pageNumber="436">Thoracophorus zicsii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A6EE4107FF8ABB20FE428641399F2E1C" box="[413,479,584,610]" pageId="5" pageNumber="436" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8F16206EFF8ABB20FF48866339082EFD" blockId="5.[151,479,584,643]" box="[151,328,618,643]" pageId="5" pageNumber="436">
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<figureCitation id="17923CEBFF8ABB20FF48866338A22EFD" box="[151,226,618,643]" captionStart="FIGURES 1 – 3" captionStartId="3.[151,264,1398,1421]" captionTargetBox="[156,1428,385,1371]" captionTargetId="figure@3.[151,1436,375,1377]" captionTargetPageId="3" captionText="FIGURES 1 – 3. Thoracophorus filum (1: from Belize, 1 ’: from Costa Rica); T. verhaaghi (2); T. zicsii (3); aedeagus in lateral aspect (A), antenna (B); scale bar: 0.1 mm." httpUri="https://zenodo.org/record/238529/files/figure.png" pageId="5" pageNumber="436">Figs. 3</figureCitation>
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A, B, 9D
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</paragraph>
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</subSubSection>
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<subSubSection id="C7B373E5FF8ABB20FF4886BB3B822E91" pageId="5" pageNumber="436" type="materials_examined">
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<paragraph id="8F16206EFF8ABB20FF4886BB3B822E91" blockId="5.[151,1436,690,1939]" pageId="5" pageNumber="436">
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<emphasis id="BDDDFC7CFF8ABB20FF4886BB39062EB5" bold="true" box="[151,326,690,715]" pageId="5" pageNumber="436">
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<typeStatus id="50129ECCFF8ABB20FF4886BB38922EB5" box="[151,210,690,715]" pageId="5" pageNumber="436">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="50129ECCFF8ABB20FE9186BA39A52EB5" box="[334,485,691,715]" pageId="5" pageNumber="436" type="holotype">HOLOTYPE</typeStatus>
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. male,
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<emphasis id="BDDDFC7CFF8ABB20FDE986BB3AE62EB5" bold="true" box="[566,678,690,715]" pageId="5" pageNumber="436">
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<collectingCountry id="F7BE60FEFF8ABB20FDE986BB3AE12EB5" box="[566,673,690,715]" name="Ecuador" pageId="5" pageNumber="436">Ecuador</collectingCountry>
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:
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</emphasis>
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Prov. Pichincha,
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<quantity id="48518D8BFF8ABB20FCB186BA3BF62EB5" box="[878,950,690,715]" metricMagnitude="4" metricUnit="m" metricValue="7.1" pageId="5" pageNumber="436" unit="km" value="71.0">71 km</quantity>
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from Quito to Santo
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<collectingCountry id="F7BE60FEFF8ABB20FB7486BB3D582EB5" box="[1195,1304,690,715]" name="Dominican Republic" pageId="5" pageNumber="436">Domingo</collectingCountry>
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, moss from lakeshore, near the fall,
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<date id="FB1706AEFF8ABB20FE7B86DE3A572E91" box="[420,535,727,751]" pageId="5" pageNumber="436" value="1988-04-24">24.4.1988</date>
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, leg. Cs. Csuzdi & A. Ziksi (
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<collectionCode id="E9B8B8ABFF8ABB20FC8586DE3BF62E91" box="[858,950,727,751]" pageId="5" pageNumber="436">NHMH</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C7B373E5FF8ABB20FF1886F33C2F2FDD" pageId="5" pageNumber="436" type="diagnosis">
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<paragraph id="8F16206EFF8ABB20FF1886F33C2F2FDD" blockId="5.[151,1436,690,1939]" pageId="5" pageNumber="436">
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<emphasis id="BDDDFC7CFF8ABB20FF1886F339012F6D" bold="true" box="[199,321,762,787]" pageId="5" pageNumber="436">Diagnosis.</emphasis>
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The species is intermediately sized compared to
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<taxonomicName id="48A95BEDFF8ABB20FCB386F53C112F6D" authority="Sharp, 1887" authorityName="Sharp" authorityYear="1887" box="[876,1105,762,787]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="436" phylum="Arthropoda" rank="species" species="filum">
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<emphasis id="BDDDFC7CFF8ABB20FCB386F53B822F6D" box="[876,962,762,787]" italics="true" pageId="5" pageNumber="436">T. filum</emphasis>
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<bibRefCitation id="EB385D9FFF8ABB20FC1686F33C112F6D" author="Sharp" box="[969,1105,762,787]" pageId="5" pageNumber="436" refString="Sharp, D. (1887) Fam. Staphylinidae. In: Sharp, D. (Ed.), Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2). Taylor & Francis, London, pp. 145 - 824. [plates. 5 - 19]" type="journal article" year="1887">Sharp, 1887</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="48A95BEDFF8ABB20FB5686F53D562F6D" box="[1161,1302,762,787]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="436" phylum="Arthropoda" rank="species" species="verhaaghi">
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<emphasis id="BDDDFC7CFF8ABB20FB5686F53D562F6D" box="[1161,1302,762,787]" italics="true" pageId="5" pageNumber="436">T. verhaaghi</emphasis>
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</taxonomicName>
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. In contrast to both species, colouration is light brown, but not mainly orange as in the last two species. It mostly resembles
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<taxonomicName id="48A95BEDFF8ABB20FA58871639312F25" authority="Sharp, 1887" authorityName="Sharp" authorityYear="1887" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="436" phylum="Arthropoda" rank="species" species="filum">
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<emphasis id="BDDDFC7CFF8ABB20FA58871638912F25" italics="true" pageId="5" pageNumber="436">T. filum</emphasis>
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<bibRefCitation id="EB385D9FFF8ABB20FF3F874B39312F25" author="Sharp" box="[224,369,834,859]" pageId="5" pageNumber="436" refString="Sharp, D. (1887) Fam. Staphylinidae. In: Sharp, D. (Ed.), Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2). Taylor & Francis, London, pp. 145 - 824. [plates. 5 - 19]" type="journal article" year="1887">
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Sharp,
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||
<quantity id="48518D8BFF8ABB20FEE6874A39312F25" box="[313,369,835,859]" metricMagnitude="1" metricUnit="m" metricValue="4.79298" pageId="5" pageNumber="436" unit="in" value="1887.0">1887</quantity>
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</bibRefCitation>
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||
</taxonomicName>
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in its slender shape, but the elytra are distinctly longer than wide, whereas they are approximately quadrate in
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<taxonomicName id="48A95BEDFF8ABB20FE19876E3AEF2F01" authority="Sharp, 1887" authorityName="Sharp" authorityYear="1887" box="[454,687,870,895]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="436" phylum="Arthropoda" rank="species" species="filum">
|
||
<emphasis id="BDDDFC7CFF8ABB20FE19876E3A5C2F01" box="[454,540,870,895]" italics="true" pageId="5" pageNumber="436">T. filum</emphasis>
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||
<bibRefCitation id="EB385D9FFF8ABB20FDFB876F3AEF2F01" author="Sharp" box="[548,687,870,895]" pageId="5" pageNumber="436" refString="Sharp, D. (1887) Fam. Staphylinidae. In: Sharp, D. (Ed.), Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2). Taylor & Francis, London, pp. 145 - 824. [plates. 5 - 19]" type="journal article" year="1887">Sharp, 1887</bibRefCitation>
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||
</taxonomicName>
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. The aedeagus also resembles that of
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<taxonomicName id="48A95BEDFF8ABB20FB85876E3D022F01" authority="Sharp, 1887" authorityName="Sharp" authorityYear="1887" box="[1114,1346,870,895]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="436" phylum="Arthropoda" rank="species" species="filum">
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<emphasis id="BDDDFC7CFF8ABB20FB85876E3CF02F01" box="[1114,1200,870,895]" italics="true" pageId="5" pageNumber="436">T. filum</emphasis>
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||
<bibRefCitation id="EB385D9FFF8ABB20FB68876F3D022F01" author="Sharp" box="[1207,1346,870,895]" pageId="5" pageNumber="436" refString="Sharp, D. (1887) Fam. Staphylinidae. In: Sharp, D. (Ed.), Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2). Taylor & Francis, London, pp. 145 - 824. [plates. 5 - 19]" type="journal article" year="1887">Sharp, 1887</bibRefCitation>
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</taxonomicName>
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, but the base of the endophallus is more slender and not circular such as in
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<taxonomicName id="48A95BEDFF8ABB20FC5B87853C2A2FDD" authority="Sharp, 1887" authorityName="Sharp" authorityYear="1887" box="[900,1130,906,931]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="436" phylum="Arthropoda" rank="species" species="filum">
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<emphasis id="BDDDFC7CFF8ABB20FC5B87853B992FDD" box="[900,985,906,931]" italics="true" pageId="5" pageNumber="436">T. filum</emphasis>
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<bibRefCitation id="EB385D9FFF8ABB20FC3F87833C2A2FDD" author="Sharp" box="[992,1130,906,931]" pageId="5" pageNumber="436" refString="Sharp, D. (1887) Fam. Staphylinidae. In: Sharp, D. (Ed.), Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2). Taylor & Francis, London, pp. 145 - 824. [plates. 5 - 19]" type="journal article" year="1887">Sharp, 1887</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C7B373E5FF8ABB20FF1887A73A8C2B35" pageId="5" pageNumber="436" type="description">
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<paragraph id="8F16206EFF8ABB20FF1887A73A672F95" blockId="5.[151,1436,690,1939]" pageId="5" pageNumber="436">
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<emphasis id="BDDDFC7CFF8ABB20FF1887A739192FB9" bold="true" box="[199,345,942,967]" pageId="5" pageNumber="436">Description.</emphasis>
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Length:
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||
<quantity id="48518D8BFF8ABB20FE1D87A73A5D2FB8" box="[450,541,942,967]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="436" unit="mm" value="1.6">1.6 mm</quantity>
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. Colouration: totally light brown, head and abdomen slightly darker; legs and antennae slightly lighter, yellowish.
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</paragraph>
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<paragraph id="8F16206EFF8ABB20FF1887FF3C052804" blockId="5.[151,1436,690,1939]" pageId="5" pageNumber="436">
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||
Head:
|
||
<quantity id="48518D8BFF8ABB20FECB87FE393C2870" box="[276,380,1015,1039]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="436" unit="mm" value="0.2">0.20 mm</quantity>
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||
long,
|
||
<quantity id="48518D8BFF8ABB20FE1587FF3A712870" box="[458,561,1014,1039]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="436" unit="mm" value="0.26">0.26 mm</quantity>
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||
wide; eyes large and prominent; twice as long as temples; temples shortly rounded to neck; neck distinctly separated from vertex; sides of fore-head margined; beginning at middle of eyes and continued to front margin of clypeus; space between margin slightly depressed; pair of central swelling indistinct, flat; without punctation; isodiametric microsculpture deep; surface matte.
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||
</paragraph>
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<paragraph id="8F16206EFF8ABB20FF18808E3A6B292D" blockId="5.[151,1436,690,1939]" pageId="5" pageNumber="436">Antenna slightly shorter than head and pronotum combined; first and second antennomere thicker than following four antennomeres and nearly equal in length; second antennomere oval; third conical antennomere approximately half as long as second; antennomeres four to six slightly increasing in width; each more or less quadrate; antennomeres seven to eleven distinctly wider than preceding antennomeres and increasing in width; seventh antennomere one forth wider than sixth; tenth antennomere nearly twice as wide as sixth; fivth to eleventh antennomere indistinctly pubescent.</paragraph>
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<paragraph id="8F16206EFF8ABB20FF188156389D2A78" blockId="5.[151,1436,690,1939]" pageId="5" pageNumber="436">
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Pronotum:
|
||
<quantity id="48518D8BFF8ABB20FE9B815739E72908" box="[324,423,1374,1399]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="436" unit="mm" value="0.26">0.26 mm</quantity>
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||
long,
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||
<quantity id="48518D8BFF8ABB20FE3281563A102908" box="[493,592,1375,1399]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="5" pageNumber="436" unit="mm" value="0.29">0.29 mm</quantity>
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||
wide; widest close to middle; slightly narrowed to anterior margin; anterior margin convex; anterior angles obtuse; sides in posterior half concave; deeply emarginate in front of rectangular posterior angles; lateral margin very indistinctly undulate; nearly smooth; central dorsal depression and swellings indistinct, flat; without punctation; isodiametric microsculpture deep; partly, meshes slightly elongate; surface matte.
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||
</paragraph>
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||
<paragraph id="8F16206EFF8ABB20FF18821B39E22AE8" blockId="5.[151,1436,690,1939]" pageId="5" pageNumber="436">
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||
Elytra:
|
||
<quantity id="48518D8BFF8ABB20FEC4821A39C02A54" box="[283,384,1555,1579]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="5" pageNumber="436" unit="mm" value="0.44">0.44 mm</quantity>
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||
long,
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||
<quantity id="48518D8BFF8ABB20FE17821B3A6D2A54" box="[456,557,1554,1579]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="5" pageNumber="436" unit="mm" value="0.36">0.36 mm</quantity>
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||
wide; parallel; shoulders distinct; shortly rounded to straight anterior margin; sides in posterior forth narrowed to posterior margin; posterior margin with short scaly setae; carinae very weakly developed; nearly absent; suture line nearly absent, too; without punctation; microsculpture deep; meshes slightly elongate; surface matte.
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||
</paragraph>
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||
<paragraph id="8F16206EFF8ABB20FF1882AB3C502AA1" blockId="5.[151,1436,690,1939]" pageId="5" pageNumber="436">Abdomen with anterior segements depressed at base; microsculpture deep; meshes slightly elongate; slightly less matte than fore-body; each segment with few scaly setae in transverse line.</paragraph>
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<paragraph id="8F16206EFF8ABB20FF1882E33A8C2B35" blockId="5.[151,1436,690,1939]" pageId="5" pageNumber="436">Aedeagus smoothly rounded between basal and apical lobe; apex obtuse; widely rounded; endophallus slender at base; ending in long filiform ductus; paramera only very shortly exceeding central lobe; apex with broad transparent area; diagonal row of sensillae at base.</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C7B373E5FF8ABB20FF18835F3A052BED" pageId="5" pageNumber="436" type="etymology">
|
||
<paragraph id="8F16206EFF8ABB20FF18835F3A052BED" blockId="5.[151,1436,690,1939]" pageId="5" pageNumber="436">
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<emphasis id="BDDDFC7CFF8ABB20FF18835F390E2B11" bold="true" box="[199,334,1878,1903]" pageId="5" pageNumber="436">Etymology.</emphasis>
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The species name derived from one of the collectors, A. Zicsi, who collected soil organisms on several expeditions to the Neotropics..
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</paragraph>
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</subSubSection>
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||
</treatment>
|
||
</document> |