treatments-xml/data/03/CF/2F/03CF2F21C626A5287DA9FA83FD1FFD37.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.4759173" ID-GBIF-Dataset="11a4efe9-83ec-450e-9361-9657cfe342bd" ID-GBIF-Taxon="182068098" ID-ISSN="1854-0392" ID-Zenodo-Dep="4759173" checkinTime="1620934097176" checkinUser="felipe" docAuthor="Bottorff, Richard L." docDate="2007" docId="03CF2F21C626A5287DA9FA83FD1FFD37" docLanguage="en" docName="Illiesia.3.6.46-52.pdf" docOrigin="Illiesia 3 (6)" docStyle="DocumentStyle:1DEF1196F206C1C297CF0540A5C6768F.1:Illesia.2005-2008.journal_article" docStyleId="1DEF1196F206C1C297CF0540A5C6768F" docStyleName="Illesia.2005-2008.journal_article" docStyleVersion="1" docTitle="Cosumnoperla sequoia Bottorff 2007, sp. n." docType="treatment" docVersion="6" lastPageId="6" lastPageNumber="51" masterDocId="FFF65759C626A52E7E69FFB2FF91FFE1" masterDocTitle="Cosumnoperla Sequoia, A New Species Of Stonefly From The Sierra Nevada, California (Plecoptera: Perlodidae: Isoperlinae)" masterLastPageNumber="52" masterPageNumber="46" pageId="0" pageNumber="46" updateTime="1643579788470" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>Cosumnoperla Sequoia, A New Species Of Stonefly From The Sierra Nevada, California (Plecoptera: Perlodidae: Isoperlinae)</mods:title>
</mods:titleInfo>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bottorff, Richard L.</mods:namePart>
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<mods:title>Illiesia</mods:title>
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<mods:date>2007</mods:date>
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<mods:number>2007-06-26</mods:number>
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<mods:number>3</mods:number>
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<mods:number>6</mods:number>
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<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4758699</mods:identifier>
<mods:identifier type="GBIF-Dataset">11a4efe9-83ec-450e-9361-9657cfe342bd</mods:identifier>
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<taxonomicName authority="Bottorff, 2007" authorityName="Bottorff" authorityYear="2007" box="[960,1223,1329,1353]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="0" pageNumber="46" phylum="Arthropoda" rank="species" species="sequoia" status="sp. n.">Cosumnoperla sequoia</taxonomicName>
,
</emphasis>
<taxonomicNameLabel box="[1235,1297,1329,1353]" pageId="0" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
</subSubSection>
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<paragraph blockId="0.[1059,1183,1364,1388]" box="[1059,1183,1364,1388]" pageId="0" pageNumber="46">
(
<figureCitation box="[1067,1176,1364,1388]" captionStart-0="Figs" captionStart-1="Fig" captionStart-2="Figs" captionStartId-0="1.[184,234,1566,1590]" captionStartId-1="2.[184,223,1497,1521]" captionStartId-2="3.[184,234,1876,1900]" captionTargetBox-0="[234,1317,294,1480]" captionTargetBox-1="[184,809,801,1428]" captionTargetBox-2="[896,1274,1310,1808]" captionTargetId-1="figure-455@2.[184,809,798,1428]" captionTargetId-2="figure-103@3.[890,1280,1303,1818]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="Figs. 1‐5. Cosumnoperla sequoia adult male. 1. Head‐pronotum, dorsal; 2. Terminalia (segments 8‐10, abdomen extended), dorsal; 3. Terminalia (segments 8‐10, abdomen contracted), right lateral; 4. Variant of reflexed process, 10th tergum, dorsal; 5. Aedeagus, right lateral (arrow shows location of median indention); Scale lines = 0.5 mm." captionText-1="Fig. 6. Cosumnoperla sequoia adult female. 6a. Terminalia (segments 8‐10), ventral; 6b‐c. Variants of subgenital plate, ventral; Scale lines = 0.5 mm." captionText-2="Figs. 7‐12. Cosumnoperla sequoia larva. 7. Head‐pronotum, dorsal; 8. Thorax, ventral; 9. Right front leg, dorsal; 10. Abdomen (segments 1‐10), dorsal; 11. Male terminalia (segments 8‐10), ventral; 12. Female terminalia (segments 8‐10), ventral; Scale lines = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4758701" figureDoi-1="http://doi.org/10.5281/zenodo.4758705" figureDoi-2="http://doi.org/10.5281/zenodo.4758707" httpUri-0="https://zenodo.org/record/4758701/files/figure.png" httpUri-1="https://zenodo.org/record/4758705/files/figure.png" httpUri-2="https://zenodo.org/record/4758707/files/figure.png" pageId="0" pageNumber="46">Figs. 1‐12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="46" type="materials_examined">
<paragraph blockId="0.[812,1404,1433,1976]" pageId="0" pageNumber="46">
<materialsCitation ID-GBIF-Occurrence="3111448301" collectingDate="1990-06-05" collectingDateMax="1990-06-05" collectingDateMin="1963-05-31" collectionCode="R, USNM" collectorName="L. Bottorff" country="United States of America" county="Fresno County" elevation="1050" latitude="36.812866" location="Redwood Creek" longLatPrecision="1" longitude="-118.835365" municipality="Sequoia National Forest" pageId="0" pageNumber="46" specimenCount="1" specimenCount-female="6" specimenCount-male="1" stateProvince="California" typeStatus="holotype">
<emphasis bold="true" box="[812,1039,1433,1457]" pageId="0" pageNumber="46">Material examined.</emphasis>
<typeStatus box="[1044,1152,1433,1457]" pageId="0" pageNumber="46">Holotype</typeStatus>
<specimenCount box="[1157,1176,1433,1457]" pageId="0" pageNumber="46" type="male">♂</specimenCount>
, allotype ♀, and 4♂, 3♀, and 7 larval paratypes from the 
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, 
<collectingRegion box="[812,929,1503,1527]" country="United States of America" name="California" pageId="0" pageNumber="46">California</collectingRegion>
, 
<collectingCounty box="[947,1126,1503,1527]" pageId="0" pageNumber="46">Fresno County</collectingCounty>
, unnamed intermittent creek (Latitude 
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36 
<emphasis box="[1019,1029,1536,1561]" italics="true" pageId="0" pageNumber="46">°</emphasis>
48.772’ N
</geoCoordinate>
, Longitude 
<geoCoordinate degrees="118" direction="west" minutes="50.122" orientation="longitude" pageId="0" pageNumber="46" precision="1" value="-118.835365">
118 
<emphasis box="[1311,1321,1536,1561]" italics="true" pageId="0" pageNumber="46">°</emphasis>
50.122’ W
</geoCoordinate>
, Elevation 
<quantity box="[963,1044,1572,1596]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="0" pageNumber="46" unit="m" value="1050.0">
<elevation box="[963,1044,1572,1596]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="0" pageNumber="46" unit="m" value="1050.0">1050 m</elevation>
</quantity>
), west of 
<location LSID="urn:lsid:plazi:treatment:03CF2F21C626A5287DA9FA83FD1FFD37:8EB9C8ECC626A52E7A15F996FAA7F9DD" box="[1148,1334,1572,1596]" country="United States of America" county="Fresno County" latitude="36.812866" longLatPrecision="1" longitude="-118.835365" municipality="Sequoia National Forest" name="Redwood Creek" pageId="0" pageNumber="46" stateProvince="California">Redwood Creek</location>
, 
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<locationDeviation box="[1343,1403,1572,1596]" metricMagnitude="4" metricUnit="m" metricValue="1.609344" pageId="0" pageNumber="46" unit="mi" value="10.0">10 mi</locationDeviation>
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[
<quantity box="[819,899,1606,1630]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="0" pageNumber="46" unit="km" value="16.0">
<locationDeviation box="[819,899,1606,1630]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="0" pageNumber="46" unit="km" value="16.0">16 km</locationDeviation>
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] west of 
<location LSID="urn:lsid:plazi:treatment:03CF2F21C626A5287DA9FA83FD1FFD37:8EB9C8ECC626A52E7A69F9F4FB0CF9BE" box="[1024,1181,1606,1631]" country="United States of America" county="Fresno County" latitude="36.812866" longLatPrecision="1" longitude="-118.835365" municipality="Sequoia National Forest" name="Cedar Grove" pageId="0" pageNumber="46" stateProvince="California">Cedar Grove</location>
, 
<location LSID="urn:lsid:plazi:treatment:03CF2F21C626A5287DA9FA83FD1FFD37:8EB9C8ECC626A52E7AC6F9F4FCE4F961" country="United States of America" county="Fresno County" latitude="36.812866" longLatPrecision="1" longitude="-118.835365" municipality="Sequoia National Forest" name="Sequoia National Forest" pageId="0" pageNumber="46" stateProvince="California">Sequoia National Forest</location>
, 
<date box="[901,1044,1641,1665]" pageId="0" pageNumber="46" value="1990-06-05">
<collectingDate box="[901,1044,1641,1665]" pageId="0" pageNumber="46" value="1990-06-05">5 June 1990</collectingDate>
</date>
, 
<collectionCode box="[1060,1080,1641,1664]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="0" pageNumber="46">R</collectionCode>
. 
<collectorName box="[1096,1219,1641,1665]" pageId="0" pageNumber="46">L. Bottorff</collectorName>
(Holotype and allotype 
<collectionCode box="[910,992,1676,1700]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="0" pageNumber="46" type="Museum">USNM</collectionCode>
,
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3111448302" collectingDate="1963-05-31" collectingDateMax="1990-06-05" collectingDateMin="1963-05-31" collectionCode="USNM, RLB, R" collectorName="Hwy &amp; NE Wilsonia &amp; L. Bottorff &amp; Kings Canyon Road &amp; C. P. Alexander &amp; Kings Canyon, Rt. &amp; Horseshoe Bend" country="United States of America" county="Fresno County" elevation="1000" latitude="36.812866" location="Fresno County" longLatPrecision="1" longitude="-118.835365" municipality="Sequoia National Forest" pageId="0" pageNumber="46" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="California" typeStatus="paratype">
<typeStatus box="[1001,1114,1676,1700]" pageId="0" pageNumber="46">paratypes</typeStatus>
<collectionCode box="[1119,1201,1676,1700]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="0" pageNumber="46" type="Museum">USNM</collectionCode>
, 
<collectionCode box="[1210,1263,1676,1700]" collectionName="RLB" pageId="0" pageNumber="46">RLB</collectionCode>
). Additional 
<typeStatus box="[812,928,1710,1734]" pageId="0" pageNumber="46">paratypes</typeStatus>
: 
<collectingCountry box="[962,1018,1710,1734]" name="United States of America" pageId="0" pageNumber="46">USA</collectingCountry>
, 
<collectingRegion box="[1052,1169,1710,1734]" country="United States of America" name="California" pageId="0" pageNumber="46">California</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:03CF2F21C626A5287DA9FA83FD1FFD37:8EB9C8ECC626A52E7ADDF91CFAE9F927" box="[1204,1400,1710,1734]" country="United States of America" county="Fresno County" latitude="36.812866" longLatPrecision="1" longitude="-118.835365" municipality="Sequoia National Forest" name="Fresno County" pageId="0" pageNumber="46" stateProvince="California">Fresno County</location>
: 
<location LSID="urn:lsid:plazi:treatment:03CF2F21C626A5287DA9FA83FD1FFD37:8EB9C8ECC626A52E7D45F963FC63F908" box="[812,1010,1745,1769]" country="United States of America" county="Fresno County" latitude="36.812866" longLatPrecision="1" longitude="-118.835365" municipality="Sequoia National Forest" name="Lockwood Creek" pageId="0" pageNumber="46" stateProvince="California">Lockwood Creek</location>
at 
<collectorName box="[1047,1104,1745,1769]" pageId="0" pageNumber="46">Hwy</collectorName>
180 (Elevation 
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<elevation box="[1285,1369,1745,1769]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="0" pageNumber="46" unit="m" value="1000.0">1000 m</elevation>
</quantity>
), 
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.2874752" pageId="0" pageNumber="46" unit="mi" value="8.0">
<locationDeviation metricMagnitude="4" metricUnit="m" metricValue="1.2874752" pageId="0" pageNumber="46" unit="mi" value="8.0">8 mi</locationDeviation>
</quantity>
[
<quantity box="[856,930,1779,1803]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="0" pageNumber="46" unit="km" value="13.0">
<locationDeviation box="[856,930,1779,1803]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="0" pageNumber="46" unit="km" value="13.0">13 km</locationDeviation>
</quantity>
] 
<collectorName box="[944,1092,1779,1803]" pageId="0" pageNumber="46">NE Wilsonia</collectorName>
, 
<collectingMunicipality box="[1102,1380,1779,1803]" pageId="0" pageNumber="46">Sequoia National Forest</collectingMunicipality>
, 
<date pageId="0" pageNumber="46" value="1990-06-05">
<collectingDate pageId="0" pageNumber="46" value="1990-06-05">5 June 1990</collectingDate>
</date>
(
<specimenCount box="[943,973,1813,1838]" pageId="0" pageNumber="46" type="male">1♂</specimenCount>
), 
<collectionCode box="[997,1017,1814,1837]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="0" pageNumber="46">R</collectionCode>
. 
<collectorName box="[1030,1149,1814,1838]" pageId="0" pageNumber="46">L. Bottorff</collectorName>
(
<collectionCode box="[1165,1219,1814,1838]" collectionName="RLB" pageId="0" pageNumber="46">RLB</collectionCode>
); 
<collectorName pageId="0" pageNumber="46">Kings Canyon Road</collectorName>
, waterfall at 3,500’ [
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<elevation box="[1097,1183,1848,1872]" metricMagnitude="3" metricUnit="m" metricValue="1.067" pageId="0" pageNumber="46" unit="m" value="1067.0">1067 m</elevation>
</quantity>
], 
<date box="[1201,1342,1848,1873]" pageId="0" pageNumber="46" value="1963-05-31">
<collectingDate box="[1201,1342,1848,1873]" pageId="0" pageNumber="46" value="1963-05-31">31 May 1963</collectingDate>
</date>
(
<specimenCount box="[1358,1388,1848,1872]" pageId="0" pageNumber="46" type="male">1♂</specimenCount>
), 
<collectorName box="[812,1002,1883,1907]" pageId="0" pageNumber="46">C. P. Alexander</collectorName>
(
<collectionCode box="[1021,1105,1883,1907]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="0" pageNumber="46" type="Museum">USNM</collectionCode>
); 
<collectorName box="[1128,1347,1883,1907]" pageId="0" pageNumber="46">Kings Canyon, Rt.</collectorName>
180, waterfall above 
<collectorName box="[999,1190,1917,1941]" pageId="0" pageNumber="46">Horseshoe Bend</collectorName>
, 3,500’ [
<quantity box="[1282,1370,1917,1941]" metricMagnitude="3" metricUnit="m" metricValue="1.067" pageId="0" pageNumber="46" unit="m" value="1067.0">
<elevation box="[1282,1370,1917,1941]" metricMagnitude="3" metricUnit="m" metricValue="1.067" pageId="0" pageNumber="46" unit="m" value="1067.0">1067 m</elevation>
</quantity>
], 
<date pageId="0" pageNumber="46" value="1963-06-01">
<collectingDate pageId="0" pageNumber="46" value="1963-06-01">1 June 1963</collectingDate>
</date>
(
<specimenCount box="[958,991,1951,1976]" pageId="0" pageNumber="46" type="male">2♂</specimenCount>
, 
<specimenCount box="[1016,1047,1951,1976]" pageId="0" pageNumber="46" type="female">2♀</specimenCount>
), 
<collectorName box="[1081,1283,1952,1976]" pageId="0" pageNumber="46">C. P. Alexander</collectorName>
(
<collectionCode box="[1308,1392,1952,1976]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="0" pageNumber="46" type="Museum">USNM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<paragraph blockId="1.[1360,1391,838,893]" box="[1360,1391,838,893]" pageId="1" pageNumber="47">
<emphasis bold="true" box="[1360,1391,838,893]" pageId="1" pageNumber="47">2</emphasis>
</paragraph>
<paragraph blockId="1.[1360,1391,1382,1437]" box="[1360,1391,1382,1437]" pageId="1" pageNumber="47">
<emphasis bold="true" box="[1360,1391,1382,1437]" pageId="1" pageNumber="47">3</emphasis>
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4758701" ID-Zenodo-Dep="4758701" httpUri="https://zenodo.org/record/4758701/files/figure.png" pageId="1" pageNumber="47" startId="1.[184,234,1566,1590]" targetBox="[234,1317,294,1480]" targetPageId="1">
<paragraph blockId="1.[184,1404,1565,1693]" pageId="1" pageNumber="47">
Figs. 1‐5. 
<taxonomicName authorityName="Bottorff" authorityYear="2007" box="[288,521,1566,1590]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="47" phylum="Arthropoda" rank="species" species="sequoia">
<emphasis box="[288,521,1566,1590]" italics="true" pageId="1" pageNumber="47">Cosumnoperla sequoia</emphasis>
</taxonomicName>
adult male. 1. Head‐ pronotum, dorsal; 2. Terminalia (segments 8‐10, abdomen extended), dorsal; 3. Terminalia (segments 8‐10, abdomen contracted), right lateral; 4. Variant of reflexed process, 10 
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tergum, dorsal; 5. Aedeagus, right lateral (arrow shows location of median indention); Scale lines = 0.5 mm.
</paragraph>
</caption>
<subSubSection lastPageId="4" lastPageNumber="50" pageId="1" pageNumber="47" type="description">
<paragraph blockId="1.[184,776,1773,1970]" lastBlockId="2.[812,1404,288,726]" lastPageId="2" lastPageNumber="48" pageId="1" pageNumber="47">
<emphasis bold="true" box="[184,251,1773,1797]" pageId="1" pageNumber="47">Male.</emphasis>
Body uniform brown, length 8.1‐9.0 mm. Head brown, width 1.9‐2.0 mm (measured to outside margins of compound eyes), dark M‐ shaped mark anterior to median ocellus, interocellar area darker brown (
<figureCitation box="[276,340,1911,1936]" captionStart="Figs" captionStartId="1.[184,234,1566,1590]" captionTargetBox="[234,1317,294,1480]" captionTargetPageId="1" captionText="Figs. 1‐5. Cosumnoperla sequoia adult male. 1. Head‐pronotum, dorsal; 2. Terminalia (segments 8‐10, abdomen extended), dorsal; 3. Terminalia (segments 8‐10, abdomen contracted), right lateral; 4. Variant of reflexed process, 10th tergum, dorsal; 5. Aedeagus, right lateral (arrow shows location of median indention); Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4758701" httpUri="https://zenodo.org/record/4758701/files/figure.png" pageId="1" pageNumber="47">Fig. 1</figureCitation>
). Pronotum uniform brown, anterior corners darker, lateral edges white, disks with rugosities. Macropterous, forewing length 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="7.449999999999999" metricValueMax="7.8" metricValueMin="7.1" pageId="1" pageNumber="47" unit="mm" value="7.449999999999999" valueMax="7.8" valueMin="7.1">7.1‐7.8 mm</quantity>
, most wing cells and veins dark brown, costa and subcosta veins white, cells between costa and radius white. Supra‐ anal process bulbous and membranous. Paraprocts reduced and lightly sclerotized, lying under supra‐ anal process. Tergum 9 and 8 with medial and bipartite elevated patches (respectively) of dense stout golden spinulae (
<figureCitation captionStart="Figs" captionStartId="1.[184,234,1566,1590]" captionTargetBox="[234,1317,294,1480]" captionTargetPageId="1" captionText="Figs. 1‐5. Cosumnoperla sequoia adult male. 1. Head‐pronotum, dorsal; 2. Terminalia (segments 8‐10, abdomen extended), dorsal; 3. Terminalia (segments 8‐10, abdomen contracted), right lateral; 4. Variant of reflexed process, 10th tergum, dorsal; 5. Aedeagus, right lateral (arrow shows location of median indention); Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4758701" httpUri="https://zenodo.org/record/4758701/files/figure.png" pageId="2" pageNumber="48">Figs. 2, 3</figureCitation>
). Tergum 10 posterior margin with a shallow median notch (slightly wider in some specimens, 
<figureCitation captionStart="Figs" captionStartId="1.[184,234,1566,1590]" captionTargetBox="[234,1317,294,1480]" captionTargetPageId="1" captionText="Figs. 1‐5. Cosumnoperla sequoia adult male. 1. Head‐pronotum, dorsal; 2. Terminalia (segments 8‐10, abdomen extended), dorsal; 3. Terminalia (segments 8‐10, abdomen contracted), right lateral; 4. Variant of reflexed process, 10th tergum, dorsal; 5. Aedeagus, right lateral (arrow shows location of median indention); Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4758701" httpUri="https://zenodo.org/record/4758701/files/figure.png" pageId="2" pageNumber="48">Fig. 4</figureCitation>
); reflexed narrow process bearing two pairs of stout spines, the anterior pair pointing forward, the posterior pair pointing dorsally; fine stiff golden setae densely covering reflexed process (not shown in drawings). Aedeagus spherical with bulbous apex and lateral basal folds; small median cavity or indention on posterior surface (this may be a narrow invaginated lobe that failed to extend when the aedeagus was extruded); posterior and lateral band of dense unpigmented stout spinulae, posterior and lateral patches of medium golden spinulae at base of aedeagus, anterior medial patch of long golden spinulae (
<figureCitation box="[923,987,356,381]" captionStart="Figs" captionStartId="1.[184,234,1566,1590]" captionTargetBox="[234,1317,294,1480]" captionTargetPageId="1" captionText="Figs. 1‐5. Cosumnoperla sequoia adult male. 1. Head‐pronotum, dorsal; 2. Terminalia (segments 8‐10, abdomen extended), dorsal; 3. Terminalia (segments 8‐10, abdomen contracted), right lateral; 4. Variant of reflexed process, 10th tergum, dorsal; 5. Aedeagus, right lateral (arrow shows location of median indention); Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4758701" httpUri="https://zenodo.org/record/4758701/files/figure.png" pageId="2" pageNumber="48">Fig. 5</figureCitation>
). Basal cercal segments (1‐5) densely covered with stiff golden setae, these most abundant on the mesal and dorsal surfaces. Vesicle absent.
</paragraph>
<paragraph blockId="2.[812,1404,288,726]" pageId="2" pageNumber="48">
<emphasis bold="true" box="[812,904,461,485]" pageId="2" pageNumber="48">Female.</emphasis>
Body coloration and general appearance similar to male, but of larger size (body length 
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.055" metricValueMax="1.26" metricValueMin="0.85" pageId="2" pageNumber="48" unit="mm" value="10.55" valueMax="12.6" valueMin="8.5">8.512.6 mm</quantity>
, head width 
<quantity box="[1079,1213,530,554]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.6" metricValueMin="2.2" pageId="2" pageNumber="48" unit="mm" value="2.4" valueMax="2.6" valueMin="2.2">2.2‐2.6 mm</quantity>
). Macropterous, forewing length 
<quantity box="[1028,1183,564,588]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.4" metricValueMin="8.6" pageId="2" pageNumber="48" unit="mm" value="9.5" valueMax="10.4" valueMin="8.6">8.6‐10.4 mm</quantity>
. Subgenital plate elongated, nearly reaching posterior margin of 10 
<superScript attach="none" box="[1389,1403,599,613]" fontSize="6" pageId="2" pageNumber="48">th</superScript>
sternum, width narrowing from base to a shallowly notched apex, lateral sides irregularly indented, plate projecting ventrally from body axis (
<figureCitation box="[1249,1391,702,726]" captionStart="Fig" captionStartId="2.[184,223,1497,1521]" captionTargetBox="[184,809,801,1428]" captionTargetId="figure-455@2.[184,809,798,1428]" captionTargetPageId="2" captionText="Fig. 6. Cosumnoperla sequoia adult female. 6a. Terminalia (segments 8‐10), ventral; 6b‐c. Variants of subgenital plate, ventral; Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4758705" httpUri="https://zenodo.org/record/4758705/files/figure.png" pageId="2" pageNumber="48">Figs. 6a,b, c</figureCitation>
).
</paragraph>
<paragraph blockId="2.[1282,1351,1133,1188]" box="[1282,1351,1133,1188]" pageId="2" pageNumber="48">
<emphasis bold="true" box="[1282,1351,1133,1188]" pageId="2" pageNumber="48">6b</emphasis>
</paragraph>
<paragraph blockId="2.[846,1341,1398,1453]" box="[846,1341,1398,1453]" pageId="2" pageNumber="48">
<emphasis bold="true" box="[846,1341,1398,1453]" pageId="2" pageNumber="48">6a 6c</emphasis>
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4758705" ID-Zenodo-Dep="4758705" httpUri="https://zenodo.org/record/4758705/files/figure.png" pageId="2" pageNumber="48" startId="2.[184,223,1497,1521]" targetBox="[184,809,801,1428]" targetPageId="2">
<paragraph blockId="2.[184,1403,1496,1555]" pageId="2" pageNumber="48">
Fig. 6. 
<taxonomicName authorityName="Bottorff" authorityYear="2007" box="[256,489,1497,1521]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="sequoia">
<emphasis box="[256,489,1497,1521]" italics="true" pageId="2" pageNumber="48">Cosumnoperla sequoia</emphasis>
</taxonomicName>
adult female. 6a. Terminalia (segments 8‐10), ventral; 6b‐ c. Variants of subgenital plate, ventral; Scale lines = 0.5 mm.
</paragraph>
</caption>
<paragraph blockId="2.[184,776,1635,1970]" pageId="2" pageNumber="48">
<emphasis bold="true" box="[184,235,1635,1659]" pageId="2" pageNumber="48">Egg.</emphasis>
Oblong (circular cross‐ section). Length 700 μm. Width 400 μm. Color olive green. Collar and eclosion line absent. Chorion covered with irregular quadrangular or hexagonal follicle cell impressions (FCIs) bordered by raised walls, FCI floors flat. Each FCI typically surrounded by 6 other FCIs (range, 5‐7). Micropyles not seen. Egg microstructure similar to that of 
<taxonomicName authorityName="Szczytko &amp; R.L.Bottorff" authorityYear="1987" box="[264,402,1877,1901]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="hypocrena">
<emphasis box="[264,402,1877,1901]" italics="true" pageId="2" pageNumber="48">C. hypocrena</emphasis>
</taxonomicName>
(see Figures 
<quantity box="[551,641,1877,1901]" metricMagnitude="-1" metricUnit="m" metricValue="5.207" metricValueMax="5.334" metricValueMin="5.08" pageId="2" pageNumber="48" unit="in" value="20.5" valueMax="21.0" valueMin="20.0">20‐21 in</quantity>
Szczytko &amp; Bottorff 1987).
</paragraph>
<paragraph blockId="2.[184,776,1635,1970]" lastBlockId="4.[812,1404,288,830]" lastPageId="4" lastPageNumber="50" pageId="2" pageNumber="48">
<emphasis bold="true" box="[184,257,1946,1970]" pageId="2" pageNumber="48">Larva.</emphasis>
General body color medium brown; pigment pattern similar to that of 
<taxonomicName authorityName="Szczytko &amp; R.L.Bottorff" authorityYear="1987" box="[1113,1253,1635,1659]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="hypocrena">
<emphasis box="[1113,1253,1635,1659]" italics="true" pageId="2" pageNumber="48">C. hypocrena</emphasis>
</taxonomicName>
(Szczytko &amp; Bottorff 1987; Stewart &amp; Stark 2002). Body length of mature larvae, 
<quantity box="[999,1105,1704,1728]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="2" pageNumber="48" unit="mm" value="11.1">11.1 mm</quantity>
male and 
<quantity box="[1245,1404,1704,1728]" metricMagnitude="-2" metricUnit="m" metricValue="1.2550000000000001" metricValueMax="1.3800000000000001" metricValueMin="1.1300000000000001" pageId="2" pageNumber="48" unit="mm" value="12.55" valueMax="13.8" valueMin="11.3">11.3‐13.8 mm</quantity>
female. Head width 
<quantity box="[1040,1165,1738,1762]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="2.4" metricValueMin="1.8" pageId="2" pageNumber="48" unit="mm" value="2.1" valueMax="2.4" valueMin="1.8">1.8‐2.4 mm</quantity>
. Head mostly brown, with light M‐ shaped mark anterior to median ocellus, ocellar triangle with a median oval white spot, rectangular white spots lateral to ocellar triangle, occiput with reticulate light patches and an irregular row of spinules (
<figureCitation box="[1008,1074,1911,1935]" captionStart="Figs" captionStartId="3.[184,234,1876,1900]" captionTargetBox="[896,1274,1310,1808]" captionTargetId="figure-103@3.[890,1280,1303,1818]" captionTargetPageId="3" captionText="Figs. 7‐12. Cosumnoperla sequoia larva. 7. Head‐pronotum, dorsal; 8. Thorax, ventral; 9. Right front leg, dorsal; 10. Abdomen (segments 1‐10), dorsal; 11. Male terminalia (segments 8‐10), ventral; 12. Female terminalia (segments 8‐10), ventral; Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4758707" httpUri="https://zenodo.org/record/4758707/files/figure.png" pageId="2" pageNumber="48">Fig. 7</figureCitation>
). Mandibles with 6 teeth, 5 
<superScript attach="none" box="[1389,1403,1912,1926]" fontSize="6" pageId="2" pageNumber="48">th</superScript>
tooth serrated; marginal brush of long stiff setae; tuft of marginal bristles below 4 
<superScript attach="left" box="[614,628,288,302]" fontSize="6" pageId="4" pageNumber="50">th</superScript>
tooth (dense and expanded on left mandible, sparse on right mandible); ventral sparse setal patch extending from 1 
<superScript attach="left" box="[197,209,392,406]" fontSize="6" pageId="4" pageNumber="50">st</superScript>
tooth to mandible base (
<figureCitation box="[488,615,391,415]" captionStart="Figs" captionStartId="5.[184,234,1220,1244]" captionTargetBox="[211,1359,671,1169]" captionTargetId="figure-518@5.[547,931,664,1148]" captionTargetPageId="5" captionText="Figs. 15‐16. Cosumnoperla sequoia larva. 15. Right (a) and left (b) mandibles, ventral; 16. Right lacinia, ventral." figureDoi="http://doi.org/10.5281/zenodo.4758711" httpUri="https://zenodo.org/record/4758711/files/figure.png" pageId="4" pageNumber="50">Figs. 15a, b</figureCitation>
). Lacinia with 2 large teeth; apical tooth about twice as long as the subapical tooth; 4‐6 setae between apical and subapical teeth; lacinial margin below subapical tooth with 6‐8 stout setae and then 10‐12 thin setae, sub‐ marginal row of 4‐6 stout setae (
<figureCitation box="[673,763,564,588]" captionStart="Figs" captionStartId="5.[184,234,1220,1244]" captionTargetBox="[211,1359,671,1169]" captionTargetId="figure-518@5.[547,931,664,1148]" captionTargetPageId="5" captionText="Figs. 15‐16. Cosumnoperla sequoia larva. 15. Right (a) and left (b) mandibles, ventral; 16. Right lacinia, ventral." figureDoi="http://doi.org/10.5281/zenodo.4758711" httpUri="https://zenodo.org/record/4758711/files/figure.png" pageId="4" pageNumber="50">Fig. 16</figureCitation>
). Pronotum with brown disks and white rugosities, lateral margins white. All thoracic sterna with distinct medial sclerites covering and extending beyond furcal pits (
<figureCitation box="[401,464,702,726]" captionStart="Figs" captionStartId="3.[184,234,1876,1900]" captionTargetBox="[896,1274,1310,1808]" captionTargetId="figure-103@3.[890,1280,1303,1818]" captionTargetPageId="3" captionText="Figs. 7‐12. Cosumnoperla sequoia larva. 7. Head‐pronotum, dorsal; 8. Thorax, ventral; 9. Right front leg, dorsal; 10. Abdomen (segments 1‐10), dorsal; 11. Male terminalia (segments 8‐10), ventral; 12. Female terminalia (segments 8‐10), ventral; Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4758707" httpUri="https://zenodo.org/record/4758707/files/figure.png" pageId="4" pageNumber="50">Fig. 8</figureCitation>
). Prosternal sclerite extends forward from furcal pits to anterior margin of prosternum, sclerite swollen in mid‐ segment and then narrowed anteriorly. Mesosternal Y‐ arms meet posterior corners of furcal pits, transverse suture connects anterior corners of furcal pits; sclerite extends laterally beyond Y‐ arms, lateral margin irregular or sinuous. Gills absent. Femora and tibia with dense short setae, scattered spinules, and fringed with light silky hairs (
<figureCitation box="[1179,1249,426,450]" captionStart="Figs" captionStartId="3.[184,234,1876,1900]" captionTargetBox="[896,1274,1310,1808]" captionTargetId="figure-103@3.[890,1280,1303,1818]" captionTargetPageId="3" captionText="Figs. 7‐12. Cosumnoperla sequoia larva. 7. Head‐pronotum, dorsal; 8. Thorax, ventral; 9. Right front leg, dorsal; 10. Abdomen (segments 1‐10), dorsal; 11. Male terminalia (segments 8‐10), ventral; 12. Female terminalia (segments 8‐10), ventral; Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4758707" httpUri="https://zenodo.org/record/4758707/files/figure.png" pageId="4" pageNumber="50">Fig. 9</figureCitation>
). Abdominal terga brown, most segments with a pair of large submedial and lateral light spots (overall, gives the appearance of longitudinal light and dark striping); each tergum with a transverse row of 6 dark dots, 2 mesally and 2 each laterally (
<figureCitation box="[1186,1273,598,623]" captionStart="Figs" captionStartId="3.[184,234,1876,1900]" captionTargetBox="[896,1274,1310,1808]" captionTargetId="figure-103@3.[890,1280,1303,1818]" captionTargetPageId="3" captionText="Figs. 7‐12. Cosumnoperla sequoia larva. 7. Head‐pronotum, dorsal; 8. Thorax, ventral; 9. Right front leg, dorsal; 10. Abdomen (segments 1‐10), dorsal; 11. Male terminalia (segments 8‐10), ventral; 12. Female terminalia (segments 8‐10), ventral; Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4758707" httpUri="https://zenodo.org/record/4758707/files/figure.png" pageId="4" pageNumber="50">Fig. 10</figureCitation>
). Posterior margin of male abdominal sterna 8 and 9 entire, with a continuous setal row (
<figureCitation box="[1084,1162,667,692]" captionStart="Figs" captionStartId="3.[184,234,1876,1900]" captionTargetBox="[896,1274,1310,1808]" captionTargetId="figure-103@3.[890,1280,1303,1818]" captionTargetPageId="3" captionText="Figs. 7‐12. Cosumnoperla sequoia larva. 7. Head‐pronotum, dorsal; 8. Thorax, ventral; 9. Right front leg, dorsal; 10. Abdomen (segments 1‐10), dorsal; 11. Male terminalia (segments 8‐10), ventral; 12. Female terminalia (segments 8‐10), ventral; Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4758707" httpUri="https://zenodo.org/record/4758707/files/figure.png" pageId="4" pageNumber="50">Fig. 11</figureCitation>
). Posterior margin of female abdominal sternum 8 sclerotized medially and slightly protruding, with a median setal gap (
<figureCitation box="[820,900,771,795]" captionStart="Figs" captionStartId="3.[184,234,1876,1900]" captionTargetBox="[896,1274,1310,1808]" captionTargetId="figure-103@3.[890,1280,1303,1818]" captionTargetPageId="3" captionText="Figs. 7‐12. Cosumnoperla sequoia larva. 7. Head‐pronotum, dorsal; 8. Thorax, ventral; 9. Right front leg, dorsal; 10. Abdomen (segments 1‐10), dorsal; 11. Male terminalia (segments 8‐10), ventral; 12. Female terminalia (segments 8‐10), ventral; Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4758707" httpUri="https://zenodo.org/record/4758707/files/figure.png" pageId="4" pageNumber="50">Fig. 12</figureCitation>
). Cercal segments with a posterior whorl of short setae, dorsal fringe of long hairs absent.
</paragraph>
</subSubSection>
<paragraph blockId="3.[737,799,1725,1780]" box="[737,799,1725,1780]" pageId="3" pageNumber="49">11</paragraph>
<paragraph blockId="3.[1297,1359,1725,1780]" box="[1297,1359,1725,1780]" pageId="3" pageNumber="49">12</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4758707" ID-Zenodo-Dep="4758707" httpUri="https://zenodo.org/record/4758707/files/figure.png" pageId="3" pageNumber="49" startId="3.[184,234,1876,1900]" targetBox="[896,1274,1310,1808]" targetPageId="3">
<paragraph blockId="3.[184,1403,1876,1970]" pageId="3" pageNumber="49">
Figs. 7‐12. 
<taxonomicName authorityName="Bottorff" authorityYear="2007" box="[302,535,1876,1900]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="sequoia">
<emphasis box="[302,535,1876,1900]" italics="true" pageId="3" pageNumber="49">Cosumnoperla sequoia</emphasis>
</taxonomicName>
larva. 7. Head‐ pronotum, dorsal; 8. Thorax, ventral; 9. Right front leg, dorsal; 10. Abdomen (segments 1‐10), dorsal; 11. Male terminalia (segments 8‐10), ventral; 12. Female terminalia (segments 8‐10), ventral; Scale lines = 0.5 mm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4758709" ID-Zenodo-Dep="4758709" httpUri="https://zenodo.org/record/4758709/files/figure.png" pageId="4" pageNumber="50" startId="4.[184,234,1427,1451]" targetBox="[174,1390,935,1383]" targetPageId="4">
<paragraph blockId="4.[184,1404,1427,1486]" pageId="4" pageNumber="50">
Figs. 13‐14. 
<taxonomicName authorityName="Szczytko &amp; R.L.Bottorff" authorityYear="1987" box="[329,599,1427,1451]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="hypocrena">
<emphasis box="[329,599,1427,1451]" italics="true" pageId="4" pageNumber="50">Cosumnoperla hypocrena</emphasis>
</taxonomicName>
adult male. 13. Terminalia (segments 9‐10), dorsal; 14. Terminalia (segments 9‐10), right lateral; Scale lines = 0.5 mm.
</paragraph>
</caption>
<subSubSection lastPageId="5" lastPageNumber="51" pageId="4" pageNumber="50" type="diagnosis">
<paragraph blockId="4.[184,776,1566,1970]" lastBlockId="4.[812,1404,1566,1970]" pageId="4" pageNumber="50">
<emphasis bold="true" box="[184,309,1566,1590]" pageId="4" pageNumber="50">Diagnosis.</emphasis>
<taxonomicName authorityName="Bottorff" authorityYear="2007" box="[331,580,1566,1590]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="sequoia">
<emphasis box="[331,580,1566,1590]" italics="true" pageId="4" pageNumber="50">Cosumnoperla sequoia</emphasis>
</taxonomicName>
shares several diagnostic features with 
<taxonomicName authorityName="Szczytko &amp; R.L.Bottorff" authorityYear="1987" box="[461,597,1601,1625]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="hypocrena">
<emphasis box="[461,597,1601,1625]" italics="true" pageId="4" pageNumber="50">C. hypocrena</emphasis>
</taxonomicName>
. Adult males of both species have (1) a reflexed process with an anterior pair of prominent spines on the 10 
<superScript attach="left" box="[665,679,1670,1684]" fontSize="6" pageId="4" pageNumber="50">th</superScript>
tergum, (2) a median patch of stout spinulae on the 9 
<superScript attach="none" box="[761,775,1705,1719]" fontSize="6" pageId="4" pageNumber="50">th</superScript>
tergum, (3) a membranous aedeagus, (4) a bulbous membranous supra‐ anal process, (5) weakly sclerotized flat paraprocts, and (6) vesicle absent. Adult females have an extended apically‐ notched subgenital plate. Larvae of both species have (1) abdominal longitudinal stripes and dark dots and (2) a sinuate mesosternal sclerite that extends laterally beyond the Y‐ arms. Eggs have a punctuate microstructure and are large, green, oblong, and collarless. Both species are restricted to intermittent streams in the Sierra 
<collectingRegion box="[1052,1144,1669,1693]" country="United States of America" name="Nevada" pageId="4" pageNumber="50">Nevada</collectingRegion>
, 
<collectingRegion box="[1156,1274,1669,1693]" country="United States of America" name="California" pageId="4" pageNumber="50">California</collectingRegion>
.
</paragraph>
<paragraph blockId="4.[812,1404,1566,1970]" lastBlockId="5.[812,1404,288,623]" lastPageId="5" lastPageNumber="51" pageId="4" pageNumber="50">
<taxonomicName authorityName="Bottorff" authorityYear="2007" box="[852,1082,1704,1728]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="sequoia">
<emphasis box="[852,1082,1704,1728]" italics="true" pageId="4" pageNumber="50">Cosumnoperla sequoia</emphasis>
</taxonomicName>
adults and larvae differ from 
<taxonomicName authorityName="Szczytko &amp; R.L.Bottorff" authorityYear="1987" box="[812,961,1738,1762]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="hypocrena">
<emphasis box="[812,961,1738,1762]" italics="true" pageId="4" pageNumber="50">C. hypocrena</emphasis>
</taxonomicName>
by their smaller size and several morphological features: (1) adult pronotum – 
<taxonomicName authorityName="Bottorff" authorityYear="2007" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="sequoia">
<emphasis italics="true" pageId="4" pageNumber="50">C. sequoia</emphasis>
</taxonomicName>
uniform brown, lateral margins white; 
<taxonomicName authorityName="Szczytko &amp; R.L.Bottorff" authorityYear="1987" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="hypocrena">
<emphasis italics="true" pageId="4" pageNumber="50">C. hypocrena</emphasis>
</taxonomicName>
median light stripe, (2) wings – 
<taxonomicName authorityName="Bottorff" authorityYear="2007" box="[1295,1404,1842,1866]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="sequoia">
<emphasis box="[1295,1404,1842,1866]" italics="true" pageId="4" pageNumber="50">C. sequoia</emphasis>
</taxonomicName>
uniform dark brown except for white cells and veins between costa and radius; 
<taxonomicName authorityName="Szczytko &amp; R.L.Bottorff" authorityYear="1987" box="[1141,1284,1911,1935]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="hypocrena">
<emphasis box="[1141,1284,1911,1935]" italics="true" pageId="4" pageNumber="50">C. hypocrena</emphasis>
</taxonomicName>
cells and veins uniform dark brown, (3) 10 
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tergum of male adults – 
<taxonomicName authorityName="Bottorff" authorityYear="2007" box="[294,408,288,311]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="sequoia">
<emphasis box="[294,408,288,311]" italics="true" pageId="5" pageNumber="51">C. sequoia</emphasis>
</taxonomicName>
narrow reflexed process with mesoposterior notch and two posterior spines; 
<taxonomicName authorityName="Szczytko &amp; R.L.Bottorff" authorityYear="1987" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="hypocrena">
<emphasis italics="true" pageId="5" pageNumber="51">C. hypocrena</emphasis>
</taxonomicName>
broad reflexed triangular process with mesoposterior entire and posterior spines absent (
<figureCitation box="[192,325,426,450]" captionStart="Figs" captionStartId="4.[184,234,1427,1451]" captionTargetBox="[174,1390,935,1383]" captionTargetId="figure-544@4.[174,689,935,1383]" captionTargetPageId="4" captionText="Figs. 13‐14. Cosumnoperla hypocrena adult male. 13. Terminalia (segments 9‐10), dorsal; 14. Terminalia (segments 9‐10), right lateral; Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4758709" httpUri="https://zenodo.org/record/4758709/files/figure.png" pageId="5" pageNumber="51">Figs. 13, 14</figureCitation>
), (4) terga 8 and 9 of male adults – 
<taxonomicName authorityName="Bottorff" authorityYear="2007" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="sequoia">
<emphasis italics="true" pageId="5" pageNumber="51">C. sequoia</emphasis>
</taxonomicName>
prominent swollen patches of golden stout spinulae; 
<taxonomicName authorityName="Szczytko &amp; R.L.Bottorff" authorityYear="1987" box="[303,449,495,519]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="hypocrena">
<emphasis box="[303,449,495,519]" italics="true" pageId="5" pageNumber="51">C. hypocrena</emphasis>
</taxonomicName>
small unswollen patch of spinulae on 9 
<superScript attach="left" box="[340,354,530,544]" fontSize="6" pageId="5" pageNumber="51">th</superScript>
tergum, few spinulae on 8 
<superScript attach="left" box="[669,683,530,544]" fontSize="6" pageId="5" pageNumber="51">th</superScript>
tergum (
<figureCitation box="[192,316,564,588]" captionStart="Figs" captionStartId="4.[184,234,1427,1451]" captionTargetBox="[174,1390,935,1383]" captionTargetId="figure-544@4.[174,689,935,1383]" captionTargetPageId="4" captionText="Figs. 13‐14. Cosumnoperla hypocrena adult male. 13. Terminalia (segments 9‐10), dorsal; 14. Terminalia (segments 9‐10), right lateral; Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4758709" httpUri="https://zenodo.org/record/4758709/files/figure.png" pageId="5" pageNumber="51">Figs. 13, 14</figureCitation>
), (5) basal cercal segments of male adults – 
<taxonomicName authorityName="Bottorff" authorityYear="2007" box="[202,308,599,622]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="sequoia">
<emphasis box="[202,308,599,622]" italics="true" pageId="5" pageNumber="51">C. sequoia</emphasis>
</taxonomicName>
dense long setae; 
<taxonomicName authorityName="Szczytko &amp; R.L.Bottorff" authorityYear="1987" box="[508,643,598,622]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="hypocrena">
<emphasis box="[508,643,598,622]" italics="true" pageId="5" pageNumber="51">C. hypocrena</emphasis>
</taxonomicName>
sparse long setae, (6) subgenital plate of female adults – 
<taxonomicName authorityName="Bottorff" authorityYear="2007" box="[1299,1404,288,311]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="sequoia">
<emphasis box="[1299,1404,288,311]" italics="true" pageId="5" pageNumber="51">C. sequoia</emphasis>
</taxonomicName>
long and narrow with irregular sides and shallow apical notch; 
<taxonomicName authorityName="Szczytko &amp; R.L.Bottorff" authorityYear="1987" box="[979,1124,357,381]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="hypocrena">
<emphasis box="[979,1124,357,381]" italics="true" pageId="5" pageNumber="51">C. hypocrena</emphasis>
</taxonomicName>
short and broad with smooth sides and deep apical notch, (7) prosternal sclerite of larvae ‐ 
<taxonomicName authorityName="Bottorff" authorityYear="2007" box="[1015,1119,426,449]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="sequoia">
<emphasis box="[1015,1119,426,449]" italics="true" pageId="5" pageNumber="51">C. sequoia</emphasis>
</taxonomicName>
swollen anterior to furcal pits; 
<taxonomicName authorityName="Szczytko &amp; R.L.Bottorff" authorityYear="1987" box="[866,1002,460,484]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="hypocrena">
<emphasis box="[866,1002,460,484]" italics="true" pageId="5" pageNumber="51">C. hypocrena</emphasis>
</taxonomicName>
uniformly narrow anterior to furcal pits. The oblong collarless green egg of 
<taxonomicName authorityName="Bottorff" authorityYear="2007" box="[1270,1378,495,518]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="sequoia">
<emphasis box="[1270,1378,495,518]" italics="true" pageId="5" pageNumber="51">C. sequoia</emphasis>
</taxonomicName>
is slightly smaller than that of 
<emphasis box="[1148,1295,529,553]" italics="true" pageId="5" pageNumber="51">
<taxonomicName authorityName="Szczytko &amp; R.L.Bottorff" authorityYear="1987" box="[1148,1291,529,553]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="hypocrena">C. hypocrena</taxonomicName>
,
</emphasis>
but both eggs have similar microstructure and are some of the largest stonefly eggs reported.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4758711" ID-Zenodo-Dep="4758711" box="[184,1404,1220,1244]" httpUri="https://zenodo.org/record/4758711/files/figure.png" pageId="5" pageNumber="51" startId="5.[184,234,1220,1244]" targetBox="[211,1359,671,1169]" targetPageId="5">
<paragraph blockId="5.[184,1404,1220,1244]" box="[184,1404,1220,1244]" pageId="5" pageNumber="51">
Figs. 15‐16. 
<taxonomicName authorityName="Bottorff" authorityYear="2007" box="[316,550,1220,1244]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="sequoia">
<emphasis box="[316,550,1220,1244]" italics="true" pageId="5" pageNumber="51">Cosumnoperla sequoia</emphasis>
</taxonomicName>
larva. 15. Right (a) and left (b) mandibles, ventral; 16. Right lacinia, ventral.
</paragraph>
</caption>
<subSubSection pageId="5" pageNumber="51" type="discussion">
<paragraph blockId="5.[184,776,1324,1970]" lastBlockId="5.[812,1404,1324,1971]" pageId="5" pageNumber="51">
Several characters clearly place 
<taxonomicName authorityName="Bottorff" authorityYear="2007" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="sequoia">
<emphasis italics="true" pageId="5" pageNumber="51">Cosumnoperla sequoia</emphasis>
</taxonomicName>
within the Isoperlinae – (1) smaller adult size than most Perlodinae, (2) reduced membranous supra‐ anal process, (3) simple unsclerotized paraprocts of male adults, (4) uncleft 10 
<superScript attach="left" box="[640,654,1463,1477]" fontSize="6" pageId="5" pageNumber="51">th</superScript>
tergum of male adults, (5) larvae with longitudinal abdominal stripes, and (6) larval submental and thoracic gills absent. Adult males of Isoperlinae often have various spinulae patches, dorsal swellings, or modifications of the 9 
<superScript attach="left" box="[270,284,1635,1649]" fontSize="6" pageId="5" pageNumber="51">th</superScript>
and 10 
<superScript attach="left" box="[368,382,1635,1649]" fontSize="6" pageId="5" pageNumber="51">th</superScript>
terga. Although not all characters clearly separate the Isoperlinae from Perlodinae, the preponderance of traits do distinguish these two subfamilies. 
<taxonomicName box="[346,497,1738,1762]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis box="[346,497,1738,1762]" italics="true" pageId="5" pageNumber="51">Cosumnoperla</emphasis>
</taxonomicName>
and the monotypic Isoperlinae genera may be transitional forms between the Perlodinae and 
<taxonomicName authorityName="Banks" authorityYear="1906" box="[507,589,1807,1831]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis box="[507,589,1807,1831]" italics="true" pageId="5" pageNumber="51">Isoperla</emphasis>
</taxonomicName>
since they share some characters with both taxonomic groups. In contrast with 
<emphasis box="[386,475,1877,1901]" italics="true" pageId="5" pageNumber="51">
<taxonomicName authorityName="Banks" authorityYear="1906" box="[386,471,1877,1901]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="genus">Isoperla</taxonomicName>
,
</emphasis>
the adult males of 
<taxonomicName box="[184,335,1911,1935]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis box="[184,335,1911,1935]" italics="true" pageId="5" pageNumber="51">Cosumnoperla</emphasis>
</taxonomicName>
and the monotypic Isoperlinae genera have various processes, lobes, ridges, or recurved hooks on tergum 10 and lightly sclerotized paraprocts. Within the Nearctic Isoperlinae, only 
<emphasis box="[812,1207,1393,1417]" italics="true" pageId="5" pageNumber="51">
<taxonomicName authorityName="Bottorff" authorityYear="2007" box="[812,1050,1393,1417]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="sequoia">Cosumnoperla sequoia</taxonomicName>
, 
<taxonomicName baseAuthorityName="Newman" baseAuthorityYear="1839" box="[1062,1203,1393,1417]" class="Insecta" family="Perlodidae" genus="Clioperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="clio">Clioperla clio</taxonomicName>
,
</emphasis>
and 
<taxonomicName baseAuthorityName="Hoppe" baseAuthorityYear="1938" class="Insecta" family="Perlodidae" genus="Cascadoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="trictura">
<emphasis italics="true" pageId="5" pageNumber="51">Cascadoperla trictura</emphasis>
</taxonomicName>
have slightly to deeply notched 10 
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terga.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="51" type="etymology">
<paragraph blockId="5.[812,1404,1324,1971]" pageId="5" pageNumber="51">
<emphasis bold="true" box="[812,946,1462,1486]" pageId="5" pageNumber="51">Etymology.</emphasis>
This species is named after the 
<typeStatus box="[1353,1404,1463,1487]" pageId="5" pageNumber="51">type</typeStatus>
locality, which is near the impressive groves of Sequoia big trees (
<taxonomicName box="[1044,1337,1531,1555]" class="Pinopsida" family="Cupressaceae" genus="Sequoiadendron" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="51" phylum="Tracheophyta" rank="species" species="giganteum">
<emphasis box="[1044,1337,1531,1555]" italics="true" pageId="5" pageNumber="51">Sequoiadendron giganteum</emphasis>
</taxonomicName>
) and Sequoia National Park.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="51" type="distribution">
<paragraph blockId="5.[812,1404,1324,1971]" pageId="5" pageNumber="51">
<emphasis bold="true" box="[812,966,1600,1624]" pageId="5" pageNumber="51">Distribution.</emphasis>
This species is presently known from the 
<typeStatus box="[855,906,1636,1660]" pageId="5" pageNumber="51">type</typeStatus>
locality and a nearby creek in the southern Sierra 
<collectingRegion box="[886,978,1670,1694]" country="United States of America" name="Nevada" pageId="5" pageNumber="51">Nevada</collectingRegion>
, 
<collectingRegion box="[989,1107,1670,1694]" country="United States of America" name="California" pageId="5" pageNumber="51">California</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="52" pageId="5" pageNumber="51" type="biology_ecology">
<paragraph blockId="5.[812,1404,1324,1971]" lastBlockId="6.[184,776,286,726]" lastPageId="6" lastPageNumber="52" pageId="5" pageNumber="51">
<emphasis bold="true" box="[812,1011,1704,1729]" pageId="5" pageNumber="51">Biological Notes.</emphasis>
When the 
<typeStatus box="[1130,1231,1705,1729]" pageId="5" pageNumber="51">holotype</typeStatus>
and 
<typeStatus box="[1285,1377,1705,1729]" pageId="5" pageNumber="51">allotype</typeStatus>
of 
<taxonomicName authorityName="Bottorff" authorityYear="2007" box="[812,923,1740,1763]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="sequoia">
<emphasis box="[812,923,1740,1763]" italics="true" pageId="5" pageNumber="51">C. sequoia</emphasis>
</taxonomicName>
were collected in 1990, the type locality stream was dry, except at one location where a small trickle of water seeped down the vertical face of a granite bedrock cliff (
<quantity box="[1049,1103,1843,1867]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="51" unit="m" value="10.0">10 m</quantity>
) and temporarily collected in a small shallow pool (
<quantity box="[1102,1145,1878,1902]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="51" unit="m" value="1.0">1 m</quantity>
diameter) at the base, before sinking into the channel substrate. Mature larvae inhabited this one pool (water temperature, 16.3 
<emphasis box="[229,239,286,311]" italics="true" pageId="6" pageNumber="52">°</emphasis>
C) and adults were collected on nearby vegetation. Obviously, these stoneflies were collected at the very end of the normal winter‐ spring flow period of this intermittent stream. 
<taxonomicName authorityName="Bottorff" authorityYear="2007" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="52" phylum="Arthropoda" rank="species" species="sequoia">
<emphasis italics="true" pageId="6" pageNumber="52">Cosumnoperla sequoia</emphasis>
</taxonomicName>
was absent from a nearby perennial stream, Redwood Creek, which instead had a typical assemblage of Sierra stoneflies, including some semivoltine 
<taxonomicName box="[325,422,529,553]" class="Insecta" family="Perlidae" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="52" phylum="Arthropoda" rank="family">Perlidae</taxonomicName>
. The distributions of 
<taxonomicName authorityName="Bottorff" authorityYear="2007" box="[667,776,530,553]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="52" phylum="Arthropoda" rank="species" species="sequoia">
<emphasis box="[667,776,530,553]" italics="true" pageId="6" pageNumber="52">C. sequoia</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Szczytko &amp; R.L.Bottorff" authorityYear="1987" box="[233,368,564,588]" class="Insecta" family="Perlodidae" genus="Cosumnoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="52" phylum="Arthropoda" rank="species" species="hypocrena">
<emphasis box="[233,368,564,588]" italics="true" pageId="6" pageNumber="52">C. hypocrena</emphasis>
</taxonomicName>
are confined to intermittent streams at low to medium elevations (
<quantity box="[550,686,598,623]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="15.0" metricValueMin="3.0" pageId="6" pageNumber="52" unit="m" value="900.0" valueMax="1500.0" valueMin="300.0">300‐1500 m</quantity>
) of the Sierra 
<collectingRegion box="[273,366,633,657]" country="United States of America" name="Nevada" pageId="6" pageNumber="52">Nevada</collectingRegion>
. Based upon presently known distributions, these two species are separated by about 
<quantity box="[256,338,702,726]" metricMagnitude="5" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="52" unit="km" value="250.0">250 km</quantity>
along this mountain range.
</paragraph>
</subSubSection>
</treatment>
</document>