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<document id="4591A41F9C6B9CEAC2D1F3A422E5F579" ID-CLB-Dataset="171436" ID-DOI="10.11646/zootaxa.5196.3.4" ID-GBIF-Dataset="32c56a4e-265a-4dee-86e7-1703ceb40035" ID-ISSN="1175-5326" ID-Zenodo-Dep="7235117" ID-ZooBank="443E7AE5-6131-48CC-9F1E-271DDA16B028" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666361560357" checkinUser="plazi" docAuthor="Mikhaljova, Elena V." docDate="2022" docId="03CF2E1BFFB4F976FF498EF004ACFDB4" docLanguage="en" docName="zootaxa.5169.3.4.pdf" docOrigin="Zootaxa 5196 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nepalella labolensis Mikhaljova 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="398" masterDocId="FFF65663FFBCF97CFFDE8E67003DFF89" masterDocTitle="Contributions to the millipede fauna of China: five new species of the genus Nepalella Shear, 1979 (Diplopoda, Chordeumatida, Megalotylidae)" masterLastPageNumber="406" masterPageNumber="388" pageNumber="396" updateTime="1699408116830" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="376E2570830C56EB68FDFF338A929A83">Contributions to the millipede fauna of China: five new species of the genus Nepalella Shear, 1979 (Diplopoda, Chordeumatida, Megalotylidae)</mods:title>
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<mods:name id="8A96DF4E5ADF9D1897702D343A98A057" type="personal">
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<mods:namePart id="95F1BE4429F74B52E9EE7510BEC74CF3">Mikhaljova, Elena V.</mods:namePart>
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<mods:title id="5A95A5719F74F64B2C462457E0935CA5">Zootaxa</mods:title>
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<mods:date id="039740AF1BE9039B3D2408BA5E4C4A5B">2022</mods:date>
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<mods:number id="8C2C5B2ADC3C089638276FA6EADDE78A">2022-10-20</mods:number>
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<treatment id="03CF2E1BFFB4F976FF498EF004ACFDB4" ID-DOI="http://doi.org/10.5281/zenodo.7235170" ID-GBIF-Taxon="203130374" ID-Zenodo-Dep="7235170" LSID="urn:lsid:plazi:treatment:03CF2E1BFFB4F976FF498EF004ACFDB4" httpUri="http://treatment.plazi.org/id/03CF2E1BFFB4F976FF498EF004ACFDB4" lastPageId="10" lastPageNumber="398" pageId="8" pageNumber="396">
<subSubSection id="C37CCC86FFB4F974FF498EF001CDFF38" box="[151,496,151,178]" pageId="8" pageNumber="396" type="nomenclature">
<paragraph id="8BD99F0DFFB4F974FF498EF001CDFF38" blockId="8.[151,496,151,213]" box="[151,496,151,178]" pageId="8" pageNumber="396">
<heading id="D0912861FFB4F974FF498EF001CDFF38" bold="true" box="[151,496,151,178]" fontSize="11" level="1" pageId="8" pageNumber="396" reason="1">
<emphasis id="B912431FFFB4F974FF498EF001CDFF38" bold="true" box="[151,496,151,178]" pageId="8" pageNumber="396">
<taxonomicName id="4C66E48EFFB4F974FF498EF001B6FF3B" authority="Mikhaljova, 2022" authorityName="Mikhaljova" authorityYear="2022" box="[151,395,151,178]" class="Diplopoda" family="Lankasomatidae" genus="Nepalella" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="396" phylum="Arthropoda" rank="species" species="labolensis" status="sp. nov.">
<emphasis id="B912431FFFB4F974FF498EF001B6FF3B" bold="true" box="[151,395,151,178]" italics="true" pageId="8" pageNumber="396">Nepalella labolensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A221FE64FFB4F974FE4C8EF001CDFF38" box="[402,496,151,177]" pageId="8" pageNumber="396" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37CCC86FFB4F974FF498EDC0128FF5C" box="[151,277,187,213]" pageId="8" pageNumber="396" type="description">
<paragraph id="8BD99F0DFFB4F974FF498EDC0128FF5C" blockId="8.[151,496,151,213]" box="[151,277,187,213]" pageId="8" pageNumber="396">
<figureCitation id="135D8388FFB4F974FF498EDC0128FF5C" box="[151,277,187,213]" captionStart-0="FIGURES 3338" captionStart-1="FIGURES 3942" captionStart-2="FIGURES 4346" captionStartId-0="8.[151,264,1903,1927]" captionStartId-1="9.[151,264,1163,1187]" captionStartId-2="10.[151,264,1453,1477]" captionTargetBox-0="[174,1412,484,1879]" captionTargetBox-1="[151,1435,181,1139]" captionTargetBox-2="[154,1431,600,1420]" captionTargetId-0="figure-114@8.[174,1412,484,1879]" captionTargetId-1="figure-30@9.[151,1435,181,1139]" captionTargetId-2="figure-213@10.[151,1436,597,1430]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURES 3338. Nepalella labolensis sp. nov., male holotype (ZMUM). 33. Tarsal papillae of leg 3. 34. Leg 3, frontal view. 35. Claw of leg 11, caudal view. 36. Femoral protuberance of leg 3. 37. Basal part of leg pair 11, caudal view. 38. Leg pair 11, caudal view. Abbreviations: d, digitiform process; f, femoral protuberance. Scales: 10 μm (Fig. 38), 20 μm (Figs 33, 35, 36), 100 μm (Figs 34, 37)." captionText-1="FIGURES 3942. Nepalella labolensis sp. nov., male holotype (ZMUM). 39. Basal part of leg pair 10, caudal view. 40.Anterior gonopods, frontal view (apex of coxite on the left broken off). 41. Posterior gonopods, frontal view. 42. Posterior gonopods, caudal view. Abbreviations: a, angiocoxite; b, bush-like structure; bb, basal blade; bl, anterior colpocoxital blade; cxi, coxite; lp, flattened process; mb, mesal blade; ml, median lobe; mp, mesal process. Scales: 100 μm." captionText-2="FIGURES 4346. Nepalella labolensis sp. nov., male holotype (ZMUM). 43, 44. Anterior gonopods, caudal and frontal views, respectively. 45. Posterior gonopods, caudal view. 46. Posterior gonopods, frontal view. Abbreviations: a, angiocoxite; b, bush-like structure; bb, basal blade; bl, anterior colpocoxital blade; lp, flattened process; mb, mesal blade; mp, mesal process. Scales: 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7235139" figureDoi-1="http://doi.org/10.5281/zenodo.7235141" figureDoi-2="http://doi.org/10.5281/zenodo.7235143" httpUri-0="https://zenodo.org/record/7235139/files/figure.png" httpUri-1="https://zenodo.org/record/7235141/files/figure.png" httpUri-2="https://zenodo.org/record/7235143/files/figure.png" pageId="8" pageNumber="396">Figs 3346</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37CCC86FFB4F974FF498E9A01ABFEE9" pageId="8" pageNumber="396" type="materials_examined">
<paragraph id="8BD99F0DFFB4F974FF498E9A01ABFEE9" blockId="8.[151,1436,253,461]" pageId="8" pageNumber="396">
<emphasis id="B912431FFFB4F974FF498E9A01BFFE9E" bold="true" box="[151,386,253,279]" pageId="8" pageNumber="396">Material examined.</emphasis>
<materialsCitation id="3B0E9550FFB4F974FE558E9901ABFEE9" ID-GBIF-Occurrence="3944419304" collectingDate="2015-06-16" collectionCode="ZMUM" collectorName="I. Belousov &amp; I. Kabak &amp; G. Davidian" country="China" elevation="3890" location="Laboluo Village" municipality="Luozhuahe" pageId="8" pageNumber="396" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54DD21AFFFB4F974FE558E9901C5FE91" box="[395,504,254,280]" pageId="8" pageNumber="396" type="holotype">Holotype</typeStatus>
: male (
<collectionCode id="ED7707C8FFB4F974FD958E990295FE91" box="[587,680,254,280]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="8" pageNumber="396">ZMUM</collectionCode>
),
<collectingCountry id="F371DF9DFFB4F974FD638E99033EFE91" box="[701,771,254,280]" name="China" pageId="8" pageNumber="396">China</collectingCountry>
,
<collectingRegion id="49A251EFFFB4F974FCD08E990359FE91" box="[782,868,254,280]" country="China" name="Yunnan" pageId="8" pageNumber="396">Yunnan</collectingRegion>
[Province],
<collectorName id="2693FADBFFB4F974FC2D8E990469FE91" box="[1011,1108,254,280]" pageId="8" pageNumber="396">N Weixi</collectorName>
, left tributary of
<collectingMunicipality id="6BBD0577FFB4F974FAC78E9905A5FE91" box="[1305,1432,254,280]" pageId="8" pageNumber="396">Luozhuahe</collectingMunicipality>
, [Yezhi Town], 6.45 km
<collectorName id="2693FADBFFB4F974FE7A8F4502AFFEB5" box="[420,658,290,316]" pageId="8" pageNumber="396">
NW
<location id="8EB9C9D6FFB4F974FE078F4502AFFEB5" LSID="urn:lsid:plazi:treatment:03CF2E1BFFB4F976FF498EF004ACFDB4:8EB9C9D6FFB4F974FE078F4502AFFEB5" box="[473,658,290,316]" country="China" municipality="Luozhuahe" name="Laboluo Village" pageId="8" pageNumber="396" stateProvince="Yunnan">Laboluo Village</location>
</collectorName>
, 27˚4349 N, 98˚5653 E, H=
<quantity id="4C9E32E8FFB4F974FC2C8F450476FEB5" box="[1010,1099,290,316]" metricMagnitude="3" metricUnit="m" metricValue="3.8899999999999997" pageId="8" pageNumber="396" unit="m" value="3890.0">
<elevation id="004B783EFFB4F974FC2C8F450476FEB5" box="[1010,1099,290,316]" metricMagnitude="3" metricUnit="m" metricValue="3.8899999999999997" pageId="8" pageNumber="396" unit="m" value="3890.0">3890 m</elevation>
</quantity>
,
<date id="FFD8B9CDFFB4F974FB898F4504E4FEB4" box="[1111,1241,290,317]" pageId="8" pageNumber="396" value="2015-06-16">
<collectingDate id="EF9C4025FFB4F974FB898F4504E4FEB4" box="[1111,1241,290,317]" pageId="8" pageNumber="396" value="2015-06-16">16.06.2015</collectingDate>
</date>
, leg.
<collectorName id="2693FADBFFB4F974FACB8F4505A5FEB5" box="[1301,1432,290,316]" pageId="8" pageNumber="396">I. Belousov</collectorName>
, I.
<collectorName id="2693FADBFFB4F974FF718F21011CFEE9" box="[175,289,326,352]" pageId="8" pageNumber="396">Kabak, G.</collectorName>
Davidian.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37CCC86FFB4F974FF198F0E052FFE45" pageId="8" pageNumber="396" type="diagnosis">
<paragraph id="8BD99F0DFFB4F974FF198F0E052FFE45" blockId="8.[151,1436,253,461]" pageId="8" pageNumber="396">
<emphasis id="B912431FFFB4F974FF198F0E017FFE0A" bold="true" box="[199,322,361,387]" pageId="8" pageNumber="396">Diagnosis.</emphasis>
Differs from congeners mainly by the posterior gonopod colpocoxite with a long, apical, flattened and caudally curved process (
<emphasis id="B912431FFFB4F974FE388FE901C0FE21" bold="true" box="[486,509,398,424]" pageId="8" pageNumber="396">lp</emphasis>
) couple with a subapical, thin, somewhat curved, mesal process (
<emphasis id="B912431FFFB4F974FB3C8FE90534FE21" bold="true" box="[1250,1289,398,424]" pageId="8" pageNumber="396">mp</emphasis>
) and a bushlike structure (
<emphasis id="B912431FFFB4F974FEE48FD50177FE45" bold="true" box="[314,330,434,460]" pageId="8" pageNumber="396">b</emphasis>
) occupying the entire length of the colpocoxite posterior surface (
<figureCitation id="135D8388FFB4F974FBF08FD5048DFE45" box="[1070,1200,434,460]" captionStart="FIGURES 3942" captionStartId="9.[151,264,1163,1187]" captionTargetBox="[151,1435,181,1139]" captionTargetId="figure-30@9.[151,1435,181,1139]" captionTargetPageId="9" captionText="FIGURES 3942. Nepalella labolensis sp. nov., male holotype (ZMUM). 39. Basal part of leg pair 10, caudal view. 40.Anterior gonopods, frontal view (apex of coxite on the left broken off). 41. Posterior gonopods, frontal view. 42. Posterior gonopods, caudal view. Abbreviations: a, angiocoxite; b, bush-like structure; bb, basal blade; bl, anterior colpocoxital blade; cxi, coxite; lp, flattened process; mb, mesal blade; ml, median lobe; mp, mesal process. Scales: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7235141" httpUri="https://zenodo.org/record/7235141/files/figure.png" pageId="8" pageNumber="396">Figs 41, 42</figureCitation>
,
<figureCitation id="135D8388FFB4F974FB658FD50538FE45" box="[1211,1285,434,461]" captionStart="FIGURES 4346" captionStartId="10.[151,264,1453,1477]" captionTargetBox="[154,1431,600,1420]" captionTargetId="figure-213@10.[151,1436,597,1430]" captionTargetPageId="10" captionText="FIGURES 4346. Nepalella labolensis sp. nov., male holotype (ZMUM). 43, 44. Anterior gonopods, caudal and frontal views, respectively. 45. Posterior gonopods, caudal view. 46. Posterior gonopods, frontal view. Abbreviations: a, angiocoxite; b, bush-like structure; bb, basal blade; bl, anterior colpocoxital blade; lp, flattened process; mb, mesal blade; mp, mesal process. Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7235143" httpUri="https://zenodo.org/record/7235143/files/figure.png" pageId="8" pageNumber="396">45, 46</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF19CF85FFB4F974FF4989080149F87A" ID-DOI="http://doi.org/10.5281/zenodo.7235139" ID-Zenodo-Dep="7235139" httpUri="https://zenodo.org/record/7235139/files/figure.png" pageId="8" pageNumber="396" startId="8.[151,264,1903,1927]" targetBox="[174,1412,484,1879]" targetPageId="8">
<paragraph id="8BD99F0DFFB4F974FF4989080149F87A" blockId="8.[151,1437,1903,2036]" pageId="8" pageNumber="396">
<emphasis id="B912431FFFB4F974FF4989080169F80E" bold="true" box="[151,340,1903,1927]" pageId="8" pageNumber="396">FIGURES 3338.</emphasis>
<taxonomicName id="4C66E48EFFB4F974FE8589080216F80E" authority="Mikhaljova, 2022" authorityName="Mikhaljova" authorityYear="2022" box="[347,555,1903,1927]" class="Diplopoda" family="Lankasomatidae" genus="Nepalella" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="396" phylum="Arthropoda" rank="species" species="labolensis" status="sp. nov.">
<emphasis id="B912431FFFB4F974FE8589080216F80E" box="[347,555,1903,1927]" italics="true" pageId="8" pageNumber="396">Nepalella labolensis</emphasis>
</taxonomicName>
<emphasis id="B912431FFFB4F974FDEF890802BFF80E" bold="true" box="[561,642,1903,1927]" pageId="8" pageNumber="396">
<taxonomicNameLabel id="A221FE64FFB4F974FDEF890802BFF80E" box="[561,642,1903,1927]" pageId="8" pageNumber="396" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype (ZMUM).
<emphasis id="B912431FFFB4F974FC4D89080391F80E" bold="true" box="[915,940,1903,1927]" pageId="8" pageNumber="396">33</emphasis>
. Tarsal papillae of leg 3.
<emphasis id="B912431FFFB4F974FB70890804FAF80E" bold="true" box="[1198,1223,1903,1927]" pageId="8" pageNumber="396">34</emphasis>
. Leg 3, frontal view.
<emphasis id="B912431FFFB4F974FF4989F4008DF822" bold="true" box="[151,176,1939,1963]" pageId="8" pageNumber="396">35</emphasis>
. Claw of leg 11, caudal view.
<emphasis id="B912431FFFB4F974FE3889F401C2F822" bold="true" box="[486,511,1939,1963]" pageId="8" pageNumber="396">36</emphasis>
. Femoral protuberance of leg 3.
<emphasis id="B912431FFFB4F974FC9389F4035BF822" bold="true" box="[845,870,1939,1963]" pageId="8" pageNumber="396">37</emphasis>
. Basal part of leg pair 11, caudal view.
<emphasis id="B912431FFFB4F974FB2589F40529F822" bold="true" box="[1275,1300,1939,1963]" pageId="8" pageNumber="396">38</emphasis>
. Leg pair 11, caudal view.
<emphasis id="B912431FFFB4F974FEC389D001EEF846" bold="true" box="[285,467,1975,1999]" pageId="8" pageNumber="396">Abbreviations: d</emphasis>
, digitiform process;
<emphasis id="B912431FFFB4F974FD7789D0028CF846" bold="true" box="[681,689,1975,1999]" pageId="8" pageNumber="396">f</emphasis>
, femoral protuberance. Scales: 10 μm (Fig. 38), 20 μm (Figs 33, 35, 36), 100 μm (Figs 34, 37).
</paragraph>
</caption>
<caption id="DF19CF85FFB5F975FF498AEC047BFA99" ID-DOI="http://doi.org/10.5281/zenodo.7235141" ID-Zenodo-Dep="7235141" httpUri="https://zenodo.org/record/7235141/files/figure.png" pageId="9" pageNumber="397" startId="9.[151,264,1163,1187]" targetBox="[151,1435,181,1139]" targetPageId="9">
<paragraph id="8BD99F0DFFB5F975FF498AEC047BFA99" blockId="9.[151,1436,1163,1296]" pageId="9" pageNumber="397">
<emphasis id="B912431FFFB5F975FF498AEC016EFB2A" bold="true" box="[151,339,1163,1187]" pageId="9" pageNumber="397">FIGURES 3942.</emphasis>
<taxonomicName id="4C66E48EFFB5F975FE898AEC021BFB2A" authority="Mikhaljova, 2022" authorityName="Mikhaljova" authorityYear="2022" box="[343,550,1163,1187]" class="Diplopoda" family="Lankasomatidae" genus="Nepalella" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="397" phylum="Arthropoda" rank="species" species="labolensis" status="sp. nov.">
<emphasis id="B912431FFFB5F975FE898AEC021BFB2A" box="[343,550,1163,1187]" italics="true" pageId="9" pageNumber="397">Nepalella labolensis</emphasis>
</taxonomicName>
<emphasis id="B912431FFFB5F975FDF58AEC0247FB2A" bold="true" box="[555,634,1163,1187]" pageId="9" pageNumber="397">
<taxonomicNameLabel id="A221FE64FFB5F975FDF58AEC0247FB2A" box="[555,634,1163,1187]" pageId="9" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype (ZMUM).
<emphasis id="B912431FFFB5F975FC5A8AEC03A0FB2A" bold="true" box="[900,925,1163,1187]" pageId="9" pageNumber="397">39</emphasis>
. Basal part of leg pair 10, caudal view.
<emphasis id="B912431FFFB5F975FAFD8AEC0501FB2A" bold="true" box="[1315,1340,1163,1187]" pageId="9" pageNumber="397">40</emphasis>
. Anterior gonopods, frontal view (apex of coxite on the left broken off).
<emphasis id="B912431FFFB5F975FCFD8AC80301FB4E" bold="true" box="[803,828,1199,1223]" pageId="9" pageNumber="397">41</emphasis>
. Posterior gonopods, frontal view.
<emphasis id="B912431FFFB5F975FB768AC804FCFB4E" bold="true" box="[1192,1217,1199,1223]" pageId="9" pageNumber="397">42</emphasis>
. Posterior gonopods, caudal view.
<emphasis id="B912431FFFB5F975FEC58AB401F2FB62" bold="true" box="[283,463,1235,1259]" pageId="9" pageNumber="397">Abbreviations: a</emphasis>
, angiocoxite;
<emphasis id="B912431FFFB5F975FD818AB40250FB62" bold="true" box="[607,621,1235,1259]" pageId="9" pageNumber="397">b</emphasis>
, bush-like structure;
<emphasis id="B912431FFFB5F975FC9B8AB4035CFB62" bold="true" box="[837,865,1235,1259]" pageId="9" pageNumber="397">bb</emphasis>
, basal blade;
<emphasis id="B912431FFFB5F975FC338AB4043FFB62" bold="true" box="[1005,1026,1235,1259]" pageId="9" pageNumber="397">bl</emphasis>
, anterior colpocoxital blade;
<emphasis id="B912431FFFB5F975FAF28AB40576FB62" bold="true" box="[1324,1355,1235,1259]" pageId="9" pageNumber="397">cxi</emphasis>
, coxite;
<emphasis id="B912431FFFB5F975FF498A900091FA86" bold="true" box="[151,172,1271,1295]" pageId="9" pageNumber="397">lp</emphasis>
, flattened process;
<emphasis id="B912431FFFB5F975FEAF8A9001A9FA86" bold="true" box="[369,404,1271,1295]" pageId="9" pageNumber="397">mb</emphasis>
, mesal blade;
<emphasis id="B912431FFFB5F975FDF98A90027EFA86" bold="true" box="[551,579,1271,1295]" pageId="9" pageNumber="397">ml</emphasis>
, median lobe;
<emphasis id="B912431FFFB5F975FD078A9002C1FA86" bold="true" box="[729,764,1271,1295]" pageId="9" pageNumber="397">mp</emphasis>
, mesal process. Scales: 100 μm.
</paragraph>
</caption>
<subSubSection id="C37CCC86FFB5F976FF198B5A01ACFD90" lastPageId="10" lastPageNumber="398" pageId="9" pageNumber="397" type="description">
<paragraph id="8BD99F0DFFB5F975FF198B5A01F5FA4A" blockId="9.[151,1437,1341,2015]" pageId="9" pageNumber="397">
<emphasis id="B912431FFFB5F975FF198B5A0164FADE" bold="true" box="[199,345,1341,1367]" pageId="9" pageNumber="397">Description.</emphasis>
Male. Length in alcohol about
<quantity id="4C9E32E8FFB5F975FD648B5A0336FAD1" box="[698,779,1341,1368]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="9" pageNumber="397" unit="mm" value="27.0">27 mm</quantity>
, width about 3.5 mm with paraterga. Coloration in alcohol bright brown dorsally, with light brown blurry spot on each paratergite. Venter and basal parts of legs pallid. Distal parts of legs marbled brownish. Head and collum marbled brown. Antennae brown. Eye patches black. Posterior gonopod telopodites white.
</paragraph>
<paragraph id="8BD99F0DFFB5F975FF198BAA03DFF96A" blockId="9.[151,1437,1341,2015]" pageId="9" pageNumber="397">
Body with 30 rings (including telson). Head sparsely setose. Antennae very long and slender. Eye patches triangular, with about 30 ommatidia. Collum semicircular. Both collum and ring 2 narrower than head with genae. Ring 2 somewhat wider than collum. Body width gradually increasing until ring 7, body parallel-sided on rings 820(21), thereafter gradually tapering. Paraterga beginning on ring 2, well developed on rings 624, poorly developed on ring 25, reduced on ring 26, onward missing. Metazonital macrochaetae in a transverse row on rings 28 and 29, like an extended (to different degrees) triangle on preceding rings. Caudolateral (exterior after
<bibRefCitation id="EFF7E2FCFFB5F975FA8E88E600EDF936" author="Spelda, J." pageId="9" pageNumber="397" pagination="29 - 48" refId="ref11464" refString="Spelda, J. (2001) Review of the millipede genus Pterygophorosoma Verhoeff, 1897 (Diplopoda, Chordeumatida, Craspedosomatidae). Andrias, 15, 29 - 48." type="journal article" year="2001">Spelda 2001</bibRefCitation>
) macrochaetae longest, anterolateral (median after
<bibRefCitation id="EFF7E2FCFFB5F975FCCE88C203A0F936" author="Spelda, J." box="[784,925,1701,1727]" pageId="9" pageNumber="397" pagination="29 - 48" refId="ref11464" refString="Spelda, J. (2001) Review of the millipede genus Pterygophorosoma Verhoeff, 1897 (Diplopoda, Chordeumatida, Craspedosomatidae). Andrias, 15, 29 - 48." type="journal article" year="2001">Spelda 2001</bibRefCitation>
) and medial (interior after
<bibRefCitation id="EFF7E2FCFFB5F975FB1388C20564F936" author="Spelda, J." box="[1229,1369,1701,1727]" pageId="9" pageNumber="397" pagination="29 - 48" refId="ref11464" refString="Spelda, J. (2001) Review of the millipede genus Pterygophorosoma Verhoeff, 1897 (Diplopoda, Chordeumatida, Craspedosomatidae). Andrias, 15, 29 - 48." type="journal article" year="2001">Spelda 2001</bibRefCitation>
) ones subequal in length. All macrochaetae pointed. Axial suture well-developed.
</paragraph>
<paragraph id="8BD99F0DFFB5F975FF198889042CF8FA" blockId="9.[151,1437,1341,2015]" pageId="9" pageNumber="397">
Legs long and slender. Leg pairs 1 and 2 typically reduced in size, with usual tarsal brushes. Other pregonopodal legs just barely enlarged. Starting from legs 3, walking legs with small, abundant, as funnels with operculum, tarsal papillae (
<figureCitation id="135D8388FFB5F975FEDD89520167F8C6" box="[259,346,1845,1871]" captionStart="FIGURES 3338" captionStartId="8.[151,264,1903,1927]" captionTargetBox="[174,1412,484,1879]" captionTargetId="figure-114@8.[174,1412,484,1879]" captionTargetPageId="8" captionText="FIGURES 3338. Nepalella labolensis sp. nov., male holotype (ZMUM). 33. Tarsal papillae of leg 3. 34. Leg 3, frontal view. 35. Claw of leg 11, caudal view. 36. Femoral protuberance of leg 3. 37. Basal part of leg pair 11, caudal view. 38. Leg pair 11, caudal view. Abbreviations: d, digitiform process; f, femoral protuberance. Scales: 10 μm (Fig. 38), 20 μm (Figs 33, 35, 36), 100 μm (Figs 34, 37)." figureDoi="http://doi.org/10.5281/zenodo.7235139" httpUri="https://zenodo.org/record/7235139/files/figure.png" pageId="9" pageNumber="397">Fig. 33</figureCitation>
); their quantity gradually decrease toward end of body. Legs of midbody with tarsal papillae occupying about 2/3 of the tarsus length. Hindmost legs without tarsal papillae.
</paragraph>
<paragraph id="8BD99F0DFFB5F975FF19891A00D3F856" blockId="9.[151,1437,1341,2015]" pageId="9" pageNumber="397">
Claws invariably long, sabre-shaped. Claws of pregonopodal and postgonopodal legs, including legs 10 and 11, at base with two small additional claws dorsally only (
<figureCitation id="135D8388FFB5F975FCFE89C60349F835" box="[800,884,1953,1980]" captionStart="FIGURES 3338" captionStartId="8.[151,264,1903,1927]" captionTargetBox="[174,1412,484,1879]" captionTargetId="figure-114@8.[174,1412,484,1879]" captionTargetPageId="8" captionText="FIGURES 3338. Nepalella labolensis sp. nov., male holotype (ZMUM). 33. Tarsal papillae of leg 3. 34. Leg 3, frontal view. 35. Claw of leg 11, caudal view. 36. Femoral protuberance of leg 3. 37. Basal part of leg pair 11, caudal view. 38. Leg pair 11, caudal view. Abbreviations: d, digitiform process; f, femoral protuberance. Scales: 10 μm (Fig. 38), 20 μm (Figs 33, 35, 36), 100 μm (Figs 34, 37)." figureDoi="http://doi.org/10.5281/zenodo.7235139" httpUri="https://zenodo.org/record/7235139/files/figure.png" pageId="9" pageNumber="397">Fig. 35</figureCitation>
). Femora 3 and 4 with low protuberance (
<emphasis id="B912431FFFB5F975FA9789C6056FF832" bold="true" box="[1353,1362,1953,1979]" pageId="9" pageNumber="397">f</emphasis>
) (
<figureCitation id="135D8388FFB5F975FAB489C600DCF856" captionStart="FIGURES 3338" captionStartId="8.[151,264,1903,1927]" captionTargetBox="[174,1412,484,1879]" captionTargetId="figure-114@8.[174,1412,484,1879]" captionTargetPageId="8" captionText="FIGURES 3338. Nepalella labolensis sp. nov., male holotype (ZMUM). 33. Tarsal papillae of leg 3. 34. Leg 3, frontal view. 35. Claw of leg 11, caudal view. 36. Femoral protuberance of leg 3. 37. Basal part of leg pair 11, caudal view. 38. Leg pair 11, caudal view. Abbreviations: d, digitiform process; f, femoral protuberance. Scales: 10 μm (Fig. 38), 20 μm (Figs 33, 35, 36), 100 μm (Figs 34, 37)." figureDoi="http://doi.org/10.5281/zenodo.7235139" httpUri="https://zenodo.org/record/7235139/files/figure.png" pageId="9" pageNumber="397">Figs 34, 36</figureCitation>
).
</paragraph>
<paragraph id="8BD99F0DFFB6F976FF198EF00475FF5C" blockId="10.[151,1437,151,573]" pageId="10" pageNumber="398">
Legs 10 and 11 with coxal glands (
<figureCitation id="135D8388FFB6F976FD8A8EF00291FF38" box="[596,684,151,177]" captionStart="FIGURES 3338" captionStartId="8.[151,264,1903,1927]" captionTargetBox="[174,1412,484,1879]" captionTargetId="figure-114@8.[174,1412,484,1879]" captionTargetPageId="8" captionText="FIGURES 3338. Nepalella labolensis sp. nov., male holotype (ZMUM). 33. Tarsal papillae of leg 3. 34. Leg 3, frontal view. 35. Claw of leg 11, caudal view. 36. Femoral protuberance of leg 3. 37. Basal part of leg pair 11, caudal view. 38. Leg pair 11, caudal view. Abbreviations: d, digitiform process; f, femoral protuberance. Scales: 10 μm (Fig. 38), 20 μm (Figs 33, 35, 36), 100 μm (Figs 34, 37)." figureDoi="http://doi.org/10.5281/zenodo.7235139" httpUri="https://zenodo.org/record/7235139/files/figure.png" pageId="10" pageNumber="398">Figs 38</figureCitation>
,
<figureCitation id="135D8388FFB6F976FD668EF002E5FF38" box="[696,728,151,177]" captionStart="FIGURES 3942" captionStartId="9.[151,264,1163,1187]" captionTargetBox="[151,1435,181,1139]" captionTargetId="figure-30@9.[151,1435,181,1139]" captionTargetPageId="9" captionText="FIGURES 3942. Nepalella labolensis sp. nov., male holotype (ZMUM). 39. Basal part of leg pair 10, caudal view. 40.Anterior gonopods, frontal view (apex of coxite on the left broken off). 41. Posterior gonopods, frontal view. 42. Posterior gonopods, caudal view. Abbreviations: a, angiocoxite; b, bush-like structure; bb, basal blade; bl, anterior colpocoxital blade; cxi, coxite; lp, flattened process; mb, mesal blade; ml, median lobe; mp, mesal process. Scales: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7235141" httpUri="https://zenodo.org/record/7235141/files/figure.png" pageId="10" pageNumber="398">39</figureCitation>
). Prefemur 11 with a basal, digitiform, partly papillate process (
<emphasis id="B912431FFFB6F976FF7F8EDC008CFF5C" bold="true" box="[161,177,187,213]" pageId="10" pageNumber="398">d</emphasis>
) (
<figureCitation id="135D8388FFB6F976FF168EDC0176FF5C" box="[200,331,187,213]" captionStart="FIGURES 3338" captionStartId="8.[151,264,1903,1927]" captionTargetBox="[174,1412,484,1879]" captionTargetId="figure-114@8.[174,1412,484,1879]" captionTargetPageId="8" captionText="FIGURES 3338. Nepalella labolensis sp. nov., male holotype (ZMUM). 33. Tarsal papillae of leg 3. 34. Leg 3, frontal view. 35. Claw of leg 11, caudal view. 36. Femoral protuberance of leg 3. 37. Basal part of leg pair 11, caudal view. 38. Leg pair 11, caudal view. Abbreviations: d, digitiform process; f, femoral protuberance. Scales: 10 μm (Fig. 38), 20 μm (Figs 33, 35, 36), 100 μm (Figs 34, 37)." figureDoi="http://doi.org/10.5281/zenodo.7235139" httpUri="https://zenodo.org/record/7235139/files/figure.png" pageId="10" pageNumber="398">Figs 37, 38</figureCitation>
). Sternum 11 frontally with a longitudinal roller widened at its base.
</paragraph>
<paragraph id="8BD99F0DFFB6F976FF198EB80231FE7C" blockId="10.[151,1437,151,573]" pageId="10" pageNumber="398">
Anterior gonopods reduced, sternum with a median lobe (
<emphasis id="B912431FFFB6F976FC8F8EB8034DFF70" bold="true" box="[849,880,223,249]" pageId="10" pageNumber="398">ml</emphasis>
) with the rectangular base and subquadrate apex; coxites (
<emphasis id="B912431FFFB6F976FF278F650126FE95" bold="true" box="[249,283,258,284]" pageId="10" pageNumber="398">cxi</emphasis>
) of medium length, curved, hornlike (
<figureCitation id="135D8388FFB6F976FD0C8F640316FE94" box="[722,811,259,285]" captionStart="FIGURES 3942" captionStartId="9.[151,264,1163,1187]" captionTargetBox="[151,1435,181,1139]" captionTargetId="figure-30@9.[151,1435,181,1139]" captionTargetPageId="9" captionText="FIGURES 3942. Nepalella labolensis sp. nov., male holotype (ZMUM). 39. Basal part of leg pair 10, caudal view. 40.Anterior gonopods, frontal view (apex of coxite on the left broken off). 41. Posterior gonopods, frontal view. 42. Posterior gonopods, caudal view. Abbreviations: a, angiocoxite; b, bush-like structure; bb, basal blade; bl, anterior colpocoxital blade; cxi, coxite; lp, flattened process; mb, mesal blade; ml, median lobe; mp, mesal process. Scales: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7235141" httpUri="https://zenodo.org/record/7235141/files/figure.png" pageId="10" pageNumber="398">Figs 40</figureCitation>
,
<figureCitation id="135D8388FFB6F976FCE68F6403B9FE94" box="[824,900,259,285]" captionStart="FIGURES 4346" captionStartId="10.[151,264,1453,1477]" captionTargetBox="[154,1431,600,1420]" captionTargetId="figure-213@10.[151,1436,597,1430]" captionTargetPageId="10" captionText="FIGURES 4346. Nepalella labolensis sp. nov., male holotype (ZMUM). 43, 44. Anterior gonopods, caudal and frontal views, respectively. 45. Posterior gonopods, caudal view. 46. Posterior gonopods, frontal view. Abbreviations: a, angiocoxite; b, bush-like structure; bb, basal blade; bl, anterior colpocoxital blade; lp, flattened process; mb, mesal blade; mp, mesal process. Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7235143" httpUri="https://zenodo.org/record/7235143/files/figure.png" pageId="10" pageNumber="398">43, 44</figureCitation>
). Posterior gonopod colpocoxite erect, apically with a long, flattened process (
<emphasis id="B912431FFFB6F976FE328F40023EFEC8" bold="true" box="[492,515,295,321]" pageId="10" pageNumber="398">lp</emphasis>
) curved caudolaterad; lateral edge of the process
<emphasis id="B912431FFFB6F976FBF48F40047CFEC8" bold="true" box="[1066,1089,295,321]" pageId="10" pageNumber="398">lp</emphasis>
curved caudad as a blade (
<figureCitation id="135D8388FFB6F976FAB48F40008BFEEC" captionStart="FIGURES 3942" captionStartId="9.[151,264,1163,1187]" captionTargetBox="[151,1435,181,1139]" captionTargetId="figure-30@9.[151,1435,181,1139]" captionTargetPageId="9" captionText="FIGURES 3942. Nepalella labolensis sp. nov., male holotype (ZMUM). 39. Basal part of leg pair 10, caudal view. 40.Anterior gonopods, frontal view (apex of coxite on the left broken off). 41. Posterior gonopods, frontal view. 42. Posterior gonopods, caudal view. Abbreviations: a, angiocoxite; b, bush-like structure; bb, basal blade; bl, anterior colpocoxital blade; cxi, coxite; lp, flattened process; mb, mesal blade; ml, median lobe; mp, mesal process. Scales: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7235141" httpUri="https://zenodo.org/record/7235141/files/figure.png" pageId="10" pageNumber="398">Figs 42</figureCitation>
,
<figureCitation id="135D8388FFB6F976FF1E8F2D00DDFEEC" box="[192,224,330,357]" captionStart="FIGURES 4346" captionStartId="10.[151,264,1453,1477]" captionTargetBox="[154,1431,600,1420]" captionTargetId="figure-213@10.[151,1436,597,1430]" captionTargetPageId="10" captionText="FIGURES 4346. Nepalella labolensis sp. nov., male holotype (ZMUM). 43, 44. Anterior gonopods, caudal and frontal views, respectively. 45. Posterior gonopods, caudal view. 46. Posterior gonopods, frontal view. Abbreviations: a, angiocoxite; b, bush-like structure; bb, basal blade; bl, anterior colpocoxital blade; lp, flattened process; mb, mesal blade; mp, mesal process. Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7235143" httpUri="https://zenodo.org/record/7235143/files/figure.png" pageId="10" pageNumber="398">45</figureCitation>
). The entire length of the colpocoxite posterior surface occupied by a narrow, longitudinal, bush-like structure (
<emphasis id="B912431FFFB6F976FF7F8F08008CFE00" bold="true" box="[161,177,367,393]" pageId="10" pageNumber="398">b</emphasis>
); angiocoxite (
<emphasis id="B912431FFFB6F976FE838F080156FE00" bold="true" box="[349,363,367,393]" pageId="10" pageNumber="398">a</emphasis>
) with a mesal blade (
<emphasis id="B912431FFFB6F976FDBD8F0802B7FE00" bold="true" box="[611,650,367,393]" pageId="10" pageNumber="398">mb</emphasis>
) and basal blade (
<emphasis id="B912431FFFB6F976FC858F080347FE00" bold="true" box="[859,890,367,393]" pageId="10" pageNumber="398">bb</emphasis>
) curved caudad. In frontal view the colpocoxite with a tongue-like, wrinkled blade (
<emphasis id="B912431FFFB6F976FDF38FF40279FE24" bold="true" box="[557,580,403,429]" pageId="10" pageNumber="398">bl</emphasis>
) along its length and a slender subapical mesal process (
<emphasis id="B912431FFFB6F976FB158FF404CFFE24" bold="true" box="[1227,1266,403,429]" pageId="10" pageNumber="398">mp</emphasis>
) (
<figureCitation id="135D8388FFB6F976FAD58FF4055EFE24" box="[1291,1379,403,429]" captionStart="FIGURES 3942" captionStartId="9.[151,264,1163,1187]" captionTargetBox="[151,1435,181,1139]" captionTargetId="figure-30@9.[151,1435,181,1139]" captionTargetPageId="9" captionText="FIGURES 3942. Nepalella labolensis sp. nov., male holotype (ZMUM). 39. Basal part of leg pair 10, caudal view. 40.Anterior gonopods, frontal view (apex of coxite on the left broken off). 41. Posterior gonopods, frontal view. 42. Posterior gonopods, caudal view. Abbreviations: a, angiocoxite; b, bush-like structure; bb, basal blade; bl, anterior colpocoxital blade; cxi, coxite; lp, flattened process; mb, mesal blade; ml, median lobe; mp, mesal process. Scales: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7235141" httpUri="https://zenodo.org/record/7235141/files/figure.png" pageId="10" pageNumber="398">Figs 41</figureCitation>
,
<figureCitation id="135D8388FFB6F976FAAE8FF405ADFE24" box="[1392,1424,403,429]" captionStart="FIGURES 4346" captionStartId="10.[151,264,1453,1477]" captionTargetBox="[154,1431,600,1420]" captionTargetId="figure-213@10.[151,1436,597,1430]" captionTargetPageId="10" captionText="FIGURES 4346. Nepalella labolensis sp. nov., male holotype (ZMUM). 43, 44. Anterior gonopods, caudal and frontal views, respectively. 45. Posterior gonopods, caudal view. 46. Posterior gonopods, frontal view. Abbreviations: a, angiocoxite; b, bush-like structure; bb, basal blade; bl, anterior colpocoxital blade; lp, flattened process; mb, mesal blade; mp, mesal process. Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7235143" httpUri="https://zenodo.org/record/7235143/files/figure.png" pageId="10" pageNumber="398">46</figureCitation>
). Angiocoxite (
<emphasis id="B912431FFFB6F976FEED8FD0017CFE58" bold="true" box="[307,321,439,465]" pageId="10" pageNumber="398">a</emphasis>
) convex, pear-shaped on anterior face. Posterior gonopod telopodites large; telopoditomere 1 very densely setose on its frontal face.
</paragraph>
<paragraph id="8BD99F0DFFB6F976FF198F9801ACFD90" blockId="10.[151,1437,151,573]" box="[199,401,511,537]" pageId="10" pageNumber="398">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C37CCC86FFB6F976FF198C4404ACFDB4" box="[199,1169,547,573]" pageId="10" pageNumber="398" type="etymology">
<paragraph id="8BD99F0DFFB6F976FF198C4404ACFDB4" blockId="10.[151,1437,151,573]" box="[199,1169,547,573]" pageId="10" pageNumber="398">
<emphasis id="B912431FFFB6F976FF198C440173FDB4" bold="true" box="[199,334,547,573]" pageId="10" pageNumber="398">Etymology.</emphasis>
The specific epithet refers to the locus typicus, Laboluo village, adjective.
</paragraph>
</subSubSection>
</treatment>
</document>