treatments-xml/data/03/C8/87/03C887BAFF915C46CC990EE434CEF867.xml

<document id="5E3B2AFB19541A3B74BE088044C000A4" ID-DOI="10.11646/zootaxa.3710.1.3" ID-GBIF-Dataset="722a4393-0aa7-424a-8a29-31f04e75048d" ID-ISSN="1175-5326" ID-Zenodo-Dep="223802" ID-ZooBank="39E00CE7-8897-462D-B367-B2C1E9C3CF9B" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460675573267" checkinUser="plazi" docAuthor="Pérez-González, Sergio, Zaballos, Juan P. &amp; Ghannem, Samir" docDate="2013" docId="03C887BAFF915C46CC990EE434CEF867" docLanguage="en" docName="zt03710p060.pdf" docOrigin="Zootaxa 3710 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Typhlocharis mixta Pérez-González, Zaballos &amp; Ghannem, 2013, n. sp." docType="treatment" docVersion="7" masterDocId="FFF1FFC2FF945C4DCC0E085E3152FFED" masterDocTitle="Intraspecific variation in Typhlocharis Dieck, 1869 (Coleoptera, Carabidae, Anillini): the case of two new species of the baetica group" updateTime="1698349748274" updateUser="plazi">
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<mods:title id="7D632107B2F8D2834C1F32726F778A4F">Intraspecific variation in Typhlocharis Dieck, 1869 (Coleoptera, Carabidae, Anillini): the case of two new species of the baetica group</mods:title>
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<mods:namePart id="958C9A434CC5E5FB91BEBD7E420B37FE">Pérez-González, Sergio</mods:namePart>
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<mods:namePart id="46ECCC4E01B740B58CA4604DC246F1C1">Zaballos, Juan P.</mods:namePart>
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<mods:namePart id="F221F0A36279202FB65BF2CA11DFACA4">Ghannem, Samir</mods:namePart>
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<subSubSection id="C37B6527FF915C48CC990EE431AAF918" pageId="5" type="nomenclature">
<paragraph id="8BDE36ACFF915C48CC990EE43098F939" blockId="5.[151,458,1722,1781]" box="[151,458,1722,1748]" pageId="5">
<heading id="D09681C0FF915C48CC990EE43098F939" bold="true" box="[151,458,1722,1748]" fontSize="11" level="1" pageId="5" reason="1">
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<taxonomicName id="4C614D2FFF915C48CC990EE430D2F939" ID-CoL="59VK8" box="[151,384,1722,1748]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="mixta" status="sp. nov.">
<emphasis id="B915EABEFF915C48CC990EE430D2F939" bold="true" box="[151,384,1722,1748]" italics="true" pageId="5">Typhlocharis mixta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22657C5FF915C48CD860EE43098F939" box="[392,458,1722,1748]" pageId="5" rank="species">n. sp.</taxonomicNameLabel>
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</paragraph>
<paragraph id="8BDE36ACFF915C48CC990E8231AAF918" blockId="5.[151,458,1722,1781]" box="[151,248,1756,1781]" pageId="5">
<figureCitation id="135A2A29FF915C48CC990E82318CF918" box="[151,222,1756,1781]" captionStart="FIGURE 4" captionStartId="7.[151,250,1949,1971]" captionTargetBox="[169,1419,214,1921]" captionTargetId="figure@7.[151,1436,193,1928]" captionTargetPageId="7" captionText="FIGURE 4. Typhlocharis mixta n. sp. – habitus: ventral (male: left half; female: right half) and dorsal (male) views (abdomen not drawn in dorsal view)." httpUri="https://zenodo.org/record/223806/files/figure.png" pageId="5">Figs 4</figureCitation>
, 
<figureCitation id="135A2A29FF915C48CCE40E8231AAF918" box="[234,248,1756,1781]" captionStart="FIGURE 5" captionStartId="8.[151,250,1010,1032]" captionTargetBox="[167,1413,189,974]" captionTargetId="figure@8.[151,1435,173,988]" captionTargetPageId="8" captionText="FIGURE 5. Typhlocharis mixta n. sp. (A, B) Aedeagus: (A) median lobe (dorsal view); (B) median lobe and parameres (lateral view). (C) Female genitalia (ventral view)." httpUri="https://zenodo.org/record/223807/files/figure.png" pageId="5">5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37B6527FF915C48CC990F7A328BF81C" pageId="5" type="materials_examined">
<paragraph id="8BDE36ACFF915C48CC990F7A328BF81C" blockId="5.[151,1437,1828,2033]" pageId="5">
<emphasis id="B915EABEFF915C48CC990F7A3077F8D0" bold="true" box="[151,293,1828,1853]" pageId="5">
<typeStatus id="54DA880EFF915C48CC990F7A3180F8D0" box="[151,210,1828,1853]" pageId="5">Type</typeStatus>
series.
</emphasis>
<typeStatus id="54DA880EFF915C48CD230F7A30CAF8D0" box="[301,408,1828,1853]" pageId="5" type="holotype">Holotype</typeStatus>
, ♂, 
<collectingCountry id="F376763CFF915C48CDC50F7B334EF8D0" box="[459,540,1829,1853]" name="Spain" pageId="5">SPAIN</collectingCountry>
, Badajoz, Puerto de los Reventones (
<quantity id="4C999B49FF915C48CFC40F7B3540F8D0" box="[970,1042,1828,1853]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="5" unit="km" value="11.0">11 km</quantity>
NE Fregenal de la Sierra), 
<date id="FFDF106CFF915C48C95C0F7B3180F88C" pageId="5" value="2012-11-22">
22- 
<quantity id="4C999B49FF915C48C9740F7B3180F88C" metricMagnitude="6" metricUnit="m" metricValue="1.0062555" metricValueMax="2.0125" metricValueMin="1.1E-5" pageId="5" unit="m" value="1006255.5" valueMax="2012500.0" valueMin="11.0">11- 2012</quantity>
</date>
, 500 m (
<geoCoordinate id="EE55506BFF915C48CD300F1730E4F88D" box="[318,438,1864,1889]" direction="north" orientation="latitude" pageId="5" precision="925" value="38.25">38º 15’ N</geoCoordinate>
, 
<geoCoordinate id="EE55506BFF915C48CDCB0F16336CF88D" box="[453,574,1864,1889]" direction="west" orientation="longitude" pageId="5" precision="925" value="-6.5666666">06º 34’ W</geoCoordinate>
<emphasis id="B915EABEFF915C48CE300F173315F88D" box="[574,583,1865,1888]" italics="true" pageId="5">)</emphasis>
, J. P. Zaballos, S. Pérez &amp; S. Ghannem leg. (Coll. J.P. Zaballos, UCM). 
<typeStatus id="54DA880EFF915C48CC990F33305AF868" box="[151,264,1901,1925]" pageId="5" type="paratype">Paratypes</typeStatus>
: 659♂♂, 616 ♀♀ same data as 
<typeStatus id="54DA880EFF915C48CE7E0F323387F868" box="[624,725,1900,1925]" pageId="5" type="holotype">holotype</typeStatus>
; 
<specimenCount id="9D67FD25FF915C48CEEC0F333257F868" box="[738,773,1901,1925]" pageId="5" type="generic">1♂</specimenCount>
same locality as 
<typeStatus id="54DA880EFF915C48CFC70F32357CF868" box="[969,1070,1900,1925]" pageId="5" type="holotype">holotype</typeStatus>
, 
<date id="FFDF106CFF915C48C8370F3335EDF868" box="[1081,1215,1901,1925]" pageId="5" value="2012-04-24">24-04-2012</date>
, J. P. Zaballos &amp; S. Pérez leg.; 10♂♂, 11 ♀♀ 
<collectingCountry id="F376763CFF915C48CDB70FCF3358F844" box="[441,522,1937,1961]" name="Spain" pageId="5">SPAIN</collectingCountry>
, Badajoz, Valverde de Burguillos (
<quantity id="4C999B49FF915C48CF830FCF328EF844" box="[909,988,1936,1961]" metricMagnitude="3" metricUnit="m" metricValue="4.4" pageId="5" unit="km" value="4.4">4.4 km</quantity>
NE), 
<date id="FFDF106CFF915C48C8110FCF35F6F844" box="[1055,1188,1937,1961]" pageId="5" value="2012-11-22">
22- 
<quantity id="4C999B49FF915C48C84B0FCF35F6F844" box="[1093,1188,1937,1961]" metricMagnitude="6" metricUnit="m" metricValue="1.0062494999999998" metricValueMax="2.012488" metricValueMin="1.1E-5" pageId="5" unit="m" value="1006249.5" valueMax="2012488.0" valueMin="11.0">11-2012</quantity>
</date>
, 488m (
<geoCoordinate id="EE55506BFF915C48C8F30FCF343EF845" box="[1277,1388,1937,1961]" direction="north" orientation="latitude" pageId="5" precision="925" value="38.35">38º 21’ N</geoCoordinate>
, 
<geoCoordinate id="EE55506BFF915C48C9790FCE31B0F820" direction="west" orientation="longitude" pageId="5" precision="925" value="-6.5166664">06º 31’ W</geoCoordinate>
), J. P. Zaballos, S. Pérez &amp; S. Ghannem leg. (Coll. J.P. Zaballos and Coll. S. Pérez-González, UCM). 4♂♂, 4 ♀♀ same data as 
<typeStatus id="54DA880EFF915C48CD7F0F863081F81C" box="[369,467,2008,2033]" pageId="5" type="holotype">holotype</typeStatus>
(MNCN, Madrid, Catalogue number: 11612).
</paragraph>
</subSubSection>
<subSubSection id="C37B6527FF925C4BCCC908C93519FE88" pageId="6" type="diagnosis">
<paragraph id="8BDE36ACFF925C4BCCC908C93519FE88" blockId="6.[151,1437,151,1905]" pageId="6">
<emphasis id="B915EABEFF925C4BCCC908C93013FF5D" bold="true" box="[199,321,151,176]" pageId="6">Diagnosis.</emphasis>
Small endogean anophthalmous beetle, with narrow and subrectangular body covered by microreticulate integument and scattered pubescence. Vertex with 
<emphasis id="B915EABEFF925C4BCF7108E33543FF38" box="[895,1041,188,213]" italics="true" pageId="6">pars stridens</emphasis>
. Clypeus with a distinctive median tooth. Pronotum with a wide medial hiatus. Elytra faintly serrated in lateral margins and apical margin with three to six pairs of small denticles, highly irregular. Umbilicate series with six setae (4 + 2). Abdomen with ventral foveae only in females. Pseudotetramerous tarsi in all the legs. Aedeagus as in 
<figureCitation id="135A2A29FF925C4BCFBB097932AAFEAD" box="[949,1016,295,320]" captionStart="FIGURE 5" captionStartId="8.[151,250,1010,1032]" captionTargetBox="[167,1413,189,974]" captionTargetId="figure@8.[151,1435,173,988]" captionTargetPageId="8" captionText="FIGURE 5. Typhlocharis mixta n. sp. (A, B) Aedeagus: (A) median lobe (dorsal view); (B) median lobe and parameres (lateral view). (C) Female genitalia (ventral view)." httpUri="https://zenodo.org/record/223807/files/figure.png" pageId="6">Fig. 5</figureCitation>
, ‘plough-shaped’ endophallic sclerite and broad parameres. Female genitalia as in 
<figureCitation id="135A2A29FF925C4BCE840912339FFE88" box="[650,717,332,357]" captionStart="FIGURE 5" captionStartId="8.[151,250,1010,1032]" captionTargetBox="[167,1413,189,974]" captionTargetId="figure@8.[151,1435,173,988]" captionTargetPageId="8" captionText="FIGURE 5. Typhlocharis mixta n. sp. (A, B) Aedeagus: (A) median lobe (dorsal view); (B) median lobe and parameres (lateral view). (C) Female genitalia (ventral view)." httpUri="https://zenodo.org/record/223807/files/figure.png" pageId="6">Fig. 5</figureCitation>
, with subcylindrical spermatheca.
</paragraph>
</subSubSection>
<subSubSection id="C37B6527FF925C4BCCC9093133EFF90D" pageId="6" type="description">
<paragraph id="8BDE36ACFF925C4BCCC9093130CFFD8D" blockId="6.[151,1437,151,1905]" pageId="6">
<emphasis id="B915EABEFF925C4BCCC90931300BFE65" bold="true" box="[199,345,367,392]" pageId="6">Description.</emphasis>
Length 
<quantity id="4C999B49FF925C4BCDB7092E330CFE6A" box="[441,606,368,392]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.49" metricValueMin="1.21" pageId="6" unit="mm" value="1.35" valueMax="1.49" valueMin="1.21">1.21–1.49 mm</quantity>
(males), 
<quantity id="4C999B49FF925C4BCEC509313222FE6A" box="[715,880,367,392]" metricMagnitude="-3" metricUnit="m" metricValue="1.435" metricValueMax="1.6" metricValueMin="1.27" pageId="6" unit="mm" value="1.435" valueMax="1.6" valueMin="1.27">1.27–1.60 mm</quantity>
(females) (
<figureCitation id="135A2A29FF925C4BCFFD09313568FE65" box="[1011,1082,367,392]" captionStart="FIGURE 4" captionStartId="7.[151,250,1949,1971]" captionTargetBox="[169,1419,214,1921]" captionTargetId="figure@7.[151,1436,193,1928]" captionTargetPageId="7" captionText="FIGURE 4. Typhlocharis mixta n. sp. – habitus: ventral (male: left half; female: right half) and dorsal (male) views (abdomen not drawn in dorsal view)." httpUri="https://zenodo.org/record/223806/files/figure.png" pageId="6">Fig. 4</figureCitation>
). Head: wider (
<quantity id="4C999B49FF925C4BC8E2092E34C6FE65" box="[1260,1428,368,392]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" metricValueMax="3.4" metricValueMin="2.4" pageId="6" unit="mm" value="0.29" valueMax="0.34" valueMin="0.24">0.24–0.34 mm</quantity>
) than long (
<quantity id="4C999B49FF925C4BCD1B09CA3093FE41" box="[277,449,404,428]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" metricValueMax="3.0" metricValueMin="2.2" pageId="6" unit="mm" value="0.26" valueMax="0.3" valueMin="0.22">0.22–0.30 mm</quantity>
), labrum subrectangular to slightly notched, rounded edges and triangular sclerotized area with middle button. Clypeus with a median tooth in the anterior margin. Right mandible with a prominent, pointy, terebral tooth on inner margin. Left mandible without teeth, but inner margin with sharp edge. Ligula with very prominent middle lobe and long paraglossae. Wide gula, approximately twice as long as wide, the lateral sutures may appear fainter. Rest of cephalic characters and cephalic chaetotaxy, including antennae, as described for 
<taxonomicName id="4C614D2FFF925C4BCCB00A17300AFD8D" box="[190,344,583,608]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="acutangula" status="sp. nov.">
<emphasis id="B915EABEFF925C4BCCB00A17300AFD8D" box="[190,344,583,608]" italics="true" pageId="6">T. acutangula</emphasis>
</taxonomicName>
<emphasis id="B915EABEFF925C4BCD510A1930CFFD8D" bold="true" box="[351,413,583,608]" pageId="6">
<taxonomicNameLabel id="A22657C5FF925C4BCD510A1930CFFD8D" box="[351,413,583,608]" pageId="6" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BDE36ACFF925C4BCCC90A3332B7FC6D" blockId="6.[151,1437,151,1905]" pageId="6">
Pronotum: subrectangular, longer (
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) than wide (
<quantity id="4C999B49FF925C4BCF950A32357EFD69" box="[923,1068,620,644]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="6" unit="mm" value="0.35" valueMax="0.4" valueMin="0.3">0.3–0.4 mm</quantity>
), slightly narrowed in posterior region. Proportions of pronotum present a wide range of variation (
<figureCitation id="135A2A29FF925C4BCF9E0AD13287FD45" box="[912,981,655,680]" captionStart="FIGURE 6" captionStartId="9.[151,250,1091,1113]" captionTargetBox="[152,1432,199,1090]" captionTargetId="figure@9.[148,1433,183,1090]" captionTargetPageId="9" captionText="FIGURE 6. Range of intraspecific variation in pronotum of Typhlocharis mixta n. sp.: 1 – 6 main morphotypes, 7 – 10 asymmetries, malformations and teratologies: 1, 2, subrectangular pronotum with smoothly curved lateral margins and well marked denticles; 3, 4, rectangular pronotum, with straight lateral margins and high number of well marked denticles; 5, 6, subsquare, with smoothly curved lateral margins and low number of blunt denticles; 7, Asymmetric posterior angles, note the different aspect of pointy and blunt angles; 8, symmetrically widened lateral margins; 9, asymmetric malformed margins; 10, symmetrically notched lateral margins." httpUri="https://zenodo.org/record/223808/files/figure.png" pageId="6">Fig. 6</figureCitation>
). Wide medial hiatus (moderately wider than three adjacent intersetae spaces). Lateral margins with denticles near posterior angles highly variable in shape and number, from two or three, irregular and faint to four or five, well marked and serrated. The posterior angles are also variable from pointy and protruding to blunt and low (
<figureCitation id="135A2A29FF925C4BCF430AA232C2FCF8" box="[845,912,764,789]" captionStart="FIGURE 6" captionStartId="9.[151,250,1091,1113]" captionTargetBox="[152,1432,199,1090]" captionTargetId="figure@9.[148,1433,183,1090]" captionTargetPageId="9" captionText="FIGURE 6. Range of intraspecific variation in pronotum of Typhlocharis mixta n. sp.: 1 – 6 main morphotypes, 7 – 10 asymmetries, malformations and teratologies: 1, 2, subrectangular pronotum with smoothly curved lateral margins and well marked denticles; 3, 4, rectangular pronotum, with straight lateral margins and high number of well marked denticles; 5, 6, subsquare, with smoothly curved lateral margins and low number of blunt denticles; 7, Asymmetric posterior angles, note the different aspect of pointy and blunt angles; 8, symmetrically widened lateral margins; 9, asymmetric malformed margins; 10, symmetrically notched lateral margins." httpUri="https://zenodo.org/record/223808/files/figure.png" pageId="6">Fig. 6</figureCitation>
). Other characters and chaetotaxy are the same as for 
<taxonomicName id="4C614D2FFF925C4BCCD20B7F3024FCD5" box="[220,374,799,824]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="acutangula" status="sp. nov.">
<emphasis id="B915EABEFF925C4BCCD20B7F3024FCD5" box="[220,374,799,824]" italics="true" pageId="6">T. acutangula</emphasis>
</taxonomicName>
<emphasis id="B915EABEFF925C4BCD720B4130E9FCD5" bold="true" box="[380,443,799,824]" pageId="6">
<taxonomicNameLabel id="A22657C5FF925C4BCD720B4130E9FCD5" box="[380,443,799,824]" pageId="6" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, except a pattern of three pairs of setae, parallel to posterior margin 
<emphasis id="B915EABEFF925C4BC8B90B41346BFCD5" bold="true" box="[1207,1337,799,824]" pageId="6">[s-l-l / l-l-s]</emphasis>
, the row of small, thin setae of anterior and posterior margins (filiform) and the short discal pubescence, arranged in four pairs of longitudinal rows. Prosternum as described for 
<taxonomicName id="4C614D2FFF925C4BCF0B0B3732CDFC6D" box="[773,927,871,896]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="acutangula" status="sp. nov.">
<emphasis id="B915EABEFF925C4BCF0B0B3732CDFC6D" box="[773,927,871,896]" italics="true" pageId="6">T. acutangula</emphasis>
</taxonomicName>
<emphasis id="B915EABEFF925C4BCFA80B3932B7FC6D" bold="true" box="[934,997,871,896]" pageId="6">
<taxonomicNameLabel id="A22657C5FF925C4BCFA80B3932B7FC6D" box="[934,997,871,896]" pageId="6" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BDE36ACFF925C4BCCC90BD23009FB28" blockId="6.[151,1437,151,1905]" pageId="6">
Elytra: approximately two times longer (
<quantity id="4C999B49FF925C4BCEA20BD2320AFC49" box="[684,856,908,933]" metricMagnitude="-4" metricUnit="m" metricValue="7.45" metricValueMax="8.5" metricValueMin="6.4" pageId="6" unit="mm" value="0.745" valueMax="0.85" valueMin="0.64">0.64–0.85 mm</quantity>
) than wide (
<quantity id="4C999B49FF925C4BCFF80BD235F6FC49" box="[1014,1188,908,932]" metricMagnitude="-4" metricUnit="m" metricValue="3.65" metricValueMax="4.1" metricValueMin="3.2" pageId="6" unit="mm" value="0.365" valueMax="0.41" valueMin="0.32">0.32–0.41 mm</quantity>
), subparallel. Lateral margins with 18–21 subtriangular denticles, faint and progressively smoother towards posterior, but still marked in last third. Apical margin smoothly rounded, with three to six pairs of small denticles (showing a notable range of individual variation, but mostly combinations of four and five pairs) (
<figureCitation id="135A2A29FF925C4BCFA40BA932BFFBFD" box="[938,1005,1015,1040]" captionStart="FIGURE 7" captionStartId="10.[151,250,1860,1882]" captionTargetBox="[186,1413,214,1834]" captionTargetId="figure@10.[151,1435,193,1854]" captionTargetPageId="10" captionText="FIGURE 7. Range of intraspecific variation in elytra of Typhlocharis mixta n. sp.: (A) Elytral apex: 1 – 3 general morphotypes: 1, lower number of denticles; 2, higher number of denticles; 3, enlarged inner pair of denticles; 4 – 6 double or triple denticles: 4, large double denticles; 5, bifid, fused double denticle; 6, fused triple denticle; 7 – 9, strong asymmetries. (B) Graphic showing the abundance of every observed combination of apical denticles, expressed as “ denticles in left elytra / denticles in right elytra ”. Note the majority of the series present combinations of four and five pairs of denticles. “? ” indicates missing data (e. g. damaged elytra). Total number of specimens: 1306." httpUri="https://zenodo.org/record/223809/files/figure.png" pageId="6">Fig. 7</figureCitation>
A, B). Transverse scutellar organ with straight to slightly subtriangular margin and pair of ‘buttonholes’ present near base of elytra. Chaetotaxy: umbilicate series formed by anterior group with four setae and posterior group with two setae (
<figureCitation id="135A2A29FF925C4BC8CF0C613455FBB5" box="[1217,1287,1087,1112]" captionStart="FIGURE 4" captionStartId="7.[151,250,1949,1971]" captionTargetBox="[169,1419,214,1921]" captionTargetId="figure@7.[151,1436,193,1928]" captionTargetPageId="7" captionText="FIGURE 4. Typhlocharis mixta n. sp. – habitus: ventral (male: left half; female: right half) and dorsal (male) views (abdomen not drawn in dorsal view)." httpUri="https://zenodo.org/record/223806/files/figure.png" pageId="6">Fig. 4</figureCitation>
). One pair of scutellar setae. No apparent discal setae, but the population of Puerto de los Reventones is prone to have one pair of anterior “pseudodiscals” (i.e. pubescent setae longer than the surrounding pubescence). Rest of characters as in 
<taxonomicName id="4C614D2FFF925C4BC9890CD73047FB28" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="acutangula" status="sp. nov.">
<emphasis id="B915EABEFF925C4BC9890CD73047FB28" italics="true" pageId="6">T. acutangula</emphasis>
</taxonomicName>
<emphasis id="B915EABEFF925C4BCD120CF23009FB28" bold="true" box="[284,347,1196,1221]" pageId="6">
<taxonomicNameLabel id="A22657C5FF925C4BCD120CF23009FB28" box="[284,347,1196,1221]" pageId="6" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BDE36ACFF925C4BCCC90C8E33F2FA95" blockId="6.[151,1437,151,1905]" pageId="6">Legs: similar in both sexes. Intermetacoxal space not widened. Metacoxae with smoothly rounded “flap”. Rounded metatrochanters, metatibiae with moderately dilated distal region and a short perpendicular seta in the inner side. Inner side of pro- and mesofemora covered with small protuberances, absent in metafemora. Pseudotetramerous tarsi in all the legs, tarsomeres 4º and 5º retain a faint articular membrane, but are partially fused and not capable of a normal articulation.</paragraph>
<paragraph id="8BDE36ACFF925C4BCCC90DDA33EFF90D" blockId="6.[151,1437,151,1905]" pageId="6">
Abdomen: as described for 
<taxonomicName id="4C614D2FFF925C4BCDF50DDB33C6FA70" box="[507,660,1412,1437]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="acutangula" status="sp. nov.">
<emphasis id="B915EABEFF925C4BCDF50DDB33C6FA70" box="[507,660,1412,1437]" italics="true" pageId="6">T. acutangula</emphasis>
</taxonomicName>
<emphasis id="B915EABEFF925C4BCE950DDA338BFA70" bold="true" box="[667,729,1412,1437]" pageId="6">
<taxonomicNameLabel id="A22657C5FF925C4BCE950DDA338BFA70" box="[667,729,1412,1437]" pageId="6" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, but with deep ventral foveae in the first and second ventrite of females, absent in males. Last ventrite with a sexually dimorphic pattern of five to six pairs of setae in posterior margin: 
<emphasis id="B915EABEFF925C4BCCF00D923375FA08" bold="true" box="[254,551,1484,1509]" pageId="6">l-s-s-l-s-m / m-(s)-l-s-s-l</emphasis>
(
<figureCitation id="135A2A29FF925C4BCE4E0D9233DDFA09" box="[576,655,1484,1509]" captionStart="FIGURE 4" captionStartId="7.[151,250,1949,1971]" captionTargetBox="[169,1419,214,1921]" captionTargetId="figure@7.[151,1436,193,1928]" captionTargetPageId="7" captionText="FIGURE 4. Typhlocharis mixta n. sp. – habitus: ventral (male: left half; female: right half) and dorsal (male) views (abdomen not drawn in dorsal view)." httpUri="https://zenodo.org/record/223806/files/figure.png" pageId="6">Fig. 4</figureCitation>
). Aedeagus: sickle-shaped median lobe (length: 
<quantity id="4C999B49FF925C4BC8E50D923409FA09" box="[1259,1371,1484,1508]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="6" unit="mm" value="0.22">0.22 mm</quantity>
) and “bottleneck-shaped”, smoothly rounded apex, slightly bent to right (anatomically oriented) in dorsal view (
<figureCitation id="135A2A29FF925C4BC9610DB131F7F9C0" captionStart="FIGURE 5" captionStartId="8.[151,250,1010,1032]" captionTargetBox="[167,1413,189,974]" captionTargetId="figure@8.[151,1435,173,988]" captionTargetPageId="8" captionText="FIGURE 5. Typhlocharis mixta n. sp. (A, B) Aedeagus: (A) median lobe (dorsal view); (B) median lobe and parameres (lateral view). (C) Female genitalia (ventral view)." httpUri="https://zenodo.org/record/223807/files/figure.png" pageId="6">Fig. 5</figureCitation>
A,B). Endophallus with ‘plough-shaped’ sclerites, as drawn in 
<figureCitation id="135A2A29FF925C4BCF6B0E4A32F5F9C0" box="[869,935,1556,1581]" captionStart="FIGURE 5" captionStartId="8.[151,250,1010,1032]" captionTargetBox="[167,1413,189,974]" captionTargetId="figure@8.[151,1435,173,988]" captionTargetPageId="8" captionText="FIGURE 5. Typhlocharis mixta n. sp. (A, B) Aedeagus: (A) median lobe (dorsal view); (B) median lobe and parameres (lateral view). (C) Female genitalia (ventral view)." httpUri="https://zenodo.org/record/223807/files/figure.png" pageId="6">Fig. 5</figureCitation>
B. Short and broad parameres, both with two medium-sized apical setae (
<figureCitation id="135A2A29FF925C4BCDDD0E69334BF9BD" box="[467,537,1591,1616]" captionStart="FIGURE 5" captionStartId="8.[151,250,1010,1032]" captionTargetBox="[167,1413,189,974]" captionTargetId="figure@8.[151,1435,173,988]" captionTargetPageId="8" captionText="FIGURE 5. Typhlocharis mixta n. sp. (A, B) Aedeagus: (A) median lobe (dorsal view); (B) median lobe and parameres (lateral view). (C) Female genitalia (ventral view)." httpUri="https://zenodo.org/record/223807/files/figure.png" pageId="6">Fig. 5</figureCitation>
B). Female genitalia adjusts to the description of Vigna-Taglianti (1972). Long tubular gonocoxites, with double apical setae. Lateral setae absent, but lateral pores in the gonocoxites. Short gonoduct with two regions, thinner proximally (diameter 
<quantity id="4C999B49FF925C4BCF3F0EDE32FBF975" box="[817,937,1664,1688]" metricMagnitude="-6" metricUnit="m" metricValue="3.0" pageId="6" unit="mm" value="0.003">0.003 mm</quantity>
) and thicker distally (diameter 
<quantity id="4C999B49FF925C4BC9190EDE34C2F975" box="[1303,1424,1664,1688]" metricMagnitude="-6" metricUnit="m" metricValue="8.0" pageId="6" unit="mm" value="0.008">0.008 mm</quantity>
). Spermatheca subcylindrical and short, “barrel-shaped” (length: 
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). Conical spermathecal gland (length: 
<quantity id="4C999B49FF925C4BCC990E96305FF90D" box="[151,269,1736,1760]" metricMagnitude="-5" metricUnit="m" metricValue="1.4" pageId="6" unit="mm" value="0.014">0.014 mm</quantity>
), sclerotized in distal region (
<figureCitation id="135A2A29FF925C4BCE590E9933CBF90D" box="[599,665,1735,1760]" captionStart="FIGURE 5" captionStartId="8.[151,250,1010,1032]" captionTargetBox="[167,1413,189,974]" captionTargetId="figure@8.[151,1435,173,988]" captionTargetPageId="8" captionText="FIGURE 5. Typhlocharis mixta n. sp. (A, B) Aedeagus: (A) median lobe (dorsal view); (B) median lobe and parameres (lateral view). (C) Female genitalia (ventral view)." httpUri="https://zenodo.org/record/223807/files/figure.png" pageId="6">Fig. 5</figureCitation>
C).
</paragraph>
</subSubSection>
<subSubSection id="C37B6527FF925C45CCC90EB23060FA61" lastPageId="8" pageId="6" type="etymology">
<paragraph id="8BDE36ACFF925C4BCCC90EB23401F89D" blockId="6.[151,1437,151,1905]" pageId="6">
<emphasis id="B915EABEFF925C4BCCC90EB2301CF8E8" bold="true" box="[199,334,1772,1797]" pageId="6">Etymology.</emphasis>
The name of the species plays a double meaning, referring to the high intraspecific variation that gives a mixture of body shapes and sizes to the populations. On the other hand, it refers to the presence of certain morphologic traits (such as the tooth in clypeus, the trend to diffuse gula or the pseudotetramery) unexpected for members of 
<taxonomicName id="4C614D2FFF925C4BCD2D0F093027F89D" box="[291,373,1879,1904]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="baetica">
<emphasis id="B915EABEFF925C4BCD2D0F093027F89D" box="[291,373,1879,1904]" italics="true" pageId="6">baetica</emphasis>
</taxonomicName>
group, but known in other species groups, like mixed pieces in different jigsaw puzzles.
</paragraph>
<caption id="DF1E6624FF935C4ACC990FC330F0F83F" httpUri="https://zenodo.org/record/223806/files/figure.png" pageId="7" targetBox="[169,1419,214,1921]" targetPageId="7">
<paragraph id="8BDE36ACFF935C4ACC990FC330F0F83F" blockId="7.[151,1436,1949,2002]" pageId="7">
<emphasis id="B915EABEFF935C4ACC990FC33041F85E" bold="true" box="[151,275,1949,1971]" pageId="7">FIGURE 4.</emphasis>
<taxonomicName id="4C614D2FFF935C4ACD170FC3308EF859" box="[281,476,1949,1972]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="mixta" status="sp. nov.">
<emphasis id="B915EABEFF935C4ACD170FC3308EF859" box="[281,476,1949,1972]" italics="true" pageId="7">Typhlocharis mixta</emphasis>
</taxonomicName>
<emphasis id="B915EABEFF935C4ACDED0FC0334EF85E" bold="true" box="[483,540,1950,1971]" pageId="7">
<taxonomicNameLabel id="A22657C5FF935C4ACDED0FC0334EF85E" box="[483,540,1950,1971]" pageId="7" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
– habitus: ventral (male: left half; female: right half) and dorsal (male) views (abdomen not drawn in dorsal view).
</paragraph>
</caption>
<caption id="DF1E6624FF9C5C45CC990BAC331BFBCA" httpUri="https://zenodo.org/record/223807/files/figure.png" pageId="8" targetBox="[167,1413,189,974]" targetPageId="8">
<paragraph id="8BDE36ACFF9C5C45CC990BAC331BFBCA" blockId="8.[151,1435,1010,1063]" pageId="8">
<emphasis id="B915EABEFF9C5C45CC990BAC3040FBEA" bold="true" box="[151,274,1010,1032]" pageId="8">FIGURE 5.</emphasis>
<taxonomicName id="4C614D2FFF9C5C45CD160BAC3088FBE5" box="[280,474,1010,1032]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="mixta" status="sp. nov.">
<emphasis id="B915EABEFF9C5C45CD160BAC3088FBE5" box="[280,474,1010,1032]" italics="true" pageId="8">Typhlocharis mixta</emphasis>
</taxonomicName>
<emphasis id="B915EABEFF9C5C45CDEE0BAC334AFBEA" bold="true" box="[480,536,1010,1031]" pageId="8">
<taxonomicNameLabel id="A22657C5FF9C5C45CDEE0BAC334AFBEA" box="[480,536,1010,1031]" pageId="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(A, B) Aedeagus: (A) median lobe (dorsal view); (B) median lobe and parameres (lateral view). (C) Female genitalia (ventral view).
</paragraph>
</caption>
<paragraph id="8BDE36ACFF9C5C45CCC90C0D3060FA61" blockId="8.[151,1436,1107,2032]" pageId="8">
<emphasis id="B915EABEFF9C5C45CCC90C0D3078FB81" bold="true" box="[199,298,1107,1132]" pageId="8">Habitat.</emphasis>
<taxonomicName id="4C614D2FFF9C5C45CD3B0C0D3341FB81" box="[309,531,1107,1132]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="mixta" status="sp. nov.">
<emphasis id="B915EABEFF9C5C45CD3B0C0D3341FB81" box="[309,531,1107,1132]" italics="true" pageId="8">Typhlocharis mixta</emphasis>
</taxonomicName>
<emphasis id="B915EABEFF9C5C45CE110C0D3331FB81" bold="true" box="[543,611,1107,1132]" pageId="8">
<taxonomicNameLabel id="A22657C5FF9C5C45CE110C0D3331FB81" box="[543,611,1107,1132]" pageId="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is found in two localities separated 
<quantity id="4C999B49FF9C5C45C8170C0A3537FB81" box="[1049,1125,1107,1132]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="8" unit="km" value="13.0">13 km</quantity>
(
<figureCitation id="135A2A29FF9C5C45C8770C0D3592FB81" box="[1145,1216,1107,1132]" captionStart="FIGURE 1" captionStartId="1.[151,250,1654,1676]" captionTargetBox="[151,1436,733,1632]" captionTargetId="figure@1.[151,1436,733,1633]" captionTargetPageId="1" captionText="FIGURE 1. Map of the distribution range of baetica group: 1, location of Typhlocharis acutangula n. sp.; 2, location of Typhlocharis mixta n. sp. Black circles represent the distribution of the other species of the group. Dashed lines indicate political frontiers." httpUri="https://zenodo.org/record/223803/files/figure.png" pageId="8">Fig. 1</figureCitation>
): 1) Puerto de los Reventones, in open prairie environment with low hills and valleys crossed by seasonal streams. Vegetation mainly composed by low herbaceous plants, grasses and thistles, with scattered broom bushes (
<emphasis id="B915EABEFF9C5C45C8A00CC335A4FB59" box="[1198,1270,1181,1204]" italics="true" pageId="8">Retam</emphasis>
Raf.). Higher grasses and rushes (
<taxonomicName id="4C614D2FFF9C5C45CD7B0C9E3091FB3A" box="[373,451,1216,1239]" class="Liliopsida" family="Juncaceae" genus="Juncus" kingdom="Plantae" order="Poales" pageId="8" phylum="Tracheophyta" rank="genus">
<emphasis id="B915EABEFF9C5C45CD7B0C9E3091FB3A" box="[373,451,1216,1239]" italics="true" pageId="8">Juncus</emphasis>
</taxonomicName>
L.) appears near the stream. Dark, brownish soil, with scarce large boulders of gneisses and shales (Proterozoic, Valverde Formation). This location was sampled four times (March, April, June and November), with the best humidity range in the last sampling. 2) Valverde de Burguillos, in a small meadow surrounded by cultures and open holm oak forest. Vegetation is akin to the former locality, with the exception of holm oaks. Brown to reddish soil, rich in clays, with embedded small to medium sized granite boulders (Proterozoic).
</paragraph>
</subSubSection>
<subSubSection id="C37B6527FF9C5C46CCC90DC93434FC48" lastPageId="11" pageId="8" type="description">
<paragraph id="8BDE36ACFF9C5C45CCC90DC93419FA39" blockId="8.[151,1436,1107,2032]" pageId="8">
<emphasis id="B915EABEFF9C5C45CCC90DC9301FFA5D" bold="true" box="[199,333,1431,1456]" pageId="8">Variability.</emphasis>
There are not significant differences between the populations of Puerto de los Reventones (1285 specimens) and Valverde de Burguillos (
<specimenCount id="9D67FD25FF9C5C45CE520DE233ABFA39" box="[604,761,1467,1492]" pageId="8" type="generic">21 specimens</specimenCount>
), both have the same range of intraspecific variation.
</paragraph>
<paragraph id="8BDE36ACFF9C5C45CCC90D813044F989" blockId="8.[151,1436,1107,2032]" pageId="8">
The species is morphologically very diverse in proportions and size, from 
<quantity id="4C999B49FF9C5C45C8290DBE359EFA1A" box="[1063,1228,1503,1528]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="8" unit="mm" value="1.4" valueMax="1.6" valueMin="1.2">1.2 to 1.6 mm</quantity>
in length, but the presence of a tooth in clypeus, umbilicate series 4+2, the perpendicular setae on distal end of metatibiae, pseudotetramerous tarsi, ventral foveae in females, genitalia and the wide hiatus are virtually constant within the population.
</paragraph>
<paragraph id="8BDE36ACFF9C5C45CCC90E31332AF8F5" blockId="8.[151,1436,1107,2032]" pageId="8">
The terebral tooth is usually very prominent but in some specimens is blunt, low or almost absent. The tooth of clypeus ranges from blunt to very pointy (but always present). The lateral sutures of the gula are very faint in about 20% of the series, resembling the “difusse gula” morphology observed in 
<emphasis id="B915EABEFF9C5C45CFDB0EE93526F93D" box="[981,1140,1719,1744]" italics="true" pageId="8">quadridentata</emphasis>
group (Pérez-González &amp; Zaballos 2013c). There is slight variation in the pronouncement of epilobes. Two specimens have a widened 
<emphasis id="B915EABEFF9C5C45C9640E8331A2F8F5" italics="true" pageId="8">pars stridens</emphasis>
, two times wider than the average.
</paragraph>
<paragraph id="8BDE36ACFF9C5C45CCC90F7A3474F81D" blockId="8.[151,1436,1107,2032]" pageId="8">
Pronotum and elytra have the higher rates of variability. Pronotum proportions can be certainly different, with three main morphs: subrectangular, smoothly curved lateral margins and 3-4 well marked posterolateral denticles (
<figureCitation id="135A2A29FF9C5C45CC910F35318CF869" box="[159,222,1899,1924]" captionStart="FIGURE 6" captionStartId="9.[151,250,1091,1113]" captionTargetBox="[152,1432,199,1090]" captionTargetId="figure@9.[148,1433,183,1090]" captionTargetPageId="9" captionText="FIGURE 6. Range of intraspecific variation in pronotum of Typhlocharis mixta n. sp.: 1 – 6 main morphotypes, 7 – 10 asymmetries, malformations and teratologies: 1, 2, subrectangular pronotum with smoothly curved lateral margins and well marked denticles; 3, 4, rectangular pronotum, with straight lateral margins and high number of well marked denticles; 5, 6, subsquare, with smoothly curved lateral margins and low number of blunt denticles; 7, Asymmetric posterior angles, note the different aspect of pointy and blunt angles; 8, symmetrically widened lateral margins; 9, asymmetric malformed margins; 10, symmetrically notched lateral margins." httpUri="https://zenodo.org/record/223808/files/figure.png" pageId="8">Fig 6</figureCitation>
: 1,2), the most common; rectangular, straight lateral margins and 4-5 (even 
<quantity id="4C999B49FF9C5C45C83A0F35350DF869" box="[1076,1119,1899,1924]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="8" unit="in" value="6.0">6 in</quantity>
one specimen) well marked denticles (
<figureCitation id="135A2A29FF9C5C45CD040FD1301BF845" box="[266,329,1935,1960]" captionStart="FIGURE 6" captionStartId="9.[151,250,1091,1113]" captionTargetBox="[152,1432,199,1090]" captionTargetId="figure@9.[148,1433,183,1090]" captionTargetPageId="9" captionText="FIGURE 6. Range of intraspecific variation in pronotum of Typhlocharis mixta n. sp.: 1 – 6 main morphotypes, 7 – 10 asymmetries, malformations and teratologies: 1, 2, subrectangular pronotum with smoothly curved lateral margins and well marked denticles; 3, 4, rectangular pronotum, with straight lateral margins and high number of well marked denticles; 5, 6, subsquare, with smoothly curved lateral margins and low number of blunt denticles; 7, Asymmetric posterior angles, note the different aspect of pointy and blunt angles; 8, symmetrically widened lateral margins; 9, asymmetric malformed margins; 10, symmetrically notched lateral margins." httpUri="https://zenodo.org/record/223808/files/figure.png" pageId="8">Fig 6</figureCitation>
: 3,4) and subsquare, with 2-3 blunt and irregular denticles (
<figureCitation id="135A2A29FF9C5C45CFE20FD1357EF845" box="[1004,1068,1935,1960]" captionStart="FIGURE 6" captionStartId="9.[151,250,1091,1113]" captionTargetBox="[152,1432,199,1090]" captionTargetId="figure@9.[148,1433,183,1090]" captionTargetPageId="9" captionText="FIGURE 6. Range of intraspecific variation in pronotum of Typhlocharis mixta n. sp.: 1 – 6 main morphotypes, 7 – 10 asymmetries, malformations and teratologies: 1, 2, subrectangular pronotum with smoothly curved lateral margins and well marked denticles; 3, 4, rectangular pronotum, with straight lateral margins and high number of well marked denticles; 5, 6, subsquare, with smoothly curved lateral margins and low number of blunt denticles; 7, Asymmetric posterior angles, note the different aspect of pointy and blunt angles; 8, symmetrically widened lateral margins; 9, asymmetric malformed margins; 10, symmetrically notched lateral margins." httpUri="https://zenodo.org/record/223808/files/figure.png" pageId="8">Fig 6</figureCitation>
: 5,6). Asymmetric pronotum are abundant, and they affect the number of denticles or the pronouncement of the posterior angle (
<figureCitation id="135A2A29FF9C5C45C8FF0FED3460F821" box="[1265,1330,1971,1996]" captionStart="FIGURE 6" captionStartId="9.[151,250,1091,1113]" captionTargetBox="[152,1432,199,1090]" captionTargetId="figure@9.[148,1433,183,1090]" captionTargetPageId="9" captionText="FIGURE 6. Range of intraspecific variation in pronotum of Typhlocharis mixta n. sp.: 1 – 6 main morphotypes, 7 – 10 asymmetries, malformations and teratologies: 1, 2, subrectangular pronotum with smoothly curved lateral margins and well marked denticles; 3, 4, rectangular pronotum, with straight lateral margins and high number of well marked denticles; 5, 6, subsquare, with smoothly curved lateral margins and low number of blunt denticles; 7, Asymmetric posterior angles, note the different aspect of pointy and blunt angles; 8, symmetrically widened lateral margins; 9, asymmetric malformed margins; 10, symmetrically notched lateral margins." httpUri="https://zenodo.org/record/223808/files/figure.png" pageId="8">Fig 6</figureCitation>
: 7). Five specimens have aberrant pronotums with widened, notched or asymmetric lateral margins (
<figureCitation id="135A2A29FF9C5C45C89C0F893583F81D" box="[1170,1233,2007,2032]" captionStart="FIGURE 6" captionStartId="9.[151,250,1091,1113]" captionTargetBox="[152,1432,199,1090]" captionTargetId="figure@9.[148,1433,183,1090]" captionTargetPageId="9" captionText="FIGURE 6. Range of intraspecific variation in pronotum of Typhlocharis mixta n. sp.: 1 – 6 main morphotypes, 7 – 10 asymmetries, malformations and teratologies: 1, 2, subrectangular pronotum with smoothly curved lateral margins and well marked denticles; 3, 4, rectangular pronotum, with straight lateral margins and high number of well marked denticles; 5, 6, subsquare, with smoothly curved lateral margins and low number of blunt denticles; 7, Asymmetric posterior angles, note the different aspect of pointy and blunt angles; 8, symmetrically widened lateral margins; 9, asymmetric malformed margins; 10, symmetrically notched lateral margins." httpUri="https://zenodo.org/record/223808/files/figure.png" pageId="8">Fig 6</figureCitation>
: 8–10).
</paragraph>
<caption id="DF1E6624FF9D5C44CC990C1D3370FB1F" httpUri="https://zenodo.org/record/223808/files/figure.png" pageId="9" targetBox="[152,1432,199,1090]" targetPageId="9">
<paragraph id="8BDE36ACFF9D5C44CC990C1D3370FB1F" blockId="9.[151,1436,1091,1266]" pageId="9">
<emphasis id="B915EABEFF9D5C44CC990C1D304BFBB5" bold="true" box="[151,281,1091,1113]" pageId="9">FIGURE 6.</emphasis>
Range of intraspecific variation in pronotum of 
<taxonomicName id="4C614D2FFF9D5C44CF390C1D3552FBB4" box="[823,1024,1091,1113]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="mixta" status="sp. nov.">
<emphasis id="B915EABEFF9D5C44CF390C1D3552FBB4" box="[823,1024,1091,1113]" italics="true" pageId="9">Typhlocharis mixta</emphasis>
</taxonomicName>
<emphasis id="B915EABEFF9D5C44C8020C1D351EFBB5" bold="true" box="[1036,1100,1091,1112]" pageId="9">
<taxonomicNameLabel id="A22657C5FF9D5C44C8020C1D351EFBB5" box="[1036,1100,1091,1112]" pageId="9" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: 1–6 main morphotypes, 7–10 asymmetries, malformations and teratologies: 1,2, subrectangular pronotum with smoothly curved lateral margins and well marked denticles; 3,4, rectangular pronotum, with straight lateral margins and high number of well marked denticles; 5,6, subsquare, with smoothly curved lateral margins and low number of blunt denticles; 7, Asymmetric posterior angles, note the different aspect of pointy and blunt angles; 8, symmetrically widened lateral margins; 9, asymmetric malformed margins; 10, symmetrically notched lateral margins.
</paragraph>
</caption>
<paragraph id="8BDE36ACFF9D5C44CCC90D7E30D6FA2A" blockId="9.[151,1436,1311,1843]" pageId="9">
Transverse scutellar organ varies from straight to subtriangular and the margins can be irregular or smooth. Lateral denticles of elytra can be more or less marked. Apical denticles of elytra appear in a wide range of combinations and shapes (
<figureCitation id="135A2A29FF9D5C44CDCB0D39335FFA92" box="[453,525,1383,1408]" captionStart="FIGURE 7" captionStartId="10.[151,250,1860,1882]" captionTargetBox="[186,1413,214,1834]" captionTargetId="figure@10.[151,1435,193,1854]" captionTargetPageId="10" captionText="FIGURE 7. Range of intraspecific variation in elytra of Typhlocharis mixta n. sp.: (A) Elytral apex: 1 – 3 general morphotypes: 1, lower number of denticles; 2, higher number of denticles; 3, enlarged inner pair of denticles; 4 – 6 double or triple denticles: 4, large double denticles; 5, bifid, fused double denticle; 6, fused triple denticle; 7 – 9, strong asymmetries. (B) Graphic showing the abundance of every observed combination of apical denticles, expressed as “ denticles in left elytra / denticles in right elytra ”. Note the majority of the series present combinations of four and five pairs of denticles. “? ” indicates missing data (e. g. damaged elytra). Total number of specimens: 1306." httpUri="https://zenodo.org/record/223809/files/figure.png" pageId="9">Fig. 7</figureCitation>
): the most common are 4/4 (17% of the series), 4/5 (15%), 5/4 (15%), and 5/5 (22%), but they range from 2/3 to 7/7. All the observed combinations and their abundance within the population are expressed in 
<figureCitation id="135A2A29FF9D5C44CD270DF13039FA2A" box="[297,363,1455,1480]" captionStart="FIGURE 7" captionStartId="10.[151,250,1860,1882]" captionTargetBox="[186,1413,214,1834]" captionTargetId="figure@10.[151,1435,193,1854]" captionTargetPageId="10" captionText="FIGURE 7. Range of intraspecific variation in elytra of Typhlocharis mixta n. sp.: (A) Elytral apex: 1 – 3 general morphotypes: 1, lower number of denticles; 2, higher number of denticles; 3, enlarged inner pair of denticles; 4 – 6 double or triple denticles: 4, large double denticles; 5, bifid, fused double denticle; 6, fused triple denticle; 7 – 9, strong asymmetries. (B) Graphic showing the abundance of every observed combination of apical denticles, expressed as “ denticles in left elytra / denticles in right elytra ”. Note the majority of the series present combinations of four and five pairs of denticles. “? ” indicates missing data (e. g. damaged elytra). Total number of specimens: 1306." httpUri="https://zenodo.org/record/223809/files/figure.png" pageId="9">Fig. 7</figureCitation>
B.
</paragraph>
<paragraph id="8BDE36ACFF9D5C44CCC90D8D32D6F9BA" blockId="9.[151,1436,1311,1843]" pageId="9">
The inner pair, closer to the elytral suture is especially developed and bigger than the other denticles (
<figureCitation id="135A2A29FF9D5C44C9330D8D34D3FA06" box="[1341,1409,1491,1516]" captionStart="FIGURE 7" captionStartId="10.[151,250,1860,1882]" captionTargetBox="[186,1413,214,1834]" captionTargetId="figure@10.[151,1435,193,1854]" captionTargetPageId="10" captionText="FIGURE 7. Range of intraspecific variation in elytra of Typhlocharis mixta n. sp.: (A) Elytral apex: 1 – 3 general morphotypes: 1, lower number of denticles; 2, higher number of denticles; 3, enlarged inner pair of denticles; 4 – 6 double or triple denticles: 4, large double denticles; 5, bifid, fused double denticle; 6, fused triple denticle; 7 – 9, strong asymmetries. (B) Graphic showing the abundance of every observed combination of apical denticles, expressed as “ denticles in left elytra / denticles in right elytra ”. Note the majority of the series present combinations of four and five pairs of denticles. “? ” indicates missing data (e. g. damaged elytra). Total number of specimens: 1306." httpUri="https://zenodo.org/record/223809/files/figure.png" pageId="9">Fig. 7</figureCitation>
A: 3) in some specimens, resembling those of 
<taxonomicName id="4C614D2FFF9D5C44CE860DA935BFF9E2" authority="Perez-Gonzalez &amp; Zaballos, 2013" authorityName="Perez-Gonzalez &amp; Zaballos" authorityYear="2013" box="[648,1261,1527,1552]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="tertia">
<emphasis id="B915EABEFF9D5C44CE860DA93232F9E2" box="[648,864,1527,1552]" italics="true" pageId="9">Typhlocharis tertia</emphasis>
Pérez-González &amp; Zaballos, 2013
</taxonomicName>
. About 14% of the series (
<specimenCount id="9D67FD25FF9D5C44CD1B0E453092F9D9" box="[277,448,1563,1588]" pageId="9" type="generic">145 specimens</specimenCount>
) have bifid or double denticles and 
<specimenCount id="9D67FD25FF9D5C44CF6B0E4232A3F9D9" box="[869,1009,1563,1588]" pageId="9" type="generic">7 specimens</specimenCount>
show triple denticles (
<figureCitation id="135A2A29FF9D5C44C8F50E453410F9DE" box="[1275,1346,1563,1588]" captionStart="FIGURE 7" captionStartId="10.[151,250,1860,1882]" captionTargetBox="[186,1413,214,1834]" captionTargetId="figure@10.[151,1435,193,1854]" captionTargetPageId="10" captionText="FIGURE 7. Range of intraspecific variation in elytra of Typhlocharis mixta n. sp.: (A) Elytral apex: 1 – 3 general morphotypes: 1, lower number of denticles; 2, higher number of denticles; 3, enlarged inner pair of denticles; 4 – 6 double or triple denticles: 4, large double denticles; 5, bifid, fused double denticle; 6, fused triple denticle; 7 – 9, strong asymmetries. (B) Graphic showing the abundance of every observed combination of apical denticles, expressed as “ denticles in left elytra / denticles in right elytra ”. Note the majority of the series present combinations of four and five pairs of denticles. “? ” indicates missing data (e. g. damaged elytra). Total number of specimens: 1306." httpUri="https://zenodo.org/record/223809/files/figure.png" pageId="9">Fig. 7</figureCitation>
A: 4-6). Asymmetries (
<figureCitation id="135A2A29FF9D5C44CD350E61302DF9B5" box="[315,383,1599,1624]" captionStart="FIGURE 7" captionStartId="10.[151,250,1860,1882]" captionTargetBox="[186,1413,214,1834]" captionTargetId="figure@10.[151,1435,193,1854]" captionTargetPageId="10" captionText="FIGURE 7. Range of intraspecific variation in elytra of Typhlocharis mixta n. sp.: (A) Elytral apex: 1 – 3 general morphotypes: 1, lower number of denticles; 2, higher number of denticles; 3, enlarged inner pair of denticles; 4 – 6 double or triple denticles: 4, large double denticles; 5, bifid, fused double denticle; 6, fused triple denticle; 7 – 9, strong asymmetries. (B) Graphic showing the abundance of every observed combination of apical denticles, expressed as “ denticles in left elytra / denticles in right elytra ”. Note the majority of the series present combinations of four and five pairs of denticles. “? ” indicates missing data (e. g. damaged elytra). Total number of specimens: 1306." httpUri="https://zenodo.org/record/223809/files/figure.png" pageId="9">Fig. 7</figureCitation>
A: 7–9) and damaged elytra are also common.
</paragraph>
<paragraph id="8BDE36ACFF9D5C44CCC90E3B3017F8DF" blockId="9.[151,1436,1311,1843]" pageId="9">
Variations in patterns of chaetotaxy occur in labium, prebasilar, last ventrite and 
<emphasis id="B915EABEFF9D5C44C84B0E3B347AF993" box="[1093,1320,1637,1662]" italics="true" pageId="9">sensilla coeloconica</emphasis>
of the 11th antennomere. It is remarkable that the ventral 
<emphasis id="B915EABEFF9D5C44CECC0ED432F9F94E" box="[706,939,1674,1699]" italics="true" pageId="9">sensilla coeloconica</emphasis>
in 5º and 6º antennomeres also varies: 
<specimenCount id="9D67FD25FF9D5C44C9800ED5305EF92B" pageId="9" type="generic">7 specimens</specimenCount>
have only 
<emphasis id="B915EABEFF9D5C44CD880EF3333BF92B" box="[390,617,1709,1734]" italics="true" pageId="9">sensilla coeloconica</emphasis>
in the 5º antennomere and 
<specimenCount id="9D67FD25FF9D5C44CF960EF03570F92B" box="[920,1058,1709,1734]" pageId="9" type="generic">3 specimens</specimenCount>
have them in antennomeres 5º, 6º and 7º. It is the first evidence of variation in this feature within the same species, and the presence of 
<emphasis id="B915EABEFF9D5C44C9480E8C304CF8E3" italics="true" pageId="9">sensilla coeloconica</emphasis>
in 5º antennomere but not in 6º has been reported only in 
<taxonomicName id="4C614D2FFF9D5C44CFB20EAB3013F8DF" authority="Perez-Gonzalez &amp; Zaballos, 2013" authorityName="Perez-Gonzalez &amp; Zaballos" authorityYear="2013" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="tetramera">
<emphasis id="B915EABEFF9D5C44CFB20EAB3596F8E3" box="[956,1220,1781,1806]" italics="true" pageId="9">Typhlocharis tetramera</emphasis>
Pérez-González &amp; Zaballos, 2013
</taxonomicName>
.
</paragraph>
<caption id="DF1E6624FF9E5C47CC990F1A33F3F81E" httpUri="https://zenodo.org/record/223809/files/figure.png" pageId="10" targetBox="[186,1413,214,1834]" targetPageId="10">
<paragraph id="8BDE36ACFF9E5C47CC990F1A33F3F81E" blockId="10.[151,1437,1860,2035]" pageId="10">
<emphasis id="B915EABEFF9E5C47CC990F1A3041F8B4" bold="true" box="[151,275,1860,1882]" pageId="10">FIGURE 7.</emphasis>
Range of intraspecific variation in elytra of 
<taxonomicName id="4C614D2FFF9E5C47CEDA0F1A32C4F8B7" box="[724,918,1860,1882]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="mixta" status="sp. nov.">
<emphasis id="B915EABEFF9E5C47CEDA0F1A32C4F8B7" box="[724,918,1860,1882]" italics="true" pageId="10">Typhlocharis mixta</emphasis>
</taxonomicName>
<emphasis id="B915EABEFF9E5C47CF920F1A3287F8B4" bold="true" box="[924,981,1860,1881]" pageId="10">
<taxonomicNameLabel id="A22657C5FF9E5C47CF920F1A3287F8B4" box="[924,981,1860,1881]" pageId="10" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: (A) Elytral apex: 1–3 general morphotypes: 1, lower number of denticles; 2, higher number of denticles; 3, enlarged inner pair of denticles; 4–6 double or triple denticles: 4, large double denticles; 5, bifid, fused double denticle; 6, fused triple denticle; 7–9, strong asymmetries. (B) Graphic showing the abundance of every observed combination of apical denticles, expressed as “denticles in left elytra/denticles in right elytra”. Note the majority of the series present combinations of four and five pairs of denticles. “?” indicates missing data (e.g. damaged elytra). Total number of specimens: 1306.
</paragraph>
</caption>
<paragraph id="8BDE36ACFF9F5C46CCC908C93254FF15" blockId="11.[151,1436,151,933]" pageId="11">
Finally, there are several specimens with teratologies, like antennomere fusion (
<specimenCount id="9D67FD25FF9F5C46C84608C93587FF5D" box="[1096,1237,151,176]" pageId="11" type="generic">5 specimens</specimenCount>
, with fused 3º-4º, 5º-6º, 9º-10º and 10º-11º antennomeres respectively), or malformed mesotibiae (
<specimenCount id="9D67FD25FF9F5C46C84F08E23597FF38" box="[1089,1221,188,213]" pageId="11" type="generic">1 specimen</specimenCount>
, associated with a malformed pronotum, possible result of a failed moult).
</paragraph>
<paragraph id="8BDE36ACFF9F5C46CCC9095A33B3FE65" blockId="11.[151,1436,151,933]" pageId="11">
<emphasis id="B915EABEFF9F5C46CCC9095A306FFEF0" bold="true" box="[199,317,260,285]" pageId="11">Affinities.</emphasis>
<taxonomicName id="4C614D2FFF9F5C46CD47095A3377FEF1" box="[329,549,260,285]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="mixta" status="sp. nov.">
<emphasis id="B915EABEFF9F5C46CD47095A3377FEF1" box="[329,549,260,285]" italics="true" pageId="11">Typhlocharis mixta</emphasis>
</taxonomicName>
<emphasis id="B915EABEFF9F5C46CE3C095A3325FEF0" bold="true" box="[562,631,260,285]" pageId="11">
<taxonomicNameLabel id="A22657C5FF9F5C46CE3C095A3325FEF0" box="[562,631,260,285]" pageId="11" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
presents the diagnostic characters of 
<taxonomicName id="4C614D2FFF9F5C46C832095A35DDFEF0" box="[1084,1167,260,285]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="baetica">
<emphasis id="B915EABEFF9F5C46C832095A35DDFEF0" box="[1084,1167,260,285]" italics="true" pageId="11">baetica</emphasis>
</taxonomicName>
group, defined by the presence of several pairs of apical denticles. Affinities with other species of the group are the umbilicate series of 4+2, shared with 
<taxonomicName id="4C614D2FFF9F5C46CD56091333A1FE89" authority="Zaballos &amp; Jeanne, 1987" authorityName="Zaballos &amp; Jeanne" authorityYear="1987" box="[344,755,332,357]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="pacensis">
<emphasis id="B915EABEFF9F5C46CD5609133086FE89" box="[344,468,333,356]" italics="true" pageId="11">T. pacensis</emphasis>
Zaballos &amp; Jeanne, 1987
</taxonomicName>
; 
<taxonomicName id="4C614D2FFF9F5C46CEF109133599FE89" authority="Zaballos &amp; Banda, 2001" authorityName="Zaballos &amp; Banda" authorityYear="2001" box="[767,1227,332,357]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="furnayulensis">
<emphasis id="B915EABEFF9F5C46CEF1091332E3FE88" box="[767,945,332,357]" italics="true" pageId="11">T. furnayulensis</emphasis>
Zaballos &amp; Banda, 2001
</taxonomicName>
; 
<taxonomicName id="4C614D2FFF9F5C46C8D90913341CFE88" box="[1239,1358,332,357]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="secunda">
<emphasis id="B915EABEFF9F5C46C8D90913341CFE88" box="[1239,1358,332,357]" italics="true" pageId="11">T. secunda</emphasis>
</taxonomicName>
Perez- González &amp; Zaballos, 2013 and 
<taxonomicName id="4C614D2FFF9F5C46CE0F092F33C8FE65" box="[513,666,367,392]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="acutangula" status="sp. nov.">
<emphasis id="B915EABEFF9F5C46CE0F092F33C8FE65" box="[513,666,367,392]" italics="true" pageId="11">T. acutangula</emphasis>
</taxonomicName>
<emphasis id="B915EABEFF9F5C46CEAC093133B3FE65" bold="true" box="[674,737,367,392]" pageId="11">
<taxonomicNameLabel id="A22657C5FF9F5C46CEAC093133B3FE65" box="[674,737,367,392]" pageId="11" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BDE36ACFF9F5C46CCC909CA353AFDD1" blockId="11.[151,1436,151,933]" pageId="11">
The protuberances in femora also appears in 
<taxonomicName id="4C614D2FFF9F5C46CEDB09CB32B6FE41" authority="Ehlers, 1883" authorityName="Ehlers" authorityYear="1883" box="[725,996,404,429]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="baetica">
<emphasis id="B915EABEFF9F5C46CEDB09CB3215FE40" box="[725,839,404,429]" italics="true" pageId="11">T. baetica</emphasis>
Ehlers, 1883
</taxonomicName>
; 
<taxonomicName id="4C614D2FFF9F5C46CFFD09CB3181FE3D" authority="Perez-Gonzalez &amp; Zaballos, 2013" authorityName="Perez-Gonzalez &amp; Zaballos" authorityYear="2013" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="tertia">
<emphasis id="B915EABEFF9F5C46CFFD09CB351DFE41" box="[1011,1103,405,428]" italics="true" pageId="11">T. tertia</emphasis>
Perez-González &amp; Zaballos, 2013
</taxonomicName>
; 
<taxonomicName id="4C614D2FFF9F5C46CCD309E73394FE3D" authority="Perez-Gonzalez &amp; Zaballos, 2013" authorityName="Perez-Gonzalez &amp; Zaballos" authorityYear="2013" box="[221,710,439,464]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="quarta">
<emphasis id="B915EABEFF9F5C46CCD309E73016FE3D" box="[221,324,441,464]" italics="true" pageId="11">T. quarta</emphasis>
Perez-González &amp; Zaballos, 2013
</taxonomicName>
and 
<taxonomicName id="4C614D2FFF9F5C46CEF209E732C4FE3D" box="[764,918,439,464]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="acutangula" status="sp. nov.">
<emphasis id="B915EABEFF9F5C46CEF209E732C4FE3D" box="[764,918,439,464]" italics="true" pageId="11">T. acutangula</emphasis>
</taxonomicName>
<emphasis id="B915EABEFF9F5C46CF9209E93289FE3D" bold="true" box="[924,987,439,464]" pageId="11">
<taxonomicNameLabel id="A22657C5FF9F5C46CF9209E93289FE3D" box="[924,987,439,464]" pageId="11" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
; the high number and serrated aspect of posterolateral denticles of pronotum are also typical of 
<taxonomicName id="4C614D2FFF9F5C46CF3C098332F5FE18" box="[818,935,476,501]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="baetica">
<emphasis id="B915EABEFF9F5C46CF3C098332F5FE18" box="[818,935,476,501]" italics="true" pageId="11">T. baetica</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C614D2FFF9F5C46CFE709833504FE19" box="[1001,1110,477,500]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="quarta">
<emphasis id="B915EABEFF9F5C46CFE709833504FE19" box="[1001,1110,477,500]" italics="true" pageId="11">T. quarta</emphasis>
</taxonomicName>
, but many specimens have pronotum with few and less defined denticles akin to that of 
<taxonomicName id="4C614D2FFF9F5C46CF430A5F32F5FDF5" box="[845,935,513,536]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="tertia">
<emphasis id="B915EABEFF9F5C46CF430A5F32F5FDF5" box="[845,935,513,536]" italics="true" pageId="11">T. tertia</emphasis>
</taxonomicName>
and the other species of 
<taxonomicName id="4C614D2FFF9F5C46C8CB09A1344AFDF5" box="[1221,1304,511,536]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="baetica">
<emphasis id="B915EABEFF9F5C46C8CB09A1344AFDF5" box="[1221,1304,511,536]" italics="true" pageId="11">baetica</emphasis>
</taxonomicName>
group. The broad parameres are unusual within the group and strongly resembles those of 
<emphasis id="B915EABEFF9F5C46C8060A7B353AFDD1" box="[1032,1128,549,572]" italics="true" pageId="11">
<taxonomicName id="4C614D2FFF9F5C46C8060A7B3536FDD1" box="[1032,1124,549,572]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="tertia">T. tertia</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BDE36ACFF9F5C46CCC90A173597FD45" blockId="11.[151,1436,151,933]" pageId="11">
<taxonomicName id="4C614D2FFF9F5C46CCC90A17339BFD8D" authority="Perez-Gonzalez &amp; Zaballos, 2013" authorityName="Perez-Gonzalez &amp; Zaballos" authorityYear="2013" box="[199,713,583,608]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="secunda">
<emphasis id="B915EABEFF9F5C46CCC90A173012FD8D" box="[199,320,583,608]" italics="true" pageId="11">T. secunda</emphasis>
Perez-González &amp; Zaballos, 2013
</taxonomicName>
is very similar: both species shares the same umbilicate series, size range, overall shape of pronotum posterolateral denticles and presence (not always in 
<taxonomicName id="4C614D2FFF9F5C46C8BD0A333442FD69" box="[1203,1296,621,644]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="B915EABEFF9F5C46C8BD0A333442FD69" box="[1203,1296,621,644]" italics="true" pageId="11">T. mixta</emphasis>
</taxonomicName>
) of anterior “pseudodiscal” setae. However, the new species is well differentiated by its diagnostic features.
</paragraph>
<paragraph id="8BDE36ACFF9F5C46CCC90AEA347AFCD5" blockId="11.[151,1436,151,933]" pageId="11">
The tooth of clypeus, subcylindrical spermatheca, metatibial perpendicular short seta and the pseudotetramerous tarsi found in 
<taxonomicName id="4C614D2FFF9F5C46CE020A873334FD1D" box="[524,614,729,752]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="mixta" status="sp. nov.">
<emphasis id="B915EABEFF9F5C46CE020A873334FD1D" box="[524,614,729,752]" italics="true" pageId="11">T. mixta</emphasis>
</taxonomicName>
<emphasis id="B915EABEFF9F5C46CE630A8933FEFD1D" bold="true" box="[621,684,727,752]" pageId="11">
<taxonomicNameLabel id="A22657C5FF9F5C46CE630A8933FEFD1D" box="[621,684,727,752]" pageId="11" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
are unique of this species within 
<taxonomicName id="4C614D2FFF9F5C46C82C0A893527FD1D" box="[1058,1141,727,752]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="baetica">
<emphasis id="B915EABEFF9F5C46C82C0A893527FD1D" box="[1058,1141,727,752]" italics="true" pageId="11">baetica</emphasis>
</taxonomicName>
group, but appear in other 
<taxonomicName id="4C614D2FFF9F5C46CC990AA2307BFCF8" box="[151,297,764,789]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="genus">
<emphasis id="B915EABEFF9F5C46CC990AA2307BFCF8" box="[151,297,764,789]" italics="true" pageId="11">Typhlocharis</emphasis>
</taxonomicName>
species. The tooth of clypeus is the definitory trait of the six species of 
<taxonomicName id="4C614D2FFF9F5C46C87E0AA335EDFCF9" box="[1136,1215,765,788]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="gomezi">
<emphasis id="B915EABEFF9F5C46C87E0AA335EDFCF9" box="[1136,1215,765,788]" italics="true" pageId="11">gomezi</emphasis>
</taxonomicName>
group (Zaballos &amp; Farinós 1995; Zaballos &amp; Ruiz-Tapiador 1997) which also have a short perpendicular seta in metatibiae.
</paragraph>
<paragraph id="8BDE36ACFF9F5C46CCC90B1A3434FC48" blockId="11.[151,1436,151,933]" pageId="11">
The trend of diffuse gula in some specimens, the variability of ventral 
<emphasis id="B915EABEFF9F5C46CFD50B1A35EDFCB0" box="[987,1215,836,861]" italics="true" pageId="11">sensilla coeloconica</emphasis>
in antennomeres 5º and 6º and the pseudotetramerous tarsi are features previously observed only in species of 
<emphasis id="B915EABEFF9F5C46C8BC0B393403FC6D" box="[1202,1361,871,896]" italics="true" pageId="11">quadridentata</emphasis>
group (Pérez-González &amp; Zaballos 2013c). In both cases the possible homology of these characters remains unclear.
</paragraph>
</subSubSection>
<subSubSection id="C37B6527FF9F5C46CC990BA934CEF867" pageId="11" type="key">
<paragraph id="8BDE36ACFF9F5C46CC990BA9355DFBFD" blockId="11.[151,1039,1015,1040]" box="[151,1039,1015,1040]" pageId="11">
<emphasis id="B915EABEFF9F5C46CC990BA93356FBFD" bold="true" box="[151,516,1015,1040]" pageId="11">
Key to species of 
<taxonomicName id="4C614D2FFF9F5C46CD6A0BA930E4FBFD" box="[356,438,1015,1040]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="baetica">
<emphasis id="B915EABEFF9F5C46CD6A0BA930E4FBFD" bold="true" box="[356,438,1015,1040]" italics="true" pageId="11">baetica</emphasis>
</taxonomicName>
group
</emphasis>
(update of Pérez 
<emphasis id="B915EABEFF9F5C46CECD0BA9339EFBFD" bold="true" box="[707,716,1015,1040]" pageId="11">-</emphasis>
González &amp; Zaballos 2013a)
</paragraph>
<key id="FFF0619DFF9F5C46CC990C6134CEF867" pageId="11">
<keyStep id="30952E09FF9F5C46CC990C6134C9FB9F" pageId="11">
<paragraph id="8BDE36ACFF9F5C46CC990C6134CEFBB9" blockId="11.[151,1436,1087,1930]" box="[151,1436,1087,1108]" pageId="11">
<keyLead id="30909599FF9F5C46CC990C6134CEFBB9" box="[151,1436,1087,1108]" pageId="11">
1. Elytra with umbilicate series 4+4...................................... 
<taxonomicName id="4C614D2FFF9F5C46CFF70C6134CEFBB9" ID-CoL="59VKK" authority="Perez-Gonzalez &amp; Zaballos, 2013" authorityName="Perez-Gonzalez &amp; Zaballos" authorityYear="2013" box="[1017,1436,1087,1108]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="quarta">
<emphasis id="B915EABEFF9F5C46CFF70C613503FBB9" box="[1017,1105,1087,1108]" italics="true" pageId="11">T. quarta</emphasis>
Pérez-González &amp; Zaballos, 2013
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDE36ACFF9F5C46CC990C0334C9FB9F" blockId="11.[151,1436,1087,1930]" box="[151,1435,1117,1138]" pageId="11">
<keyLead id="30909599FF9F5C46CC990C0334C9FB9F" box="[151,1435,1117,1138]" pageId="11">- Elytra with umbilicate series different to 4+4.............................................................. 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30952E09FF9F5C46CC990C2434C9FB41" pageId="11">
<paragraph id="8BDE36ACFF9F5C46CC990C2434C9FB62" blockId="11.[151,1436,1087,1930]" box="[151,1435,1146,1167]" pageId="11">
<keyLead id="30909599FF9F5C46CC990C2434C9FB62" box="[151,1435,1146,1167]" pageId="11">2. Elytra with umbilicate series 4+3........................................................................ 3</keyLead>
</paragraph>
<paragraph id="8BDE36ACFF9F5C46CC990CC934C9FB41" blockId="11.[151,1436,1087,1930]" box="[151,1435,1175,1196]" pageId="11">
<keyLead id="30909599FF9F5C46CC990CC934C9FB41" box="[151,1435,1175,1196]" pageId="11">- Elytra with umbilicate series 4+2 or 4+1.................................................................. 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30952E09FF9F5C46CC990CEB34C7FB0A" pageId="11">
<paragraph id="8BDE36ACFF9F5C46CC990CEB34C7FB27" blockId="11.[151,1436,1087,1930]" box="[151,1429,1205,1226]" pageId="11">
<keyLead id="30909599FF9F5C46CC990CEB34C7FB27" box="[151,1429,1205,1226]" pageId="11">
3. Inner side of femora with protuberances, warty surface. Length&gt; 
<quantity id="4C999B49FF9F5C46CF560CEB32F3FB27" box="[856,929,1205,1226]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="11" unit="mm" value="1.4">1.4 mm</quantity>
....................................... 4
</keyLead>
</paragraph>
<paragraph id="8BDE36ACFF9F5C46CC990C8C34C7FB0A" blockId="11.[151,1436,1087,1930]" box="[151,1429,1234,1255]" pageId="11">
<keyLead id="30909599FF9F5C46CC990C8C34C7FB0A" box="[151,1429,1234,1255]" pageId="11">
- Inner side of femora without protuberances, smooth surface. Length &lt;
<quantity id="4C999B49FF9F5C46CF8F0C8C329BFB0A" box="[897,969,1234,1255]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="11" unit="mm" value="1.4">1.4 mm</quantity>
.................................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30952E09FF9F5C46CC990CB134CEFACF" pageId="11">
<paragraph id="8BDE36ACFF9F5C46CC990CB134C4FAE9" blockId="11.[151,1436,1087,1930]" box="[151,1430,1263,1284]" pageId="11">
<keyLead id="30909599FF9F5C46CC990CB134C4FAE9" box="[151,1430,1263,1284]" pageId="11">
4. Apical margin of elytra with 6–7 pairs of small denticles. Length 1.5......................... 
<taxonomicName id="4C614D2FFF9F5C46C8B60CB134C4FAE9" authority="Ehlers, 1883" authorityName="Ehlers" authorityYear="1883" box="[1208,1430,1263,1284]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="baetica">
<emphasis id="B915EABEFF9F5C46C8B60CB13445FAE9" box="[1208,1303,1263,1284]" italics="true" pageId="11">T. baetica</emphasis>
Ehlers, 1883
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDE36ACFF9F5C46CC990D5334CEFACF" blockId="11.[151,1436,1087,1930]" box="[151,1436,1293,1314]" pageId="11">
<keyLead id="30909599FF9F5C46CC990D5334CEFACF" box="[151,1436,1293,1314]" pageId="11">
- Apical margin of elytra with 3 pairs of large denticles. Length 
<quantity id="4C999B49FF9F5C46CF360D5332D3FACF" box="[824,897,1293,1314]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="11" unit="mm" value="1.7">1.7 mm</quantity>
.......... 
<taxonomicName id="4C614D2FFF9F5C46C80B0D5334CEFACF" ID-CoL="59VKV" authority="Perez-Gonzalez &amp; Zaballos, 2013" authorityName="Perez-Gonzalez &amp; Zaballos" authorityYear="2013" box="[1029,1436,1293,1314]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="tertia">
<emphasis id="B915EABEFF9F5C46C80B0D533503FACF" box="[1029,1105,1293,1314]" italics="true" pageId="11">T. tertia</emphasis>
Pérez-González &amp; Zaballos, 2013
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30952E09FF9F5C46CC990D7434CEFA7A" pageId="11">
<paragraph id="8BDE36ACFF9F5C46CC990D7434CEFAB1" blockId="11.[151,1436,1087,1930]" pageId="11">
<keyLead id="30909599FF9F5C46CC990D7434CEFAB1" pageId="11">
5. Elytra with lateral denticles marked in all their length. Very faint or absent ventral foveae. Length 
<quantity id="4C999B49FF9F5C46C8800D743585FAD2" box="[1166,1239,1322,1343]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="11" unit="mm" value="1.1">1.1 mm</quantity>
........................................................................................ 
<taxonomicName id="4C614D2FFF9F5C46C8480D1934CEFAB1" ID-CoL="59VJ4" authority="Zaballos &amp; Banda, 2001" authorityName="Zaballos &amp; Banda" authorityYear="2001" box="[1094,1436,1351,1372]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="aguirrei">
<emphasis id="B915EABEFF9F5C46C8480D1935F9FAB1" box="[1094,1195,1351,1372]" italics="true" pageId="11">T. aguirrei</emphasis>
Zaballos &amp; Banda, 2001
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDE36ACFF9F5C46CC990D3B34CEFA7A" blockId="11.[151,1436,1087,1930]" pageId="11">
<keyLead id="30909599FF9F5C46CC990D3B34CEFA7A" pageId="11">
- Elytra with lateral denticles marked only in the first third of their length. Ventral foveae well developed in both sexes. Length 
<quantity id="4C999B49FF9F5C46CCEF0DDC3078FA7A" box="[225,298,1410,1431]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="11" unit="mm" value="1.3">1.3 mm</quantity>
<taxonomicName id="4C614D2FFF9F5C46C8430DDC34CEFA7A" ID-CoL="59VK6" authority="Zaballos &amp; Banda, 2001" authorityName="Zaballos &amp; Banda" authorityYear="2001" box="[1101,1436,1410,1431]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="matiasi">
<emphasis id="B915EABEFF9F5C46C8430DDC35FEFA7A" box="[1101,1196,1410,1431]" italics="true" pageId="11">T. matiasi</emphasis>
Zaballos &amp; Banda, 2001
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30952E09FF9F5C46CC990DC134C9FA3F" pageId="11">
<paragraph id="8BDE36ACFF9F5C46CC990DC134C9FA59" blockId="11.[151,1436,1087,1930]" box="[151,1435,1439,1460]" pageId="11">
<keyLead id="30909599FF9F5C46CC990DC134C9FA59" box="[151,1435,1439,1460]" pageId="11">6. Elytra with umbilicate series 4+2........................................................................ 7</keyLead>
</paragraph>
<paragraph id="8BDE36ACFF9F5C46CC990DE334C9FA3F" blockId="11.[151,1436,1087,1930]" box="[151,1435,1469,1490]" pageId="11">
<keyLead id="30909599FF9F5C46CC990DE334C9FA3F" box="[151,1435,1469,1490]" pageId="11">- Elytra with umbilicate series 4+1....................................................................... 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="30952E09FF9F5C46CC990D8434C9F9E1" pageId="11">
<paragraph id="8BDE36ACFF9F5C46CC990D8434CEFA1D" blockId="11.[151,1436,1087,1930]" box="[151,1436,1498,1520]" pageId="11">
<keyLead id="30909599FF9F5C46CC990D8434CEFA1D" box="[151,1436,1498,1520]" pageId="11">
7. Clypeus with a median tooth. Ventral foveae in females. Length 
<quantity id="4C999B49FF9F5C46CF450D8432EFFA02" box="[843,957,1498,1519]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="11" unit="mm" value="1.4" valueMax="1.6" valueMin="1.2">1.2–1.6 mm</quantity>
........................... 
<taxonomicName id="4C614D2FFF9F5C46C91C0D84340DFA02" ID-CoL="59VK8" box="[1298,1375,1498,1519]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="mixta" status="sp. nov.">
<emphasis id="B915EABEFF9F5C46C91C0D84340DFA02" box="[1298,1375,1498,1519]" italics="true" pageId="11">T. mixta</emphasis>
</taxonomicName>
<emphasis id="B915EABEFF9F5C46C9680D8434CEFA1D" bold="true" box="[1382,1436,1498,1520]" pageId="11">
<taxonomicNameLabel id="A22657C5FF9F5C46C9680D8434CEFA1D" box="[1382,1436,1498,1520]" pageId="11" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BDE36ACFF9F5C46CC990DA934C9F9E1" blockId="11.[151,1436,1087,1930]" box="[151,1435,1527,1548]" pageId="11">
<keyLead id="30909599FF9F5C46CC990DA934C9F9E1" box="[151,1435,1527,1548]" pageId="11">- Clypeus with straight margin, without median tooth........................................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30952E09FF9F5C46CC990E4B34C9F989" pageId="11">
<paragraph id="8BDE36ACFF9F5C46CC990E4B34CEF9A5" blockId="11.[151,1436,1087,1930]" pageId="11">
<keyLead id="30909599FF9F5C46CC990E4B34CEF9A5" pageId="11">
8. Metatibiae with a pointy dentiform projection in the distal end. Ventral foveae absent. Length 
<quantity id="4C999B49FF9F5C46C8680E4B358BF9C7" box="[1126,1241,1557,1578]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="11" unit="mm" value="1.2" valueMax="1.3" valueMin="1.1">1.1–1.3 mm</quantity>
..................................................................................................... 
<taxonomicName id="4C614D2FFF9F5C46C8D20E6C340DF9AA" ID-CoL="59VJ3" box="[1244,1375,1586,1607]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="acutangula" status="sp. nov.">
<emphasis id="B915EABEFF9F5C46C8D20E6C340DF9AA" box="[1244,1375,1586,1607]" italics="true" pageId="11">T. acutangula</emphasis>
</taxonomicName>
<emphasis id="B915EABEFF9F5C46C9680E6C34CEF9A5" bold="true" box="[1382,1436,1586,1608]" pageId="11">
<taxonomicNameLabel id="A22657C5FF9F5C46C9680E6C34CEF9A5" box="[1382,1436,1586,1608]" pageId="11" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BDE36ACFF9F5C46CC990E1134C9F989" blockId="11.[151,1436,1087,1930]" box="[151,1435,1615,1636]" pageId="11">
<keyLead id="30909599FF9F5C46CC990E1134C9F989" box="[151,1435,1615,1636]" pageId="11">- Metatibiae without dentiform projection.................................................................... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="30952E09FF9F5C46CC990E3334C9F972" pageId="11">
<paragraph id="8BDE36ACFF9F5C46CC990E3334CEF96F" blockId="11.[151,1436,1087,1930]" box="[151,1436,1645,1666]" pageId="11">
<keyLead id="30909599FF9F5C46CC990E3334CEF96F" box="[151,1436,1645,1666]" pageId="11">
9. Elytra with 1 pair of discal setae in the anterior third. Length 
<quantity id="4C999B49FF9F5C46CF220E333224F96F" box="[812,886,1645,1666]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="11" unit="mm" value="1.4">1.4 mm</quantity>
.......... 
<taxonomicName id="4C614D2FFF9F5C46CFE50E3334CEF96F" ID-CoL="59VKR" authority="Perez-Gonzalez &amp; Zaballos, 2013" authorityName="Perez-Gonzalez &amp; Zaballos" authorityYear="2013" box="[1003,1436,1645,1666]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="secunda">
<emphasis id="B915EABEFF9F5C46CFE50E333503F96F" box="[1003,1105,1645,1666]" italics="true" pageId="11">T. secunda</emphasis>
Pérez-González &amp; Zaballos, 2013
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDE36ACFF9F5C46CC990ED434C9F972" blockId="11.[151,1436,1087,1930]" box="[151,1435,1674,1695]" pageId="11">
<keyLead id="30909599FF9F5C46CC990ED434C9F972" box="[151,1435,1674,1695]" pageId="11">
- Elytra without discal setae. Length &lt;
<quantity id="4C999B49FF9F5C46CE4C0ED433D9F972" box="[578,651,1674,1695]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="11" unit="mm" value="1.1">1.1 mm</quantity>
.............................................................. 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30952E09FF9F5C46CC990EF934CEF8F9" pageId="11">
<paragraph id="8BDE36ACFF9F5C46CC990EF934CEF937" blockId="11.[151,1436,1087,1930]" pageId="11">
<keyLead id="30909599FF9F5C46CC990EF934CEF937" pageId="11">
10. Apical margin of elytra with 4–5 pairs of small denticles. Diffuse endophallus sclerites. Length 
<quantity id="4C999B49FF9F5C46C8780EF93589F951" box="[1142,1243,1703,1724]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="11" unit="mm" value="0.95" valueMax="1.0" valueMin="0.9">0.9–1 mm</quantity>
........................................................................................ 
<taxonomicName id="4C614D2FFF9F5C46C8330E9B34CEF937" ID-CoL="59VKC" authority="Zaballos &amp; Jeanne, 1987" authorityName="Zaballos &amp; Jeanne" authorityYear="1987" box="[1085,1436,1733,1754]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="pacensis">
<emphasis id="B915EABEFF9F5C46C8330E9B35F5F937" box="[1085,1191,1733,1754]" italics="true" pageId="11">T. pacensis</emphasis>
Zaballos &amp; Jeanne, 1987
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDE36ACFF9F5C46CC990EBC34CEF8F9" blockId="11.[151,1436,1087,1930]" pageId="11">
<keyLead id="30909599FF9F5C46CC990EBC34CEF8F9" pageId="11">
- Apical margin of elytra with 5–7 pairs of small apical denticles. “Y-shaped” endophallus sclerites. Length 
<quantity id="4C999B49FF9F5C46C8D30EBC3474F91A" box="[1245,1318,1762,1783]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="11" unit="mm" value="1.1">1.1 mm</quantity>
............................................................................. 
<taxonomicName id="4C614D2FFF9F5C46C81C0EA134CEF8F9" ID-CoL="59VJT" authority="Zaballos &amp; Banda, 2001" authorityName="Zaballos &amp; Banda" authorityYear="2001" box="[1042,1436,1791,1812]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="furnayulensis">
<emphasis id="B915EABEFF9F5C46C81C0EA135F9F8F9" box="[1042,1195,1791,1812]" italics="true" pageId="11">T. furnayulensis</emphasis>
Zaballos &amp; Banda, 2001
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30952E09FF9F5C46CC990F4334CEF867" pageId="11">
<paragraph id="8BDE36ACFF9F5C46CC990F4334CEF8A2" blockId="11.[151,1436,1087,1930]" pageId="11">
<keyLead id="30909599FF9F5C46CC990F4334CEF8A2" pageId="11">
11. Elytra with lateral denticles marked in all their length. Thin, “Y-shaped” endophallus sclerites. Subspheric-irregular spermath- eca. Length 
<quantity id="4C999B49FF9F5C46CD590F6430F2F8A2" box="[343,416,1850,1871]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="11" unit="mm" value="1.1">1.1 mm</quantity>
<taxonomicName id="4C614D2FFF9F5C46C83E0F6434CEF8A2" ID-CoL="59VK7" authority="Zaballos &amp; Banda, 2001" authorityName="Zaballos &amp; Banda" authorityYear="2001" box="[1072,1436,1850,1871]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="millenaria">
<emphasis id="B915EABEFF9F5C46C83E0F6435F9F8A2" box="[1072,1195,1850,1871]" italics="true" pageId="11">T. millenaria</emphasis>
Zaballos &amp; Banda, 2001
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDE36ACFF9F5C46CC990F0934CEF867" blockId="11.[151,1436,1087,1930]" pageId="11">
<keyLead id="30909599FF9F5C46CC990F0934CEF867" pageId="11">
- Elytra with lateral denticles marked only in the first third of their length. Thick, stick-shaped endophallus. Reniform spermath- eca. Length 
<quantity id="4C999B49FF9F5C46CD590F2B30F2F867" box="[343,416,1909,1930]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="11" unit="mm" value="1.1">1.1 mm</quantity>
................................................. 
<taxonomicName id="4C614D2FFF9F5C46C80E0F2B34CEF867" ID-CoL="59VKH" authority="Perez-Gonzalez &amp; Zaballos, 2013" authorityName="Perez-Gonzalez &amp; Zaballos" authorityYear="2013" box="[1024,1436,1909,1930]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="prima">
<emphasis id="B915EABEFF9F5C46C80E0F2B351DF867" box="[1024,1103,1909,1930]" italics="true" pageId="11">T. prima</emphasis>
Pérez-González &amp; Zaballos, 2013
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>