283 lines
19 KiB
XML
283 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.87.76085" ID-GBIF-Dataset="ac841941-ab0d-438b-ad40-26bd8e43e5f3" ID-PMC="PMC8873192" ID-Pensoft-Pub="1314-4049-87-133" ID-Pensoft-UUID="3E84DC4D23DD55D1BD51C48DF991EADD" ID-PubMed="35221753" ModsDocID="1314-4049-87-133" checkinTime="1645139634519" checkinUser="pensoft" docAuthor="Zhang, Guang-Zhi, Yang, He-Tong, Zhang, Xin-Jian, Zhou, Fang-Yuan, Wu, Xiao-Qing, Xie, Xue-Ying, Zhao, Xiao-Yan & Zhou, Hong-Zi" docDate="2022" docId="DE284B67F38459E092E805DCBFD3BCE4" docLanguage="en" docName="MycoKeys 87: 133-157" docOrigin="MycoKeys 87" docPubDate="2022-02-17" docSource="http://dx.doi.org/10.3897/mycokeys.87.76085" docTitle="Trichoderma nordicum G. Z. Zhang 2022, sp. nov." docType="treatment" docVersion="4" id="3E84DC4D23DD55D1BD51C48DF991EADD" lastPageNumber="133" masterDocId="3E84DC4D23DD55D1BD51C48DF991EADD" masterDocTitle="Five new species of Trichoderma from moist soils in China" masterLastPageNumber="157" masterPageNumber="133" pageNumber="133" updateTime="1668136798648" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Five new species of Trichoderma from moist soils in China</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zhang, Guang-Zhi</mods:namePart>
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<mods:affiliation>Qilu University of Technology (Shandong Academy of Sciences), Ecology Institute, Shandong Provincial Key Laboratory for Applied Microbiology, Jinan 250103, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yang, He-Tong</mods:namePart>
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<mods:affiliation>Qilu University of Technology (Shandong Academy of Sciences), Ecology Institute, Shandong Provincial Key Laboratory for Applied Microbiology, Jinan 250103, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zhang, Xin-Jian</mods:namePart>
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<mods:affiliation>Qilu University of Technology (Shandong Academy of Sciences), Ecology Institute, Shandong Provincial Key Laboratory for Applied Microbiology, Jinan 250103, China</mods:affiliation>
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<mods:nameIdentifier type="email">zhangxj@sdas.org</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zhou, Fang-Yuan</mods:namePart>
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<mods:affiliation>Qilu University of Technology (Shandong Academy of Sciences), Ecology Institute, Shandong Provincial Key Laboratory for Applied Microbiology, Jinan 250103, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wu, Xiao-Qing</mods:namePart>
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<mods:affiliation>Qilu University of Technology (Shandong Academy of Sciences), Ecology Institute, Shandong Provincial Key Laboratory for Applied Microbiology, Jinan 250103, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Xie, Xue-Ying</mods:namePart>
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<mods:affiliation>Qilu University of Technology (Shandong Academy of Sciences), Ecology Institute, Shandong Provincial Key Laboratory for Applied Microbiology, Jinan 250103, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zhao, Xiao-Yan</mods:namePart>
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<mods:affiliation>Qilu University of Technology (Shandong Academy of Sciences), Ecology Institute, Shandong Provincial Key Laboratory for Applied Microbiology, Jinan 250103, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zhou, Hong-Zi</mods:namePart>
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<mods:affiliation>Qilu University of Technology (Shandong Academy of Sciences), Ecology Institute, Shandong Provincial Key Laboratory for Applied Microbiology, Jinan 250103, China</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>MycoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-02-17</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>87</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>133</mods:start>
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<mods:end>157</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/mycokeys.87.76085</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.87.76085</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-4049-87-133</mods:identifier>
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<mods:identifier type="Pensoft-UUID">3E84DC4D23DD55D1BD51C48DF991EADD</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="193397160" LSID="urn:lsid:plazi:treatment:DE284B67F38459E092E805DCBFD3BCE4" httpUri="http://treatment.plazi.org/id/DE284B67F38459E092E805DCBFD3BCE4" lastPageNumber="133" pageId="0" pageNumber="133">
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<subSubSection pageId="0" pageNumber="133" type="nomenclature">
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||
<paragraph pageId="0" pageNumber="133">
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<taxonomicName LSID="DE284B67-F384-59E0-92E8-05DCBFD3BCE4" authority="G. Z. Zhang" authorityName="G. Z. Zhang" authorityYear="2022" class="Sordariomycetes" family="Hypocreaceae" genus="Trichoderma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Trichoderma nordicum" order="Hypocreales" pageId="0" pageNumber="133" phylum="Ascomycota" rank="species" species="nordicum" status="sp. nov.">Trichoderma nordicum G.Z. Zhang</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="133">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="133" type="description">
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<paragraph pageId="0" pageNumber="133">
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Trichoderma nordicum A-D cultures on different media at 25 ° C after 10 days (A on PDA B on MEA C on CMD D on SNA) E-G, I, J conidiophores and phialides H conidia. Notes: E on PDA F-J on MEA A-D from WT 13001 E-J from WT 61001. Scale bars: 10 μm (E-J)." figureDoi="10.3897/mycokeys.87.76085.figure5" httpUri="https://binary.pensoft.net/fig/647558" pageId="0" pageNumber="133">Fig. 5</figureCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="0" pageNumber="133" type="etymology">
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<paragraph pageId="0" pageNumber="133">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="133">
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<normalizedToken originalValue="“nord”">"nord"</normalizedToken>
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means found in the north of China.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="133" type="holotype">
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<paragraph pageId="0" pageNumber="133">Holotype.</paragraph>
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<paragraph pageId="0" pageNumber="133">China, Beijing, Yu-yuan-tan Park, 43 m (altitude), isolated from soil, 27 October 2016, G.Z. Zhang (Holotype WT 13001), ex-type culture ACCC 39713.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="133" type="diagnosis">
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<paragraph pageId="0" pageNumber="133">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="133">
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Phylogenetically
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<taxonomicName authorityName="G. Z. Zhang" authorityYear="2022" class="Sordariomycetes" family="Hypocreaceae" genus="Trichoderma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Trichoderma nordicum" order="Hypocreales" pageId="0" pageNumber="133" phylum="Ascomycota" rank="species" species="nordicum">
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<emphasis italics="true" pageId="0" pageNumber="133">Trichoderma nordicum</emphasis>
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</taxonomicName>
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is related to
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<taxonomicName lsidName="T. paratroviride" pageId="0" pageNumber="133" rank="species" species="paratroviride">
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<emphasis italics="true" pageId="0" pageNumber="133">T. paratroviride</emphasis>
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</taxonomicName>
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, but the sequence similarities of
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<emphasis italics="true" pageId="0" pageNumber="133">rpb2</emphasis>
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and
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<emphasis italics="true" pageId="0" pageNumber="133">tef1</emphasis>
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-α were 98.15% and 94.43%, respectively. That does not meet the
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<emphasis italics="true" pageId="0" pageNumber="133">sp</emphasis>
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∃!(
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<emphasis italics="true" pageId="0" pageNumber="133">rpb2</emphasis>
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99≅
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<emphasis italics="true" pageId="0" pageNumber="133">tef1</emphasis>
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97) standard for
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<taxonomicName lsidName="T. paratroviride" pageId="0" pageNumber="133" rank="species" species="paratroviride">
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<emphasis italics="true" pageId="0" pageNumber="133">T. paratroviride</emphasis>
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</taxonomicName>
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or other known
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<taxonomicName class="Sordariomycetes" family="Hypocreaceae" genus="Trichoderma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Trichoderma" order="Hypocreales" pageId="0" pageNumber="133" phylum="Ascomycota" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="133">Trichoderma</emphasis>
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</taxonomicName>
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species. Morphologically, conidiophores of
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<taxonomicName lsidName="T. paratroviride" pageId="0" pageNumber="133" rank="species" species="paratroviride">
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<emphasis italics="true" pageId="0" pageNumber="133">T. paratroviride</emphasis>
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</taxonomicName>
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consisting of a main axis and often distantly-spaced side branches, not re-branching. Conidiophores of
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<taxonomicName lsidName="T. nordicum" pageId="0" pageNumber="133" rank="species" species="nordicum">
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<emphasis italics="true" pageId="0" pageNumber="133">T. nordicum</emphasis>
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</taxonomicName>
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are branched in a more complex manner; conidia are larger than those of
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<taxonomicName lsidName="T. paratroviride" pageId="0" pageNumber="133" rank="species" species="paratroviride">
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<emphasis italics="true" pageId="0" pageNumber="133">T. paratroviride</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="133" type="teleomorph">
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<paragraph pageId="0" pageNumber="133">Teleomorph.</paragraph>
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<paragraph pageId="0" pageNumber="133">Unknown.</paragraph>
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<paragraph pageId="0" pageNumber="133">
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Growth optimal at 25 °C, slow or limited at 30 °C, absent at 35 °C. Colonies grew fast on PDA, CMD and MEA and slow on SNA. Colony radius after 72 h at 25 °C 67-71 mm on PDA, 68-71 mm on CMD, 51-55 mm on MEA and 21-24 mm on SNA. Aerial mycelia sparse on PDA after 72 h at 25 °C under 12 h photoperiod and conidiation developed within 48 h beginning at the inoculation point and progressed around, grey-white at first and slowly turning green. Diffusing pigment or distinctive odour absent. Aerial mycelia sparse and flocculence on MEA after 72 h at 20 °C under 12 h photoperiod. Conidia developed within 48 h beginning near the colony margin on MEA, grey-white at first and slowly turning green, transparent liquid secreted. Aerial mycelia few on SNA and CMD after 72 h at 25 °C, conidia formed around the inoculation point and in distinct concentric rings after 96 h under 12 h photoperiod on SNA and CMD, diffusing pigment not produced. Conidiophores and branches narrow and flexuous, tending to be regularly verticillate forming a pyramidal structure, each branch terminating in a cruciate whorl of up to five phialides. Phialides, lageniform, (6.2-)7.2-10.3(-12.9)
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<normalizedToken originalValue="×">x</normalizedToken>
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(2.6-)2.9-3.2(-3.4)
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(mean = 8.8
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<normalizedToken originalValue="×">x</normalizedToken>
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3.1
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<normalizedToken originalValue="μm">μm</normalizedToken>
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), 1.6-2.3
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(mean = 1.9
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<normalizedToken originalValue="μm">μm</normalizedToken>
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) near the base; phialide length/width ratio (2.1-)2.4-3.4(-4.3) (mean = 2.9). On PDA, phialides curved, distinguished from those on other media. Conidia, globose to obovoidal, (4.1-)4.4-4.8(-5.0)
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<normalizedToken originalValue="×">x</normalizedToken>
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(4.0-)4.1-4.4(-4.6)
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(mean = 4.6
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<normalizedToken originalValue="×">x</normalizedToken>
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4.3
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<normalizedToken originalValue="μm">μm</normalizedToken>
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), length/width ratio 1.0-1.2 (mean = 1.1). Chlamydospores sometimes present, (8.7-)9.8
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<normalizedToken originalValue="×">x</normalizedToken>
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10.4(-12.5)
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<normalizedToken originalValue="μm">μm</normalizedToken>
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.
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</paragraph>
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<caption doi="10.3897/mycokeys.87.76085.figure5" httpUri="https://binary.pensoft.net/fig/647558" pageId="0" pageNumber="133" start="Figure 5" startId="F5">
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<paragraph pageId="0" pageNumber="133">
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<emphasis bold="true" pageId="0" pageNumber="133">Figure 5.</emphasis>
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<taxonomicName authorityName="G. Z. Zhang" authorityYear="2022" class="Sordariomycetes" family="Hypocreaceae" genus="Trichoderma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Trichoderma nordicum" order="Hypocreales" pageId="0" pageNumber="133" phylum="Ascomycota" rank="species" species="nordicum">
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<emphasis italics="true" pageId="0" pageNumber="133">Trichoderma nordicum</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="133">A-D</emphasis>
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cultures on different media at 25 °C after 10 days (
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<emphasis bold="true" pageId="0" pageNumber="133">A</emphasis>
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on PDA
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<emphasis bold="true" pageId="0" pageNumber="133">B</emphasis>
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on MEA
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<emphasis bold="true" pageId="0" pageNumber="133">C</emphasis>
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on CMD
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<emphasis bold="true" pageId="0" pageNumber="133">D</emphasis>
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on SNA)
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<emphasis bold="true" pageId="0" pageNumber="133">E-G, I, J</emphasis>
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conidiophores and phialides
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<emphasis bold="true" pageId="0" pageNumber="133">H</emphasis>
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conidia. Notes:
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<emphasis bold="true" pageId="0" pageNumber="133">E</emphasis>
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on PDA
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<emphasis bold="true" pageId="0" pageNumber="133">F-J</emphasis>
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on MEA
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<emphasis bold="true" pageId="0" pageNumber="133">A-D</emphasis>
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from WT13001
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<emphasis bold="true" pageId="0" pageNumber="133">E-J</emphasis>
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from WT61001. Scale bars: 10
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(
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<emphasis bold="true" pageId="0" pageNumber="133">E-J</emphasis>
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).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="133" type="distribution">
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<paragraph pageId="0" pageNumber="133">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="133">China, Beijing and Hebei.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="133" type="additional specimen examined">
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<paragraph pageId="0" pageNumber="133">Additional specimen examined.</paragraph>
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<paragraph pageId="0" pageNumber="133">
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China. Hebei, Bai-yang Lake, 19 m (altitude), isolated from soil, 15 September 2016,
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<emphasis italics="true" pageId="0" pageNumber="133">J.S. Li</emphasis>
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(WT 61001).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="133" type="notes">
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<paragraph pageId="0" pageNumber="133">Notes.</paragraph>
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<paragraph pageId="0" pageNumber="133">
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Phylogenetically,
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<taxonomicName authorityName="G. Z. Zhang" authorityYear="2022" class="Sordariomycetes" family="Hypocreaceae" genus="Trichoderma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Trichoderma nordicum" order="Hypocreales" pageId="0" pageNumber="133" phylum="Ascomycota" rank="species" species="nordicum">
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<emphasis italics="true" pageId="0" pageNumber="133">Trichoderma nordicum</emphasis>
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</taxonomicName>
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is related to
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<taxonomicName lsidName="T. paratroviride" pageId="0" pageNumber="133" rank="species" species="paratroviride">
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<emphasis italics="true" pageId="0" pageNumber="133">T. paratroviride</emphasis>
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</taxonomicName>
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(Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phylogenetic tree, based on the Maximum Likelihood analysis of the rpb 2 (left; InL = - 5930.92) and tef 1 - α (right; InL = - 7681.95) dataset. Bootstrap values of Maximum Likelihood (left) and Maximum Parsimony (right) above 50 % are indicated at the nodes. The tree is rooted with Protocrea illinoensis TFC 9698 and P. farinose CPK 3144. New species proposed here are indicated in bold. The type strains are indicated with an asterisk (*) after the strain number. Results of the pairwise sequence similarity are illustrated on the dashed lines between the query strain and its closely-related species (arrows point to the reference strains)." figureDoi="10.3897/mycokeys.87.76085.figure1" httpUri="https://binary.pensoft.net/fig/647554" pageId="0" pageNumber="133">1</figureCitation>
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), but the sequence similarities of
|
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<emphasis italics="true" pageId="0" pageNumber="133">rpb2</emphasis>
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and
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<emphasis italics="true" pageId="0" pageNumber="133">tef1</emphasis>
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-α were 98.15% and 94.43%, respectively. That does not meet the
|
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<emphasis italics="true" pageId="0" pageNumber="133">sp</emphasis>
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∃!(
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<emphasis italics="true" pageId="0" pageNumber="133">rpb2</emphasis>
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99≅
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<emphasis italics="true" pageId="0" pageNumber="133">tef1</emphasis>
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97) standard for
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<taxonomicName lsidName="T. paratroviride" pageId="0" pageNumber="133" rank="species" species="paratroviride">
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<emphasis italics="true" pageId="0" pageNumber="133">T. paratroviride</emphasis>
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</taxonomicName>
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or other known
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<taxonomicName class="Sordariomycetes" family="Hypocreaceae" genus="Trichoderma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Trichoderma" order="Hypocreales" pageId="0" pageNumber="133" phylum="Ascomycota" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="133">Trichoderma</emphasis>
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</taxonomicName>
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species. Morphologically, conidiophores of
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<taxonomicName lsidName="T. paratroviride" pageId="0" pageNumber="133" rank="species" species="paratroviride">
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<emphasis italics="true" pageId="0" pageNumber="133">T. paratroviride</emphasis>
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</taxonomicName>
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consist of a main axis and often distantly-spaced side branches, not re-branching. Conidiophores of
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<taxonomicName lsidName="T. nordicum" pageId="0" pageNumber="133" rank="species" species="nordicum">
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<emphasis italics="true" pageId="0" pageNumber="133">T. nordicum</emphasis>
|
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</taxonomicName>
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are branched in a more complex manner; conidia are larger than those of
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<taxonomicName lsidName="T. paratroviride" pageId="0" pageNumber="133" rank="species" species="paratroviride">
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<emphasis italics="true" pageId="0" pageNumber="133">T. paratroviride</emphasis>
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</taxonomicName>
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(
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<bibRefCitation DOI="https://doi.org/10.1016/j.simyco.2014.11.001" author="Jaklitsch, WM" journalOrPublisher="Studies in Mycology" pageId="0" pageNumber="133" pagination="1 - 87" refId="B30" refString="Jaklitsch, WM, Voglmayr, H, 2015. Biodiversity of Trichoderma (Hypocreaceae) in Southern Europe and Macaronesia. Studies in Mycology 80: 1 - 87, DOI: https://doi.org/10.1016/j.simyco.2014.11.001" title="Biodiversity of Trichoderma (Hypocreaceae) in Southern Europe and Macaronesia." url="https://doi.org/10.1016/j.simyco.2014.11.001" volume="80" year="2015">Jaklitsch and Voglmayr 2015</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |