treatments-xml/data/DD/4B/64/DD4B641B111F4510FF495555FCB1FEFD.xml

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<document id="85C276565340355B7A4F2B0EB84E4C2E" ID-CLB-Dataset="46992" ID-DOI="10.5281/zenodo.201054" ID-GBIF-Dataset="7f030caf-c242-4e4f-acf6-0f6c5fa403c6" ID-ISSN="1175-5326" ID-Zenodo-Dep="201054" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460342089644" checkinUser="plazi" docAuthor="Stipala, Jan, Lutzmánn, Nicola, Malonza, Patrick K., Borghesio, Luca, Wilkinson, Paul, Godley, Brendan &amp; Evans, Matthew R." docDate="2011" docId="DD4B641B111F4510FF495555FCB1FEFD" docLanguage="en" docName="zt03002p016.pdf" docOrigin="Zootaxa 3002" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Trioceros nyirit Stipala, Lutzmánn, Malonza, Borghesio, Wilkinson, Godley &amp; Evans, 2011, sp. nov." docType="treatment" docVersion="9" lastPageNumber="11" masterDocId="21721C631119451AFFDE5339FFE3FF99" masterDocTitle="A new species of chameleon (Sauria: Chamaeleonidae) from the highlands of northwest Kenya" masterLastPageNumber="16" masterPageNumber="1" pageNumber="7" updateTime="1732486442677" updateUser="ExternalLinkService">
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<mods:title id="492F92DEE0249EE92ECB1C722F0D5CD3">A new species of chameleon (Sauria: Chamaeleonidae) from the highlands of northwest Kenya</mods:title>
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<mods:namePart id="50B74D03AE3E44CEEF2979427271D511">Stipala, Jan</mods:namePart>
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<mods:namePart id="250B90D96D8EB95FC607F40FA603C0B0">Lutzmánn, Nicola</mods:namePart>
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<mods:namePart id="A4444EF269856160BDB393410195CA36">Malonza, Patrick K.</mods:namePart>
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<mods:namePart id="A8081B0440FE4BD2E60D15B456A21C66">Borghesio, Luca</mods:namePart>
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<mods:namePart id="4C23640E8C29DF2E323F437BF256691E">Wilkinson, Paul</mods:namePart>
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<mods:namePart id="E61209811B63AB9F17696BACA88D652C">Godley, Brendan</mods:namePart>
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<mods:namePart id="2D9BCD6E4F21E23A6C598F2D87D3392D">Evans, Matthew R.</mods:namePart>
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<mods:date id="50BAE58620EBC366EF2746C276D93ED0">2011</mods:date>
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<treatment id="DD4B641B111F4510FF495555FCB1FEFD" ID-DOI="http://doi.org/10.5281/zenodo.5658789" ID-GBIF-Taxon="119406639" ID-Zenodo-Dep="5658789" LSID="urn:lsid:plazi:treatment:DD4B641B111F4510FF495555FCB1FEFD" httpUri="http://treatment.plazi.org/id/DD4B641B111F4510FF495555FCB1FEFD" lastPageId="10" lastPageNumber="11" pageId="6" pageNumber="7" scope_family="Chamaeleonidae">
<subSubSection id="1DF88686111F451CFF495555FE56F93E" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="555DD50D111F451CFF495555FE54F91F" blockId="6.[151,439,1644,1703]" box="[151,439,1644,1670]" pageId="6" pageNumber="7">
<heading id="0E156261111F451CFF495555FE54F91F" bold="true" box="[151,439,1644,1670]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<emphasis id="6796091F111F451CFF495555FE54F91F" bold="true" box="[151,439,1644,1670]" pageId="6" pageNumber="7">
<taxonomicName id="92E2AE8E111F451CFF495555FEB1F91F" ID-CoL="58VQV" box="[151,338,1644,1670]" class="Reptilia" family="Chamaeleonidae" genus="Trioceros" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="nyirit" status="sp. nov.">
<emphasis id="6796091F111F451CFF495555FEB1F91F" bold="true" box="[151,338,1644,1670]" italics="true" pageId="6" pageNumber="7">Trioceros nyirit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7CA5B464111F451CFE875555FE54F91F" box="[345,439,1644,1670]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="555DD50D111F451CFF4955B6FE56F93E" blockId="6.[151,439,1644,1703]" box="[151,437,1679,1703]" pageId="6" pageNumber="7">
(
<figureCitation id="CDD9C988111F451CFF4055B6FEEAF93E" box="[158,265,1679,1703]" captionStart="FIGURE 2" captionStartId="4.[151,250,1874,1897]" captionTargetBox="[183,1399,1116,1847]" captionTargetId="figure@4.[173,1408,1108,1868]" captionTargetPageId="4" captionText="FIGURE 2. Typical head shape of male T. nyirit sp. nov. and comparison with other morphologically similar or genetically closely related species from the bitaeniatus group. Two specimens of T. hoehnelii are included to show intra-specific variation. A— T. nyirit sp. nov., holotype, Mtelo massif, B— T. hoehnelii, Elgeyo Escarpment; C— T. sternfeldi, Mt. Meru; D— T. schubotzi, Mt. Kenya; E— T. hoehnelii, Mt. Elgon; F— T. narraioca, Mt. Kulal." httpUri="https://zenodo.org/record/201056/files/figure.png" pageId="6" pageNumber="7">Figures 2</figureCitation>
B and 5A &amp; B)
</paragraph>
</subSubSection>
<subSubSection id="1DF88686111F451CFF4955EEFEBBF877" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="555DD50D111F451CFF4955EEFDA6F88F" blockId="6.[151,1436,1751,2030]" pageId="6" pageNumber="7">
<emphasis id="6796091F111F451CFF4955EEFEEBF976" bold="true" box="[151,264,1751,1775]" pageId="6" pageNumber="7">
<typeStatus id="8A596BAF111F451CFF4955EEFEE7F976" box="[151,260,1751,1775]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
.
</emphasis>
NMK-L3166/1, adult male, Gatau Pass (
<quantity id="921A78E8111F451CFD1355EEFCFCF976" box="[717,799,1751,1775]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="7" unit="m" value="2500.0">2500m</quantity>
), Mtelo massif, northwest
<collectingCountry id="2DF5959D111F451CFB9D55E1FB6EF969" box="[1091,1165,1752,1776]" name="Kenya" pageId="6" pageNumber="7">Kenya</collectingCountry>
, collected by Jan Stipala and Joash Nyamache on
<date id="215CF3CD111F451CFE7955C7FDA2F88F" box="[423,577,1787,1814]" pageId="6" pageNumber="7" value="2006-07-21">21st July 2006</date>
.
</paragraph>
<paragraph id="555DD50D111F451CFF19541BFF17F81B" blockId="6.[151,1436,1751,2030]" pageId="6" pageNumber="7">
<emphasis id="6796091F111F451CFF19541BFEA0F8A3" bold="true" box="[199,323,1826,1850]" pageId="6" pageNumber="7">
<typeStatus id="8A596BAF111F451CFF19541BFEDDF8A3" box="[199,318,1826,1850]" pageId="6" pageNumber="7" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="43E41E84111F451CFE95541BFE42F8A3" box="[331,417,1826,1850]" pageId="6" pageNumber="7" type="male">4 males</specimenCount>
and
<specimenCount id="43E41E84111F451CFE04541BFDAAF8A3" box="[474,585,1826,1850]" pageId="6" pageNumber="7" type="female">4 females</specimenCount>
. NMK-L3166/2, male, NMK-L3166/ 3-4, females, southern slopes of Mtelo massif (
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); NMK-L2990/1 &amp; 5, males, NMK-L2990/ 2-3, females, Gatau Pass, Mtelo massif (
<quantity id="921A78E8111F451CFA85547FFF0BF81B" metricMagnitude="3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="6" pageNumber="7" unit="m" value="2350.0" valueMax="2500.0" valueMin="2200.0">2200- 2500m</quantity>
).
</paragraph>
<paragraph id="555DD50D111F451CFF1954B7FEBBF877" blockId="6.[151,1436,1751,2030]" pageId="6" pageNumber="7">
<emphasis id="6796091F111F451CFF1954B7FE4AF83F" bold="true" box="[199,425,1933,1958]" pageId="6" pageNumber="7">Non-type material.</emphasis>
NMK-L2998/1-5, males, NMK-L2998/6-11, females, Kaptalamwa and Kapiego (
<quantity id="921A78E8111F451CFA8554B7FF04F853" metricMagnitude="3" metricUnit="m" metricValue="2.95" metricValueMax="3.0" metricValueMin="2.9" pageId="6" pageNumber="7" unit="m" value="2950.0" valueMax="3000.0" valueMin="2900.0">2900- 3000m</quantity>
), Cherangani Hills;
<collectionCode id="33F34DC8111F451CFE07548AFDCFF853" box="[473,556,1971,1994]" pageId="6" pageNumber="7">BMNH</collectionCode>
1969.2588-9,
<collectionCode id="33F34DC8111F451CFD05548AFCCDF853" box="[731,814,1971,1994]" pageId="6" pageNumber="7">BMNH</collectionCode>
1969.2591,
<collectionCode id="33F34DC8111F451CFC18548AFBFAF853" box="[966,1049,1971,1994]" pageId="6" pageNumber="7">BMNH</collectionCode>
1969.2595-6, Sondang (
<quantity id="921A78E8111F451CFAE1548BFA73F853" box="[1343,1424,1970,1994]" metricMagnitude="3" metricUnit="m" metricValue="3.05" pageId="6" pageNumber="7" unit="m" value="3050.0">3050m</quantity>
), Cherangani Hills.
</paragraph>
</subSubSection>
<subSubSection id="1DF88686111E451DFF1953AEFA9BFE84" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="555DD50D111E451DFF1953AEFA9BFE84" blockId="7.[151,1437,151,1940]" pageId="7" pageNumber="8">
<emphasis id="6796091F111E451DFF1953AEFEDDFF29" bold="true" box="[199,318,151,176]" pageId="7" pageNumber="8">Diagnosis.</emphasis>
A medium sized chameleon (largest male
<quantity id="921A78E8111E451DFCFC53AEFC95FF36" box="[802,886,151,175]" metricMagnitude="-1" metricUnit="m" metricValue="1.91" pageId="7" pageNumber="8" unit="mm" value="191.0">191mm</quantity>
TL, largest female
<quantity id="921A78E8111E451DFB8B53AEFB5EFF36" box="[1109,1213,151,175]" metricMagnitude="-1" metricUnit="m" metricValue="1.805" pageId="7" pageNumber="8" unit="mm" value="180.5">180.5mm</quantity>
TL) with heterogeneous body scalation; short, forward-pointing rostral process covered in enlarged scales; straight to weakly curved parietal crest, rising gradually posteriorly to meet the lateral crests to form a prominent, moderately steep triangular casque; This combination of characters differentiates
<taxonomicName id="92E2AE8E111E451DFD01523CFCD0FE85" box="[735,819,261,284]" class="Reptilia" family="Chamaeleonidae" genus="Trioceros" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="nyirit" status="sp. nov.">
<emphasis id="6796091F111E451DFD01523CFCD0FE85" box="[735,819,261,284]" italics="true" pageId="7" pageNumber="8">T. nyirit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7CA5B464111E451DFCE2523DFC70FE85" box="[828,915,260,284]" pageId="7" pageNumber="8" rank="species">
<emphasis id="6796091F111E451DFCE2523DFC70FE85" bold="true" box="[828,915,260,284]" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
from other species in the
<taxonomicName id="92E2AE8E111E451DFB70523DFAC4FE85" box="[1198,1319,260,284]" class="Reptilia" family="Chamaeleonidae" genus="Trioceros" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="bitaeniatus">
<emphasis id="6796091F111E451DFB70523DFAC4FE85" box="[1198,1319,260,284]" italics="true" pageId="7" pageNumber="8">bitaeniatus</emphasis>
</taxonomicName>
-group.
</paragraph>
</subSubSection>
<subSubSection id="1DF88686111E451DFF19521EFCD3FB39" pageId="7" pageNumber="8" type="description">
<paragraph id="555DD50D111E451DFF19521EFEF3FDC6" blockId="7.[151,1437,151,1940]" pageId="7" pageNumber="8">
<emphasis id="6796091F111E451DFF19521EFE05FED9" bold="true" box="[199,486,295,320]" pageId="7" pageNumber="8">
Description of
<typeStatus id="8A596BAF111E451DFEA55211FE02FED9" box="[379,481,296,320]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
.
</emphasis>
Total length
<quantity id="921A78E8111E451DFD5C521EFD35FEA6" box="[642,726,295,319]" metricMagnitude="-1" metricUnit="m" metricValue="1.65" pageId="7" pageNumber="8" unit="mm" value="165.0">165mm</quantity>
(SVL
<quantity id="921A78E8111E451DFCF7521EFC90FEA6" box="[809,883,295,319]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="8" unit="mm" value="80.0">80mm</quantity>
, tail
<quantity id="921A78E8111E451DFC71521EFC19FEA6" box="[943,1018,295,319]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" pageId="7" pageNumber="8" unit="mm" value="85.0">85mm</quantity>
). The scalation is heterogenous with rounded or oval convex tubercles of different sizes scattered on the body and legs and tail. There are two clearly defined horizontal rows of enlarged tubercles on each flank, larger than those elsewhere on the body, the lower row extending between the front and hind legs, the second row higher on the flank. The scales on the belly are fine and relatively homogenous. The dorsal crest is well-developed along the entire length of the dorsal keel and extends onto the proximal two thirds of the tail, decreasing in size posteriorly. It is composed of enlarged conical tubercles, formed into sequential groups of three or four scales, the scales within each group increasing in size posteriorly, the most posterior scale much larger than the one before it. The ventral crest consists of short conical tubercles and is contiguous with the gular crest.
</paragraph>
<paragraph id="555DD50D111E451DFF1B5155FE6FFB96" blockId="7.[151,1437,151,1940]" pageId="7" pageNumber="8">
The head has prominent tubercular crests. The supra-orbital crests are continuous with the lateral crests, which rise posteriorly to meet the parietal crest forming a distinct, raised triangular casque. The parietal crest is straight and rises gradually posteriorly. It is forked anteriorly, the branches extending anterio-laterally to contact the supra-orbital crests. The scales on the casque are large, flat and polygonal. The scales between the supra-orbital crests are smaller than on top of the casque and moderately convex. A row of five strongly convex scales form a median ridge between the orbits and is distinct from the flat scales around it. The supra-orbital crests are contiguous with the canthi rostrales and merge anteriorly to form a short, forward-pointing rostral projection, which is covered in enlarged scales and extends beyond the the edge of the snout by
<quantity id="921A78E8111E451DFD5B505EFD21FCE6" box="[645,706,871,895]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="8" unit="mm" value="1.0">1mm</quantity>
. The sides of the head are covered in moderately enlarged convex tubercles. The scales on the eyelids are fine and weakly heterogenous. The gular region has several longitudinal rows of strongly convex tubercles that border a distinct gular crest. The gular crest is composed of conical scales, short anteriorly, becoming longer under the posterior half of the head, the longest scales about equal in length to the diameter of the eye opening.
</paragraph>
<paragraph id="555DD50D111E451DFF1B5725FCDAFBE5" blockId="7.[151,1437,151,1940]" pageId="7" pageNumber="8">
The hemipenis is structurally similar to that of
<taxonomicName id="92E2AE8E111E451DFD135724FB84FBAD" authority="Bohme &amp; Klaver 1986" authorityName="Bohme &amp; Klaver" authorityYear="1986" box="[717,1127,1052,1076]" class="Reptilia" family="Chamaeleonidae" genus="Trioceros" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="hoehnelii">
<emphasis id="6796091F111E451DFD135724FCACFBAD" box="[717,847,1052,1076]" italics="true" pageId="7" pageNumber="8">T. hoehnelii</emphasis>
(Böhme &amp; Klaver 1986)
</taxonomicName>
but differs, the two pairs of cogwheels being more strongy serrated. The parietal peritoneum shows only light pigmentation and is not extensive as seen in other members of the
<taxonomicName id="92E2AE8E111E451DFE04575DFDB0FBE5" box="[474,595,1124,1148]" class="Reptilia" family="Chamaeleonidae" genus="Trioceros" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="bitaeniatus">
<emphasis id="6796091F111E451DFE04575DFDB0FBE5" box="[474,595,1124,1148]" italics="true" pageId="7" pageNumber="8">bitaeniatus</emphasis>
</taxonomicName>
group (
<bibRefCitation id="3173A8FC111E451DFD76575DFCCEFBE5" author="Bauer" box="[680,813,1124,1148]" pageId="7" pageNumber="8" refString="Bauer, A. (1997) Peritoneal pigmentation and generic allocation in the Chamaeleonidae. African Journal of Herpetology, 46 (2), 117 - 123." type="journal article" year="1997">Bauer 1997</bibRefCitation>
).
</paragraph>
<paragraph id="555DD50D111E451DFF1957BEFCD3FB39" blockId="7.[151,1437,151,1940]" box="[199,816,1159,1184]" pageId="7" pageNumber="8">
<emphasis id="6796091F111E451DFF1957BEFE34FB39" bold="true" box="[199,471,1159,1184]" pageId="7" pageNumber="8">Colour in preservation.</emphasis>
The
<typeStatus id="8A596BAF111E451DFDD157B1FD8CFB39" box="[527,623,1160,1184]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
is uniform black.
</paragraph>
</subSubSection>
<subSubSection id="1DF88686111E451DFF195792FC5FF80D" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="555DD50D111E451DFF195792FC02FACD" blockId="7.[151,1437,151,1940]" pageId="7" pageNumber="8">
<emphasis id="6796091F111E451DFF195792FD3EFB5D" bold="true" box="[199,733,1195,1220]" pageId="7" pageNumber="8">
Variation in
<typeStatus id="8A596BAF111E451DFE865795FE2FFB5D" box="[344,460,1196,1220]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
and non-type material.
</emphasis>
The following snout-vent length (SVL) and tail length measurements were taken from living specimens, some of which were preserved and deposited in the
<collectionCode id="33F34DC8111E451DFB1957E8FAE8FB71" box="[1223,1291,1233,1256]" httpUri="http://grbio.org/cool/h6fg-emd8" name="National Museums of Kenya" pageId="7" pageNumber="8">NMK</collectionCode>
. Mean SVL: males =
<quantity id="921A78E8111E451DFF2B57CDFEACFA95" box="[245,335,1268,1292]" metricMagnitude="-2" metricUnit="m" metricValue="7.64" pageId="7" pageNumber="8" unit="mm" value="76.4">76.4mm</quantity>
± 8.5 (n = 27). Females =
<quantity id="921A78E8111E451DFDA957CDFD32FA95" box="[631,721,1268,1292]" metricMagnitude="-2" metricUnit="m" metricValue="7.29" pageId="7" pageNumber="8" unit="mm" value="72.9">72.9mm</quantity>
±7.1 (n = 35). Largest male:
<quantity id="921A78E8111E451DFBCB57CDFB8AFA95" box="[1045,1129,1268,1292]" metricMagnitude="-1" metricUnit="m" metricValue="1.91" pageId="7" pageNumber="8" unit="mm" value="191.0">191mm</quantity>
total length (SVL =
<quantity id="921A78E8111E451DFA9157CDFA7BFA95" box="[1359,1432,1268,1292]" metricMagnitude="-2" metricUnit="m" metricValue="9.1" pageId="7" pageNumber="8" unit="mm" value="91.0">91mm</quantity>
, tail =
<quantity id="921A78E8111E451DFF035621FED7FAA9" box="[221,308,1304,1328]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="8" unit="mm" value="100.0">100mm</quantity>
. Largest female:
<quantity id="921A78E8111E451DFE275621FDADFAA9" box="[505,590,1304,1328]" metricMagnitude="-1" metricUnit="m" metricValue="1.6" pageId="7" pageNumber="8" unit="mm" value="160.0">160mm</quantity>
total length (SVL =
<quantity id="921A78E8111E451DFC9F5621FC6AFAA9" box="[833,905,1304,1328]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" pageId="7" pageNumber="8" unit="mm" value="85.0">85mm</quantity>
, tail
<quantity id="921A78E8111E451DFC1B5621FBF3FAA9" box="[965,1040,1304,1328]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="7" pageNumber="8" unit="mm" value="75.0">75mm</quantity>
). Morphometric measurements and inter-orbital scale count of
<typeStatus id="8A596BAF111E451DFE635604FE0FFACC" box="[445,492,1341,1365]" pageId="7" pageNumber="8">type</typeStatus>
and non-type specimens are given in Table 3.
</paragraph>
<paragraph id="555DD50D111E451DFF1B5659FDAFF9B5" blockId="7.[151,1437,151,1940]" pageId="7" pageNumber="8">
Several morphological characters were quite variable. Body scales in some individuals are fine, granular and almost homogenous except for the two rows of tubercles, which in some individuals can be indistinct, especially the lower row. Gular crest development is variable in both Mtelo and Cherangani populations, being very short in some individuals (see
<figureCitation id="CDD9C988111E451DFE9356F5FE4FFA7D" box="[333,428,1484,1508]" captionStart="FIGURE 4" captionStartId="6.[151,250,1410,1433]" captionTargetBox="[232,1354,680,1374]" captionTargetId="figure@6.[217,1369,663,1386]" captionTargetPageId="6" captionText="FIGURE 4. Phylogeny generated from Maximum Parsimony analysis, based on partial 16 S and ND 4 mtDNA markers. Above branches = MP bootstrap support values. Below branches = Bayesian posterior probabilities. Scale bar indicates the number of nucleotide substitutions." httpUri="https://zenodo.org/record/201058/files/figure.png" pageId="7" pageNumber="8">Figure 4</figureCitation>
B) and long in others, equivalent in length to some specimens of
<taxonomicName id="92E2AE8E111E451DFB5856F4FAE6FA7D" box="[1158,1285,1484,1508]" class="Reptilia" family="Chamaeleonidae" genus="Trioceros" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="hoehnelii">
<emphasis id="6796091F111E451DFB5856F4FAE6FA7D" box="[1158,1285,1484,1508]" italics="true" pageId="7" pageNumber="8">T. hoehnelii</emphasis>
</taxonomicName>
. The parietal crest is straight or weakly curved, however it never reaches the extent of strong curvature nor rises steeply anteriorly as seen in
<taxonomicName id="92E2AE8E111E451DFED9552CFE6BF9B5" box="[263,392,1556,1580]" class="Reptilia" family="Chamaeleonidae" genus="Trioceros" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="hoehnelii">
<emphasis id="6796091F111E451DFED9552CFE6BF9B5" box="[263,392,1556,1580]" italics="true" pageId="7" pageNumber="8">T. hoehnelii</emphasis>
</taxonomicName>
and
<taxonomicName id="92E2AE8E111E451DFE63552CFDA6F9B5" box="[445,581,1557,1580]" class="Reptilia" family="Chamaeleonidae" genus="Trioceros" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="narraioca">
<emphasis id="6796091F111E451DFE63552CFDA6F9B5" box="[445,581,1557,1580]" italics="true" pageId="7" pageNumber="8">T. narraioca</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="555DD50D111E451DFF19550EFA93F901" blockId="7.[151,1437,151,1940]" pageId="7" pageNumber="8">
<emphasis id="6796091F111E451DFF19550EFE88F9C9" bold="true" box="[199,363,1591,1616]" pageId="7" pageNumber="8">Colour in life.</emphasis>
<typeStatus id="8A596BAF111E451DFEAD5501FE3AF9C9" box="[371,473,1592,1616]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
was uniform bright green with an indistinct wash of turquoise bands on the body, legs and tail. The eye is bright green with a dark horizontal line passing through it. The dorsal crest and adjacent scales on the dorsal keel, the head crests and rostral projection are dark red. The toes and soles of the feet are bright yellow.
</paragraph>
<paragraph id="555DD50D111E451DFF1B559DFA7BF8B1" blockId="7.[151,1437,151,1940]" pageId="7" pageNumber="8">Other males were also typically uniform bright green, the lower flanks, gular region, legs and tail a lighter turquoise-green. The head crests, rostral projection, dorsal crest and adjacent scales on the dorsal keel vary from red to black. When stressed, excited or basking, in males a darker pattern of broad vertical bars appears on the flanks, the rows of flank tubercles remaining light green. Bright yellow toes and soles of the feet were only seen in Mtelo males.</paragraph>
<paragraph id="555DD50D111E451DFF1B540DFC5FF80D" blockId="7.[151,1437,151,1940]" pageId="7" pageNumber="8">Females were more variable in colour than males. Some females were uniform green, while others were shades of light and dark brown, the body with several broad bands, the flank tubercles and throat cream-white or yellow. The head and dorsal crests and rostral projection were red in most individuals.</paragraph>
</subSubSection>
<subSubSection id="1DF8868611104513FF1B53A1FDC7FF4D" pageId="9" pageNumber="10" type="etymology">
<paragraph id="555DD50D11104513FF1B53A1FDC7FF4D" blockId="9.[151,1436,151,538]" pageId="9" pageNumber="10">
<emphasis id="6796091F11104513FF1B53A1FEA8FF29" bold="true" box="[197,331,152,176]" pageId="9" pageNumber="10">Etymology.</emphasis>
Named after the Pokot word for chameleon,
<taxonomicName id="92E2AE8E11104513FC8553A0FC74FF29" box="[859,919,153,176]" class="Reptilia" family="Chamaeleonidae" genus="Trioceros" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="nyirit">
<emphasis id="6796091F11104513FC8553A0FC74FF29" box="[859,919,153,176]" italics="true" pageId="9" pageNumber="10">nyirit</emphasis>
</taxonomicName>
. The distribution of
<taxonomicName id="92E2AE8E11104513FB5753A0FB00FF29" box="[1161,1251,153,176]" class="Reptilia" family="Chamaeleonidae" genus="Trioceros" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="nyirit" status="sp. nov.">
<emphasis id="6796091F11104513FB5753A0FB00FF29" box="[1161,1251,153,176]" italics="true" pageId="9" pageNumber="10">T. nyirit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7CA5B46411104513FB3353A1FAAAFF29" box="[1261,1353,152,176]" pageId="9" pageNumber="10" rank="species">
<emphasis id="6796091F11104513FB3353A1FAAAFF29" bold="true" box="[1261,1353,152,176]" pageId="9" pageNumber="10">sp. nov.</emphasis>
</taxonomicNameLabel>
occurs mainly within the Pokot tribal area.
</paragraph>
</subSubSection>
<subSubSection id="1DF8868611104510FF1953E6FCB1FEFD" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="distribution">
<paragraph id="555DD50D11104513FF1953E6FABEFD83" blockId="9.[151,1436,151,538]" pageId="9" pageNumber="10">
<emphasis id="6796091F11104513FF1953E6FE83FF61" bold="true" box="[199,352,223,248]" pageId="9" pageNumber="10">Distribution.</emphasis>
<taxonomicName id="92E2AE8E11104513FEB353D8FDC3FF61" box="[365,544,225,248]" class="Reptilia" family="Chamaeleonidae" genus="Trioceros" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="nyirit" status="sp. nov.">
<emphasis id="6796091F11104513FEB353D8FDC3FF61" box="[365,544,225,248]" italics="true" pageId="9" pageNumber="10">Trioceros nyirit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7CA5B46411104513FDF053D9FD6DFF61" box="[558,654,224,248]" pageId="9" pageNumber="10" rank="species">
<emphasis id="6796091F11104513FDF053D9FD6DFF61" bold="true" box="[558,654,224,248]" pageId="9" pageNumber="10">sp. nov.</emphasis>
</taxonomicNameLabel>
was recorded between
<quantity id="921A78E811104513FC7353E6FBDFFF6E" box="[941,1084,223,247]" metricMagnitude="3" metricUnit="m" metricValue="3.025" metricValueMax="3.15" metricValueMin="2.9" pageId="9" pageNumber="10" unit="m" value="3025.0" valueMax="3150.0" valueMin="2900.0">2900-3150m</quantity>
on the Cherangani Hills and between
<quantity id="921A78E811104513FEDB523DFE77FE85" box="[261,404,260,284]" metricMagnitude="3" metricUnit="m" metricValue="2.6985" metricValueMax="3.121" metricValueMin="2.276" pageId="9" pageNumber="10" unit="m" value="2698.5" valueMax="3121.0" valueMin="2276.0">2276-3121m</quantity>
on the Mtelo massif. At lower elevations other chameleon species (
<taxonomicName id="92E2AE8E11104513FB22523CFA83FE85" box="[1276,1376,260,284]" class="Reptilia" family="Chamaeleonidae" genus="Trioceros" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="ellioti">
<emphasis id="6796091F11104513FB22523CFA83FE85" box="[1276,1376,260,284]" italics="true" pageId="9" pageNumber="10">T. ellioti</emphasis>
</taxonomicName>
and
<taxonomicName id="92E2AE8E11104513FF495210FEA4FED9" box="[151,327,297,320]" class="Reptilia" family="Chamaeleonidae" genus="Trioceros" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="conirostratus">
<emphasis id="6796091F11104513FF495210FEA4FED9" box="[151,327,297,320]" italics="true" pageId="9" pageNumber="10">T. conirostratus</emphasis>
</taxonomicName>
) were abundant, suggesting that
<taxonomicName id="92E2AE8E11104513FD605210FCF6FED9" box="[702,789,297,320]" class="Reptilia" family="Chamaeleonidae" genus="Trioceros" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="nyirit" status="sp. nov.">
<emphasis id="6796091F11104513FD605210FCF6FED9" box="[702,789,297,320]" italics="true" pageId="9" pageNumber="10">T. nyirit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7CA5B46411104513FCC05211FC9AFED9" box="[798,889,296,320]" pageId="9" pageNumber="10" rank="species">
<emphasis id="6796091F11104513FCC05211FC9AFED9" bold="true" box="[798,889,296,320]" pageId="9" pageNumber="10">sp. nov.</emphasis>
</taxonomicNameLabel>
is restricted to higher elevations. In the Cherangani Hills
<taxonomicName id="92E2AE8E11104513FED15274FE8BFEFD" box="[271,360,333,356]" class="Reptilia" family="Chamaeleonidae" genus="Trioceros" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="nyirit" status="sp. nov.">
<emphasis id="6796091F11104513FED15274FE8BFEFD" box="[271,360,333,356]" italics="true" pageId="9" pageNumber="10">T. nyirit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7CA5B46411104513FEAC5275FE2EFEFD" box="[370,461,332,356]" pageId="9" pageNumber="10" rank="species">
<emphasis id="6796091F11104513FEAC5275FE2EFEFD" bold="true" box="[370,461,332,356]" pageId="9" pageNumber="10">sp. nov.</emphasis>
</taxonomicNameLabel>
appears to be widespread as specimens were collected from Sondang (
<quantity id="921A78E811104513FB245275FAA8FEFD" box="[1274,1355,332,356]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="10" unit="m" value="3000.0">3000m</quantity>
) in the north and Cheptongoi Hills (
<quantity id="921A78E811104513FE0B5256FDC5FE1E" box="[469,550,367,391]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="10" unit="m" value="3000.0">3000m</quantity>
) to the south of the range. We did not sample between
<quantity id="921A78E811104513FB595256FB36FE1E" box="[1159,1237,367,391]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="10" unit="m" value="2300.0">2300m</quantity>
and
<quantity id="921A78E811104513FAD45256FAB4FE1E" box="[1290,1367,367,391]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="9" pageNumber="10" unit="m" value="2900.0">2900m</quantity>
in the Cherangani Hills but
<taxonomicName id="92E2AE8E11104513FE5652ACFE3CFE35" box="[392,479,405,428]" class="Reptilia" family="Chamaeleonidae" genus="Trioceros" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="nyirit" status="sp. nov.">
<emphasis id="6796091F11104513FE5652ACFE3CFE35" box="[392,479,405,428]" italics="true" pageId="9" pageNumber="10">T. nyirit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7CA5B46411104513FE3752ADFDA0FE35" box="[489,579,404,428]" pageId="9" pageNumber="10" rank="species">
<emphasis id="6796091F11104513FE3752ADFDA0FE35" bold="true" box="[489,579,404,428]" pageId="9" pageNumber="10">sp. nov.</emphasis>
</taxonomicNameLabel>
is likely to occur below
<quantity id="921A78E811104513FC8452ADFC4BFE35" box="[858,936,404,428]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="9" pageNumber="10" unit="m" value="2900.0">2900m</quantity>
given its elevation range on Mtelo. We estimated the distribution of
<taxonomicName id="92E2AE8E11104513FE6B5280FDEEFE49" box="[437,525,441,464]" class="Reptilia" family="Chamaeleonidae" genus="Trioceros" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="nyirit" status="sp. nov.">
<emphasis id="6796091F11104513FE6B5280FDEEFE49" box="[437,525,441,464]" italics="true" pageId="9" pageNumber="10">T. nyirit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7CA5B46411104513FDC85281FD93FE49" box="[534,624,440,464]" pageId="9" pageNumber="10" rank="species">
<emphasis id="6796091F11104513FDC85281FD93FE49" bold="true" box="[534,624,440,464]" pageId="9" pageNumber="10">sp. nov.</emphasis>
</taxonomicNameLabel>
using maximum and minimum elevation values from a GPS recorded at collecting localities and using Google Earth. The predicted range size of
<taxonomicName id="92E2AE8E11104513FC0F52E4FBCAFE6D" box="[977,1065,477,500]" class="Reptilia" family="Chamaeleonidae" genus="Trioceros" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="nyirit" status="sp. nov.">
<emphasis id="6796091F11104513FC0F52E4FBCAFE6D" box="[977,1065,477,500]" italics="true" pageId="9" pageNumber="10">T. nyirit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7CA5B46411104513FBEC52E5FB6EFE6D" box="[1074,1165,476,500]" pageId="9" pageNumber="10" rank="species">
<emphasis id="6796091F11104513FBEC52E5FB6EFE6D" bold="true" box="[1074,1165,476,500]" pageId="9" pageNumber="10">sp. nov.</emphasis>
</taxonomicNameLabel>
in the Cherangani Hills (assuming that all sites&gt;
<quantity id="921A78E811104513FE73513BFE1FFD83" box="[429,508,514,538]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="9" pageNumber="10" unit="m" value="2900.0">2900m</quantity>
are potential habitat) is
<quantity id="921A78E811104513FCD5513BFCBBFD83" box="[779,856,514,538]" metricMagnitude="5" metricUnit="m" metricValue="7.54" pageId="9" pageNumber="10" unit="km" value="754.0">754km</quantity>
2 and on the Mtelo massif is
<quantity id="921A78E811104513FB48513BFB35FD83" box="[1174,1238,514,538]" metricMagnitude="4" metricUnit="m" metricValue="9.3" pageId="9" pageNumber="10" unit="km" value="93.0">93km</quantity>
2 (&gt;
<quantity id="921A78E811104513FB23513BFAB2FD83" box="[1277,1361,514,538]" metricMagnitude="3" metricUnit="m" metricValue="2.276" pageId="9" pageNumber="10" unit="m" value="2276.0">2276m</quantity>
).
</paragraph>
<caption id="019D858511104513FF4950F7FAF4FC7C" box="[151,1303,974,997]" httpUri="https://zenodo.org/record/201059/files/figure.png" pageId="9" pageNumber="10" targetBox="[226,1360,599,949]" targetPageId="9">
<paragraph id="555DD50D11104513FF4950F7FAF4FC7C" blockId="9.[151,1303,974,997]" box="[151,1303,974,997]" pageId="9" pageNumber="10">
<emphasis id="6796091F11104513FF4950F7FEF1FC7D" bold="true" box="[151,274,974,997]" pageId="9" pageNumber="10">FIGURE 5.</emphasis>
A. Adult male, holotype,
<taxonomicName id="92E2AE8E11104513FDC550E9FD88FC7C" box="[539,619,976,997]" class="Reptilia" family="Chamaeleonidae" genus="Trioceros" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="nyirit" status="sp. nov.">
<emphasis id="6796091F11104513FDC550E9FD88FC7C" box="[539,619,976,997]" italics="true" pageId="9" pageNumber="10">T. nyirit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7CA5B46411104513FDAF50F6FD22FC7D" box="[625,705,975,996]" pageId="9" pageNumber="10" rank="species">
<emphasis id="6796091F11104513FDAF50F6FD22FC7D" bold="true" box="[625,705,975,996]" pageId="9" pageNumber="10">sp. nov.</emphasis>
</taxonomicNameLabel>
, Gatau Pass, Mtelo massif. B. Adult female, same locality.
</paragraph>
</caption>
<caption id="019D858511104513FF495408FCBEF8FF" httpUri="https://zenodo.org/record/201060/files/figure.png" pageId="9" pageNumber="10" targetBox="[217,1369,1052,1820]" targetPageId="9">
<paragraph id="555DD50D11104513FF495408FCBEF8FF" blockId="9.[151,1436,1841,1894]" pageId="9" pageNumber="10">
<emphasis id="6796091F11104513FF495408FEF7F8DE" bold="true" box="[151,276,1841,1864]" pageId="9" pageNumber="10">FIGURE 6.</emphasis>
Habitat of
<taxonomicName id="92E2AE8E11104513FE52540AFE3DF8D1" box="[396,478,1843,1864]" class="Reptilia" family="Chamaeleonidae" genus="Trioceros" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="nyirit" status="sp. nov.">
<emphasis id="6796091F11104513FE52540AFE3DF8D1" box="[396,478,1843,1864]" italics="true" pageId="9" pageNumber="10">T. nyirit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7CA5B46411104513FE38540BFDDAF8DE" box="[486,569,1842,1863]" pageId="9" pageNumber="10" rank="species">
<emphasis id="6796091F11104513FE38540BFDDAF8DE" bold="true" box="[486,569,1842,1863]" pageId="9" pageNumber="10">sp. nov.</emphasis>
</taxonomicNameLabel>
on the Mtelo massif. Specimens were found in the ericaceous zone (foreground) at 3000m and in shrubs adjacent to the forest at 2800m (middle ground).
</paragraph>
</caption>
<paragraph id="555DD50D11104513FF1B54AAFC68F86A" blockId="9.[151,1436,1939,2035]" pageId="9" pageNumber="10">
<emphasis id="6796091F11104513FF1B54AAFECAF832" bold="true" box="[197,297,1939,1963]" pageId="9" pageNumber="10">Ecology.</emphasis>
On the Mtelo massif,
<taxonomicName id="92E2AE8E11104513FDFF54ADFD94F832" box="[545,631,1940,1963]" class="Reptilia" family="Chamaeleonidae" genus="Trioceros" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="nyirit" status="sp. nov.">
<emphasis id="6796091F11104513FDFF54ADFD94F832" box="[545,631,1940,1963]" italics="true" pageId="9" pageNumber="10">T. nyirit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7CA5B46411104513FDA154AAFD3BF832" box="[639,728,1939,1963]" pageId="9" pageNumber="10" rank="species">
<emphasis id="6796091F11104513FDA154AAFD3BF832" bold="true" box="[639,728,1939,1963]" pageId="9" pageNumber="10">sp. nov.</emphasis>
</taxonomicNameLabel>
was found on shrubs at the edges of cleared afromontane forest and also in the ericaceous zone above the forest. In the Cherangani Hills, specimens were found on shrubs and small trees at the edge of fields, on hedges and roadside vegetation.
</paragraph>
<paragraph id="555DD50D11134510FF1953AEFCB1FEFD" blockId="10.[151,1437,151,356]" pageId="10" pageNumber="11">
<emphasis id="6796091F11134510FF1953AEFE58FF29" bold="true" box="[199,443,151,176]" pageId="10" pageNumber="11">Conservation status.</emphasis>
<collectingCountry id="2DF5959D11134510FE1F53A1FE03FF29" box="[449,480,152,176]" name="American Samoa" pageId="10" pageNumber="11">As</collectingCountry>
well as occurring in ericaceous afroalpine vegetation and shrubs at the forest edge,
<taxonomicName id="92E2AE8E11134510FA5953A0FF30FF4D" class="Reptilia" family="Chamaeleonidae" genus="Trioceros" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="nyirit" status="sp. nov.">
<emphasis id="6796091F11134510FA5953A0FF30FF4D" italics="true" pageId="10" pageNumber="11">T. nyirit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7CA5B46411134510FF045385FED7FF4D" box="[218,308,188,212]" pageId="10" pageNumber="11" rank="species">
<emphasis id="6796091F11134510FF045385FED7FF4D" bold="true" box="[218,308,188,212]" pageId="10" pageNumber="11">sp. nov.</emphasis>
</taxonomicNameLabel>
were also collected in disturbed habitats in agricultural landscapes and appears to be relatively abundant in these areas. This suggests that despite its limited distribution it does not seem to be threatened by anthropogenic activities. However, deforestation and conversion of natural habitats to agriculture on the Mtelo massif continues to reduce forest cover (John Yoposiwa, pers. comm.) and rates of habitat change are believed to be high also in the Cherangani (
<bibRefCitation id="3173A8FC11134510FE7E5275FDFCFEFD" author="Wass" box="[416,543,332,356]" pageId="10" pageNumber="11" refString="Wass, P. (Ed) (1995) Kenya's indigenous forests. Status, management and conservation. The World Conservation Union, Gland, Switzerland." type="book" year="1995">Wass 1995</bibRefCitation>
;
<bibRefCitation id="3173A8FC11134510FDF45275FCA4FEFD" author="Akotsi" box="[554,839,332,356]" pageId="10" pageNumber="11" refString="Akotsi, E. &amp; Gachanja, M. (2004) Changes in forest cover in Kenya's five ' Water towers' 2000 - 2003. Kenya Forests Working Group, Nairobi." type="book chapter" year="2004">Akotsi &amp; Gachanja 2004</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>