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<document id="32DAB184CDF949B81517DC9BB59516E4" ID-DOI="10.5281/zenodo.5340977" ID-ISSN="0374-1036" ID-Zenodo-Dep="5340977" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630188923135" checkinUser="felipe" docAuthor="Zettel, Herbert &amp; Gapud, Victor P." docDate="2008" docId="03BBD93AFFDC4B7056FF3F96FB93FC0B" docLanguage="en" docName="ActaEntMusNatPra.2008.48.2.319-329.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 48 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Orthosaldula Gapud 1986" docType="treatment" docVersion="4" lastPageNumber="321" masterDocId="FF82A142FFDD4B72573B3D66FFAAFFD8" masterDocTitle="Notes on the endemic Philippine genus Orthosaldula (Hemiptera: Heteroptera: Saldidae), with descriptions of two new species" masterLastPageNumber="329" masterPageNumber="319" pageNumber="320" updateTime="1699180942148" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="119F3188A167D49D16643E515907881A">Notes on the endemic Philippine genus Orthosaldula (Hemiptera: Heteroptera: Saldidae), with descriptions of two new species</mods:title>
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<mods:namePart id="BBC498A8E1B849512C96E19D65150D5C">Zettel, Herbert</mods:namePart>
<mods:affiliation id="125973D0CE8CAF55F8B4CAAA476DAE98">Natural History Museum, International Research Institute for Entomology, Burgring 7, A- 1010 Vienna, Austria; e-mail: herbert. zettel @ nhm-wien. ac. at</mods:affiliation>
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<mods:namePart id="C9EE94AEC3BD40324BC2B47429D89501">Gapud, Victor P.</mods:namePart>
<mods:affiliation id="263DED54AF08DFCFB6B0D778E5C1E5CC">Crop Protection Cluster, University of the Philippines, Los Baños, College, Laguna 4031, The Philippines; e-mail: vicgap @ laguna. net</mods:affiliation>
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<mods:title id="25A1BE0C5F224499DE811EA5B2A89060">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<treatment id="03BBD93AFFDC4B7056FF3F96FB93FC0B" ID-DOI="http://doi.org/10.5281/zenodo.5443053" ID-Zenodo-Dep="5443053" LSID="urn:lsid:plazi:treatment:03BBD93AFFDC4B7056FF3F96FB93FC0B" httpUri="http://treatment.plazi.org/id/03BBD93AFFDC4B7056FF3F96FB93FC0B" lastPageId="2" lastPageNumber="321" pageId="1" pageNumber="320">
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<paragraph id="8BAD682CFFDC4B7356FF3F96FD47FCD1" blockId="1.[452,749,752,777]" box="[452,749,752,777]" pageId="1" pageNumber="320">
<heading id="D0E5DF40FFDC4B7356FF3F96FD47FCD1" bold="true" box="[452,749,752,777]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="320" reason="3">
<taxonomicName id="4C1213AFFFDC4B7356FF3F96FD47FCD1" ID-CoL="63H9D" authority="Gapud, 1986" authorityName="Gapud" authorityYear="1986" box="[452,749,752,777]" class="Insecta" family="Saldidae" genus="Orthosaldula" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="320" phylum="Arthropoda" rank="genus">
<emphasis id="B966B43EFFDC4B7356FF3F96FD47FCD1" bold="true" box="[452,749,752,777]" pageId="1" pageNumber="320">
<emphasis id="B966B43EFFDC4B7356FF3F96FDFFFCD1" bold="true" box="[452,597,752,777]" italics="true" pageId="1" pageNumber="320">Orthosaldula</emphasis>
<bibRefCitation id="EF8315DDFFDC4B7355673F97FD47FCD1" author="GAPUD V. P." box="[604,749,753,777]" pageId="1" pageNumber="320" pagination="1 - 47" refId="ref4388" refString="GAPUD V. P. 1986: Philippine Water Bugs. Guide to Philippine Flora and Fauna 8: 1 - 47." type="journal article" year="1986">Gapud, 1986</bibRefCitation>
</emphasis>
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</paragraph>
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<paragraph id="8BAD682CFFDC4B73574A3E7EFEAEFC9D" blockId="1.[113,1089,791,864]" pageId="1" pageNumber="320">
<treatmentCitationGroup id="AB024F02FFDC4B73574A3E7EFEAEFC9D" pageId="1" pageNumber="320">
<treatmentCitation id="0AB34E3DFFDC4B73574A3E7EFEDAFCF3" author="GAPUD V. P." box="[113,368,791,811]" page="7" pageId="1" pageNumber="320" year="1986">
<taxonomicName id="4C1213AFFFDC4B73574A3E7EFEDAFCF3" ID-CoL="63H9D" authority="Gapud, 1986: 7" authorityName="Gapud" authorityPageNumber="7" authorityYear="1986" box="[113,368,791,811]" class="Insecta" family="Saldidae" genus="Orthosaldula" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="320" phylum="Arthropoda" rank="genus">
<emphasis id="B966B43EFFDC4B73574A3E7EFF49FCF2" box="[113,227,792,810]" italics="true" pageId="1" pageNumber="320">Orthosaldula</emphasis>
<bibRefCitation id="EF8315DDFFDC4B7357D23E71FEDAFCF3" author="GAPUD V. P." box="[233,368,791,811]" pageId="1" pageNumber="320" pagination="1 - 47" refId="ref4388" refString="GAPUD V. P. 1986: Philippine Water Bugs. Guide to Philippine Flora and Fauna 8: 1 - 47." type="journal article" year="1986">Gapud, 1986: 7</bibRefCitation>
</taxonomicName>
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(key), 16-17 (description) (originally quoted as Gapud, 1977 [sic!], see Taxonomic notes below).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAD682CFFDC4B73574A3E2BFCEEFCB8" blockId="1.[113,1089,791,864]" box="[113,836,844,864]" pageId="1" pageNumber="320">
<emphasis id="B966B43EFFDC4B73574A3E2BFF78FC87" box="[113,210,845,863]" italics="true" pageId="1" pageNumber="320">Orthosalda</emphasis>
[sic!]:
<bibRefCitation id="EF8315DDFFDC4B73562B3E2BFE0AFCB8" author="DUDGEON D." box="[272,416,845,864]" pageId="1" pageNumber="320" refId="ref4363" refString="DUDGEON D. 1999: Tropical Asian Streams. Zoobenthos, Ecology and Conservation. Hong Kong University Press, xii + 830 pp." type="book" year="1999">DUDGEON (1999)</bibRefCitation>
: 356 (mentioned as a Philippine endemic genus).
</paragraph>
</subSubSection>
<subSubSection id="C3083BA7FFDC4B73574A3E1EFCFBFC48" box="[113,849,888,912]" pageId="1" pageNumber="320" type="type_taxon">
<paragraph id="8BAD682CFFDC4B73574A3E1EFCFBFC48" blockId="1.[113,849,888,912]" box="[113,849,888,912]" pageId="1" pageNumber="320">
<emphasis id="B966B43EFFDC4B73574A3E1EFEAFFC48" bold="true" box="[113,261,888,912]" pageId="1" pageNumber="320">
<typeStatus id="54A9D68EFFDC4B73574A3E1EFF03FC48" box="[113,169,888,912]" pageId="1" pageNumber="320">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C1213AFFFDC4B7356373E1FFD09FC48" ID-CoL="74YMK" authority="Gapud, 1986" authorityName="Gapud" authorityYear="1986" box="[268,675,888,912]" class="Insecta" family="Saldidae" genus="Orthosaldula" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="320" phylum="Arthropoda" rank="species" species="rubroalata">
<emphasis id="B966B43EFFDC4B7356373E1FFDBEFC48" box="[268,532,889,912]" italics="true" pageId="1" pageNumber="320">Orthosaldula rubroalata</emphasis>
<bibRefCitation id="EF8315DDFFDC4B7355213E1EFD09FC48" author="GAPUD V. P." box="[538,675,888,912]" pageId="1" pageNumber="320" pagination="1 - 47" refId="ref4388" refString="GAPUD V. P. 1986: Philippine Water Bugs. Guide to Philippine Flora and Fauna 8: 1 - 47." type="journal article" year="1986">Gapud, 1986</bibRefCitation>
</taxonomicName>
(by monotypy).
</paragraph>
</subSubSection>
<subSubSection id="C3083BA7FFDC4B70574A3EC1FE3DFD6B" lastPageId="2" lastPageNumber="321" pageId="1" pageNumber="320" type="diagnosis">
<paragraph id="8BAD682CFFDC4B70574A3EC1FB85FE2B" blockId="1.[113,1089,935,1535]" lastBlockId="2.[151,1127,219,980]" lastPageId="2" lastPageNumber="321" pageId="1" pageNumber="320">
<emphasis id="B966B43EFFDC4B73574A3EC1FF4CFC67" bold="true" box="[113,230,935,959]" pageId="1" pageNumber="320">Diagnosis.</emphasis>
Body small, slender. Colour (
<figureCitation id="132974A9FFDC4B73551F3EC1FD29FC67" box="[548,643,935,959]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Figs" captionStartId-0="4.[151,182,1474,1493]" captionStartId-1="5.[113,144,1533,1552]" captionStartId-2="7.[113,144,1546,1565]" captionStartId-3="8.[688,728,1493,1512]" captionTargetBox-0="[261,1015,220,1443]" captionTargetBox-1="[223,979,528,1500]" captionTargetBox-2="[238,962,411,1515]" captionTargetBox-3="[151,637,219,1587]" captionTargetId-0="figure-10@4.[260,1015,219,1443]" captionTargetId-1="figure-141@5.[223,980,527,1500]" captionTargetId-2="figure-13@7.[238,963,410,1515]" captionTargetId-3="figure-240@8.[151,637,219,1588]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="Fig. 1. Orthosaldula rubroalata Gapud, 1986, female, body length 3.3 mm. (© NHMW Hemiptera Image Collection, published with permission)." captionText-1="Fig. 2. Orthosaldula stysi sp. nov., female, body length 3.2 mm. (© NHMW Hemiptera Image Collection, published with permission)." captionText-2="Fig. 3. Orthosaldula flavonigra sp. nov., male, body length 2.8 mm. (© NHMW Hemiptera Image Collection, published with permission)." captionText-3="Figs. 4-6. Head, frontal aspect: 4 Orthosaldula rubroalata Gapud, 1986; 5 O. stysi sp. nov.; 6 O." figureDoi-0="http://doi.org/10.5281/zenodo.5340981" figureDoi-1="http://doi.org/10.5281/zenodo.5340983" figureDoi-2="http://doi.org/10.5281/zenodo.5340985" figureDoi-3="http://doi.org/10.5281/zenodo.5340987" httpUri-0="https://zenodo.org/record/5340981/files/figure.png" httpUri-1="https://zenodo.org/record/5340983/files/figure.png" httpUri-2="https://zenodo.org/record/5340985/files/figure.png" httpUri-3="https://zenodo.org/record/5340987/files/figure.png" pageId="1" pageNumber="320">Figs. 1-6</figureCitation>
) unique. Black, thorax and some areas on head with more or less metallic, bluish or violet traces. Head with yellow, orange, or reddish patches near anterodorsal eye margin and on mandibular plates; maxillary plates, anteclypeus, gular lobes, and labrum totally yellow, orange, or reddish. Metapleura pale. Forewing with ground colour red or yellow and with prominent black mark in centre, lateral areas pale. Antenna and legs chiefly pale, or with reddish traces, but procoxa always black; fourth antennomere infuscated in some specimens. Sternites
<quantity id="4CEAC5C9FFDC4B7355263901FDF1FBA7" box="[541,603,1127,1151]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.524" metricValueMin="0.508" pageId="1" pageNumber="320" unit="in" value="4.0" valueMax="6.0" valueMin="2.0">2-6 in</quantity>
both sexes with reddish hind margins, at least narrow and laterally; sternite 7 of female pale, almost transparent, except black stripe at base. Pilosity reduced, without long standing setae; short decumbent whitish pilosity present on some parts of head and thorax, most dominant on sides; hemelytron with short dark, appressed or decumbent setae. Vertex narrow. Ocelli located on distinct tubercle that is laterally delimited by deep, posteriad prolonged anteocellar furrows. Rostrum very long, reaching apex of hind coxa, third segment ca. 2.2 times as long as fourth. Antenna: antennomere 2 longest, relative lengths of antennomeres approximately as 5: 10: 9: 9, antennomeres 1 and 2 cylindrical in both sexes. Pronotum coarsely sculptured, with sides slightly concave and strongly converging anteriad, with posterior margin almost straight (slightly convex laterally, slightly concave medially); collar very narrow; callus narrow (median length less than two thirds of posterior lobe length) and domed, with impressed median line; transverse furrow separating callus from posterior lobe deeply punctured and reaching lateral margins; posterior lobe more or less domed, laterally with sharply impressed longitudinal furrows separating humeri from disk. Mesoscutum domed. Scutellum anteriorly strongly raised, posterior flattened. Hemelytron (
<figureCitation id="132974A9FFDF4B7057A43C7DFF74FEEC" box="[159,222,283,308]" captionStart="Figs" captionStartId="9.[113,152,1087,1106]" captionTargetBox="[122,1085,222,1055]" captionTargetId="figure-145@9.[113,1088,219,1061]" captionTargetPageId="9" captionText="Figs. 7-14. 7-11 Orthosaldula rubroalata Gapud, 1986: 7 right forewing (ef = embolar fracture, R = radius); 8 parandria, caudal view; 9 proctiger, dorsal view; 10 left paramere, lateral view; 11 subgenital plate of female, ventral view. 12-14 dorsal outline of pronotum, lateral view (co = collar, ca = callus, pl = posterior lobe). 12 O. rubroalata; 13 O. stysi sp. nov.; 14 O. flavonigra sp. nov. Pilosity omitted, except in Fig. 10." figureDoi="http://doi.org/10.5281/zenodo.5340989" httpUri="https://zenodo.org/record/5340989/files/figure.png" pageId="2" pageNumber="321">Fig. 7</figureCitation>
) with short embolar fracture (ef), not reaching level of apex of radius (R); membrane with four cells, innermost cell (relative to wing the posterior one) shorter and narrower than the following cell; hypocostal ridge simple, secondary costal ridge absent. Genital capsule of male with horn-shaped parandria (
<figureCitation id="132974A9FFDF4B7055393C1DFDE8FE4C" box="[514,578,379,404]" captionStart="Figs" captionStartId="9.[113,152,1087,1106]" captionTargetBox="[122,1085,222,1055]" captionTargetId="figure-145@9.[113,1088,219,1061]" captionTargetPageId="9" captionText="Figs. 7-14. 7-11 Orthosaldula rubroalata Gapud, 1986: 7 right forewing (ef = embolar fracture, R = radius); 8 parandria, caudal view; 9 proctiger, dorsal view; 10 left paramere, lateral view; 11 subgenital plate of female, ventral view. 12-14 dorsal outline of pronotum, lateral view (co = collar, ca = callus, pl = posterior lobe). 12 O. rubroalata; 13 O. stysi sp. nov.; 14 O. flavonigra sp. nov. Pilosity omitted, except in Fig. 10." figureDoi="http://doi.org/10.5281/zenodo.5340989" httpUri="https://zenodo.org/record/5340989/files/figure.png" pageId="2" pageNumber="321">Fig. 8</figureCitation>
); proctiger (
<figureCitation id="132974A9FFDF4B7055FF3C1DFCA9FE4C" box="[708,771,379,404]" captionStart="Figs" captionStartId="9.[113,152,1087,1106]" captionTargetBox="[122,1085,222,1055]" captionTargetId="figure-145@9.[113,1088,219,1061]" captionTargetPageId="9" captionText="Figs. 7-14. 7-11 Orthosaldula rubroalata Gapud, 1986: 7 right forewing (ef = embolar fracture, R = radius); 8 parandria, caudal view; 9 proctiger, dorsal view; 10 left paramere, lateral view; 11 subgenital plate of female, ventral view. 12-14 dorsal outline of pronotum, lateral view (co = collar, ca = callus, pl = posterior lobe). 12 O. rubroalata; 13 O. stysi sp. nov.; 14 O. flavonigra sp. nov. Pilosity omitted, except in Fig. 10." figureDoi="http://doi.org/10.5281/zenodo.5340989" httpUri="https://zenodo.org/record/5340989/files/figure.png" pageId="2" pageNumber="321">Fig. 9</figureCitation>
) subtrapezoidal, with moderately convex hind margin; paramere (
<figureCitation id="132974A9FFDF4B7056D53CFDFD91FE6C" box="[494,571,411,436]" captionStart="Figs" captionStartId="9.[113,152,1087,1106]" captionTargetBox="[122,1085,222,1055]" captionTargetId="figure-145@9.[113,1088,219,1061]" captionTargetPageId="9" captionText="Figs. 7-14. 7-11 Orthosaldula rubroalata Gapud, 1986: 7 right forewing (ef = embolar fracture, R = radius); 8 parandria, caudal view; 9 proctiger, dorsal view; 10 left paramere, lateral view; 11 subgenital plate of female, ventral view. 12-14 dorsal outline of pronotum, lateral view (co = collar, ca = callus, pl = posterior lobe). 12 O. rubroalata; 13 O. stysi sp. nov.; 14 O. flavonigra sp. nov. Pilosity omitted, except in Fig. 10." figureDoi="http://doi.org/10.5281/zenodo.5340989" httpUri="https://zenodo.org/record/5340989/files/figure.png" pageId="2" pageNumber="321">Fig. 10</figureCitation>
) strongly curved, with weakly developed processus sensualis. Phallus complex, containing numerous minute sclerites. Subgenital plate of female (
<figureCitation id="132974A9FFDF4B7057A43CBDFF41FE2C" box="[159,235,475,500]" captionStart="Figs" captionStartId="9.[113,152,1087,1106]" captionTargetBox="[122,1085,222,1055]" captionTargetId="figure-145@9.[113,1088,219,1061]" captionTargetPageId="9" captionText="Figs. 7-14. 7-11 Orthosaldula rubroalata Gapud, 1986: 7 right forewing (ef = embolar fracture, R = radius); 8 parandria, caudal view; 9 proctiger, dorsal view; 10 left paramere, lateral view; 11 subgenital plate of female, ventral view. 12-14 dorsal outline of pronotum, lateral view (co = collar, ca = callus, pl = posterior lobe). 12 O. rubroalata; 13 O. stysi sp. nov.; 14 O. flavonigra sp. nov. Pilosity omitted, except in Fig. 10." figureDoi="http://doi.org/10.5281/zenodo.5340989" httpUri="https://zenodo.org/record/5340989/files/figure.png" pageId="2" pageNumber="321">Fig. 11</figureCitation>
) caudally produced, in middle of hind margin with very small, indistinct lobe.
</paragraph>
<paragraph id="8BAD682CFFDF4B7057AC3C9DFE3DFD6B" blockId="2.[151,1127,219,980]" pageId="2" pageNumber="321">
<emphasis id="B966B43EFFDF4B7057AC3C9DFED2FDCC" bold="true" box="[151,376,507,532]" pageId="2" pageNumber="321">Comparative notes.</emphasis>
<taxonomicName id="4C1213AFFFDF4B7056BA3C9BFDA5FDCC" authorityName="Gapud" authorityYear="1986" box="[385,527,509,532]" class="Insecta" family="Saldidae" genus="Orthosaldula" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="B966B43EFFDF4B7056BA3C9BFDA5FDCC" box="[385,527,509,532]" italics="true" pageId="2" pageNumber="321">Orthosaldula</emphasis>
</taxonomicName>
belongs to the subfamily
<taxonomicName id="4C1213AFFFDF4B7054093C9DFB9BFDCB" authority="(GAPUD 1986)" baseAuthorityName="GAPUD" baseAuthorityYear="1986" box="[818,1073,507,532]" class="Insecta" family="Saldidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="321" phylum="Arthropoda" rank="subFamily" subFamily="Saldinae">
Saldinae (
<bibRefCitation id="EF8315DDFFDF4B7054A43C9AFB83FDCB" author="GAPUD V. P." box="[927,1065,507,532]" pageId="2" pageNumber="321" pagination="1 - 47" refId="ref4388" refString="GAPUD V. P. 1986: Philippine Water Bugs. Guide to Philippine Flora and Fauna 8: 1 - 47." type="journal article" year="1986">GAPUD 1986</bibRefCitation>
)
</taxonomicName>
. The modifications of the pronotum and the vivid colour pattern are unique. Especially noteworthy are the coarse sculpture of the pronotum, and the narrow collar and callus as compared with the long, domed, posterior lobe with almost straight hind margin, which distinguish
<taxonomicName id="4C1213AFFFDF4B7054C43F3BFF6CFD4C" authorityName="Gapud" authorityYear="1986" class="Insecta" family="Saldidae" genus="Orthosaldula" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="B966B43EFFDF4B7054C43F3BFF6CFD4C" italics="true" pageId="2" pageNumber="321">Orthosaldula</emphasis>
</taxonomicName>
from all other
<taxonomicName id="4C1213AFFFDF4B7056493F1DFE64FD4B" authorityName="Amyot &amp; Serville" authorityYear="1843" box="[370,462,635,659]" class="Insecta" family="Saldidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="321" phylum="Arthropoda" rank="family">Saldidae</taxonomicName>
in the Oriental Region. The relationships with other saldine genera are not resolved.
</paragraph>
</subSubSection>
<subSubSection id="C3083BA7FFDF4B7057AC3FDDFB93FC0B" pageId="2" pageNumber="321" type="distribution">
<paragraph id="8BAD682CFFDF4B7057AC3FDDFB93FC0B" blockId="2.[151,1127,219,980]" pageId="2" pageNumber="321">
<emphasis id="B966B43EFFDF4B7057AC3FDDFDB2FD0B" bold="true" box="[151,536,699,724]" pageId="2" pageNumber="321">Distribution and species diversity.</emphasis>
Endemic to the
<collectingCountry id="F30528BCFFDF4B7055FF3FDDFCEAFD0B" box="[708,832,699,723]" name="Philippines" pageId="2" pageNumber="321">Philippines</collectingCountry>
. Hitherto known from three islands, Luzon,
<collectingRegion id="49D6A6CEFFDF4B7056003FBAFEDDFD2C" box="[315,375,732,756]" country="Philippines" name="Southern Leyte" pageId="2" pageNumber="321">Leyte</collectingRegion>
and
<collectingRegion id="49D6A6CEFFDF4B70569C3FBDFE48FD2B" box="[423,482,731,755]" country="Philippines" name="Cebu" pageId="2" pageNumber="321">Cebu</collectingRegion>
, each home of one species.A fourth species has been collected in northern Luzon (J. T. Polhemus, pers. comm.), but was not made available for our study.
<emphasis id="B966B43EFFDF4B7057AC3E7DFEF3FCEC" bold="true" box="[151,345,795,820]" pageId="2" pageNumber="321">Taxonomic notes.</emphasis>
<bibRefCitation id="EF8315DDFFDF4B7056663E7AFE47FCEB" author="GAPUD V. P." box="[349,493,795,820]" pageId="2" pageNumber="321" pagination="1 - 47" refId="ref4388" refString="GAPUD V. P. 1986: Philippine Water Bugs. Guide to Philippine Flora and Fauna 8: 1 - 47." type="journal article" year="1986">GAPUD (1986)</bibRefCitation>
introduced the new genus
<taxonomicName id="4C1213AFFFDF4B7055C53E7BFC20FCEC" authorityName="Gapud" authorityYear="1986" box="[766,906,797,820]" class="Insecta" family="Saldidae" genus="Orthosaldula" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="B966B43EFFDF4B7055C53E7BFC20FCEC" box="[766,906,797,820]" italics="true" pageId="2" pageNumber="321">Orthosaldula</emphasis>
</taxonomicName>
and four new species names,
<taxonomicName id="4C1213AFFFDF4B7057D03E5BFE5FFC8C" authorityName="Gapud" authorityYear="1986" box="[235,501,829,852]" class="Insecta" family="Saldidae" genus="Orthosaldula" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="321" phylum="Arthropoda" rank="species" species="rubroalata">
<emphasis id="B966B43EFFDF4B7057D03E5BFE5FFC8C" box="[235,501,829,852]" italics="true" pageId="2" pageNumber="321">Orthosaldula rubroalata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1213AFFFDF4B70553F3E5BFCEFFC8B" authority="Gapud, 1986" authorityName="Gapud" authorityYear="1986" box="[516,837,827,852]" class="Insecta" family="Saldidae" genus="Saldula" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="321" phylum="Arthropoda" rank="species" species="mimica">
<emphasis id="B966B43EFFDF4B70553F3E5BFD07FC8C" box="[516,685,829,852]" italics="true" pageId="2" pageNumber="321">Saldula mimica</emphasis>
<bibRefCitation id="EF8315DDFFDF4B70558D3E5DFCEFFC8B" author="GAPUD V. P." box="[694,837,827,851]" pageId="2" pageNumber="321" pagination="1 - 47" refId="ref4388" refString="GAPUD V. P. 1986: Philippine Water Bugs. Guide to Philippine Flora and Fauna 8: 1 - 47." type="journal article" year="1986">Gapud, 1986</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C1213AFFFDF4B7054BA3E5BFE89FCAB" authority="Gapud, 1986" authorityName="Gapud" authorityYear="1986" class="Insecta" family="Saldidae" genus="Chartoscirta" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="321" phylum="Arthropoda" rank="species" species="mayona">
<emphasis id="B966B43EFFDF4B7054BA3E5BFBCCFC8C" box="[897,1126,829,852]" italics="true" pageId="2" pageNumber="321">Chartoscirta mayona</emphasis>
<bibRefCitation id="EF8315DDFFDF4B7057AC3E3DFE89FCAB" author="GAPUD V. P." box="[151,291,859,883]" pageId="2" pageNumber="321" pagination="1 - 47" refId="ref4388" refString="GAPUD V. P. 1986: Philippine Water Bugs. Guide to Philippine Flora and Fauna 8: 1 - 47." type="journal article" year="1986">
Gapud,
<quantity id="4CEAC5C9FFDF4B7057D53E3DFE89FCAB" box="[238,291,859,883]" metricMagnitude="1" metricUnit="m" metricValue="5.04444" pageId="2" pageNumber="321" unit="in" value="1986.0">1986</quantity>
</bibRefCitation>
</taxonomicName>
in
<taxonomicName id="4C1213AFFFDF4B7056773E3DFE01FCAB" authorityName="Amyot &amp; Serville" authorityYear="1843" box="[332,427,859,883]" class="Insecta" family="Saldidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="321" phylum="Arthropoda" rank="family">Saldidae</taxonomicName>
,
<taxonomicName id="4C1213AFFFDF4B7056833E3BFC93FCAB" authority="Gapud, 1986" authorityName="Gapud" authorityYear="1986" box="[440,825,859,884]" class="Insecta" family="Omaniidae" genus="Corallocoris" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="321" phylum="Arthropoda" rank="species" species="cinereus">
<emphasis id="B966B43EFFDF4B7056833E3BFD08FCAC" box="[440,674,861,884]" italics="true" pageId="2" pageNumber="321">Corallocoris cinereus</emphasis>
<bibRefCitation id="EF8315DDFFDF4B7055973E3DFC93FCAB" author="GAPUD V. P." box="[684,825,859,883]" pageId="2" pageNumber="321" pagination="1 - 47" refId="ref4388" refString="GAPUD V. P. 1986: Philippine Water Bugs. Guide to Philippine Flora and Fauna 8: 1 - 47." type="journal article" year="1986">
Gapud,
<quantity id="4CEAC5C9FFDF4B70543F3E3DFC93FCAB" box="[772,825,859,883]" metricMagnitude="1" metricUnit="m" metricValue="5.04444" pageId="2" pageNumber="321" unit="in" value="1986.0">1986</quantity>
</bibRefCitation>
</taxonomicName>
in
<taxonomicName id="4C1213AFFFDF4B70545A3E3DFC70FCAB" authorityName="Cobben" authorityYear="1970" box="[865,986,859,883]" class="Insecta" family="Omaniidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="321" phylum="Arthropoda" rank="family">Omaniidae</taxonomicName>
. By a lapsus calami, these taxa were referred to Gapud, 1977, a publication that does not exist. However, the study by
<bibRefCitation id="EF8315DDFFDF4B70561B3EFAFE19FC6B" author="GAPUD V. P." box="[288,435,923,948]" pageId="2" pageNumber="321" pagination="1 - 47" refId="ref4388" refString="GAPUD V. P. 1986: Philippine Water Bugs. Guide to Philippine Flora and Fauna 8: 1 - 47." type="journal article" year="1986">GAPUD (1986)</bibRefCitation>
provides descriptions of all new taxa and therefore all names are fully available according to the requirements by the ICZN, Third Edition (
<bibRefCitation id="EF8315DDFFDF4B7054963EDAFB87FC0B" author="ICZN" box="[941,1069,955,980]" pageId="2" pageNumber="321" refId="ref4411" refString="ICZN 1985: International Code of Zoological Nomenclature. Third edition. International Trust for Zoological Museum, London, 388 pp." type="book" year="1985">ICZN 1985</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>