treatments-xml/data/03/93/AD/0393AD2C6645FFECFF56FD0CFC3AF898.xml
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<document id="C1458B1CAA551769A429B9ACAE2C8DF7" ID="10.11646/zootaxa.4244.3.5SLASH10568" ID-CLB-Dataset="36229" ID-DOI="10.11646/zootaxa.4244.3.5" ID-GBIF-Dataset="6772faf1-98da-4f04-b15c-80aa86086dab" ID-ISSN="1175-5326" ID-Zenodo-Dep="428269" ID-ZooBank="909345AC-276E-4D8D-96CF-620FBC5EB6B4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1490085696906" checkinUser="plazi" docAuthor="Roni, Narengbam, Sarbojit, Thaosen &amp; Vishwanath, Waikhom" docDate="2017" docId="0393AD2C6645FFECFF56FD0CFC3AF898" docLanguage="en" docName="zootaxa.4244.3.5SLASH10568.pdf" docOrigin="Zootaxa 4244 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Garra clavirostris Roni, Sarbojit &amp; Vishwanath, 2017, new species" docType="treatment" docVersion="9" lastPageNumber="372" masterDocId="FFAAD5546644FFE9FFC1FFDAFFDDFF89" masterDocTitle="Garra clavirostris, a new cyprinid fish (Teleostei: Cyprinidae: Labeoninae) from the Brahmaputra drainage, India" masterLastPageNumber="376" masterPageNumber="367" pageNumber="368" updateTime="1698438209736" updateUser="ExternalLinkService">
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<mods:title id="A9A4BAEFA8EB9172F972D64B9E34EF49">Garra clavirostris, a new cyprinid fish (Teleostei: Cyprinidae: Labeoninae) from the Brahmaputra drainage, India</mods:title>
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<mods:namePart id="29FE5B57B1152417980964038637A384">Roni, Narengbam</mods:namePart>
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<mods:namePart id="E3E6A8F800B7DA346831D676E6B20D86">Sarbojit, Thaosen</mods:namePart>
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<mods:namePart id="ECEB74F081DD070B5283E27C55C22513">Vishwanath, Waikhom</mods:namePart>
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<mods:title id="DD8350B652E1E11843AA24EC29D56799">Zootaxa</mods:title>
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<mods:part id="1A8C2FDE3EDCC4CE9054AF664264BC94">
<mods:date id="F94F953D66DF33EF275195F751FC338F">2017</mods:date>
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<treatment id="0393AD2C6645FFECFF56FD0CFC3AF898" ID-DOI="http://doi.org/10.5281/zenodo.5685559" ID-GBIF-Taxon="127848125" ID-Zenodo-Dep="5685559" LSID="urn:lsid:plazi:treatment:0393AD2C6645FFECFF56FD0CFC3AF898" httpUri="http://treatment.plazi.org/id/0393AD2C6645FFECFF56FD0CFC3AF898" lastPageId="5" lastPageNumber="372" pageId="1" pageNumber="368">
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<paragraph id="8B851C3A6645FFE8FF56FD0CFDD6FD79" blockId="1.[151,523,726,786]" box="[151,523,726,752]" pageId="1" pageNumber="368">
<heading id="D0CDAB566645FFE8FF56FD0CFDD6FD79" bold="true" box="[151,523,726,752]" fontSize="11" level="1" pageId="1" pageNumber="368" reason="1">
<emphasis id="B94EC0286645FFE8FF56FD0CFDD6FD79" bold="true" box="[151,523,726,752]" pageId="1" pageNumber="368">
<taxonomicName id="4C3A67B96645FFE8FF56FD0CFEB1FD79" box="[151,364,726,752]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="368" phylum="Chordata" rank="species" species="clavirostris" status="sp. nov.">
<emphasis id="B94EC0286645FFE8FF56FD0CFEB1FD79" bold="true" box="[151,364,726,752]" italics="true" pageId="1" pageNumber="368">Garra clavirostris</emphasis>
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,
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</emphasis>
</heading>
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<paragraph id="8B851C3A6645FFE8FF56FD23FF36FC9B" blockId="1.[151,523,726,786]" box="[151,235,761,786]" pageId="1" pageNumber="368">
(
<figureCitation id="130100BF6645FFE8FF5EFD23FF3DFC9B" box="[159,224,761,786]" captionStart="FIGURE 1" captionStartId="2.[151,250,1110,1132]" captionTargetBox="[227,1356,280,1073]" captionTargetId="figure@2.[217,1369,265,1089]" captionTargetPageId="2" captionText="FIGURE 1. Garra clavirostris, sp. nov., holotype, MUMF 22004, 117.5 mm SL, male; a. lateral, b. dorsal and c. ventral views. India: Assam: Dima Hasao District: Dilaima River at Boro Chenam Village." httpUri="https://zenodo.org/record/428274/files/figure.png" pageId="1" pageNumber="368">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3204FB16645FFE8FF56FC9BFED7FBBB" pageId="1" pageNumber="368" type="materials_examined">
<paragraph id="8B851C3A6645FFE8FF56FC9BFD4EFC2B" blockId="1.[151,1437,833,2010]" pageId="1" pageNumber="368">
<materialsCitation id="3B5216676645FFE8FF56FC9BFD4EFC2B" ID-GBIF-Occurrence="1440783601" collectingDate="2015-04-19" collectionCode="MUMF" country="India" county="Dima Hasao District" elevation="401" latitude="25.300833" location="Boro Chenam Village" longLatPrecision="21" longitude="92.86806" municipality="Dilaima River" pageId="1" pageNumber="368" specimenCode="MUMF 22004, 117.5" specimenCount="1" stateProvince="Assam" typeStatus="holotype">
<emphasis id="B94EC0286645FFE8FF56FC9BFED0FCD0" bold="true" box="[151,269,833,857]" pageId="1" pageNumber="368">
<typeStatus id="5481A2986645FFE8FF56FC9BFED5FCD0" box="[151,264,833,857]" pageId="1" pageNumber="368" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCode id="DB9CB4416645FFE8FED2FC98FDDCFCD3" box="[275,513,833,858]" pageId="1" pageNumber="368">
<collectionCode id="ED2B84FF6645FFE8FED2FC98FEB4FCD0" box="[275,361,834,857]" country="India" httpUri="http://grbio.org/cool/x8gv-47he" name="Department of Life Sciences" pageId="1" pageNumber="368">MUMF</collectionCode>
<quantity id="4CC2B1DF6645FFE8FEAEFC98FDDCFCD3" box="[367,513,833,858]" metricMagnitude="4" metricUnit="m" metricValue="2.2004117499999998" pageId="1" pageNumber="368" unit="mm" value="2.20041175E7">22004, 117.5</quantity>
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mm SL, male;
<collectingCountry id="F32D5CAA6645FFE8FD6DFC9BFD37FCD3" box="[684,746,833,858]" name="India" pageId="1" pageNumber="368">India</collectingCountry>
:
<collectingRegion id="49FED2D86645FFE8FD32FC98FC99FCD3" box="[755,836,834,858]" country="India" name="Assam" pageId="1" pageNumber="368">Assam</collectingRegion>
:
<collectingCounty id="62E464B66645FFE8FC8FFC9BFBE5FCD3" box="[846,1080,833,858]" pageId="1" pageNumber="368">Dima Hasao District</collectingCounty>
:
<collectingMunicipality id="6BE186406645FFE8FB80FC9BFB02FCD3" box="[1089,1247,833,858]" pageId="1" pageNumber="368">Dilaima River</collectingMunicipality>
at
<location id="8EE54AE16645FFE8FAC0FC98FF35FCF7" LSID="urn:lsid:plazi:treatment:0393AD2C6645FFECFF56FD0CFC3AF898:8EE54AE16645FFE8FAC0FC98FF35FCF7" country="India" county="Dima Hasao District" latitude="25.300833" longLatPrecision="21" longitude="92.86806" municipality="Dilaima River" name="Boro Chenam Village" pageId="1" pageNumber="368" stateProvince="Assam">Boro Chenam Village</location>
below the confluence of
<location id="8EE54AE16645FFE8FDCBFCBFFDB9FCF7" LSID="urn:lsid:plazi:treatment:0393AD2C6645FFECFF56FD0CFC3AF898:8EE54AE16645FFE8FDCBFCBFFDB9FCF7" box="[522,612,869,894]" country="India" county="Dima Hasao District" latitude="25.300833" longLatPrecision="21" longitude="92.86806" municipality="Dilaima River" name="Dilaima" pageId="1" pageNumber="368" stateProvince="Assam">Dilaima</location>
and
<location id="8EE54AE16645FFE8FD5FFCBFFD25FCF7" LSID="urn:lsid:plazi:treatment:0393AD2C6645FFECFF56FD0CFC3AF898:8EE54AE16645FFE8FD5FFCBFFD25FCF7" box="[670,760,869,894]" country="India" county="Dima Hasao District" latitude="25.300833" longLatPrecision="21" longitude="92.86806" municipality="Dilaima River" name="Dihandi" pageId="1" pageNumber="368" stateProvince="Assam">Dihandi</location>
(Brahmaputra drainage);
<geoCoordinate id="EE0E7AFD6645FFE8FBDFFCBFFB73FCF4" box="[1054,1198,869,894]" direction="north" orientation="latitude" pageId="1" pageNumber="368" precision="15" value="25.300833">25°1803&quot; N</geoCoordinate>
,
<geoCoordinate id="EE0E7AFD6645FFE8FB7AFCBFFA95FCF4" box="[1211,1352,869,894]" direction="east" orientation="longitude" pageId="1" pageNumber="368" precision="15" value="92.86806">92°5205&quot; E</geoCoordinate>
,
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<quantity id="4CC2B1DF6645FFE8FA95FCBCFA41FCF4" box="[1364,1436,870,894]" metricMagnitude="2" metricUnit="m" metricValue="4.01" pageId="1" pageNumber="368" unit="m" value="401.0">401 m</quantity>
above sea level
</elevation>
,
<location id="8EE54AE16645FFE8FE91FC53FE74FC2B" LSID="urn:lsid:plazi:treatment:0393AD2C6645FFECFF56FD0CFC3AF898:8EE54AE16645FFE8FE91FC53FE74FC2B" box="[336,425,905,930]" country="India" county="Dima Hasao District" latitude="25.300833" longLatPrecision="21" longitude="92.86806" municipality="Dilaima River" name="Sarbojit" pageId="1" pageNumber="368" stateProvince="Assam">Sarbojit</location>
<emphasis id="B94EC0286645FFE8FE6EFC50FE3CFC28" box="[431,481,905,929]" italics="true" pageId="1" pageNumber="368">et al</emphasis>
,
<date id="FF843AFA6645FFE8FE2FFC50FD4DFC2B" box="[494,656,905,930]" pageId="1" pageNumber="368" value="2015-04-19">
<collectingDate id="EFC0C3126645FFE8FE2FFC50FD4DFC2B" box="[494,656,905,930]" pageId="1" pageNumber="368" value="2015-04-19">19 April 2015</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8B851C3A6645FFE8FF06FC77FED7FBBB" blockId="1.[151,1437,833,2010]" pageId="1" pageNumber="368">
<materialsCitation id="3B5216676645FFE8FF06FC77FED7FBBB" ID-GBIF-Occurrence="1440783602" collectingDate="2016-03-16" collectionCode="MUMF" country="India" elevation="87" latitude="25.80611" location="Diyung River" longLatPrecision="21" longitude="92.95722" municipality="Dima Hasao District" pageId="1" pageNumber="368" specimenCode="MUMF 22001-22003, 3, MUMF 22005-22013, 9, MUMF 22002, 110.4" specimenCount="1" stateProvince="Assam" typeStatus="paratype">
<emphasis id="B94EC0286645FFE8FF06FC77FE95FC4C" bold="true" box="[199,328,941,965]" pageId="1" pageNumber="368">
<typeStatus id="5481A2986645FFE8FF06FC77FE9EFC4C" box="[199,323,941,965]" pageId="1" pageNumber="368" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCode id="DB9CB4416645FFE8FE93FC74FDB1FC4F" box="[338,620,942,966]" pageId="1" pageNumber="368">
<collectionCode id="ED2B84FF6645FFE8FE93FC74FE7AFC4C" box="[338,423,942,965]" country="India" httpUri="http://grbio.org/cool/x8gv-47he" name="Department of Life Sciences" pageId="1" pageNumber="368">MUMF</collectionCode>
2200122003, 3
</specimenCode>
specimens,
<quantity id="4CC2B1DF6645FFE8FD3CFC77FC62FC4C" box="[765,959,941,966]" metricMagnitude="-1" metricUnit="m" metricValue="1.0757999999999999" metricValueMax="1.2746" metricValueMin="0.877" pageId="1" pageNumber="368" unit="mm" value="107.58" valueMax="127.46" valueMin="87.7">87.7127.46 mm</quantity>
SL,
<specimenCode id="DB9CB4416645FFE8FC3DFC74FACBFC4F" box="[1020,1302,941,966]" pageId="1" pageNumber="368">
<collectionCode id="ED2B84FF6645FFE8FC3DFC74FB8FFC4C" box="[1020,1106,942,965]" country="India" httpUri="http://grbio.org/cool/x8gv-47he" name="Department of Life Sciences" pageId="1" pageNumber="368">MUMF</collectionCode>
2200522013, 9
</specimenCode>
specimens,
<quantity id="4CC2B1DF6645FFE8FF56FC0BFE84FC60" box="[151,345,977,1002]" metricMagnitude="-2" metricUnit="m" metricValue="6.600999999999999" metricValueMax="7.773999999999999" metricValueMin="5.428" pageId="1" pageNumber="368" unit="mm" value="66.00999999999999" valueMax="77.74" valueMin="54.28">54.2877.74 mm</quantity>
SL;
<collectingCountry id="F32D5CAA6645FFE8FE54FC0BFE0EFC63" box="[405,467,977,1002]" name="India" pageId="1" pageNumber="368">India</collectingCountry>
:
<collectingRegion id="49FED2D86645FFE8FE20FC08FDEFFC63" box="[481,562,978,1002]" country="India" name="Assam" pageId="1" pageNumber="368">Assam</collectingRegion>
:
<collectingMunicipality id="6BE186406645FFE8FDFEFC0BFCECFC63" box="[575,817,977,1002]" pageId="1" pageNumber="368">Dima Hasao District</collectingMunicipality>
:
<location id="8EE54AE16645FFE8FCFFFC0BFC07FC63" LSID="urn:lsid:plazi:treatment:0393AD2C6645FFECFF56FD0CFC3AF898:8EE54AE16645FFE8FCFFFC0BFC07FC63" box="[830,986,977,1002]" country="India" latitude="25.80611" longLatPrecision="21" longitude="92.95722" municipality="Dima Hasao District" name="Diyung River" pageId="1" pageNumber="368" stateProvince="Assam">Diyung River</location>
(Brahmaputra drainage);
<geoCoordinate id="EE0E7AFD6645FFE8FAC7FC0BFA45FC60" box="[1286,1432,977,1002]" direction="north" orientation="latitude" pageId="1" pageNumber="368" precision="15" value="25.80611">25°4822&quot; N</geoCoordinate>
,
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,
<elevation id="0017FB096645FFE8FEF6FC2CFDE5FB87" box="[311,568,1013,1038]" metricMagnitude="1" metricUnit="m" metricValue="8.7" pageId="1" pageNumber="368" unit="m" value="87.0">
<quantity id="4CC2B1DF6645FFE8FEF6FC2CFEA8FB84" box="[311,373,1014,1038]" metricMagnitude="1" metricUnit="m" metricValue="8.7" pageId="1" pageNumber="368" unit="m" value="87.0">87 m</quantity>
above sea level
</elevation>
, Roni
<emphasis id="B94EC0286645FFE8FD4BFC2CFD1DFB84" box="[650,704,1013,1037]" italics="true" pageId="1" pageNumber="368">et al</emphasis>
,
<date id="FF843AFA6645FFE8FD12FC2FFC56FB87" box="[723,907,1013,1038]" pageId="1" pageNumber="368" value="2016-03-16">
<collectingDate id="EFC0C3126645FFE8FD12FC2FFC56FB87" box="[723,907,1013,1038]" pageId="1" pageNumber="368" value="2016-03-16">16 March 2016</collectingDate>
</date>
.
<specimenCode id="DB9CB4416645FFE8FC5BFC2CFB4FFB87" box="[922,1170,1014,1038]" pageId="1" pageNumber="368">
<collectionCode id="ED2B84FF6645FFE8FC5BFC2CFC32FB84" box="[922,1007,1014,1037]" country="India" httpUri="http://grbio.org/cool/x8gv-47he" name="Department of Life Sciences" pageId="1" pageNumber="368">MUMF</collectionCode>
<quantity id="4CC2B1DF6645FFE8FC3DFC2CFB4FFB87" box="[1020,1170,1014,1038]" metricMagnitude="4" metricUnit="m" metricValue="2.20021104" pageId="1" pageNumber="368" unit="mm" value="2.20021104E7">22002, 110.4</quantity>
</specimenCode>
mm SL, dissected for osteology.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3204FB16645FFE8FF06FBE6FB63FADB" pageId="1" pageNumber="368" type="diagnosis">
<paragraph id="8B851C3A6645FFE8FF06FBE6FB63FADB" blockId="1.[151,1437,833,2010]" pageId="1" pageNumber="368">
<emphasis id="B94EC0286645FFE8FF06FBE6FE9CFBDC" bold="true" box="[199,321,1084,1109]" pageId="1" pageNumber="368">Diagnosis.</emphasis>
<taxonomicName id="4C3A67B96645FFE8FE8FFBE4FDFDFBDC" box="[334,544,1085,1109]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="368" phylum="Chordata" rank="species" species="clavirostris">
<emphasis id="B94EC0286645FFE8FE8FFBE4FDFDFBDC" box="[334,544,1085,1109]" italics="true" pageId="1" pageNumber="368">Garra clavirostris</emphasis>
</taxonomicName>
is distinguished from its congeners in the Brahmaputra and the Chindwin drainages in having the following combination of characters: a transverse lobe bearing 1725 small- to large-sized multicuspid tubercles; a prominent club-shaped unilobed proboscis prominently protruding forward and overhanging the depressed rostral surface, with a row of 59 medium- to large-sized uni- to multi-cuspid tubercles on its anterior margin; the antero-ventral marginal aspect of the proboscis with 56 medium-sized uni- to multicuspid tubercles, distinctly differentiated from the depressed rostral surface by a distinct transverse groove; 8½ or 9½ branched dorsal-fin rays; 5½ branched anal-fin rays; 3334 lateral-line scales; a black spot at the upper angle of gill opening; and 45 narrow black stripes laterally, more distinct towards the caudal peduncle.
</paragraph>
</subSubSection>
<subSubSection id="C3204FB16645FFEBFF06FA86FD0DF8DA" lastPageId="2" lastPageNumber="369" pageId="1" pageNumber="368" type="description">
<paragraph id="8B851C3A6645FFE8FF06FA86FE87F88B" blockId="1.[151,1437,833,2010]" pageId="1" pageNumber="368">
<emphasis id="B94EC0286645FFE8FF06FA86FE84FAFC" bold="true" box="[199,345,1372,1397]" pageId="1" pageNumber="368">Description.</emphasis>
General body shape as shown in
<figureCitation id="130100BF6645FFE8FD12FA87FCCBFAFF" box="[723,790,1373,1398]" captionStart="FIGURE 1" captionStartId="2.[151,250,1110,1132]" captionTargetBox="[227,1356,280,1073]" captionTargetId="figure@2.[217,1369,265,1089]" captionTargetPageId="2" captionText="FIGURE 1. Garra clavirostris, sp. nov., holotype, MUMF 22004, 117.5 mm SL, male; a. lateral, b. dorsal and c. ventral views. India: Assam: Dima Hasao District: Dilaima River at Boro Chenam Village." httpUri="https://zenodo.org/record/428274/files/figure.png" pageId="1" pageNumber="368">Fig. 1</figureCitation>
. Morphometric and meristic data for the
<typeStatus id="5481A2986645FFE8FB26FA87FA94FAFF" box="[1255,1353,1373,1398]" pageId="1" pageNumber="368" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D3CD7B36645FFE8FA41FA84FEDFFA13" pageId="1" pageNumber="368" type="generic" typeStatus="paratypes">12 paratypes</specimenCount>
are listed in
<tableCitation id="C6B829816645FFE8FE5AFA5BFE2CFA13" box="[411,497,1409,1434]" captionStart="TABLE 1" captionStartId="3.[151,239,152,175]" captionTargetBox="[161,1328,202,2007]" captionTargetPageId="3" captionText="TABLE 1. Morphometric and meristic data of holotype and twelve paratypes of Garra clavirostris." httpUri="http://table.plazi.org/id/DF454CB26647FFEAFF56FF42FB61FF26" pageId="1" pageNumber="368" tableUuid="DF454CB26647FFEAFF56FF42FB61FF26">Table 1</tableCitation>
. Body elongate, compressed laterally, more compressed towards caudal penducle. Dorsal head profile rising gently over snout, slightly more convex than dorsal body profile to dorsal-fin origin. Ventral profile more or less straight to anal-fin origin. Head large, depressed, with moderately convex interorbital region, head depth less than head width and head length. Snout rounded, its transverse lobe covered with 1725 medium to large multicuspid tubercles, each demarcated posteriorly by a shallow transverse groove (
<figureCitation id="130100BF6645FFE8FAD2F9CBFA84F9A3" box="[1299,1369,1553,1578]" captionStart="FIGURE 2" captionStartId="4.[151,250,1312,1334]" captionTargetBox="[163,1425,216,1261]" captionTargetId="figure@4.[151,1436,193,1291]" captionTargetPageId="4" captionText="FIGURE 2. Snout of Garra clavirostris, a. morphology of snout showing transverse lobe, proboscis, depressed rostral surface and lateral surfaces of holotype; b. transverse lobe showing tubercles (MUMF 22001, paratype, 87.7 mm SL); c. one tubercle in b enlarged." httpUri="https://zenodo.org/record/428279/files/figure.png" pageId="1" pageNumber="368">Fig. 2</figureCitation>
a,b,c). Proboscis prominent, protruding forward up to transverse groove, with 59 medium to large uni- to multi-cuspid tubercles on anterior margin, 56 uni- to multi-cuspid tubercles on anteroventral marginal aspect, differentiated from depressed rostral surface by a distinct transverse groove; anterior region of proboscis bulging, club-shaped, overhanging depressed rostral surface (
<figureCitation id="130100BF6645FFE8FD9BF97BFD7CF933" box="[602,673,1697,1722]" captionStart="FIGURE 3" captionStartId="5.[151,250,1250,1272]" captionTargetBox="[355,1247,557,1229]" captionTargetId="figure@5.[340,1247,553,1229]" captionTargetPageId="5" captionText="FIGURE 3. Lateral view of snout of Garra clavirostris, holotype, showing shape of proboscis and the position of eye." httpUri="https://zenodo.org/record/428281/files/figure.png" pageId="1" pageNumber="368">Fig. 3</figureCitation>
). Depressed rostral surface slightly convex, sub-lachrymal groove shallow, connected to rostral-cap groove. Eyes dorsolaterally located, closer to posterior margin of opercle than to snout tip (
<figureCitation id="130100BF6645FFE8FEC6F933FE96F88B" box="[263,331,1769,1794]" captionStart="FIGURE 3" captionStartId="5.[151,250,1250,1272]" captionTargetBox="[355,1247,557,1229]" captionTargetId="figure@5.[340,1247,553,1229]" captionTargetPageId="5" captionText="FIGURE 3. Lateral view of snout of Garra clavirostris, holotype, showing shape of proboscis and the position of eye." httpUri="https://zenodo.org/record/428281/files/figure.png" pageId="1" pageNumber="368">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8B851C3A6645FFEBFF06F8D7FD6BFF5C" blockId="1.[151,1437,833,2010]" lastBlockId="2.[151,1436,151,213]" lastPageId="2" lastPageNumber="369" pageId="1" pageNumber="368">
Barbels in two pairs; rostral barbels anterolaterally located, shorter than eye diameter; maxillary barbels at the corner of mouth, much shorter than rostral barbels. Upper jaw entirely covered by well-developed rostral cap which has a highly fimbriate posterior margin, about one-sixth of its length at distal margin smooth (
<figureCitation id="130100BF6645FFE8FAF8F88FFA52F8E7" box="[1337,1423,1877,1902]" captionStart="FIGURE 4" captionStartId="6.[151,250,1740,1762]" captionTargetBox="[158,1421,250,1711]" captionTargetId="figure@6.[151,1436,193,1733]" captionTargetPageId="6" captionText="FIGURE 4. a. Oromandibular structure of Garra clavirostris, holotype; magnified images of b. rostral cap and c. lateroposterior flap (MUMF 22001, paratype, 87.7 mm SL)." httpUri="https://zenodo.org/record/428283/files/figure.png" pageId="1" pageNumber="368">Fig. 4a</figureCitation>
). Papillae on rostral cap with finger-like unculi projecting in different directions (
<figureCitation id="130100BF6645FFE8FBD2F8A3FB8AF81B" box="[1043,1111,1913,1938]" captionStart="FIGURE 4" captionStartId="6.[151,250,1740,1762]" captionTargetBox="[158,1421,250,1711]" captionTargetId="figure@6.[151,1436,193,1733]" captionTargetPageId="6" captionText="FIGURE 4. a. Oromandibular structure of Garra clavirostris, holotype; magnified images of b. rostral cap and c. lateroposterior flap (MUMF 22001, paratype, 87.7 mm SL)." httpUri="https://zenodo.org/record/428283/files/figure.png" pageId="1" pageNumber="368">Fig. 4</figureCitation>
b). Mental disc oval; papillae on anteromedian fold irregularly arranged; unculi on papillae blunt, projecting to one side (
<figureCitation id="130100BF6645FFE8FB0FF847FACAF83F" box="[1230,1303,1949,1974]" captionStart="FIGURE 4" captionStartId="6.[151,250,1740,1762]" captionTargetBox="[158,1421,250,1711]" captionTargetId="figure@6.[151,1436,193,1733]" captionTargetPageId="6" captionText="FIGURE 4. a. Oromandibular structure of Garra clavirostris, holotype; magnified images of b. rostral cap and c. lateroposterior flap (MUMF 22001, paratype, 87.7 mm SL)." httpUri="https://zenodo.org/record/428283/files/figure.png" pageId="1" pageNumber="368">Fig. 4</figureCitation>
c). Anterior marginal region of central callous pad with small, elongated, sparsely-arranged papillae; papillae of whole length of lateroposterior flap large, coarsely arranged; posteriormost margin of lateroposterior flap extending slightly beyond vertical through posterior margin of eye.
</paragraph>
<caption id="DF454CB26646FFEBFF56FB8CFC3FFB02" httpUri="https://zenodo.org/record/428274/files/figure.png" pageId="2" pageNumber="369" targetBox="[227,1356,280,1073]" targetPageId="2">
<paragraph id="8B851C3A6646FFEBFF56FB8CFC3FFB02" blockId="2.[151,1435,1110,1163]" pageId="2" pageNumber="369">
<emphasis id="B94EC0286646FFEBFF56FB8CFECBFBE5" bold="true" box="[151,278,1110,1132]" pageId="2" pageNumber="369">FIGURE 1.</emphasis>
<taxonomicName id="4C3A67B96646FFEBFEDEFB8DFE07FBE5" box="[287,474,1110,1132]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="369" phylum="Chordata" rank="species" species="clavirostris" status="sp. nov.">
<emphasis id="B94EC0286646FFEBFEDEFB8DFE07FBE5" box="[287,474,1110,1132]" italics="true" pageId="2" pageNumber="369">Garra clavirostris</emphasis>
</taxonomicName>
,
<emphasis id="B94EC0286646FFEBFE2BFB8CFDE0FBE5" bold="true" box="[490,573,1110,1132]" pageId="2" pageNumber="369">
<taxonomicNameLabel id="A27D7D536646FFEBFE2BFB8CFDE0FBE5" box="[490,573,1110,1132]" pageId="2" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, MUMF 22004, 117.5 mm SL, male;
<emphasis id="B94EC0286646FFEBFBFDFB8CFB92FBE5" bold="true" box="[1084,1103,1110,1132]" pageId="2" pageNumber="369">a.</emphasis>
lateral,
<emphasis id="B94EC0286646FFEBFB69FB8CFB61FBE5" bold="true" box="[1192,1212,1110,1132]" pageId="2" pageNumber="369">b.</emphasis>
dorsal and
<emphasis id="B94EC0286646FFEBFAFAFB8CFA91FBE5" bold="true" box="[1339,1356,1110,1132]" pageId="2" pageNumber="369">c.</emphasis>
ventral views. India: Assam: Dima Hasao District: Dilaima River at Boro Chenam Village.
</paragraph>
</caption>
<paragraph id="8B851C3A6646FFEBFF06FB68FDB4F986" blockId="2.[151,1436,1202,2019]" pageId="2" pageNumber="369">Dorsal fin with three simple and 8½9½ branched rays, last simple ray slightly shorter than head length; distal margin markedly concave; origin closer to snout tip than to caudal-fin base; first branched ray longest, tip of last branched ray not extending beyond vertical through anal-fin origin. Pectoral fin with one simple and 1415 branched rays; its length less than head length; margin subacuminate; fifth branched ray longest, not extending to base of pelvic fin. Pelvic fin with one simple and 78 branched rays, not surpassing vent when adpressed; second branched ray longest, not extending to base of anal-fin origin; origin closer to anal-fin origin than to pectoral-fin origin, inserted below third or fourth branched dorsal-fin ray; distal margin slightly convex. Anal fin short, with three simple and 5½ branched rays; first branched ray longest. Anus situated posteriorly (distance between anus and anal-fin origin 1825% of that between pelvic-fin origin and anal-fin origin). Caudal fin forked; tips of lobes pointed; lobes equal; 10+9 principal rays.</paragraph>
<paragraph id="8B851C3A6646FFEBFF06F9C0FB08F9F2" blockId="2.[151,1436,1202,2019]" pageId="2" pageNumber="369">Lateral line complete, with 3334 scales. Transverse scale rows above lateral line 4½5½; between lateral line and pelvic fin origin 3½. Predorsal scales 910; scales regularly arranged; chest and belly scaled, scales partially embedded in skin. Preanal scales 34. Scales on dorsal-fin base 67. Scales on anal fin base 45.</paragraph>
<paragraph id="8B851C3A6646FFEBFF06F95CFAC7F916" blockId="2.[151,1436,1202,2019]" box="[199,1306,1670,1695]" pageId="2" pageNumber="369">Total vertebrae 32 (1), consisting of 21 abdominal + 11 caudal vertebrae; predorsal vertebrae 9 (1).</paragraph>
<paragraph id="8B851C3A6646FFEBFF06F970FD0DF8DA" blockId="2.[151,1436,1202,2019]" pageId="2" pageNumber="369">
<emphasis id="B94EC0286646FFEBFF06F970FEC6F94B" box="[199,283,1706,1730]" italics="true" pageId="2" pageNumber="369">Colour.</emphasis>
In formalin: head, dorsum and side dark brown to dark grey. Mouth, chest and abdomen yellowish white. Dorsal fin more distinct dark-grey hue than anal, pelvic and pectoral fins. In some specimens, lower lobe of caudal fin darker than upper lobe. Four or five narrow black stripes laterally, more distinct towards caudal peduncle; yellowish white ventrally at origin of pectoral and pelvic fins. A black spot on first scale of lateral line, covering about one-third height of scale anteriorly.
</paragraph>
</subSubSection>
<subSubSection id="C3204FB16646FFEBFF06F887FB46F86A" pageId="2" pageNumber="369" type="distribution">
<paragraph id="8B851C3A6646FFEBFF06F887FB46F86A" blockId="2.[151,1436,1202,2019]" pageId="2" pageNumber="369">
<emphasis id="B94EC0286646FFEBFF06F887FEBDF8FF" bold="true" box="[199,352,1885,1910]" pageId="2" pageNumber="369">Distribution.</emphasis>
<taxonomicName id="4C3A67B96646FFEBFEA8F885FDEBF8FF" box="[361,566,1886,1910]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="369" phylum="Chordata" rank="species" species="clavirostris">
<emphasis id="B94EC0286646FFEBFEA8F885FDEBF8FF" box="[361,566,1886,1910]" italics="true" pageId="2" pageNumber="369">Garra clavirostris</emphasis>
</taxonomicName>
is presently known only from the
<typeStatus id="5481A2986646FFEBFC00F885FC2CF8FE" box="[961,1009,1887,1911]" pageId="2" pageNumber="369">type</typeStatus>
locality Dilaima River (Brahmaputra drainage), Boro Chenam Village, below the confluence of the Dilaima and Dihandi, and also from the Diyung River (Brahmaputra drainage) of Dima Hasao district,
<collectingRegion id="49FED2D86646FFEBFCC9F87DFC8AF836" box="[776,855,1959,1983]" country="India" name="Assam" pageId="2" pageNumber="369">Assam</collectingRegion>
,
<collectingCountry id="F32D5CAA6646FFEBFCA5F87CFC43F836" box="[868,926,1958,1983]" name="India" pageId="2" pageNumber="369">India</collectingCountry>
(
<figureCitation id="130100BF6646FFEBFC6FF87CFC28F836" box="[942,1013,1958,1983]" captionStart="FIGURE 5" captionStartId="7.[151,250,2003,2025]" captionTargetBox="[163,1427,823,1974]" captionTargetId="figure@7.[151,1436,805,1981]" captionTargetPageId="7" captionText="FIGURE 5. Map showing the type locality of Garra clavirostris, sp. nov.; Holotype (•) and Paratype (̊)." httpUri="https://zenodo.org/record/428290/files/figure.png" pageId="2" pageNumber="369">Fig. 5</figureCitation>
). It inhabits medium- to fast-flowing clearwater hill streams with a gravelly substrate covered in lush green algal growth (
<figureCitation id="130100BF6646FFEBFB89F810FB50F86A" box="[1096,1165,1994,2019]" captionStart="FIGURE 6" captionStartId="8.[151,250,854,876]" captionTargetBox="[217,1369,193,833]" captionTargetId="figure@8.[217,1369,193,833]" captionTargetPageId="8" captionText="FIGURE 6. Diyung River, type locality and habitat of Garra clavirostris." httpUri="https://zenodo.org/record/428291/files/figure.png" pageId="2" pageNumber="369">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF454CB26647FFEAFF56FF42FB61FF26" ID-Table-UUID="DF454CB26647FFEAFF56FF42FB61FF26" box="[151,1212,151,175]" httpUri="http://table.plazi.org/id/DF454CB26647FFEAFF56FF42FB61FF26" pageId="3" pageNumber="370" targetBox="[161,1328,202,2007]" targetIsTable="true" targetPageId="3">
<paragraph id="8B851C3A6647FFEAFF56FF42FB61FF26" blockId="3.[151,1212,151,175]" box="[151,1212,151,175]" pageId="3" pageNumber="370">
<emphasis id="B94EC0286647FFEAFF56FF42FED7FF26" bold="true" box="[151,266,152,175]" pageId="3" pageNumber="370">TABLE 1.</emphasis>
Morphometric and meristic data of holotype and twelve paratypes of
<taxonomicName id="4C3A67B96647FFEAFC32FF42FB69FF26" box="[1011,1204,151,175]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="370" phylum="Chordata" rank="species" species="clavirostris">
<emphasis id="B94EC0286647FFEAFC32FF42FB69FF26" box="[1011,1204,151,175]" italics="true" pageId="3" pageNumber="370">Garra clavirostris</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8B851C3A6647FFEAFD0DFF10FC45F85E" pageId="3" pageNumber="370">
<table id="F93AEE9A66470016FF60FF10FAEDF85E" box="[161,1328,202,2007]" gridcols="5" gridrows="43" pageId="3" pageNumber="370">
<tr id="350A1E7866470016FF60FF10FAEDFE81" box="[161,1328,202,264]" gridrow="0" pageId="3" pageNumber="370" rowspan-0="1">
<th id="76DB770466470016FD0DFF10FCF7FE81" box="[716,810,202,264]" gridcol="1" gridrow="0" pageId="3" pageNumber="370">Holotype MUMF</th>
<th id="76DB770466470016FCB3FF10FC2CFE81" box="[882,1009,202,264]" gridcol="2" gridrow="0" pageId="3" pageNumber="370">Paratypes Range</th>
<th id="76DB770466470016FBABFF10FB79FE81" box="[1130,1188,202,264]" gridcol="3" gridrow="0" pageId="3" pageNumber="370">Mean</th>
<th id="76DB770466470016FAC8FF10FAEDFE81" box="[1289,1328,202,264]" gridcol="4" gridrow="0" pageId="3" pageNumber="370">S.D</th>
</tr>
<tr id="350A1E7866470016FF60FEC1FAEDFEB8" box="[161,1328,283,305]" gridrow="1" pageId="3" pageNumber="370" rowspan-3="1" rowspan-4="1">
<th id="76DB770466470016FF60FEC1FE01FEB8" box="[161,476,283,305]" gridcol="0" gridrow="1" pageId="3" pageNumber="370">Standard length (mm)</th>
<td id="76DB770466470016FD0DFEC1FCF7FEB8" box="[716,810,283,305]" gridcol="1" gridrow="1" pageId="3" pageNumber="370">117.5</td>
<td id="76DB770466470016FCB3FEC1FC2CFEB8" box="[882,1009,283,305]" gridcol="2" gridrow="1" pageId="3" pageNumber="370">46.4127.46</td>
</tr>
<tr id="350A1E7866470016FF60FEB6FAEDFE0B" box="[161,1328,364,386]" gridrow="2" pageId="3" pageNumber="370" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="76DB770466470016FF60FEB6FE01FE0B" box="[161,476,364,386]" gridcol="0" gridrow="2" pageId="3" pageNumber="370">In % of standard length</th>
</tr>
<tr id="350A1E7866470016FF60FE4EFAEDFE23" box="[161,1328,404,426]" gridrow="3" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60FE4EFE01FE23" box="[161,476,404,426]" gridcol="0" gridrow="3" pageId="3" pageNumber="370">Body depth</th>
<td id="76DB770466470016FD0DFE4EFCF7FE23" box="[716,810,404,426]" gridcol="1" gridrow="3" pageId="3" pageNumber="370">20.8</td>
<td id="76DB770466470016FCB3FE4EFC2CFE23" box="[882,1009,404,426]" gridcol="2" gridrow="3" pageId="3" pageNumber="370">19.424.0</td>
<td id="76DB770466470016FBABFE4EFB79FE23" box="[1130,1188,404,426]" gridcol="3" gridrow="3" pageId="3" pageNumber="370">21.3</td>
<td id="76DB770466470016FAC8FE4EFAEDFE23" box="[1289,1328,404,426]" gridcol="4" gridrow="3" pageId="3" pageNumber="370">1.4</td>
</tr>
<tr id="350A1E7866470016FF60FE67FAEDFE5A" box="[161,1328,445,467]" gridrow="4" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60FE67FE01FE5A" box="[161,476,445,467]" gridcol="0" gridrow="4" pageId="3" pageNumber="370">Head length</th>
<td id="76DB770466470016FD0DFE67FCF7FE5A" box="[716,810,445,467]" gridcol="1" gridrow="4" pageId="3" pageNumber="370">24.8</td>
<td id="76DB770466470016FCB3FE67FC2CFE5A" box="[882,1009,445,467]" gridcol="2" gridrow="4" pageId="3" pageNumber="370">23.127.1</td>
<td id="76DB770466470016FBABFE67FB79FE5A" box="[1130,1188,445,467]" gridcol="3" gridrow="4" pageId="3" pageNumber="370">25.9</td>
<td id="76DB770466470016FAC8FE67FAEDFE5A" box="[1289,1328,445,467]" gridcol="4" gridrow="4" pageId="3" pageNumber="370">1.2</td>
</tr>
<tr id="350A1E7866470016FF60FE3FFAEDFE72" box="[161,1328,485,507]" gridrow="5" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60FE3FFE01FE72" box="[161,476,485,507]" gridcol="0" gridrow="5" pageId="3" pageNumber="370">Head depth at nape</th>
<td id="76DB770466470016FD0DFE3FFCF7FE72" box="[716,810,485,507]" gridcol="1" gridrow="5" pageId="3" pageNumber="370">8.4</td>
<td id="76DB770466470016FCB3FE3FFC2CFE72" box="[882,1009,485,507]" gridcol="2" gridrow="5" pageId="3" pageNumber="370">8.014.1</td>
<td id="76DB770466470016FBABFE3FFB79FE72" box="[1130,1188,485,507]" gridcol="3" gridrow="5" pageId="3" pageNumber="370">11.4</td>
<td id="76DB770466470016FAC8FE3FFAEDFE72" box="[1289,1328,485,507]" gridcol="4" gridrow="5" pageId="3" pageNumber="370">2.1</td>
</tr>
<tr id="350A1E7866470016FF60FDD4FAEDFDAD" box="[161,1328,526,548]" gridrow="6" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60FDD4FE01FDAD" box="[161,476,526,548]" gridcol="0" gridrow="6" pageId="3" pageNumber="370">Head depth at eye</th>
<td id="76DB770466470016FD0DFDD4FCF7FDAD" box="[716,810,526,548]" gridcol="1" gridrow="6" pageId="3" pageNumber="370">9.7</td>
<td id="76DB770466470016FCB3FDD4FC2CFDAD" box="[882,1009,526,548]" gridcol="2" gridrow="6" pageId="3" pageNumber="370">9.718.2</td>
<td id="76DB770466470016FBABFDD4FB79FDAD" box="[1130,1188,526,548]" gridcol="3" gridrow="6" pageId="3" pageNumber="370">14.2</td>
<td id="76DB770466470016FAC8FDD4FAEDFDAD" box="[1289,1328,526,548]" gridcol="4" gridrow="6" pageId="3" pageNumber="370">2.5</td>
</tr>
<tr id="350A1E7866470016FF60FDECFAEDFDC5" box="[161,1328,566,588]" gridrow="7" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60FDECFE01FDC5" box="[161,476,566,588]" gridcol="0" gridrow="7" pageId="3" pageNumber="370">Body width at anal-fin origin</th>
<td id="76DB770466470016FD0DFDECFCF7FDC5" box="[716,810,566,588]" gridcol="1" gridrow="7" pageId="3" pageNumber="370">14.7</td>
<td id="76DB770466470016FCB3FDECFC2CFDC5" box="[882,1009,566,588]" gridcol="2" gridrow="7" pageId="3" pageNumber="370">7.916.0</td>
<td id="76DB770466470016FBABFDECFB79FDC5" box="[1130,1188,566,588]" gridcol="3" gridrow="7" pageId="3" pageNumber="370">11.7</td>
<td id="76DB770466470016FAC8FDECFAEDFDC5" box="[1289,1328,566,588]" gridcol="4" gridrow="7" pageId="3" pageNumber="370">2.6</td>
</tr>
<tr id="350A1E7866470016FF60FD85FAEDFDFC" box="[161,1328,607,629]" gridrow="8" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60FD85FE01FDFC" box="[161,476,607,629]" gridcol="0" gridrow="8" pageId="3" pageNumber="370">Body width at dorsal-fin origin</th>
<td id="76DB770466470016FD0DFD85FCF7FDFC" box="[716,810,607,629]" gridcol="1" gridrow="8" pageId="3" pageNumber="370">19.1</td>
<td id="76DB770466470016FCB3FD85FC2CFDFC" box="[882,1009,607,629]" gridcol="2" gridrow="8" pageId="3" pageNumber="370">14.321.1</td>
<td id="76DB770466470016FBABFD85FB79FDFC" box="[1130,1188,607,629]" gridcol="3" gridrow="8" pageId="3" pageNumber="370">17.7</td>
<td id="76DB770466470016FAC8FD85FAEDFDFC" box="[1289,1328,607,629]" gridcol="4" gridrow="8" pageId="3" pageNumber="370">1.8</td>
</tr>
<tr id="350A1E7866470016FF60FD5DFAEDFD14" box="[161,1328,647,669]" gridrow="9" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60FD5DFE01FD14" box="[161,476,647,669]" gridcol="0" gridrow="9" pageId="3" pageNumber="370">Caudal peduncle length</th>
<td id="76DB770466470016FD0DFD5DFCF7FD14" box="[716,810,647,669]" gridcol="1" gridrow="9" pageId="3" pageNumber="370">11.5</td>
<td id="76DB770466470016FCB3FD5DFC2CFD14" box="[882,1009,647,669]" gridcol="2" gridrow="9" pageId="3" pageNumber="370">10.114.6</td>
<td id="76DB770466470016FBABFD5DFB79FD14" box="[1130,1188,647,669]" gridcol="3" gridrow="9" pageId="3" pageNumber="370">12.5</td>
<td id="76DB770466470016FAC8FD5DFAEDFD14" box="[1289,1328,647,669]" gridcol="4" gridrow="9" pageId="3" pageNumber="370">1.3</td>
</tr>
<tr id="350A1E7866470016FF60FD6AFAEDFD4F" box="[161,1328,688,710]" gridrow="10" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60FD6AFE01FD4F" box="[161,476,688,710]" gridcol="0" gridrow="10" pageId="3" pageNumber="370">Caudal peduncle depth</th>
<td id="76DB770466470016FD0DFD6AFCF7FD4F" box="[716,810,688,710]" gridcol="1" gridrow="10" pageId="3" pageNumber="370">11.4</td>
<td id="76DB770466470016FCB3FD6AFC2CFD4F" box="[882,1009,688,710]" gridcol="2" gridrow="10" pageId="3" pageNumber="370">2.211.5</td>
<td id="76DB770466470016FBABFD6AFB79FD4F" box="[1130,1188,688,710]" gridcol="3" gridrow="10" pageId="3" pageNumber="370">5.9</td>
<td id="76DB770466470016FAC8FD6AFAEDFD4F" box="[1289,1328,688,710]" gridcol="4" gridrow="10" pageId="3" pageNumber="370">3.8</td>
</tr>
<tr id="350A1E7866470016FF60FD02FAEDFD67" box="[161,1328,728,750]" gridrow="11" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60FD02FE01FD67" box="[161,476,728,750]" gridcol="0" gridrow="11" pageId="3" pageNumber="370">Dorsal-fin base length</th>
<td id="76DB770466470016FD0DFD02FCF7FD67" box="[716,810,728,750]" gridcol="1" gridrow="11" pageId="3" pageNumber="370">12.0</td>
<td id="76DB770466470016FCB3FD02FC2CFD67" box="[882,1009,728,750]" gridcol="2" gridrow="11" pageId="3" pageNumber="370">12.019.0</td>
<td id="76DB770466470016FBABFD02FB79FD67" box="[1130,1188,728,750]" gridcol="3" gridrow="11" pageId="3" pageNumber="370">16.8</td>
<td id="76DB770466470016FAC8FD02FAEDFD67" box="[1289,1328,728,750]" gridcol="4" gridrow="11" pageId="3" pageNumber="370">2.0</td>
</tr>
<tr id="350A1E7866470016FF60FCDBFAEDFC9E" box="[161,1328,769,791]" gridrow="12" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60FCDBFE01FC9E" box="[161,476,769,791]" gridcol="0" gridrow="12" pageId="3" pageNumber="370">Dorsal-fin length</th>
<td id="76DB770466470016FD0DFCDBFCF7FC9E" box="[716,810,769,791]" gridcol="1" gridrow="12" pageId="3" pageNumber="370">22.7</td>
<td id="76DB770466470016FCB3FCDBFC2CFC9E" box="[882,1009,769,791]" gridcol="2" gridrow="12" pageId="3" pageNumber="370">20.927.0</td>
<td id="76DB770466470016FBABFCDBFB79FC9E" box="[1130,1188,769,791]" gridcol="3" gridrow="12" pageId="3" pageNumber="370">24.2</td>
<td id="76DB770466470016FAC8FCDBFAEDFC9E" box="[1289,1328,769,791]" gridcol="4" gridrow="12" pageId="3" pageNumber="370">1.9</td>
</tr>
<tr id="350A1E7866470016FF60FCF0FAEDFCC9" box="[161,1328,810,832]" gridrow="13" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60FCF0FE01FCC9" box="[161,476,810,832]" gridcol="0" gridrow="13" pageId="3" pageNumber="370">Pectoral-fin length</th>
<td id="76DB770466470016FD0DFCF0FCF7FCC9" box="[716,810,810,832]" gridcol="1" gridrow="13" pageId="3" pageNumber="370">21.1</td>
<td id="76DB770466470016FCB3FCF0FC2CFCC9" box="[882,1009,810,832]" gridcol="2" gridrow="13" pageId="3" pageNumber="370">19.624.8</td>
<td id="76DB770466470016FBABFCF0FB79FCC9" box="[1130,1188,810,832]" gridcol="3" gridrow="13" pageId="3" pageNumber="370">21.9</td>
<td id="76DB770466470016FAC8FCF0FAEDFCC9" box="[1289,1328,810,832]" gridcol="4" gridrow="13" pageId="3" pageNumber="370">1.4</td>
</tr>
<tr id="350A1E7866470016FF60FC88FAEDFCE1" box="[161,1328,850,872]" gridrow="14" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60FC88FE01FCE1" box="[161,476,850,872]" gridcol="0" gridrow="14" pageId="3" pageNumber="370">Pelvic-fin length</th>
<td id="76DB770466470016FD0DFC88FCF7FCE1" box="[716,810,850,872]" gridcol="1" gridrow="14" pageId="3" pageNumber="370">17.4</td>
<td id="76DB770466470016FCB3FC88FC2CFCE1" box="[882,1009,850,872]" gridcol="2" gridrow="14" pageId="3" pageNumber="370">17.422.7</td>
<td id="76DB770466470016FBABFC88FB79FCE1" box="[1130,1188,850,872]" gridcol="3" gridrow="14" pageId="3" pageNumber="370">19.8</td>
<td id="76DB770466470016FAC8FC88FAEDFCE1" box="[1289,1328,850,872]" gridcol="4" gridrow="14" pageId="3" pageNumber="370">1.3</td>
</tr>
<tr id="350A1E7866470016FF60FCA1FAEDFC18" box="[161,1328,891,913]" gridrow="15" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60FCA1FE01FC18" box="[161,476,891,913]" gridcol="0" gridrow="15" pageId="3" pageNumber="370">Anal-fin base length</th>
<td id="76DB770466470016FD0DFCA1FCF7FC18" box="[716,810,891,913]" gridcol="1" gridrow="15" pageId="3" pageNumber="370">8.0</td>
<td id="76DB770466470016FCB3FCA1FC2CFC18" box="[882,1009,891,913]" gridcol="2" gridrow="15" pageId="3" pageNumber="370">6.18.5</td>
<td id="76DB770466470016FBABFCA1FB79FC18" box="[1130,1188,891,913]" gridcol="3" gridrow="15" pageId="3" pageNumber="370">7.7</td>
<td id="76DB770466470016FAC8FCA1FAEDFC18" box="[1289,1328,891,913]" gridcol="4" gridrow="15" pageId="3" pageNumber="370">0.8</td>
</tr>
<tr id="350A1E7866470016FF60FC79FAEDFC30" box="[161,1328,931,953]" gridrow="16" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60FC79FE01FC30" box="[161,476,931,953]" gridcol="0" gridrow="16" pageId="3" pageNumber="370">Anal-fin length</th>
<td id="76DB770466470016FD0DFC79FCF7FC30" box="[716,810,931,953]" gridcol="1" gridrow="16" pageId="3" pageNumber="370">17.0</td>
<td id="76DB770466470016FCB3FC79FC2CFC30" box="[882,1009,931,953]" gridcol="2" gridrow="16" pageId="3" pageNumber="370">11.019.4</td>
<td id="76DB770466470016FBABFC79FB79FC30" box="[1130,1188,931,953]" gridcol="3" gridrow="16" pageId="3" pageNumber="370">17.6</td>
<td id="76DB770466470016FAC8FC79FAEDFC30" box="[1289,1328,931,953]" gridcol="4" gridrow="16" pageId="3" pageNumber="370">2.2</td>
</tr>
<tr id="350A1E7866470016FF60FC16FAEDFC6B" box="[161,1328,972,994]" gridrow="17" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60FC16FE01FC6B" box="[161,476,972,994]" gridcol="0" gridrow="17" pageId="3" pageNumber="370">Predorsal length</th>
<td id="76DB770466470016FD0DFC16FCF7FC6B" box="[716,810,972,994]" gridcol="1" gridrow="17" pageId="3" pageNumber="370">44.4</td>
<td id="76DB770466470016FCB3FC16FC2CFC6B" box="[882,1009,972,994]" gridcol="2" gridrow="17" pageId="3" pageNumber="370">44.450.4</td>
<td id="76DB770466470016FBABFC16FB79FC6B" box="[1130,1188,972,994]" gridcol="3" gridrow="17" pageId="3" pageNumber="370">47.8</td>
<td id="76DB770466470016FAC8FC16FAEDFC6B" box="[1289,1328,972,994]" gridcol="4" gridrow="17" pageId="3" pageNumber="370">1.7</td>
</tr>
<tr id="350A1E7866470016FF60FC2EFAEDFB83" box="[161,1328,1012,1034]" gridrow="18" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60FC2EFE01FB83" box="[161,476,1012,1034]" gridcol="0" gridrow="18" pageId="3" pageNumber="370">Prepectoral length</th>
<td id="76DB770466470016FD0DFC2EFCF7FB83" box="[716,810,1012,1034]" gridcol="1" gridrow="18" pageId="3" pageNumber="370">23.2</td>
<td id="76DB770466470016FCB3FC2EFC2CFB83" box="[882,1009,1012,1034]" gridcol="2" gridrow="18" pageId="3" pageNumber="370">21.925.3</td>
<td id="76DB770466470016FBABFC2EFB79FB83" box="[1130,1188,1012,1034]" gridcol="3" gridrow="18" pageId="3" pageNumber="370">23.9</td>
<td id="76DB770466470016FAC8FC2EFAEDFB83" box="[1289,1328,1012,1034]" gridcol="4" gridrow="18" pageId="3" pageNumber="370">1.1</td>
</tr>
<tr id="350A1E7866470016FF60FBC7FAEDFBBA" box="[161,1328,1053,1075]" gridrow="19" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60FBC7FE01FBBA" box="[161,476,1053,1075]" gridcol="0" gridrow="19" pageId="3" pageNumber="370">Prepelvic length</th>
<td id="76DB770466470016FD0DFBC7FCF7FBBA" box="[716,810,1053,1075]" gridcol="1" gridrow="19" pageId="3" pageNumber="370">52.0</td>
<td id="76DB770466470016FCB3FBC7FC2CFBBA" box="[882,1009,1053,1075]" gridcol="2" gridrow="19" pageId="3" pageNumber="370">50.555.8</td>
<td id="76DB770466470016FBABFBC7FB79FBBA" box="[1130,1188,1053,1075]" gridcol="3" gridrow="19" pageId="3" pageNumber="370">52.1</td>
<td id="76DB770466470016FAC8FBC7FAEDFBBA" box="[1289,1328,1053,1075]" gridcol="4" gridrow="19" pageId="3" pageNumber="370">1.5</td>
</tr>
<tr id="350A1E7866470016FF60FB9FFAEDFBD2" box="[161,1328,1093,1115]" gridrow="20" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60FB9FFE01FBD2" box="[161,476,1093,1115]" gridcol="0" gridrow="20" pageId="3" pageNumber="370">Preanal length</th>
<td id="76DB770466470016FD0DFB9FFCF7FBD2" box="[716,810,1093,1115]" gridcol="1" gridrow="20" pageId="3" pageNumber="370">79.0</td>
<td id="76DB770466470016FCB3FB9FFC2CFBD2" box="[882,1009,1093,1115]" gridcol="2" gridrow="20" pageId="3" pageNumber="370">76.681.5</td>
<td id="76DB770466470016FBABFB9FFB79FBD2" box="[1130,1188,1093,1115]" gridcol="3" gridrow="20" pageId="3" pageNumber="370">78.4</td>
<td id="76DB770466470016FAC8FB9FFAEDFBD2" box="[1289,1328,1093,1115]" gridcol="4" gridrow="20" pageId="3" pageNumber="370">1.7</td>
</tr>
<tr id="350A1E7866470016FF60FBB4FAEDFB0D" box="[161,1328,1134,1156]" gridrow="21" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60FBB4FE01FB0D" box="[161,476,1134,1156]" gridcol="0" gridrow="21" pageId="3" pageNumber="370">Pelvic anal distance</th>
<td id="76DB770466470016FD0DFBB4FCF7FB0D" box="[716,810,1134,1156]" gridcol="1" gridrow="21" pageId="3" pageNumber="370">25.9</td>
<td id="76DB770466470016FCB3FBB4FC2CFB0D" box="[882,1009,1134,1156]" gridcol="2" gridrow="21" pageId="3" pageNumber="370">24.930.9</td>
<td id="76DB770466470016FBABFBB4FB79FB0D" box="[1130,1188,1134,1156]" gridcol="3" gridrow="21" pageId="3" pageNumber="370">27.1</td>
<td id="76DB770466470016FAC8FBB4FAEDFB0D" box="[1289,1328,1134,1156]" gridcol="4" gridrow="21" pageId="3" pageNumber="370">1.7</td>
</tr>
<tr id="350A1E7866470016FF60FB4CFAEDFB25" box="[161,1328,1174,1196]" gridrow="22" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60FB4CFE01FB25" box="[161,476,1174,1196]" gridcol="0" gridrow="22" pageId="3" pageNumber="370">Snout length</th>
<td id="76DB770466470016FD0DFB4CFCF7FB25" box="[716,810,1174,1196]" gridcol="1" gridrow="22" pageId="3" pageNumber="370">14.6</td>
<td id="76DB770466470016FCB3FB4CFC2CFB25" box="[882,1009,1174,1196]" gridcol="2" gridrow="22" pageId="3" pageNumber="370">13.616.5</td>
<td id="76DB770466470016FBABFB4CFB79FB25" box="[1130,1188,1174,1196]" gridcol="3" gridrow="22" pageId="3" pageNumber="370">15.1</td>
<td id="76DB770466470016FAC8FB4CFAEDFB25" box="[1289,1328,1174,1196]" gridcol="4" gridrow="22" pageId="3" pageNumber="370">0.8</td>
</tr>
<tr id="350A1E7866470016FF60FB65FAEDFB5C" box="[161,1328,1215,1237]" gridrow="23" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60FB65FE01FB5C" box="[161,476,1215,1237]" gridcol="0" gridrow="23" pageId="3" pageNumber="370">Eye diameter</th>
<td id="76DB770466470016FD0DFB65FCF7FB5C" box="[716,810,1215,1237]" gridcol="1" gridrow="23" pageId="3" pageNumber="370">3.4</td>
<td id="76DB770466470016FCB3FB65FC2CFB5C" box="[882,1009,1215,1237]" gridcol="2" gridrow="23" pageId="3" pageNumber="370">3.45.6</td>
<td id="76DB770466470016FBABFB65FB79FB5C" box="[1130,1188,1215,1237]" gridcol="3" gridrow="23" pageId="3" pageNumber="370">4.3</td>
<td id="76DB770466470016FAC8FB65FAEDFB5C" box="[1289,1328,1215,1237]" gridcol="4" gridrow="23" pageId="3" pageNumber="370">0.5</td>
</tr>
<tr id="350A1E7866470016FF60FB3DFAEDFB74" box="[161,1328,1255,1277]" gridrow="24" pageId="3" pageNumber="370" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="76DB770466470016FF60FB3DFE01FB74" box="[161,476,1255,1277]" gridcol="0" gridrow="24" pageId="3" pageNumber="370">
<emphasis id="B94EC0286647FFEAFF60FB3DFE51FB74" bold="true" box="[161,396,1255,1277]" pageId="3" pageNumber="370">In % of pelvic-anal di</emphasis>
s
<emphasis id="B94EC0286647FFEAFE54FB3DFE13FB74" bold="true" box="[405,462,1255,1277]" pageId="3" pageNumber="370">tance</emphasis>
</th>
</tr>
<tr id="350A1E7866470016FF60FACAFAEDFAAF" box="[161,1328,1296,1318]" gridrow="25" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60FACAFE01FAAF" box="[161,476,1296,1318]" gridcol="0" gridrow="25" pageId="3" pageNumber="370">Distance from anus to anal fin</th>
<td id="76DB770466470016FD0DFACAFCF7FAAF" box="[716,810,1296,1318]" gridcol="1" gridrow="25" pageId="3" pageNumber="370">20.0</td>
<td id="76DB770466470016FCB3FACAFC2CFAAF" box="[882,1009,1296,1318]" gridcol="2" gridrow="25" pageId="3" pageNumber="370">18.625.1</td>
<td id="76DB770466470016FBABFACAFB79FAAF" box="[1130,1188,1296,1318]" gridcol="3" gridrow="25" pageId="3" pageNumber="370">22.7</td>
<td id="76DB770466470016FAC8FACAFAEDFAAF" box="[1289,1328,1296,1318]" gridcol="4" gridrow="25" pageId="3" pageNumber="370">2.5</td>
</tr>
<tr id="350A1E7866470016FF60FAE2FAEDFAC7" box="[161,1328,1336,1358]" gridrow="26" pageId="3" pageNumber="370" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="76DB770466470016FF60FAE2FE01FAC7" box="[161,476,1336,1358]" gridcol="0" gridrow="26" pageId="3" pageNumber="370">In % of head length</th>
</tr>
<tr id="350A1E7866470016FF60FABBFAEDFAFE" box="[161,1328,1377,1399]" gridrow="27" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60FABBFE01FAFE" box="[161,476,1377,1399]" gridcol="0" gridrow="27" pageId="3" pageNumber="370">Snout length</th>
<td id="76DB770466470016FD0DFABBFCF7FAFE" box="[716,810,1377,1399]" gridcol="1" gridrow="27" pageId="3" pageNumber="370">58.9</td>
<td id="76DB770466470016FCB3FABBFC2CFAFE" box="[882,1009,1377,1399]" gridcol="2" gridrow="27" pageId="3" pageNumber="370">52.064.4</td>
<td id="76DB770466470016FBABFABBFB79FAFE" box="[1130,1188,1377,1399]" gridcol="3" gridrow="27" pageId="3" pageNumber="370">58.4</td>
<td id="76DB770466470016FAC8FABBFAEDFAFE" box="[1289,1328,1377,1399]" gridcol="4" gridrow="27" pageId="3" pageNumber="370">2.8</td>
</tr>
<tr id="350A1E7866470016FF60FA50FAEDFA29" box="[161,1328,1418,1440]" gridrow="28" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60FA50FE01FA29" box="[161,476,1418,1440]" gridcol="0" gridrow="28" pageId="3" pageNumber="370">Eye diameter</th>
<td id="76DB770466470016FD0DFA50FCF7FA29" box="[716,810,1418,1440]" gridcol="1" gridrow="28" pageId="3" pageNumber="370">14.0</td>
<td id="76DB770466470016FCB3FA50FC2CFA29" box="[882,1009,1418,1440]" gridcol="2" gridrow="28" pageId="3" pageNumber="370">14.021.4</td>
<td id="76DB770466470016FBABFA50FB79FA29" box="[1130,1188,1418,1440]" gridcol="3" gridrow="28" pageId="3" pageNumber="370">16.9</td>
<td id="76DB770466470016FAC8FA50FAEDFA29" box="[1289,1328,1418,1440]" gridcol="4" gridrow="28" pageId="3" pageNumber="370">1.8</td>
</tr>
<tr id="350A1E7866470016FF60FA68FAEDFA41" box="[161,1328,1458,1480]" gridrow="29" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60FA68FE01FA41" box="[161,476,1458,1480]" gridcol="0" gridrow="29" pageId="3" pageNumber="370">Disc width</th>
<td id="76DB770466470016FD0DFA68FCF7FA41" box="[716,810,1458,1480]" gridcol="1" gridrow="29" pageId="3" pageNumber="370">41.4</td>
<td id="76DB770466470016FCB3FA68FC2CFA41" box="[882,1009,1458,1480]" gridcol="2" gridrow="29" pageId="3" pageNumber="370">40.961.5</td>
<td id="76DB770466470016FBABFA68FB79FA41" box="[1130,1188,1458,1480]" gridcol="3" gridrow="29" pageId="3" pageNumber="370">52.1</td>
<td id="76DB770466470016FAC8FA68FAEDFA41" box="[1289,1328,1458,1480]" gridcol="4" gridrow="29" pageId="3" pageNumber="370">7.5</td>
</tr>
<tr id="350A1E7866470016FF60FA01FAEDFA78" box="[161,1328,1499,1521]" gridrow="30" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60FA01FE01FA78" box="[161,476,1499,1521]" gridcol="0" gridrow="30" pageId="3" pageNumber="370">Disc length</th>
<td id="76DB770466470016FD0DFA01FCF7FA78" box="[716,810,1499,1521]" gridcol="1" gridrow="30" pageId="3" pageNumber="370">55.1</td>
<td id="76DB770466470016FCB3FA01FC2CFA78" box="[882,1009,1499,1521]" gridcol="2" gridrow="30" pageId="3" pageNumber="370">50.965.7</td>
<td id="76DB770466470016FBABFA01FB79FA78" box="[1130,1188,1499,1521]" gridcol="3" gridrow="30" pageId="3" pageNumber="370">54.7</td>
<td id="76DB770466470016FAC8FA01FAEDFA78" box="[1289,1328,1499,1521]" gridcol="4" gridrow="30" pageId="3" pageNumber="370">4.4</td>
</tr>
<tr id="350A1E7866470016FF60F9D9FAEDF990" box="[161,1328,1539,1561]" gridrow="31" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60F9D9FE01F990" box="[161,476,1539,1561]" gridcol="0" gridrow="31" pageId="3" pageNumber="370">Central callous pad width</th>
<td id="76DB770466470016FD0DF9D9FCF7F990" box="[716,810,1539,1561]" gridcol="1" gridrow="31" pageId="3" pageNumber="370">22.2</td>
<td id="76DB770466470016FCB3F9D9FC2CF990" box="[882,1009,1539,1561]" gridcol="2" gridrow="31" pageId="3" pageNumber="370">21.934.3</td>
<td id="76DB770466470016FBABF9D9FB79F990" box="[1130,1188,1539,1561]" gridcol="3" gridrow="31" pageId="3" pageNumber="370">29.1</td>
<td id="76DB770466470016FAC8F9D9FAEDF990" box="[1289,1328,1539,1561]" gridcol="4" gridrow="31" pageId="3" pageNumber="370">4.6</td>
</tr>
<tr id="350A1E7866470016FF60F9F6FAEDF9CB" box="[161,1328,1580,1602]" gridrow="32" pageId="3" pageNumber="370">
<th id="76DB770466470016FF60F9F6FE01F9CB" box="[161,476,1580,1602]" gridcol="0" gridrow="32" pageId="3" pageNumber="370">Central callous pad length</th>
<td id="76DB770466470016FD0DF9F6FCF7F9CB" box="[716,810,1580,1602]" gridcol="1" gridrow="32" pageId="3" pageNumber="370">31.1</td>
<td id="76DB770466470016FCB3F9F6FC2CF9CB" box="[882,1009,1580,1602]" gridcol="2" gridrow="32" pageId="3" pageNumber="370">20.835.2</td>
<td id="76DB770466470016FBABF9F6FB79F9CB" box="[1130,1188,1580,1602]" gridcol="3" gridrow="32" pageId="3" pageNumber="370">26.0</td>
<td id="76DB770466470016FAC8F9F6FAEDF9CB" box="[1289,1328,1580,1602]" gridcol="4" gridrow="32" pageId="3" pageNumber="370">4.9</td>
</tr>
<tr id="350A1E7866470016FF60F98EFAEDF9E3" box="[161,1328,1620,1642]" gridrow="33" pageId="3" pageNumber="370" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="76DB770466470016FF60F98EFE01F9E3" box="[161,476,1620,1642]" gridcol="0" gridrow="33" pageId="3" pageNumber="370">Meristic counts</th>
</tr>
<tr id="350A1E7866470016FF60F9A7FAEDF91A" box="[161,1328,1661,1683]" gridrow="34" pageId="3" pageNumber="370" rowspan-3="1" rowspan-4="1">
<th id="76DB770466470016FF60F9A7FE01F91A" box="[161,476,1661,1683]" gridcol="0" gridrow="34" pageId="3" pageNumber="370">Dorsal- fin</th>
<td id="76DB770466470016FD0DF9A7FCF7F91A" box="[716,810,1661,1683]" gridcol="1" gridrow="34" pageId="3" pageNumber="370">iii/9½</td>
<td id="76DB770466470016FCB3F9A7FC2CF91A" box="[882,1009,1661,1683]" gridcol="2" gridrow="34" pageId="3" pageNumber="370">iii/8½</td>
</tr>
<tr id="350A1E7866470016FF60F97FFAEDF932" box="[161,1328,1701,1723]" gridrow="35" pageId="3" pageNumber="370" rowspan-3="1" rowspan-4="1">
<th id="76DB770466470016FF60F97FFE01F932" box="[161,476,1701,1723]" gridcol="0" gridrow="35" pageId="3" pageNumber="370">Pectoral- fin</th>
<td id="76DB770466470016FD0DF97FFCF7F932" box="[716,810,1701,1723]" gridcol="1" gridrow="35" pageId="3" pageNumber="370">i/15</td>
<td id="76DB770466470016FCB3F97FFC2CF932" box="[882,1009,1701,1723]" gridcol="2" gridrow="35" pageId="3" pageNumber="370">i/1415</td>
</tr>
<tr id="350A1E7866470016FF60F914FAEDF96D" box="[161,1328,1742,1764]" gridrow="36" pageId="3" pageNumber="370" rowspan-3="1" rowspan-4="1">
<th id="76DB770466470016FF60F914FE01F96D" box="[161,476,1742,1764]" gridcol="0" gridrow="36" pageId="3" pageNumber="370">Pelvic- fin</th>
<td id="76DB770466470016FD0DF914FCF7F96D" box="[716,810,1742,1764]" gridcol="1" gridrow="36" pageId="3" pageNumber="370">i/8</td>
<td id="76DB770466470016FCB3F914FC2CF96D" box="[882,1009,1742,1764]" gridcol="2" gridrow="36" pageId="3" pageNumber="370">i/78</td>
</tr>
<tr id="350A1E7866470016FF60F92CFAEDF885" box="[161,1328,1782,1804]" gridrow="37" pageId="3" pageNumber="370" rowspan-3="1" rowspan-4="1">
<th id="76DB770466470016FF60F92CFE01F885" box="[161,476,1782,1804]" gridcol="0" gridrow="37" pageId="3" pageNumber="370">Anal- fin</th>
<td id="76DB770466470016FD0DF92CFCF7F885" box="[716,810,1782,1804]" gridcol="1" gridrow="37" pageId="3" pageNumber="370">iii/5½</td>
<td id="76DB770466470016FCB3F92CFC2CF885" box="[882,1009,1782,1804]" gridcol="2" gridrow="37" pageId="3" pageNumber="370">iii/5½</td>
</tr>
<tr id="350A1E7866470016FF60F8C5FAEDF8BC" box="[161,1328,1823,1845]" gridrow="38" pageId="3" pageNumber="370" rowspan-3="1" rowspan-4="1">
<th id="76DB770466470016FF60F8C5FE01F8BC" box="[161,476,1823,1845]" gridcol="0" gridrow="38" pageId="3" pageNumber="370">Lateral line scales</th>
<td id="76DB770466470016FD0DF8C5FCF7F8BC" box="[716,810,1823,1845]" gridcol="1" gridrow="38" pageId="3" pageNumber="370">34</td>
<td id="76DB770466470016FCB3F8C5FC2CF8BC" box="[882,1009,1823,1845]" gridcol="2" gridrow="38" pageId="3" pageNumber="370">3334</td>
</tr>
<tr id="350A1E7866470016FF60F89DFAEDF8D4" box="[161,1328,1863,1885]" gridrow="39" pageId="3" pageNumber="370" rowspan-3="1" rowspan-4="1">
<th id="76DB770466470016FF60F89DFE01F8D4" box="[161,476,1863,1885]" gridcol="0" gridrow="39" pageId="3" pageNumber="370">Predorsal scales</th>
<td id="76DB770466470016FD0DF89DFCF7F8D4" box="[716,810,1863,1885]" gridcol="1" gridrow="39" pageId="3" pageNumber="370">10</td>
<td id="76DB770466470016FCB3F89DFC2CF8D4" box="[882,1009,1863,1885]" gridcol="2" gridrow="39" pageId="3" pageNumber="370">910</td>
</tr>
<tr id="350A1E7866470016FF60F8AAFAEDF80F" box="[161,1328,1904,1926]" gridrow="40" pageId="3" pageNumber="370" rowspan-3="1" rowspan-4="1">
<th id="76DB770466470016FF60F8AAFE01F80F" box="[161,476,1904,1926]" gridcol="0" gridrow="40" pageId="3" pageNumber="370">Preanal scales</th>
<td id="76DB770466470016FD0DF8AAFCF7F80F" box="[716,810,1904,1926]" gridcol="1" gridrow="40" pageId="3" pageNumber="370">4</td>
<td id="76DB770466470016FCB3F8AAFC2CF80F" box="[882,1009,1904,1926]" gridcol="2" gridrow="40" pageId="3" pageNumber="370">34</td>
</tr>
<tr id="350A1E7866470016FF60F842FAEDF827" box="[161,1328,1944,1966]" gridrow="41" pageId="3" pageNumber="370" rowspan-3="1" rowspan-4="1">
<th id="76DB770466470016FF60F842FE01F827" box="[161,476,1944,1966]" gridcol="0" gridrow="41" pageId="3" pageNumber="370">Dorsal fin base scales</th>
<td id="76DB770466470016FD0DF842FCF7F827" box="[716,810,1944,1966]" gridcol="1" gridrow="41" pageId="3" pageNumber="370">7</td>
<td id="76DB770466470016FCB3F842FC2CF827" box="[882,1009,1944,1966]" gridcol="2" gridrow="41" pageId="3" pageNumber="370">67</td>
</tr>
<tr id="350A1E7866470016FF60F81BFAEDF85E" box="[161,1328,1985,2007]" gridrow="42" pageId="3" pageNumber="370" rowspan-3="1" rowspan-4="1">
<th id="76DB770466470016FF60F81BFE01F85E" box="[161,476,1985,2007]" gridcol="0" gridrow="42" pageId="3" pageNumber="370">Anal fin base scales</th>
<td id="76DB770466470016FD0DF81BFCF7F85E" box="[716,810,1985,2007]" gridcol="1" gridrow="42" pageId="3" pageNumber="370">4</td>
<td id="76DB770466470016FCB3F81BFC2CF85E" box="[882,1009,1985,2007]" gridcol="2" gridrow="42" pageId="3" pageNumber="370">45</td>
</tr>
</table>
</paragraph>
<caption id="DF454CB26640FFEDFF56FAFAFED5FAFA" httpUri="https://zenodo.org/record/428279/files/figure.png" pageId="4" pageNumber="371" targetBox="[163,1425,216,1261]" targetPageId="4">
<paragraph id="8B851C3A6640FFEDFF56FAFAFED5FAFA" blockId="4.[151,1436,1312,1395]" pageId="4" pageNumber="371">
<emphasis id="B94EC0286640FFEDFF56FAFAFECEFABF" bold="true" box="[151,275,1312,1334]" pageId="4" pageNumber="371">FIGURE 2.</emphasis>
Snout of
<taxonomicName id="4C3A67B96640FFEDFEB6FAFBFDF2FABF" box="[375,559,1312,1334]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="371" phylum="Chordata" rank="species" species="clavirostris">
<emphasis id="B94EC0286640FFEDFEB6FAFBFDF2FABF" box="[375,559,1312,1334]" italics="true" pageId="4" pageNumber="371">Garra clavirostris</emphasis>
</taxonomicName>
,
<emphasis id="B94EC0286640FFEDFDFDFAFAFD92FABF" bold="true" box="[572,591,1312,1334]" pageId="4" pageNumber="371">a.</emphasis>
morphology of snout showing transverse lobe, proboscis, depressed rostral surface and lateral surfaces of holotype;
<emphasis id="B94EC0286640FFEDFE1EFAE5FE2EFADC" bold="true" box="[479,499,1343,1365]" pageId="4" pageNumber="371">b.</emphasis>
transverse lobe showing tubercles (MUMF 22001, paratype, 87.7 mm SL);
<emphasis id="B94EC0286640FFEDFB2EFAE5FADCFADC" bold="true" box="[1263,1281,1343,1365]" pageId="4" pageNumber="371">c.</emphasis>
one tubercle in b enlarged.
</paragraph>
</caption>
<subSubSection id="C3204FB16640FFEDFF06FA7AFD26FA54" pageId="4" pageNumber="371" type="etymology">
<paragraph id="8B851C3A6640FFEDFF06FA7AFD26FA54" blockId="4.[151,1436,1440,2005]" pageId="4" pageNumber="371">
<emphasis id="B94EC0286640FFEDFF06FA7AFE93FA31" bold="true" box="[199,334,1440,1464]" pageId="4" pageNumber="371">Etymology.</emphasis>
The species-name is derived from the Latin clava(club), an allusion to the shape of the proboscis; and rostrum(snout). A noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C3204FB16640FFECFF06FA3DFC2CFA0C" lastPageId="5" lastPageNumber="372" pageId="4" pageNumber="371" type="discussion">
<paragraph id="8B851C3A6640FFECFF06FA3DFA4EFE7C" blockId="4.[151,1436,1440,2005]" lastBlockId="5.[151,1437,151,501]" lastPageId="5" lastPageNumber="372" pageId="4" pageNumber="371">
<emphasis id="B94EC0286640FFEDFF06FA3DFE91F989" bold="true" box="[199,332,1511,1536]" pageId="4" pageNumber="371">Discussion.</emphasis>
The characteristics of the snout and the size and distribution patterns of the tubercles on the proboscis are of taxonomic significance in distinguishing species of the genus
<taxonomicName id="4C3A67B96640FFEDFBDBF9D7FF06F9C1" authority="Nebeshwar &amp; Vishwanath 2013" authorityName="Nebeshwar &amp; Vishwanath" authorityYear="2013" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="371" phylum="Chordata" rank="genus">
<emphasis id="B94EC0286640FFEDFBDBF9D7FBBDF9AD" box="[1050,1120,1549,1572]" italics="true" pageId="4" pageNumber="371">Garra</emphasis>
(
<bibRefCitation id="EFAB61CB6640FFEDFBB0F9D6FF09F9C1" author="Nebeshwar" pageId="4" pageNumber="371" refString="Nebeshwar, K. &amp; Vishwanath, W. (2013) Three new species of Garra (Pisces: Cyprinidae) from north-eastern India and redescription of G. gotyla. Ichthyological Exploration of Freshwaters, 24, 97 - 192." type="journal article" year="2013">Nebeshwar &amp; Vishwanath 2013</bibRefCitation>
)
</taxonomicName>
.
<taxonomicName id="4C3A67B96640FFEDFF27F9EAFE6CF9C1" box="[230,433,1584,1608]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="371" phylum="Chordata" rank="species" species="clavirostris">
<emphasis id="B94EC0286640FFEDFF27F9EAFE6CF9C1" box="[230,433,1584,1608]" italics="true" pageId="4" pageNumber="371">Garra clavirostris</emphasis>
</taxonomicName>
has a prominent proboscis and is therefore compared only with the congeners in the river drainages of northeast
<collectingCountry id="F32D5CAA6640FFEDFE59F98EFE0FF9E4" box="[408,466,1620,1645]" name="India" pageId="4" pageNumber="371">India</collectingCountry>
having a proboscis on the snout.
<taxonomicName id="4C3A67B96640FFEDFC8FF98FFBC7F9E5" box="[846,1050,1620,1644]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="371" phylum="Chordata" rank="species" species="clavirostris">
<emphasis id="B94EC0286640FFEDFC8FF98FFBC7F9E5" box="[846,1050,1620,1644]" italics="true" pageId="4" pageNumber="371">Garra clavirostris</emphasis>
</taxonomicName>
is easily distinguished from other proboscis-bearing congeners, viz.,
<taxonomicName id="4C3A67B96640FFEDFDDDF9A2FD38F919" box="[540,741,1656,1680]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="371" phylum="Chordata" rank="species" species="arunachalensis">
<emphasis id="B94EC0286640FFEDFDDDF9A2FD38F919" box="[540,741,1656,1680]" italics="true" pageId="4" pageNumber="371">G. arunachalensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A67B96640FFEDFD33F9A2FCB3F919" box="[754,878,1656,1680]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="371" phylum="Chordata" rank="species" species="birostris">
<emphasis id="B94EC0286640FFEDFD33F9A2FCB3F919" box="[754,878,1656,1680]" italics="true" pageId="4" pageNumber="371">G. birostris</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A67B96640FFEDFCBDF9A2FBD5F919" box="[892,1032,1656,1680]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="371" phylum="Chordata" rank="species" species="cornigera">
<emphasis id="B94EC0286640FFEDFCBDF9A2FBD5F919" box="[892,1032,1656,1680]" italics="true" pageId="4" pageNumber="371">G. cornigera</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A67B96640FFEDFBD7F9A2FB4BF919" box="[1046,1174,1656,1680]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="371" phylum="Chordata" rank="species" species="elongata">
<emphasis id="B94EC0286640FFEDFBD7F9A2FB4BF919" box="[1046,1174,1656,1680]" italics="true" pageId="4" pageNumber="371">G. elongata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A67B96640FFEDFB62F9A2FADAF919" box="[1187,1287,1656,1680]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="371" phylum="Chordata" rank="species" species="gotyla">
<emphasis id="B94EC0286640FFEDFB62F9A2FADAF919" box="[1187,1287,1656,1680]" italics="true" pageId="4" pageNumber="371">G. gotyla</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A67B96640FFEDFAD4F9A2FA48F919" box="[1301,1429,1656,1680]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="371" phylum="Chordata" rank="species" species="kalpangi">
<emphasis id="B94EC0286640FFEDFAD4F9A2FA48F919" box="[1301,1429,1656,1680]" italics="true" pageId="4" pageNumber="371">G. kalpangi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A67B96640FFEDFF56F947FEFBF93D" box="[151,294,1692,1716]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="371" phylum="Chordata" rank="species" species="litanensis">
<emphasis id="B94EC0286640FFEDFF56F947FEFBF93D" box="[151,294,1692,1716]" italics="true" pageId="4" pageNumber="371">G. litanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A67B96640FFEDFEF9F947FE7FF93D" box="[312,418,1693,1716]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="371" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="B94EC0286640FFEDFEF9F947FE7FF93D" box="[312,418,1693,1716]" italics="true" pageId="4" pageNumber="371">G. nasuta</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A67B96640FFEDFE79F947FD59F93D" box="[440,644,1692,1716]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="371" phylum="Chordata" rank="species" species="quadratirostris">
<emphasis id="B94EC0286640FFEDFE79F947FD59F93D" box="[440,644,1692,1716]" italics="true" pageId="4" pageNumber="371">G. quadratirostris</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A67B96640FFEDFD05F947FC94F93D" box="[708,841,1692,1716]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="371" phylum="Chordata" rank="species" species="trilobata">
<emphasis id="B94EC0286640FFEDFD05F947FC94F93D" box="[708,841,1692,1716]" italics="true" pageId="4" pageNumber="371">G. trilobata</emphasis>
</taxonomicName>
as follows. From
<taxonomicName id="4C3A67B96640FFEDFBE7F947FB28F93D" box="[1062,1269,1692,1716]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="371" phylum="Chordata" rank="species" species="arunachalensis">
<emphasis id="B94EC0286640FFEDFBE7F947FB28F93D" box="[1062,1269,1692,1716]" italics="true" pageId="4" pageNumber="371">G. arunachalensis</emphasis>
</taxonomicName>
(Brahmaputra drainage) in having the depressed rostral surface slightly convex (vs. flat), 33¯34 (vs. 35) lateral-line scales, presence (vs. absence) of anterolateral lobe of the lower lip, and absence (vs. presence) of minute tubercles and pores on the depressed rostral surface; from
<taxonomicName id="4C3A67B96640FFEDFD4DF8D2FCD7F8A9" box="[652,778,1800,1824]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="371" phylum="Chordata" rank="species" species="birostris">
<emphasis id="B94EC0286640FFEDFD4DF8D2FCD7F8A9" box="[652,778,1800,1824]" italics="true" pageId="4" pageNumber="371">G. birostris</emphasis>
</taxonomicName>
(Brahmaputra drainage) in having a club-shaped unilobed (vs. bilobed) proboscis, disc of the lower lip oval (vs. elliptical) and sublachrymal groove connected (vs. not connected) to rostral-cap groove; from
<taxonomicName id="4C3A67B96640FFEDFD92F88AFD6BF8E1" box="[595,694,1872,1896]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="371" phylum="Chordata" rank="species" species="gotyla">
<emphasis id="B94EC0286640FFEDFD92F88AFD6BF8E1" box="[595,694,1872,1896]" italics="true" pageId="4" pageNumber="371">G. gotyla</emphasis>
</taxonomicName>
(Brahamaputra drainage) in having the forward protrusion of the proboscis reaching (vs. not reaching) the transverse groove and a slightly (vs. highly) convex depressed rostral surface; from
<taxonomicName id="4C3A67B96640FFEDFEF5F843FE20F839" box="[308,509,1944,1968]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="371" phylum="Chordata" rank="species" species="quadratirostris">
<emphasis id="B94EC0286640FFEDFEF5F843FE20F839" box="[308,509,1944,1968]" italics="true" pageId="4" pageNumber="371">G. quadratirostris</emphasis>
</taxonomicName>
(Brahmaputra drainage) in having 33¯34 (vs. 37) lateral-line scales, 34 (vs. 56) preanal scales and slightly convex (vs. creased and convex) depressed rostral surface; from
<taxonomicName id="4C3A67B96640FFEDFAD3F867FA41F85D" box="[1298,1436,1980,2004]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="371" phylum="Chordata" rank="species" species="kalpangi">
<emphasis id="B94EC0286640FFEDFAD3F867FA41F85D" box="[1298,1436,1980,2004]" italics="true" pageId="4" pageNumber="371">G. kalpangi</emphasis>
</taxonomicName>
(Brahamaputra drainage) in having a well-developed (vs. weakly developed) proboscis, 4½5½ (vs. 3½) transverse scale rows above the lateral line and 1416 (vs. 1012) branched pectoral-fin rays; from
<taxonomicName id="4C3A67B96641FFECFB59FF66FAC6FF5D" box="[1176,1307,188,212]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="372" phylum="Chordata" rank="species" species="elongata">
<emphasis id="B94EC0286641FFECFB59FF66FAC6FF5D" box="[1176,1307,188,212]" italics="true" pageId="5" pageNumber="372">G. elongata</emphasis>
</taxonomicName>
(Chindwin drainage) in having prominent (vs. weakly developed) proboscis, 3334 (vs. 3940) lateral-line scales, the absence (vs. presence) of a transverse black bar on the dorsal fin and the absence (vs. presence) of a longitudinal black band on the middle of the caudal fin; from
<taxonomicName id="4C3A67B96641FFECFDFDFEF2FD1AFEC9" box="[572,711,296,320]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="372" phylum="Chordata" rank="species" species="litanensis">
<emphasis id="B94EC0286641FFECFDFDFEF2FD1AFEC9" box="[572,711,296,320]" italics="true" pageId="5" pageNumber="372">G. litanensis</emphasis>
</taxonomicName>
(Chindwin drainage) in having 3334 (vs. 32) lateral-line scales and the absence (vs. presence) of black spots along the base of the dorsal fin; from
<taxonomicName id="4C3A67B96641FFECFB47FE96FAC5FEED" box="[1158,1304,332,356]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="372" phylum="Chordata" rank="species" species="cornigera">
<emphasis id="B94EC0286641FFECFB47FE96FAC5FEED" box="[1158,1304,332,356]" italics="true" pageId="5" pageNumber="372">G. cornigera</emphasis>
</taxonomicName>
(Chindwin drainage) in having a unilobed (vs. bilobed) proboscis, the presence (vs. absence) of a black spot at the upper angle of the gill opening, slightly convex (vs. flat) depressed rostral surface and 4½5½ (vs. 3½) transverse scale rows above the lateral line; and from
<taxonomicName id="4C3A67B96641FFECFE3AFE62FDA6FE59" box="[507,635,440,464]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="372" phylum="Chordata" rank="species" species="trilobata">
<emphasis id="B94EC0286641FFECFE3AFE62FDA6FE59" box="[507,635,440,464]" italics="true" pageId="5" pageNumber="372">G. trilobata</emphasis>
</taxonomicName>
(Chindwin drainage) in having an unilobed (vs. trilobed) proboscis, the presence (vs. absence) of an anterolateral lobe of the lower lip and in having 3334 (vs. 3132) lateral-line scales.
</paragraph>
<caption id="DF454CB26641FFECFF56FB38FA94FB71" box="[151,1353,1250,1272]" httpUri="https://zenodo.org/record/428281/files/figure.png" pageId="5" pageNumber="372" targetBox="[355,1247,557,1229]" targetPageId="5">
<paragraph id="8B851C3A6641FFECFF56FB38FA94FB71" blockId="5.[151,1353,1250,1272]" box="[151,1353,1250,1272]" pageId="5" pageNumber="372">
<emphasis id="B94EC0286641FFECFF56FB38FED0FB71" bold="true" box="[151,269,1250,1272]" pageId="5" pageNumber="372">FIGURE 3</emphasis>
. Lateral view of snout of
<taxonomicName id="4C3A67B96641FFECFDD3FB39FD17FB71" box="[530,714,1250,1272]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="372" phylum="Chordata" rank="species" species="clavirostris">
<emphasis id="B94EC0286641FFECFDD3FB39FD17FB71" box="[530,714,1250,1272]" italics="true" pageId="5" pageNumber="372">Garra clavirostris</emphasis>
</taxonomicName>
, holotype, showing shape of proboscis and the position of eye.
</paragraph>
</caption>
<paragraph id="8B851C3A6641FFECFF06FAFFFC2CFA0C" blockId="5.[151,1436,1317,1810]" pageId="5" pageNumber="372">
MClelland (1838) described
<taxonomicName id="4C3A67B96641FFECFDD7FAFCFD5CFAB4" box="[534,641,1318,1341]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="372" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="B94EC0286641FFECFDD7FAFCFD5CFAB4" box="[534,641,1318,1341]" italics="true" pageId="5" pageNumber="372">G. nasuta</emphasis>
</taxonomicName>
from a single specimen from the Khasi hills and provides insufficient information for its diagnosis, except for its characteristic presence of a pit between the nares. The new species, however, is distinct from
<taxonomicName id="4C3A67B96641FFECFE75FAB4FDC3FA0C" box="[436,542,1390,1413]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="372" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="B94EC0286641FFECFE75FAB4FDC3FA0C" box="[436,542,1390,1413]" italics="true" pageId="5" pageNumber="372">G. nasuta</emphasis>
</taxonomicName>
in the absence of a pit between the nares.
</paragraph>
</subSubSection>
<subSubSection id="C3204FB16641FFECFF06FA4BFC3AF898" pageId="5" pageNumber="372" type="reference_group">
<paragraph id="8B851C3A6641FFECFF06FA4BFC3AF898" blockId="5.[151,1436,1317,1810]" pageId="5" pageNumber="372">
<taxonomicName id="4C3A67B96641FFECFF06FA4BFE45FA20" box="[199,408,1425,1449]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="372" phylum="Chordata" rank="species" species="clavirostris">
<emphasis id="B94EC0286641FFECFF06FA4BFE45FA20" box="[199,408,1425,1449]" italics="true" pageId="5" pageNumber="372">Garra clavirostris</emphasis>
</taxonomicName>
can be distinguished from the two recently-described species occurring in
<collectingCountry id="F32D5CAA6641FFECFADEFA4BFAAEFA23" box="[1311,1395,1425,1450]" name="Bhutan" pageId="5" pageNumber="372">Bhutan</collectingCountry>
,
<taxonomicName id="4C3A67B96641FFECFA45FA4BFE99FA44" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="372" phylum="Chordata" rank="species" species="bimaculacauda">
<emphasis id="B94EC0286641FFECFA45FA4BFE99FA44" italics="true" pageId="5" pageNumber="372">G. bimaculacauda</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A67B96641FFECFEBDFA6CFDAFFA44" box="[380,626,1461,1485]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="372" phylum="Chordata" rank="species" species="parastenorhynchus">
<emphasis id="B94EC0286641FFECFEBDFA6CFDAFFA44" box="[380,626,1461,1485]" italics="true" pageId="5" pageNumber="372">G. parastenorhynchus</emphasis>
</taxonomicName>
, in the absence (vs. presence) of two distinct black spots on the caudal fin, one each on the tips of the dorsal and ventral lobes). In fact these spots (see
<figureCitation id="130100BF6641FFECFBFEFA03FBA0FA78" box="[1087,1149,1497,1522]" captionStart="FIGURE 3" captionStartId="5.[151,250,1250,1272]" captionTargetBox="[355,1247,557,1229]" captionTargetId="figure@5.[340,1247,553,1229]" captionTargetPageId="5" captionText="FIGURE 3. Lateral view of snout of Garra clavirostris, holotype, showing shape of proboscis and the position of eye." httpUri="https://zenodo.org/record/428281/files/figure.png" pageId="5" pageNumber="372">Fig 3</figureCitation>
, page 118, of
<bibRefCitation id="EFAB61CB6641FFECFADEFA03FF0CF99F" author="Thoni" pageId="5" pageNumber="372" refString="Thoni, R. J., Gurung, D. B. &amp; Mayden, R. L. (2016) A new review of the genus Garra Hamilton 1822 of Bhutan, including the description of two new species and three additional records (Cypriniformes: Cyprinidae). Zootaxa, 4169 (1), 115 - 132. https: // doi. org / 10.11646 / zootaxa. 4169.1.5" type="journal article" year="2016">
Thoni
<emphasis id="B94EC0286641FFECFAABFA00FA41FA78" box="[1386,1436,1497,1521]" italics="true" pageId="5" pageNumber="372">et al</emphasis>
2016
</bibRefCitation>
) may be better described as elongated dark brown pigmentation on two or three principal rays. The new species also differs from
<taxonomicName id="4C3A67B96641FFECFE7BF9FBFD6CF9B0" box="[442,689,1569,1593]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="372" phylum="Chordata" rank="species" species="parastenorhynchus">
<emphasis id="B94EC0286641FFECFE7BF9FBFD6CF9B0" box="[442,689,1569,1593]" italics="true" pageId="5" pageNumber="372">G. parastenorhynchus</emphasis>
</taxonomicName>
in having a more pronounced (vs. less pronounced) club-shaped proboscis, the club overhanging the depressed rostral surface (vs. not overhanging); in the absence (vs. presence) of a conspicuous dark spot at the caudal-fin base, the absence (vs presence) of a series of 8 ̶9 dark-brown spots at the base of dorsal fin (as in
<bibRefCitation id="EFAB61CB6641FFECFE67F957FDB6F92F" author="Thoni" box="[422,619,1677,1702]" pageId="5" pageNumber="372" refString="Thoni, R. J., Gurung, D. B. &amp; Mayden, R. L. (2016) A new review of the genus Garra Hamilton 1822 of Bhutan, including the description of two new species and three additional records (Cypriniformes: Cyprinidae). Zootaxa, 4169 (1), 115 - 132. https: // doi. org / 10.11646 / zootaxa. 4169.1.5" type="journal article" year="2016">
Thoni
<emphasis id="B94EC0286641FFECFE30F954FDFFF92C" box="[497,546,1677,1701]" italics="true" pageId="5" pageNumber="372">et al</emphasis>
, 2016
</bibRefCitation>
, Fig. 9 and R.J. Thoni, pers. comm) and also in the tuberculation pattern on the snout.
<taxonomicName id="4C3A67B96641FFECFE81F96BFDCFF940" box="[320,530,1713,1737]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="372" phylum="Chordata" rank="species" species="clavirostris">
<emphasis id="B94EC0286641FFECFE81F96BFDCFF940" box="[320,530,1713,1737]" italics="true" pageId="5" pageNumber="372">Garra clavirostris</emphasis>
</taxonomicName>
also possesses a greater number of multicuspid tubercles on the proboscis, transverse lobe and lateral surface of the snout. The rostral lobe referred to by
<bibRefCitation id="EFAB61CB6641FFECFBF3F90FFAD9F967" box="[1074,1284,1749,1774]" pageId="5" pageNumber="372" refString="Thoni, R. J., Gurung, D. B. &amp; Mayden, R. L. (2016) A new review of the genus Garra Hamilton 1822 of Bhutan, including the description of two new species and three additional records (Cypriniformes: Cyprinidae). Zootaxa, 4169 (1), 115 - 132. https: // doi. org / 10.11646 / zootaxa. 4169.1.5" type="journal article">
Thoni
<emphasis id="B94EC0286641FFECFBBCF90CFB72F964" box="[1149,1199,1749,1773]" italics="true" pageId="5" pageNumber="372">et al</emphasis>
(2016)
</bibRefCitation>
is in fact the transverse lobe (
<bibRefCitation id="EFAB61CB6641FFECFE8FF923FD82F89B" author="Zhang" box="[334,607,1785,1810]" pageId="5" pageNumber="372" refString="Zhang, E. &amp; Kottelat, M. (2006) Akrokolioplax, a new genus of Southeast Asian labeonine fishes (Teleostei: Cyprinidae). Zootaxa, 1225, 21 - 30." type="journal article" year="2006">Zhang &amp; Kottelat, 2006</bibRefCitation>
;
<bibRefCitation id="EFAB61CB6641FFECFDABF923FC06F898" author="Nebeshwar" box="[618,987,1785,1810]" pageId="5" pageNumber="372" refString="Nebeshwar, K. &amp; Vishwanath, W. (2013) Three new species of Garra (Pisces: Cyprinidae) from north-eastern India and redescription of G. gotyla. Ichthyological Exploration of Freshwaters, 24, 97 - 192." type="journal article" year="2013">Nebeshwar &amp; Vishwanath, 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>