477 lines
73 KiB
XML
477 lines
73 KiB
XML
<document id="8BC32411772922883D11B82E52116CF0" ID-CLB-Dataset="298729" ID-DOI="10.11646/zootaxa.5471.2.6" ID-GBIF-Dataset="f3432e7e-46a0-47ef-b289-51be82e17f07" ID-ISSN="1175-5326" ID-Zenodo-Dep="12190006" ID-ZooBank="0A8FCD2C-A657-4B98-93EE-CD64E5E94FA6" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1718893067710" checkinUser="plazi" docAuthor="Felder, Darryl L. & Tavares, Marcos" docDate="2024" docId="5D1787F8FFB1FFD2FF12FA1F886FF835" docLanguage="en" docName="zootaxa.5471.2.6.pdf" docOrigin="Zootaxa 5471 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5471.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cymonomus windsorae Felder & Tavares 2024, n. sp." docType="treatment" docVersion="6" lastPageNumber="265" masterDocId="A12EFF80FFB0FFD6FF85FF9C8946FFA0" masterDocTitle="A new species of Cymonomus A. Milne-Edwards, 1880, from the deep waters of the Gulf of Mexico (Crustacea: Brachyura: Cymonomidae)" masterLastPageNumber="267" masterPageNumber="261" pageNumber="262" updateTime="1719001744511" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="15D51DC028ADA2A129CCF446F4EF22F2">A new species of Cymonomus A. Milne-Edwards, 1880, from the deep waters of the Gulf of Mexico (Crustacea: Brachyura: Cymonomidae)</mods:title>
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<mods:roleTerm id="C80967CF996FDD4A0646200B29EC72AE">Author</mods:roleTerm>
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<mods:namePart id="DE14589575B1FB4C75363947910C7AE0">Felder, Darryl L.</mods:namePart>
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<mods:nameIdentifier id="F598AEC148C6F015446860BA520E4A92" type="ORCID">0000-0001-7679-7712</mods:nameIdentifier>
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<mods:affiliation id="DC28A16786512103D82A01DC12065FC1">Department of Biology, University of Louisiana, Lafayette, Louisiana 70504 - 2451, U. S. A. & Department of Invertebrate Zoology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20560, U. S. A</mods:affiliation>
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<mods:nameIdentifier id="3CCF5DEB689F3ED1D2BCF19EAE00C233" type="email">dlf4517@louisiana.edu</mods:nameIdentifier>
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<mods:namePart id="9C1946308F840D0F0C9D0F1BE6BC6D0A">Tavares, Marcos</mods:namePart>
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<mods:affiliation id="0FAC7BF43A21BB6DE237DCB44D0C8E76">Museu de Zoologia, Universidade de São Paulo, Ipiranga 04263 - 000, São Paulo, SP, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="E139A986D37863E38073FEC039C683FD" type="email">mdst@usp.br</mods:nameIdentifier>
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<mods:title id="D7930FB6FADBFF4BC53476219FC19BFE">Zootaxa</mods:title>
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<mods:part id="FC5B2E93D32CB00CFF5F827BCBE3901E">
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<mods:date id="1C8301EBFA5597A69885A52290003C47">2024</mods:date>
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<mods:number id="01D8F10F974854BDEBF79E4D9778C0E8">2024-06-18</mods:number>
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<mods:number id="96BD18D22A331C3B80B21236CC5D0876">5471</mods:number>
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<treatment id="5D1787F8FFB1FFD2FF12FA1F886FF835" ID-DOI="http://doi.org/10.5281/zenodo.12190735" ID-GBIF-Taxon="233317635" ID-Zenodo-Dep="12190735" LSID="urn:lsid:plazi:treatment:5D1787F8FFB1FFD2FF12FA1F886FF835" httpUri="http://treatment.plazi.org/id/5D1787F8FFB1FFD2FF12FA1F886FF835" lastPageId="4" lastPageNumber="265" pageId="1" pageNumber="262">
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<subSubSection id="9DA46565FFB1FFD7FF12FA1F88BAFA3D" box="[151,508,1410,1437]" pageId="1" pageNumber="262" type="nomenclature">
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<paragraph id="D50136EEFFB1FFD7FF12FA1F88BAFA3D" blockId="1.[151,508,1410,1473]" box="[151,508,1410,1437]" pageId="1" pageNumber="262">
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<heading id="8E498182FFB1FFD7FF12FA1F88BAFA3D" bold="true" box="[151,508,1410,1437]" fontSize="11" level="1" pageId="1" pageNumber="262" reason="1">
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<emphasis id="E7CAEAFCFFB1FFD7FF12FA1F88BAFA3D" bold="true" box="[151,508,1410,1437]" pageId="1" pageNumber="262">
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<taxonomicName id="12BE4D6DFFB1FFD7FF12FA1F88F4FA3D" authority="Felder & Tavares, 2024" authorityName="Felder & Tavares" authorityYear="2024" box="[151,434,1410,1437]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="windsorae" status="sp. nov.">
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<emphasis id="E7CAEAFCFFB1FFD7FF12FA1F88F4FA3D" bold="true" box="[151,434,1410,1437]" italics="true" pageId="1" pageNumber="262">Cymonomus windsorae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="FCF95787FFB1FFD7FE3CFA1F88BAFA3D" box="[441,508,1411,1437]" pageId="1" pageNumber="262" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="9DA46565FFB1FFD7FF1BFA3B883AFA61" box="[158,380,1447,1473]" pageId="1" pageNumber="262" type="description">
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<paragraph id="D50136EEFFB1FFD7FF1BFA3B883AFA61" blockId="1.[151,508,1410,1473]" box="[158,380,1447,1473]" pageId="1" pageNumber="262">
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(
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<figureCitation id="4D852A6BFFB1FFD7FF23FA3B8864FA61" box="[166,290,1447,1473]" captionStart="FIGURE 1" captionStartId="2.[151,250,1685,1709]" captionTargetBox="[159,1430,639,1653]" captionTargetId="figure-211@2.[151,1436,630,1659]" captionTargetPageId="2" captionText="FIGURE 1. Cymonomus windsorae n. sp.: A, D, E, USNM 1546450 (= ULLZ 12698A) ovigerous female holotype; B, C, USNM 1705285 (= ULLZ 12698B) ovigerous female paratype.A, carapace and eyestalks, dorsal; B, anterior carapace, eyestalks, antennules, and antennae, dorsal; C, carapace, extended pleon, and pleotelson, dorsal; D, left third maxilliped, external; E, anterior thorax, proximal articles of third maxillipeds, first pereopod coxae, and anterior sternum, ventral. Scale bars = 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.12190008" httpUri="https://zenodo.org/record/12190008/files/figure.png" pageId="1" pageNumber="262">Figs 1A–E</figureCitation>
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,
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<figureCitation id="4D852A6BFFB1FFD7FEABFA3B8832FA61" box="[302,372,1447,1473]" captionStart="FIGURE 2" captionStartId="3.[151,250,1581,1605]" captionTargetBox="[151,1436,1065,1557]" captionTargetId="figure-339@3.[151,1436,1065,1557]" captionTargetPageId="3" captionText="FIGURE 2. Cymonomus windsorae n. sp.: A, USNM 1546450 (= ULLZ 12698A) preserved ovigerous female holotype, cw 3.9 mm, northwestern Gulf of Mexico; B–D, USNM 1705285 (= ULLZ 12698B) preserved ovigerous female paratype, cw 4.0 mm, northwestern Gulf of Mexico." figureDoi="http://doi.org/10.5281/zenodo.12190010" httpUri="https://zenodo.org/record/12190010/files/figure.png" pageId="1" pageNumber="262">2A–D</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="9DA46565FFB1FFD7FF12FA728B74F9D5" pageId="1" pageNumber="262" type="materials_examined">
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<paragraph id="D50136EEFFB1FFD7FF12FA728B74F9D5" blockId="1.[151,1437,1518,2013]" pageId="1" pageNumber="262">
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<emphasis id="E7CAEAFCFFB1FFD7FF12FA7288C4F9A8" bold="true" box="[151,386,1518,1544]" pageId="1" pageNumber="262">Material examined.</emphasis>
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<materialsCitation id="65D63CB3FFB1FFD7FE0FFA738831F9F1" ID-GBIF-Occurrence="4898005302" collectingDate="1983-11-27" collectionCode="USNM, ULLZ" collectorName="Cruise I" elevation="850" latitude="27.8" location="northwestern Gulf" longLatPrecision="929" longitude="-90.055" pageId="1" pageNumber="262" specimenCode="USNM 1546450, ULLZ 12698" specimenCount="1" stateProvince="Louisiana" typeStatus="holotype">
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<typeStatus id="0A05884CFFB1FFD7FE0FFA7388B1F9A9" box="[394,503,1519,1545]" pageId="1" pageNumber="262" type="holotype">Holotype</typeStatus>
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: Ovigerous female, cw
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<quantity id="12469B0BFFB1FFD7FC80FA738A18F9A9" box="[773,862,1519,1545]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="1" pageNumber="262" unit="mm" value="3.9">3.9 mm</quantity>
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, pcl
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<quantity id="12469B0BFFB1FFD7FC13FA728AA9F9A9" box="[918,1007,1518,1545]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="262" unit="mm" value="3.5">3.5 mm</quantity>
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,
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<specimenCode id="85189E95FFB1FFD7FC79FA738DFEF9A9" box="[1020,1208,1518,1545]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="262" type="Museum">USNM 1546450</specimenCode>
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(=
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<specimenCode id="85189E95FFB1FFD7FB67FA738C3DF9A9" box="[1250,1403,1519,1545]" collectionCode="ULLZ" country="USA" httpUri="http://grbio.org/cool/ggzt-j23h" name="University of Louisiana at Layafette, Zoological Collection" pageId="1" pageNumber="262">ULLZ 12698</specimenCode>
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A), trawl sample, LGL Stn 3,
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<collectorName id="784B5338FFB1FFD7FE48F98F8B6AF98D" box="[461,556,1555,1581]" pageId="1" pageNumber="262">Cruise I</collectorName>
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||
, off
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<collectingRegion id="177AF80CFFB1FFD7FDE2F98F8B9EF98D" box="[615,728,1555,1581]" country="United States of America" name="Louisiana" pageId="1" pageNumber="262">Louisiana</collectingRegion>
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,
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<location id="D0616035FFB1FFD7FD6DF98F8D7BF98D" LSID="urn:lsid:plazi:treatment:5D1787F8FFB1FFD2FF12FA1F886FF835:D0616035FFB1FFD7FD6DF98F8D7BF98D" box="[744,1085,1555,1581]" latitude="27.8" longLatPrecision="929" longitude="-90.055" name="northwestern Gulf" pageId="1" pageNumber="262" stateProvince="Louisiana">northwestern Gulf of Mexico</location>
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,
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<geoCoordinate id="B08A5029FFB1FFD7FBC8F98F8DF6F98D" box="[1101,1200,1555,1581]" degrees="27" direction="north" minutes="48" orientation="latitude" pageId="1" pageNumber="262" precision="925" value="27.8">27°48’N</geoCoordinate>
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,
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<geoCoordinate id="B08A5029FFB1FFD7FB3AF98F8C7DF98D" box="[1215,1339,1555,1581]" degrees="90" direction="west" minutes="03.3" orientation="longitude" pageId="1" pageNumber="262" precision="92" value="-90.055">90°03.3’W</geoCoordinate>
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,
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<quantity id="12469B0BFFB1FFD7FACEF98E8CDEF98D" box="[1355,1432,1554,1581]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="1" pageNumber="262" unit="m" value="850.0">
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<elevation id="5E93D1DDFFB1FFD7FACEF98E8CDEF98D" box="[1355,1432,1554,1581]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="1" pageNumber="262" unit="m" value="850.0">850 m</elevation>
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</quantity>
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,
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<date id="A100102EFFB1FFD7FF12F9AB8831F9F1" box="[151,375,1591,1617]" pageId="1" pageNumber="262" value="1983-11-27">
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<collectingDate id="B144E9C6FFB1FFD7FF12F9AB8831F9F1" box="[151,375,1591,1617]" pageId="1" pageNumber="262" value="1983-11-27">27 November 1983</collectingDate>
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</date>
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</materialsCitation>
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||
.
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<materialsCitation id="65D63CB3FFB1FFD7FE00F9AB8B69F9D5" ID-GBIF-Occurrence="4898005301" collectingDate="1983-11-27" collectionCode="USNM, ULLZ" collectorName="Cruise I" elevation="850" latitude="27.8" location="northwestern Gulf" longLatPrecision="929" longitude="-90.055" pageId="1" pageNumber="262" specimenCode="USNM 1705285, ULLZ 12698" specimenCount="1" stateProvince="Louisiana" typeStatus="paratype">
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<typeStatus id="0A05884CFFB1FFD7FE00F9AB88ADF9F1" box="[389,491,1591,1617]" pageId="1" pageNumber="262" type="paratype">Paratype</typeStatus>
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: Ovigerous female, cw 4.0 mm, pcl
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<quantity id="12469B0BFFB1FFD7FC14F9AB8AAAF9F1" box="[913,1004,1591,1617]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="1" pageNumber="262" unit="mm" value="3.7">3.7 mm</quantity>
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||
,
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<specimenCode id="85189E95FFB1FFD7FC7FF9AB8DF0F9F1" box="[1018,1206,1590,1617]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="262" type="Museum">USNM 1705285</specimenCode>
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||
(=
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<specimenCode id="85189E95FFB1FFD7FB67F9AB8C3BF9F1" box="[1250,1405,1591,1617]" collectionCode="ULLZ" country="USA" httpUri="http://grbio.org/cool/ggzt-j23h" name="University of Louisiana at Layafette, Zoological Collection" pageId="1" pageNumber="262">ULLZ 12698</specimenCode>
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B), collection data same as for holotype
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</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="9DA46565FFB1FFD7FF42F9E28A15F889" pageId="1" pageNumber="262" type="diagnosis">
|
||
<paragraph id="D50136EEFFB1FFD7FF42F9E28A15F889" blockId="1.[151,1437,1518,2013]" pageId="1" pageNumber="262">
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||
<emphasis id="E7CAEAFCFFB1FFD7FF42F9E28804F938" bold="true" box="[199,322,1662,1688]" pageId="1" pageNumber="262">Diagnosis.</emphasis>
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Carapace regions well marked, irregularly granulate; broad gastric region delimited laterally by distinct arched furrow. Eyestalks elongate, untapered, subcylindrical with slight dorsoventral flattening proximally, directed anteriorly, nearly straight, freely articulated at base, no evident fusion to one another or frontal margin of carapace. Cornea translucent, unfaceted, no remaining evidence of pigment, distinctly bulbous, separation from adjacent heavily sclerotized eyestalk cuticle distinctly evident.
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</paragraph>
|
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</subSubSection>
|
||
<subSubSection id="9DA46565FFB1FFD5FF42F8AE89BDFBB1" lastPageId="3" lastPageNumber="264" pageId="1" pageNumber="262" type="description">
|
||
<paragraph id="D50136EEFFB1FFD4FF42F8AE8B48FDC1" blockId="1.[151,1437,1518,2013]" lastBlockId="2.[151,1437,151,609]" lastPageId="2" lastPageNumber="263" pageId="1" pageNumber="262">
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<emphasis id="E7CAEAFCFFB1FFD7FF42F8AE881FF8EC" bold="true" box="[199,345,1842,1868]" pageId="1" pageNumber="262">Description.</emphasis>
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Carapace outline subquadrate, lateral margins sinuous, subparallel, widest posteriorly at lateral margin of slightly inflated branchial region, irregularly granulate anterolateral margin slightly extended mesially above base of outer orbital process (
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<figureCitation id="4D852A6BFFB1FFD7FDC6F8E78B82F835" box="[579,708,1915,1941]" captionStart="FIGURE 1" captionStartId="2.[151,250,1685,1709]" captionTargetBox="[159,1430,639,1653]" captionTargetId="figure-211@2.[151,1436,630,1659]" captionTargetPageId="2" captionText="FIGURE 1. Cymonomus windsorae n. sp.: A, D, E, USNM 1546450 (= ULLZ 12698A) ovigerous female holotype; B, C, USNM 1705285 (= ULLZ 12698B) ovigerous female paratype.A, carapace and eyestalks, dorsal; B, anterior carapace, eyestalks, antennules, and antennae, dorsal; C, carapace, extended pleon, and pleotelson, dorsal; D, left third maxilliped, external; E, anterior thorax, proximal articles of third maxillipeds, first pereopod coxae, and anterior sternum, ventral. Scale bars = 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.12190008" httpUri="https://zenodo.org/record/12190008/files/figure.png" pageId="1" pageNumber="262">Figs 1A–C</figureCitation>
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,
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<figureCitation id="4D852A6BFFB1FFD7FD51F8E78A5BF835" box="[724,797,1915,1941]" captionStart="FIGURE 2" captionStartId="3.[151,250,1581,1605]" captionTargetBox="[151,1436,1065,1557]" captionTargetId="figure-339@3.[151,1436,1065,1557]" captionTargetPageId="3" captionText="FIGURE 2. Cymonomus windsorae n. sp.: A, USNM 1546450 (= ULLZ 12698A) preserved ovigerous female holotype, cw 3.9 mm, northwestern Gulf of Mexico; B–D, USNM 1705285 (= ULLZ 12698B) preserved ovigerous female paratype, cw 4.0 mm, northwestern Gulf of Mexico." figureDoi="http://doi.org/10.5281/zenodo.12190010" httpUri="https://zenodo.org/record/12190010/files/figure.png" pageId="1" pageNumber="262">2A–D</figureCitation>
|
||
). Carapace regions dorsally indicated, shallow cervical groove evident; hepatic region with lateral supramarginal crest bearing irregular coarse granules and small denticles; broad gastric region slightly elevated anteriorly above level hepatic regions, delimited laterally by distinct arched furrow, frontal and fronto-orbital margins upturned, posteriorly with distinct pair of small gastric pits near median line; pattern of four shallow depressions framing intestinal region, two near posterolateral limits of gastric region, two near anterolateral limits of cardiac region; cardiac region distinctly inflated, elevated above thicken posterior margin of carapace; surfaces overall with well-separated small granules and weak rugae marking regional furrows, cervical groove, branchial grooves, also as irregular patterns on hepatic region, branchial region, and near carapacial margins. Anterolateral spine low, anteriorly directed, slightly enlarged among other acute to subacute marginal granules of anterolateral margin. Fronto-orbital margin reaching anteriorly little if any beyond anterolateral margins; width about half that of anterior carapace; outer orbital process narrowly digitiform, positioned below plane of rostrum, dorsoventrally thickened by raised granulate margins proximally, mesial margin continued from upturned fronto-orbital margin, raised margins obsolete distally. Rostrum (fractured from carapace at base, appearing crushed terminally) narrowly elongate, extending anteriorly at least as far as eyestalk corneas, lacking subrostral tubercle; rostral margins parallel at sides, divergent at base. Carapace ventrally with pterygostomian region swollen, finely granulate, forming elongate boss.
|
||
</paragraph>
|
||
<caption id="81C16666FFB2FFD4FF12F9098CDAF8B9" ID-DOI="http://doi.org/10.5281/zenodo.12190008" ID-Zenodo-Dep="12190008" httpUri="https://zenodo.org/record/12190008/files/figure.png" pageId="2" pageNumber="263" startId="2.[151,250,1685,1709]" targetBox="[159,1430,639,1653]" targetPageId="2" targetType="figure">
|
||
<paragraph id="D50136EEFFB2FFD4FF12F9098CDAF8B9" blockId="2.[151,1436,1685,1817]" pageId="2" pageNumber="263">
|
||
<emphasis id="E7CAEAFCFFB2FFD4FF12F9098850F90D" bold="true" box="[151,278,1685,1709]" pageId="2" pageNumber="263">FIGURE 1.</emphasis>
|
||
<taxonomicName id="12BE4D6DFFB2FFD4FE9BF90A8B48F90D" authority="Felder & Tavares, 2024" authorityName="Felder & Tavares" authorityYear="2024" box="[286,526,1685,1709]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="windsorae" status="sp. nov.">
|
||
<emphasis id="E7CAEAFCFFB2FFD4FE9BF90A8B48F90D" box="[286,526,1685,1709]" italics="true" pageId="2" pageNumber="263">Cymonomus windsorae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E7CAEAFCFFB2FFD4FD92F9098B15F90D" bold="true" box="[535,595,1685,1709]" pageId="2" pageNumber="263">
|
||
<taxonomicNameLabel id="FCF95787FFB2FFD4FD92F9098B15F90D" box="[535,595,1685,1709]" pageId="2" pageNumber="263" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: A, D, E, USNM 1546450 (= ULLZ 12698A) ovigerous female holotype; B, C, USNM 1705285 (= ULLZ 12698B) ovigerous female paratype.A, carapace and eyestalks, dorsal; B, anterior carapace, eyestalks, antennules, and antennae, dorsal; C, carapace, extended pleon, and pleotelson, dorsal; D, left third maxilliped, external; E, anterior thorax, proximal articles of third maxillipeds, first pereopod coxae, and anterior sternum, ventral. Scale bars = 1.0 mm
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D50136EEFFB2FFD5FF42F8F789B7FF75" blockId="2.[151,1436,1899,2034]" lastBlockId="3.[151,1437,151,1041]" lastPageId="3" lastPageNumber="264" pageId="2" pageNumber="263">
|
||
Eyestalks narrow, elongate, subcylindrical with slight dorsoventral flattening proximally, directed anteriorly, nearly straight, freely articulated at base, no evident fusion to one another or frontal margin of carapace (
|
||
<figureCitation id="4D852A6BFFB2FFD4FA9EF8138CDEF80A" box="[1307,1432,1935,1962]" captionStart="FIGURE 1" captionStartId="2.[151,250,1685,1709]" captionTargetBox="[159,1430,639,1653]" captionTargetId="figure-211@2.[151,1436,630,1659]" captionTargetPageId="2" captionText="FIGURE 1. Cymonomus windsorae n. sp.: A, D, E, USNM 1546450 (= ULLZ 12698A) ovigerous female holotype; B, C, USNM 1705285 (= ULLZ 12698B) ovigerous female paratype.A, carapace and eyestalks, dorsal; B, anterior carapace, eyestalks, antennules, and antennae, dorsal; C, carapace, extended pleon, and pleotelson, dorsal; D, left third maxilliped, external; E, anterior thorax, proximal articles of third maxillipeds, first pereopod coxae, and anterior sternum, ventral. Scale bars = 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.12190008" httpUri="https://zenodo.org/record/12190008/files/figure.png" pageId="2" pageNumber="263">Figs 1A, B</figureCitation>
|
||
,
|
||
<figureCitation id="4D852A6BFFB2FFD4FF12F82889BBF86E" box="[151,253,1972,1998]" captionStart="FIGURE 2" captionStartId="3.[151,250,1581,1605]" captionTargetBox="[151,1436,1065,1557]" captionTargetId="figure-339@3.[151,1436,1065,1557]" captionTargetPageId="3" captionText="FIGURE 2. Cymonomus windsorae n. sp.: A, USNM 1546450 (= ULLZ 12698A) preserved ovigerous female holotype, cw 3.9 mm, northwestern Gulf of Mexico; B–D, USNM 1705285 (= ULLZ 12698B) preserved ovigerous female paratype, cw 4.0 mm, northwestern Gulf of Mexico." figureDoi="http://doi.org/10.5281/zenodo.12190010" httpUri="https://zenodo.org/record/12190010/files/figure.png" pageId="2" pageNumber="263">2A, B, D</figureCitation>
|
||
); cornea reaching to just beyond midlength of antennular peduncle first article; dorsal surface of stalk with scattered granules, granules denser, stronger along mesial margin; cornea well defined, translucent, unfaceted, no remaining evidence of pigment, terminally inflated, separation from adjacent heavily sclerotized cuticle distinctly evident.
|
||
</paragraph>
|
||
<paragraph id="D50136EEFFB3FFD5FF42FF4388FCFEE1" blockId="3.[151,1437,151,1041]" pageId="3" pageNumber="264">Epistome surface produced anteriorly below rostral base, bearing several small, elongate granules, one centered on midline, transverse row of four enlarged granules between bases of antennules; flattened subtriangular lobe mesial to base of antenna.</paragraph>
|
||
<paragraph id="D50136EEFFB3FFD5FF42FED78BE0FE0D" blockId="3.[151,1437,151,1041]" pageId="3" pageNumber="264">
|
||
Antennular peduncle length (
|
||
<typeStatus id="0A05884CFFB3FFD5FD88FED78B37FEC5" box="[525,625,331,357]" pageId="3" pageNumber="264" type="paratype">paratype</typeStatus>
|
||
) about 0.82 carapace width (
|
||
<figureCitation id="4D852A6BFFB3FFD5FC30FED78D4AFEC5" box="[949,1036,331,357]" captionStart="FIGURE 1" captionStartId="2.[151,250,1685,1709]" captionTargetBox="[159,1430,639,1653]" captionTargetId="figure-211@2.[151,1436,630,1659]" captionTargetPageId="2" captionText="FIGURE 1. Cymonomus windsorae n. sp.: A, D, E, USNM 1546450 (= ULLZ 12698A) ovigerous female holotype; B, C, USNM 1705285 (= ULLZ 12698B) ovigerous female paratype.A, carapace and eyestalks, dorsal; B, anterior carapace, eyestalks, antennules, and antennae, dorsal; C, carapace, extended pleon, and pleotelson, dorsal; D, left third maxilliped, external; E, anterior thorax, proximal articles of third maxillipeds, first pereopod coxae, and anterior sternum, ventral. Scale bars = 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.12190008" httpUri="https://zenodo.org/record/12190008/files/figure.png" pageId="3" pageNumber="264">Fig. 1B</figureCitation>
|
||
); first article granulate dorsally and laterally; second and third articles mostly smooth. Antennal peduncle first to third articles minutely granulate; fourth and fifth articles minutely granulate to smooth.
|
||
</paragraph>
|
||
<paragraph id="D50136EEFFB3FFD5FF42FE2B8D90FD09" blockId="3.[151,1437,151,1041]" pageId="3" pageNumber="264">
|
||
Third maxilliped ischiobasis subquadrate, with distinct longitudinal sublateral groove, mesial margin coarsely granulate, granules dentiform to spiniform distally (
|
||
<figureCitation id="4D852A6BFFB3FFD5FD5EFE478A7EFE55" box="[731,824,475,501]" captionStart="FIGURE 1" captionStartId="2.[151,250,1685,1709]" captionTargetBox="[159,1430,639,1653]" captionTargetId="figure-211@2.[151,1436,630,1659]" captionTargetPageId="2" captionText="FIGURE 1. Cymonomus windsorae n. sp.: A, D, E, USNM 1546450 (= ULLZ 12698A) ovigerous female holotype; B, C, USNM 1705285 (= ULLZ 12698B) ovigerous female paratype.A, carapace and eyestalks, dorsal; B, anterior carapace, eyestalks, antennules, and antennae, dorsal; C, carapace, extended pleon, and pleotelson, dorsal; D, left third maxilliped, external; E, anterior thorax, proximal articles of third maxillipeds, first pereopod coxae, and anterior sternum, ventral. Scale bars = 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.12190008" httpUri="https://zenodo.org/record/12190008/files/figure.png" pageId="3" pageNumber="264">Figs 1D</figureCitation>
|
||
,
|
||
<figureCitation id="4D852A6BFFB3FFD5FCC1FE478ACCFE55" box="[836,906,475,501]" captionStart="FIGURE 2" captionStartId="3.[151,250,1581,1605]" captionTargetBox="[151,1436,1065,1557]" captionTargetId="figure-339@3.[151,1436,1065,1557]" captionTargetPageId="3" captionText="FIGURE 2. Cymonomus windsorae n. sp.: A, USNM 1546450 (= ULLZ 12698A) preserved ovigerous female holotype, cw 3.9 mm, northwestern Gulf of Mexico; B–D, USNM 1705285 (= ULLZ 12698B) preserved ovigerous female paratype, cw 4.0 mm, northwestern Gulf of Mexico." figureDoi="http://doi.org/10.5281/zenodo.12190010" httpUri="https://zenodo.org/record/12190010/files/figure.png" pageId="3" pageNumber="264">2C, D</figureCitation>
|
||
); lateral margin mostly smooth, ending distally in sharp corner; fusion of ischium and basis marked by distinct groove. Merus length near equal that of ischium, elongate, weakly bent toward midline in distal half, terminally rounded; margins bearing a few dentiform granules and short, blunt spines; external surfaces with scattered denticles. Carpus heavy, subtriangular, with sparse marginal dentiform granules, propodus subcylindrical, dactylus subconiform, weakly arched, tapering to subacute tip. Exopod reaching to distal quarter of endopod merus; exopod flagellum well developed, multi-segmented.
|
||
</paragraph>
|
||
<paragraph id="D50136EEFFB3FFD5FF42FD2F8CDAFD6D" blockId="3.[151,1437,151,1041]" box="[199,1436,691,717]" pageId="3" pageNumber="264">
|
||
Except for coxae (
|
||
<figureCitation id="4D852A6BFFB3FFD5FE11FD2F8B57FD6D" box="[404,529,691,717]" captionStart="FIGURE 2" captionStartId="3.[151,250,1581,1605]" captionTargetBox="[151,1436,1065,1557]" captionTargetId="figure-339@3.[151,1436,1065,1557]" captionTargetPageId="3" captionText="FIGURE 2. Cymonomus windsorae n. sp.: A, USNM 1546450 (= ULLZ 12698A) preserved ovigerous female holotype, cw 3.9 mm, northwestern Gulf of Mexico; B–D, USNM 1705285 (= ULLZ 12698B) preserved ovigerous female paratype, cw 4.0 mm, northwestern Gulf of Mexico." figureDoi="http://doi.org/10.5281/zenodo.12190010" httpUri="https://zenodo.org/record/12190010/files/figure.png" pageId="3" pageNumber="264">Fig. 2A–D</figureCitation>
|
||
), chelipeds and other pereopods unknown, missing from
|
||
<typeStatus id="0A05884CFFB3FFD5FB14FD2F8DB5FD6D" box="[1169,1267,691,717]" pageId="3" pageNumber="264" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="0A05884CFFB3FFD5FAACFD2F8CDEFD6D" box="[1321,1432,691,717]" pageId="3" pageNumber="264" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D50136EEFFB3FFD5FF42FD4B8DCCFCFD" blockId="3.[151,1437,151,1041]" pageId="3" pageNumber="264">
|
||
Thoracic third sternite subpentagonal, thickened granulate anterolateral margins converging to anterior point at small terminal tubercle; thickened lateral margins slightly concave, weakly divergent posteriorly, posterior width about one and one-half times median length (
|
||
<figureCitation id="4D852A6BFFB3FFD5FD02FC838B98FC99" box="[647,734,799,825]" captionStart="FIGURE 1" captionStartId="2.[151,250,1685,1709]" captionTargetBox="[159,1430,639,1653]" captionTargetId="figure-211@2.[151,1436,630,1659]" captionTargetPageId="2" captionText="FIGURE 1. Cymonomus windsorae n. sp.: A, D, E, USNM 1546450 (= ULLZ 12698A) ovigerous female holotype; B, C, USNM 1705285 (= ULLZ 12698B) ovigerous female paratype.A, carapace and eyestalks, dorsal; B, anterior carapace, eyestalks, antennules, and antennae, dorsal; C, carapace, extended pleon, and pleotelson, dorsal; D, left third maxilliped, external; E, anterior thorax, proximal articles of third maxillipeds, first pereopod coxae, and anterior sternum, ventral. Scale bars = 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.12190008" httpUri="https://zenodo.org/record/12190008/files/figure.png" pageId="3" pageNumber="264">Figs 1E</figureCitation>
|
||
,
|
||
<figureCitation id="4D852A6BFFB3FFD5FD6CFC838A48FC99" box="[745,782,799,825]" captionStart="FIGURE 2" captionStartId="3.[151,250,1581,1605]" captionTargetBox="[151,1436,1065,1557]" captionTargetId="figure-339@3.[151,1436,1065,1557]" captionTargetPageId="3" captionText="FIGURE 2. Cymonomus windsorae n. sp.: A, USNM 1546450 (= ULLZ 12698A) preserved ovigerous female holotype, cw 3.9 mm, northwestern Gulf of Mexico; B–D, USNM 1705285 (= ULLZ 12698B) preserved ovigerous female paratype, cw 4.0 mm, northwestern Gulf of Mexico." figureDoi="http://doi.org/10.5281/zenodo.12190010" httpUri="https://zenodo.org/record/12190010/files/figure.png" pageId="3" pageNumber="264">2C</figureCitation>
|
||
). Fourth sternite lateral margin thickened, forming strongly raised coarsely granulate swollen lip along concavity accommodating first pereopod coxa.
|
||
</paragraph>
|
||
<paragraph id="D50136EEFFB3FFD5FF42FCFB89BDFBB1" blockId="3.[151,1437,151,1041]" pageId="3" pageNumber="264">
|
||
Pleon with raised median longitudinal crest on pleonites and pleotelson, otherwise faintly patterned by low anastomosed weakly granular elevations and shallow sulci (
|
||
<figureCitation id="4D852A6BFFB3FFD5FCBDFC178AD5FC05" box="[824,915,907,933]" captionStart="FIGURE 1" captionStartId="2.[151,250,1685,1709]" captionTargetBox="[159,1430,639,1653]" captionTargetId="figure-211@2.[151,1436,630,1659]" captionTargetPageId="2" captionText="FIGURE 1. Cymonomus windsorae n. sp.: A, D, E, USNM 1546450 (= ULLZ 12698A) ovigerous female holotype; B, C, USNM 1705285 (= ULLZ 12698B) ovigerous female paratype.A, carapace and eyestalks, dorsal; B, anterior carapace, eyestalks, antennules, and antennae, dorsal; C, carapace, extended pleon, and pleotelson, dorsal; D, left third maxilliped, external; E, anterior thorax, proximal articles of third maxillipeds, first pereopod coxae, and anterior sternum, ventral. Scale bars = 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.12190008" httpUri="https://zenodo.org/record/12190008/files/figure.png" pageId="3" pageNumber="264">Figs 1C</figureCitation>
|
||
,
|
||
<figureCitation id="4D852A6BFFB3FFD5FC25FC178AAFFC05" box="[928,1001,907,933]" captionStart="FIGURE 2" captionStartId="3.[151,250,1581,1605]" captionTargetBox="[151,1436,1065,1557]" captionTargetId="figure-339@3.[151,1436,1065,1557]" captionTargetPageId="3" captionText="FIGURE 2. Cymonomus windsorae n. sp.: A, USNM 1546450 (= ULLZ 12698A) preserved ovigerous female holotype, cw 3.9 mm, northwestern Gulf of Mexico; B–D, USNM 1705285 (= ULLZ 12698B) preserved ovigerous female paratype, cw 4.0 mm, northwestern Gulf of Mexico." figureDoi="http://doi.org/10.5281/zenodo.12190010" httpUri="https://zenodo.org/record/12190010/files/figure.png" pageId="3" pageNumber="264">2A–D</figureCitation>
|
||
); pattern of transverse sulci evident on especially second to fourth pleonites. Pleotelson subtriangular, width slightly less than twice length, sixth pleonite and telson completely fused with no evidence of separating suture along entire width; obtuse angle of terminus rounded.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="81C16666FFB3FFD5FF12F9B188BDF92D" ID-DOI="http://doi.org/10.5281/zenodo.12190010" ID-Zenodo-Dep="12190010" httpUri="https://zenodo.org/record/12190010/files/figure.png" pageId="3" pageNumber="264" startId="3.[151,250,1581,1605]" targetBox="[151,1436,1065,1557]" targetPageId="3" targetType="figure">
|
||
<paragraph id="D50136EEFFB3FFD5FF12F9B188BDF92D" blockId="3.[151,1437,1581,1677]" pageId="3" pageNumber="264">
|
||
<emphasis id="E7CAEAFCFFB3FFD5FF12F9B18848F9E5" bold="true" box="[151,270,1581,1605]" pageId="3" pageNumber="264">FIGURE 2</emphasis>
|
||
.
|
||
<taxonomicName id="12BE4D6DFFB3FFD5FE99F9B18B4DF9E5" authority="Felder & Tavares, 2024" authorityName="Felder & Tavares" authorityYear="2024" box="[284,523,1581,1605]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="windsorae" status="sp. nov.">
|
||
<emphasis id="E7CAEAFCFFB3FFD5FE99F9B18B4DF9E5" box="[284,523,1581,1605]" italics="true" pageId="3" pageNumber="264">Cymonomus windsorae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E7CAEAFCFFB3FFD5FD97F9B18B0BF9E5" bold="true" box="[530,589,1581,1605]" pageId="3" pageNumber="264">
|
||
<taxonomicNameLabel id="FCF95787FFB3FFD5FD97F9B18B0BF9E5" box="[530,589,1581,1605]" pageId="3" pageNumber="264" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: A, USNM 1546450 (= ULLZ 12698A) preserved ovigerous female holotype, cw 3.9 mm, northwestern Gulf of Mexico; B–D, USNM 1705285 (= ULLZ 12698B) preserved ovigerous female paratype, cw 4.0 mm, northwestern Gulf of Mexico.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="9DA46565FFB3FFD5FF42F9438A88F8BD" pageId="3" pageNumber="264" type="etymology">
|
||
<paragraph id="D50136EEFFB3FFD5FF42F9438A88F8BD" blockId="3.[151,1437,1759,2038]" pageId="3" pageNumber="264">
|
||
<emphasis id="E7CAEAFCFFB3FFD5FF42F9438808F959" bold="true" box="[199,334,1759,1785]" pageId="3" pageNumber="264">Etymology.</emphasis>
|
||
This species is named for our colleague, Amanda M. Windsor, in recognition of her substantial contributions to phylogenetic and systematic studies of brachyuran crabs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9DA46565FFB3FFD5FF42F8BB88A6F8C6" pageId="3" pageNumber="264" type="distribution">
|
||
<paragraph id="D50136EEFFB3FFD5FF42F8BB88A6F8C6" blockId="3.[151,1437,1759,2038]" pageId="3" pageNumber="264">
|
||
<emphasis id="E7CAEAFCFFB3FFD5FF42F8BB88B6F8E1" bold="true" box="[199,496,1831,1857]" pageId="3" pageNumber="264">Distribution and habitat.</emphasis>
|
||
Known at present from only the
|
||
<typeStatus id="0A05884CFFB3FFD5FCD8F8B48ACBF8E2" box="[861,909,1832,1858]" pageId="3" pageNumber="264">type</typeStatus>
|
||
locality, northwestern Gulf of
|
||
<collectingRegion id="177AF80CFFB3FFD5FB67F8BB8C7CF8E1" box="[1250,1338,1831,1857]" country="Mexico" name="Mexico" pageId="3" pageNumber="264">Mexico</collectingRegion>
|
||
, western Atlantic, at a depth of
|
||
<specimenCount id="C3B8FD67FFB3FFD5FE17F8D788A6F8C6" box="[402,480,1867,1894]" count="850" pageId="3" pageNumber="264" type="generic">
|
||
<quantity id="12469B0BFFB3FFD5FE17F8D7889AF8C6" box="[402,476,1867,1894]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="3" pageNumber="264" unit="m" value="850.0">850 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9DA46565FFB3FFD2FF42F8F3886FF835" lastPageId="4" lastPageNumber="265" pageId="3" pageNumber="264" type="discussion">
|
||
<paragraph id="D50136EEFFB3FFD2FF42F8F38D1FFF75" blockId="3.[151,1437,1759,2038]" lastBlockId="4.[151,1437,151,1941]" lastPageId="4" lastPageNumber="265" pageId="3" pageNumber="264">
|
||
<emphasis id="E7CAEAFCFFB3FFD5FF42F8F3887DF829" bold="true" box="[199,315,1903,1929]" pageId="3" pageNumber="264">Remarks.</emphasis>
|
||
Specimens herewith described as
|
||
<taxonomicName id="12BE4D6DFFB3FFD5FD54F8F38A2CF829" authority="Felder & Tavares, 2024" authorityName="Felder & Tavares" authorityYear="2024" box="[721,874,1903,1929]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="windsorae" status="n. sp.">
|
||
<emphasis id="E7CAEAFCFFB3FFD5FD54F8F38A2CF829" box="[721,874,1903,1929]" italics="true" pageId="3" pageNumber="264">C. windsorae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E7CAEAFCFFB3FFD5FCF0F8F38AFEF829" bold="true" box="[885,952,1903,1929]" pageId="3" pageNumber="264">
|
||
<taxonomicNameLabel id="FCF95787FFB3FFD5FCF0F8F38AFEF829" box="[885,952,1903,1929]" pageId="3" pageNumber="264" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
were originally referred to one of us for study (DLF) by Dr. Linda Pequegnat, a consultant to LGL Ecological Research Associates, Bryan,
|
||
<collectingRegion id="177AF80CFFB3FFD5FB70F8088C7CF80E" box="[1269,1338,1940,1966]" country="United States of America" name="Texas" pageId="3" pageNumber="264">Texas</collectingRegion>
|
||
. As both specimens lacked appendages and exhibited varied degrees of damage to the rostrum, description was long deferred in hopes of making additional collections that might include intact and perhaps male specimens, or even DNA sequence-quality representatives as methods progressed. However, despite repeated collection efforts at similar depths in the northwestern Gulf of
|
||
<collectingRegion id="177AF80CFFB4FFD2FD9AFF278B31FF75" box="[543,631,187,213]" country="Mexico" name="Mexico" pageId="4" pageNumber="265">Mexico</collectingRegion>
|
||
, no additional specimens have been found.
|
||
</paragraph>
|
||
<paragraph id="D50136EEFFB4FFD2FF42FF438BBEFDB9" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="265">
|
||
Both specimens are unfortunately now in a very poor state of preservation. Initially fractured, missing appendages, and not well preserved, they were among a few specimens that became dried in the course of the
|
||
<collectionCode id="B3AFAE2BFFB4FFD2FAD0FE9F8CDBFEBD" box="[1365,1437,259,285]" country="USA" httpUri="http://grbio.org/cool/ggzt-j23h" name="University of Louisiana at Layafette, Zoological Collection" pageId="4" pageNumber="265">ULLZ</collectionCode>
|
||
collections being shipped to and temporarily stored at (without routine curation during the covid pandemic) the
|
||
<collectionCode id="B3AFAE2BFFB4FFD2FF12FED789ADFEC5" box="[151,235,331,357]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="265" type="Museum">USNM</collectionCode>
|
||
, where they are now curated with other of the former
|
||
<collectionCode id="B3AFAE2BFFB4FFD2FCCDFED78AD6FEC5" box="[840,912,331,357]" country="USA" httpUri="http://grbio.org/cool/ggzt-j23h" name="University of Louisiana at Layafette, Zoological Collection" pageId="4" pageNumber="265">ULLZ</collectionCode>
|
||
holdings. However, most of the line drawings, specimen photographs, and morphological notes were fortunately prepared by DLF prior to the specimens’ drying. Having now been rehydrated by standard protocols and returned to alcohol, essential features at very least remain recognizable in the type materials. However, both the
|
||
<typeStatus id="0A05884CFFB4FFD2FD6BFE2B8A16FE71" box="[750,848,439,465]" pageId="4" pageNumber="265" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="0A05884CFFB4FFD2FC00FE2B8AA3FE71" box="[901,997,439,465]" pageId="4" pageNumber="265" type="paratype">paratype</typeStatus>
|
||
females of
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FBE3FE2B8DBDFE71" authority="Felder & Tavares, 2024" authorityName="Felder & Tavares" authorityYear="2024" box="[1126,1275,439,465]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="windsorae" status="n. sp.">
|
||
<emphasis id="E7CAEAFCFFB4FFD2FBE3FE2B8DBDFE71" box="[1126,1275,439,465]" italics="true" pageId="4" pageNumber="265">C. windsorae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E7CAEAFCFFB4FFD2FA84FE2B8C79FE71" bold="true" box="[1281,1343,439,465]" pageId="4" pageNumber="265">
|
||
<taxonomicNameLabel id="FCF95787FFB4FFD2FA84FE2B8C79FE71" box="[1281,1343,439,465]" pageId="4" pageNumber="265" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
initially carried eggs that have now somewhat shriveled and disintegrated. The
|
||
<typeStatus id="0A05884CFFB4FFD2FC31FE478D50FE55" box="[948,1046,475,501]" pageId="4" pageNumber="265" type="holotype">holotype</typeStatus>
|
||
bore fourteen and the
|
||
<typeStatus id="0A05884CFFB4FFD2FA91FE478C32FE55" box="[1300,1396,475,501]" pageId="4" pageNumber="265" type="paratype">paratype</typeStatus>
|
||
six relatively large eggs,
|
||
<quantity id="12469B0BFFB4FFD2FE0DFE638B49FDB9" box="[392,527,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="4" pageNumber="265" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">0.9–1.1 mm</quantity>
|
||
in greatest diameter.
|
||
</paragraph>
|
||
<paragraph id="D50136EEFFB4FFD2FF42FDBF8B68FCFD" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="265">
|
||
Assignment of
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FEF7FDBF8B4EFD9D" authority="Felder & Tavares, 2024" authorityName="Felder & Tavares" authorityYear="2024" box="[370,520,547,573]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="windsorae" status="n. sp.">
|
||
<emphasis id="E7CAEAFCFFB4FFD2FEF7FDBF8B4EFD9D" box="[370,520,547,573]" italics="true" pageId="4" pageNumber="265">C. windsorae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E7CAEAFCFFB4FFD2FD8AFDBF8B0BFD9D" bold="true" box="[527,589,547,573]" pageId="4" pageNumber="265">
|
||
<taxonomicNameLabel id="FCF95787FFB4FFD2FD8AFDBF8B0BFD9D" box="[527,589,547,573]" pageId="4" pageNumber="265" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
to the genus
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FD66FDBF8A2BFD9D" authorityName="A. Milne-Edwards" authorityYear="1880" box="[739,877,547,573]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E7CAEAFCFFB4FFD2FD66FDBF8A2BFD9D" box="[739,877,547,573]" italics="true" pageId="4" pageNumber="265">Cymonomus</emphasis>
|
||
</taxonomicName>
|
||
is based upon a combination of characters, which include ocular peduncles not covered by the long slender rostrum, so that the peduncles are fully exposed in dorsal view; absence of an interantennular septum; third maxilliped merus remarkably elongated distally with the palp articulating at its subdistal inner surface and exopod with well-developed flagellum; and pleon of five pleonites and pleotelson without any trace of suture separating the sixth pleonite from the telson. This is in contrast to members of the closely related genus
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FE55FD4B8B3BFD51" authorityName="Tavares" authorityYear="1993" box="[464,637,727,753]" class="Malacostraca" family="Cymonomidae" genus="Cymonomoides" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E7CAEAFCFFB4FFD2FE55FD4B8B3BFD51" box="[464,637,727,753]" italics="true" pageId="4" pageNumber="265">Cymonomoides</emphasis>
|
||
</taxonomicName>
|
||
, in which the rostrum is vestigial or very short and all six pleonites and the telson are freely articulated, thus facilitating ready separation of
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FC2BFD678DF1FCB5" authority="Felder & Tavares, 2024" authorityName="Felder & Tavares" authorityYear="2024" box="[942,1207,763,789]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="windsorae" status="sp. nov.">
|
||
<emphasis id="E7CAEAFCFFB4FFD2FC2BFD678DF1FCB5" box="[942,1207,763,789]" italics="true" pageId="4" pageNumber="265">Cymonomus windsorae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E7CAEAFCFFB4FFD2FB45FD678C47FCB5" bold="true" box="[1216,1281,763,789]" pageId="4" pageNumber="265">
|
||
<taxonomicNameLabel id="FCF95787FFB4FFD2FB45FD678C47FCB5" box="[1216,1281,763,789]" pageId="4" pageNumber="265" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from species of
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FF3CFC838820FC99" authorityName="Tavares" authorityYear="1993" box="[185,358,799,825]" class="Malacostraca" family="Cymonomidae" genus="Cymonomoides" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E7CAEAFCFFB4FFD2FF3CFC838820FC99" box="[185,358,799,825]" italics="true" pageId="4" pageNumber="265">Cymonomoides</emphasis>
|
||
</taxonomicName>
|
||
, also known to occur in the Gulf of
|
||
<collectingRegion id="177AF80CFFB4FFD2FC92FC838A2BFC99" box="[791,877,799,825]" country="Mexico" name="Mexico" pageId="4" pageNumber="265">Mexico</collectingRegion>
|
||
and adjacent waters (Tavares 1993;
|
||
<bibRefCitation id="B12F4B1FFFB4FFD2FA92FC83880BFCFD" author="Lemaitre, R. & Bermudez, A." pageId="4" pageNumber="265" pagination="974 - 979" refId="ref4216" refString="Lemaitre, R. & Bermudez, A. (2000) A new cyclodorippoid crab of the genus Cymonomoides Tavares, 1993 (Crustacea: Decapoda: Brachyura: Cymonomidae) from the Caribbean coast of Colombia. Proceedings of the Biological Society of Washington, 113 (4), 974 - 979." type="journal article" year="2000">Lemaitre & Bermúdez 2000</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B12F4B1FFFB4FFD2FEDDFCDF8B65FCFD" author="Felder, D. L. & Alvarez, F. & Goy, J. W. & Lemaitre, R." box="[344,547,835,861]" pageId="4" pageNumber="265" refId="ref4117" refString="Felder, D. L., Alvarez, F., Goy, J. W. & Lemaitre, R. (2009) Chapter 59: Decapoda (Crustacea) of the Gulf of Mexico, with comments on the Amphionidaceain. In: Felder, D. L. & Camp, D. K. (Eds.), Gulf of Mexico Origin, Waters, and Biota. Vol. 1. Biodiversity. Texas A & M University Press, College Station, Texas, pp. 1019 - 1104." type="book" year="2009">
|
||
Felder
|
||
<emphasis id="E7CAEAFCFFB4FFD2FE22FCDF889EFCFD" box="[423,472,835,861]" italics="true" pageId="4" pageNumber="265">et al</emphasis>
|
||
. 2009
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D50136EEFFB4FFD2FF42FCFB8B02FAAD" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="265">
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FF42FCFB8896FC21" authority="Felder & Tavares, 2024" authorityName="Felder & Tavares" authorityYear="2024" box="[199,464,871,897]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="windsorae" status="sp. nov.">
|
||
<emphasis id="E7CAEAFCFFB4FFD2FF42FCFB8896FC21" box="[199,464,871,897]" italics="true" pageId="4" pageNumber="265">Cymonomus windsorae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E7CAEAFCFFB4FFD2FE5CFCFB8B5CFC21" bold="true" box="[473,538,871,897]" pageId="4" pageNumber="265">
|
||
<taxonomicNameLabel id="FCF95787FFB4FFD2FE5CFCFB8B5CFC21" box="[473,538,871,897]" pageId="4" pageNumber="265" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be readily distinguished from all other species in the genus by having an untapered rostrum with rostral margins parallel at the sides and divergent at the base and perhaps distally (though damaged there), and its unique eye morphology with well-developed bulbous terminal corneas of the nearly straight eyestalks, the smooth integumental covering of which is clearly demarcated from that of the eyestalk itself. It superficially resembles the Mediterranean
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FDFCFC6B8AC3FBB1" baseAuthorityName="Norman in Thomson" baseAuthorityYear="1873" box="[633,901,1015,1041]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="granulatus">
|
||
<emphasis id="E7CAEAFCFFB4FFD2FDFCFC6B8AC3FBB1" box="[633,901,1015,1041]" italics="true" pageId="4" pageNumber="265">Cymonomus granulatus</emphasis>
|
||
</taxonomicName>
|
||
(Norman
|
||
<emphasis id="E7CAEAFCFFB4FFD2FC7EFC6B8D57FBB0" box="[1019,1041,1015,1040]" italics="true" pageId="4" pageNumber="265">in</emphasis>
|
||
<bibRefCitation id="B12F4B1FFFB4FFD2FB9CFC6B8C7FFBB1" author="Wyville Thompson, C." box="[1049,1337,1015,1041]" pageId="4" pageNumber="265" refId="ref4834" refString="Wyville Thompson, C. (1873) The depths of the sea. An account of the general results of the dredging cruises of the H. M. SS. ' Porcupine' and ' Lightning' during the Summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F. R. S., J. Gwyn Jeffreys, F. R. S. and Dr. Wyville Thomson, F. R. S. Macmillan and Co., London, xxi + 527 pp., VIII pls. https: // doi. org / 10.5962 / bhl. title. 16330" type="book" year="1873">Wyville Thompson, 1873</bibRefCitation>
|
||
) and the Tasmanian
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FE9DFB878BF4FB95" authority="Ahyong, 2019" authorityName="Ahyong" authorityYear="2019" box="[280,690,1051,1077]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="karenae">
|
||
<emphasis id="E7CAEAFCFFB4FFD2FE9DFB878B42FB95" box="[280,516,1051,1077]" italics="true" pageId="4" pageNumber="265">Cymonomus karenae</emphasis>
|
||
<bibRefCitation id="B12F4B1FFFB4FFD2FD88FB878BF4FB95" author="Ahyong, S. T." box="[525,690,1051,1077]" pageId="4" pageNumber="265" pagination="33 - 69" refId="ref3623" refString="Ahyong, S. T. (2019) The cymonomid crabs of New Zealand and Australia (Crustacea: Brachyura: Cyclodorripoida). Records of the Australian Museum, 71 (2), 33 - 69. https: // doi. org / 10.3853 / j. 2201 - 4349.71.2019.1682" type="journal article" year="2019">Ahyong, 2019</bibRefCitation>
|
||
</taxonomicName>
|
||
, in having the transparent cornea similarly demarcated, but in the latter two species the cornea is less bulbous and in
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FD55FBA38A0CFBF9" authorityName="Ahyong" authorityYear="2019" box="[720,842,1087,1113]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="karenae">
|
||
<emphasis id="E7CAEAFCFFB4FFD2FD55FBA38A0CFBF9" box="[720,842,1087,1113]" italics="true" pageId="4" pageNumber="265">C. karenae</emphasis>
|
||
</taxonomicName>
|
||
, it is centered on the distoventral terminus of a stout, strongly divergent stalk (
|
||
<bibRefCitation id="B12F4B1FFFB4FFD2FE2EFBFF8B34FBDD" author="Ahyong, S. T." box="[427,626,1122,1149]" pageId="4" pageNumber="265" pagination="33 - 69" refId="ref3623" refString="Ahyong, S. T. (2019) The cymonomid crabs of New Zealand and Australia (Crustacea: Brachyura: Cyclodorripoida). Records of the Australian Museum, 71 (2), 33 - 69. https: // doi. org / 10.3853 / j. 2201 - 4349.71.2019.1682" type="journal article" year="2019">Ahyong 2019: 58</bibRefCitation>
|
||
, fig. 22C, D). Additionally, the two aforementioned species can be easily set apart from
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FEBFFB1B8896FB01" authority="Felder & Tavares, 2024" authorityName="Felder & Tavares" authorityYear="2024" box="[314,464,1159,1185]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="windsorae" status="n. sp.">
|
||
<emphasis id="E7CAEAFCFFB4FFD2FEBFFB1B8896FB01" box="[314,464,1159,1185]" italics="true" pageId="4" pageNumber="265">C. windsorae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E7CAEAFCFFB4FFD2FE52FB1B8B50FB01" bold="true" box="[471,534,1159,1185]" pageId="4" pageNumber="265">
|
||
<taxonomicNameLabel id="FCF95787FFB4FFD2FE52FB1B8B50FB01" box="[471,534,1159,1185]" pageId="4" pageNumber="265" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the ornamentation of the dorsal and lateral surfaces of the carapace, which is densely covered with prominent spination or coarse granulation in
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FCFBFB378D50FB65" baseAuthorityName="Norman in Thomson" baseAuthorityYear="1873" box="[894,1046,1195,1221]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="granulatus">
|
||
<emphasis id="E7CAEAFCFFB4FFD2FCFBFB378D50FB65" box="[894,1046,1195,1221]" italics="true" pageId="4" pageNumber="265">C. granulatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FBC9FB378D83FB65" authorityName="Ahyong" authorityYear="2019" box="[1100,1221,1195,1221]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="karenae">
|
||
<emphasis id="E7CAEAFCFFB4FFD2FBC9FB378D83FB65" box="[1100,1221,1195,1221]" italics="true" pageId="4" pageNumber="265">C. karenae</emphasis>
|
||
</taxonomicName>
|
||
, respectively. In
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FA06FB37884AFB49" authority="Felder & Tavares, 2024" authorityName="Felder & Tavares" authorityYear="2024" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="windsorae" status="n. sp.">
|
||
<emphasis id="E7CAEAFCFFB4FFD2FA06FB37884AFB49" italics="true" pageId="4" pageNumber="265">C. windsorae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E7CAEAFCFFB4FFD2FE96FB538815FB49" bold="true" box="[275,339,1231,1257]" pageId="4" pageNumber="265">
|
||
<taxonomicNameLabel id="FCF95787FFB4FFD2FE96FB538815FB49" box="[275,339,1231,1257]" pageId="4" pageNumber="265" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
it is instead irregularly granulate with surfaces overall bearing well-separated small granules and weak rugae marking regional furrows.
|
||
</paragraph>
|
||
<paragraph id="D50136EEFFB4FFD2FF42FA8B8AF3F9A9" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="265">
|
||
Among the three known species of
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FDD1FA8B8B98FA91" authorityName="A. Milne-Edwards" authorityYear="1880" box="[596,734,1303,1329]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E7CAEAFCFFB4FFD2FDD1FA8B8B98FA91" box="[596,734,1303,1329]" italics="true" pageId="4" pageNumber="265">Cymonomus</emphasis>
|
||
</taxonomicName>
|
||
that range into at least eastern extremes of the Gulf of
|
||
<collectingRegion id="177AF80CFFB4FFD2FAC2FA8B8CDBFA91" box="[1351,1437,1303,1329]" country="Mexico" name="Mexico" pageId="4" pageNumber="265">Mexico</collectingRegion>
|
||
(
|
||
<bibRefCitation id="B12F4B1FFFB4FFD2FF1AFAA7882BFAF5" author="Felder, D. L. & Alvarez, F. & Goy, J. W. & Lemaitre, R." box="[159,365,1339,1365]" pageId="4" pageNumber="265" refId="ref4117" refString="Felder, D. L., Alvarez, F., Goy, J. W. & Lemaitre, R. (2009) Chapter 59: Decapoda (Crustacea) of the Gulf of Mexico, with comments on the Amphionidaceain. In: Felder, D. L. & Camp, D. K. (Eds.), Gulf of Mexico Origin, Waters, and Biota. Vol. 1. Biodiversity. Texas A & M University Press, College Station, Texas, pp. 1019 - 1104." type="book" year="2009">
|
||
Felder
|
||
<emphasis id="E7CAEAFCFFB4FFD2FF75FAA7886CFAF5" box="[240,298,1339,1365]" italics="true" pageId="4" pageNumber="265">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
), the cornea and eyestalks readily separate these species from
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FBB4FAA78D8EFAF5" authority="Felder & Tavares, 2024" authorityName="Felder & Tavares" authorityYear="2024" box="[1073,1224,1339,1365]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="windsorae" status="n. sp.">
|
||
<emphasis id="E7CAEAFCFFB4FFD2FBB4FAA78D8EFAF5" box="[1073,1224,1339,1365]" italics="true" pageId="4" pageNumber="265">C. windsorae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E7CAEAFCFFB4FFD2FB55FAA78C56FAF5" bold="true" box="[1232,1296,1339,1365]" pageId="4" pageNumber="265">
|
||
<taxonomicNameLabel id="FCF95787FFB4FFD2FB55FAA78C56FAF5" box="[1232,1296,1339,1365]" pageId="4" pageNumber="265" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, as it shares neither the absence of cornea as in
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FD91FAC38A6CFAD9" authority="Chace, 1940" authorityName="Chace" authorityYear="1940" box="[532,810,1375,1401]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="rostratus">
|
||
<emphasis id="E7CAEAFCFFB4FFD2FD91FAC38BD1FAD8" box="[532,663,1375,1401]" italics="true" pageId="4" pageNumber="265">C. rostratus</emphasis>
|
||
<bibRefCitation id="B12F4B1FFFB4FFD2FD19FAC38A6CFAD9" author="Chace, F. A. Jr." box="[668,810,1375,1401]" pageId="4" pageNumber="265" pagination="1 - 67" refId="ref4065" refString="Chace, F. A. Jr. (1940) Reports on the scientific results of the Atlantis expeditions to the West Indies, under the joint auspices of the University of Havana and Harvard University. The brachyuran crabs. Torreia, Havana, 4, 1 - 67." type="journal article" year="1940">Chace, 1940</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FCE4FAC38D19FAD9" authority="Chace, 1940" authorityName="Chace" authorityYear="1940" box="[865,1119,1375,1401]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="caecus">
|
||
<emphasis id="E7CAEAFCFFB4FFD2FCE4FAC38A8AFAD8" box="[865,972,1375,1401]" italics="true" pageId="4" pageNumber="265">C. caecus</emphasis>
|
||
<bibRefCitation id="B12F4B1FFFB4FFD2FC54FAC38D19FAD9" author="Chace, F. A. Jr." box="[977,1119,1375,1401]" pageId="4" pageNumber="265" pagination="1 - 67" refId="ref4065" refString="Chace, F. A. Jr. (1940) Reports on the scientific results of the Atlantis expeditions to the West Indies, under the joint auspices of the University of Havana and Harvard University. The brachyuran crabs. Torreia, Havana, 4, 1 - 67." type="journal article" year="1940">Chace, 1940</bibRefCitation>
|
||
</taxonomicName>
|
||
, the distinct lateral curvature of the stalks as in
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FEEBFA1F8A6DFA3D" authority="A. Milne-Edwards, 1880" authorityName="A. Milne-Edwards" authorityYear="1880" box="[366,811,1411,1437]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="quadratus">
|
||
<emphasis id="E7CAEAFCFFB4FFD2FEEBFA1F8B45FA3D" box="[366,515,1411,1437]" italics="true" pageId="4" pageNumber="265">C. quadratus</emphasis>
|
||
A.
|
||
<bibRefCitation id="B12F4B1FFFB4FFD2FDB4FA1F8A6DFA3D" author="Milne-Edwards, A." box="[561,811,1411,1437]" pageId="4" pageNumber="265" pagination="1 - 68" refId="ref4272" refString="Milne-Edwards, A. (1880) Etudes preliminaires sur les crustaces. In: Reports on the Results of Dredging, under the Supervision of Alexander Agassiz, in the Gulf of Mexico, and in the Caribbean Sea, 1877, ' 78, ' 79, by the United States Coast Survey Steamer " Blake, " Lieut. - Commander C. D. Sigsbee, U. S. N., and Commander J. R. Bartlett, U. S. N., Commanding, VIII. Bulletin of the Museum of Comparative Zoology, Harvard, 8 (1), 1 - 68." type="journal article" year="1880">Milne-Edwards, 1880</bibRefCitation>
|
||
</taxonomicName>
|
||
, nor the strong taper toward the apices as found in
|
||
<emphasis id="E7CAEAFCFFB4FFD2FA06FA1F88B0FA61" italics="true" pageId="4" pageNumber="265">
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FA06FA1F89A1FA60" authorityName="Chace" authorityYear="1940" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="caecus">C. caecus</taxonomicName>
|
||
.
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FF74FA3B88B0FA61" authority="Felder & Tavares, 2024" authorityName="Felder & Tavares" authorityYear="2024" box="[241,502,1447,1473]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="windsorae" status="sp. nov.">Cymonomus windsorae</taxonomicName>
|
||
</emphasis>
|
||
<emphasis id="E7CAEAFCFFB4FFD2FE79FA3B8B7CFA61" bold="true" box="[508,570,1447,1473]" pageId="4" pageNumber="265">
|
||
<taxonomicNameLabel id="FCF95787FFB4FFD2FE79FA3B8B7CFA61" box="[508,570,1447,1473]" pageId="4" pageNumber="265" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be further separated from
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FC0BFA3B8D54FA60" authorityName="Chace" authorityYear="1940" box="[910,1042,1447,1473]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="rostratus">
|
||
<emphasis id="E7CAEAFCFFB4FFD2FC0BFA3B8D54FA60" box="[910,1042,1447,1473]" italics="true" pageId="4" pageNumber="265">C. rostratus</emphasis>
|
||
</taxonomicName>
|
||
by the anterolateral shoulder of the carapace that appears more produced or angular in dorsal perspective, often forming a rounded, sometimes spined corner as opposed to a more broadly rounded margin as in
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FCACFA738AE8F9A8" authorityName="Chace" authorityYear="1940" box="[809,942,1519,1545]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="rostratus">
|
||
<emphasis id="E7CAEAFCFFB4FFD2FCACFA738AE8F9A8" box="[809,942,1519,1545]" italics="true" pageId="4" pageNumber="265">C. rostratus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D50136EEFFB4FFD2FF42F98F886FF835" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="265">
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FF42F98F8817F98D" authorityName="A. Milne-Edwards" authorityYear="1880" box="[199,337,1555,1581]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E7CAEAFCFFB4FFD2FF42F98F8817F98D" box="[199,337,1555,1581]" italics="true" pageId="4" pageNumber="265">Cymonomus</emphasis>
|
||
</taxonomicName>
|
||
is essentially a deep-water group currently consisting of 45 species, of which 33 are Indo-West Pacific and 12 Atlanto-Mediterranean/East Pacific in distribution. It also includes one species from the Late Eocene of
|
||
<collectingCountry id="ADA9767EFFB4FFD2FF3DF9C7885DF9D5" box="[184,283,1627,1653]" name="Hungary" pageId="4" pageNumber="265">Hungary</collectingCountry>
|
||
,
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FEADF9C78B91F9D5" authority="Muller & Collins, 1991" authorityName="Muller & Collins" authorityYear="1991" box="[296,727,1627,1653]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="primitivus">
|
||
<emphasis id="E7CAEAFCFFB4FFD2FEADF9C788FBF9D4" box="[296,445,1627,1653]" italics="true" pageId="4" pageNumber="265">C. primitivus</emphasis>
|
||
<bibRefCitation id="B12F4B1FFFB4FFD2FE43F9C78B91F9D5" author="Muller, P. & Collins, J. S. H." box="[454,727,1627,1653]" pageId="4" pageNumber="265" pagination="47 - 92" refId="ref4490" refString="Muller, P. & Collins, J. S. H. (1991) Late Eocene coral-associated decapods (Crustacea) from Hungary. Contributions to Tertiary and Quaternary Geology, 28 (2 - 3), 47 - 92." type="journal article" year="1991">Müller & Collins, 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
. A number of descriptive and distributional accounts over the last two decades have markedly expanded documented diversity of
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FC09F9E38D50F939" authorityName="A. Milne-Edwards" authorityYear="1880" box="[908,1046,1663,1689]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E7CAEAFCFFB4FFD2FC09F9E38D50F939" box="[908,1046,1663,1689]" italics="true" pageId="4" pageNumber="265">Cymonomus</emphasis>
|
||
</taxonomicName>
|
||
in Indo-Pacific waters (
|
||
<bibRefCitation id="B12F4B1FFFB4FFD2FAA6F9E38860F91D" author="Ahyong, S. T. & Brown, D. E." pageId="4" pageNumber="265" pagination="1363 - 1374" refId="ref3677" refString="Ahyong, S. T. & Brown, D. E. (2003) New species of Cymonomus from southeastern Australia (Brachyura, Cymonomidae) with a key to the Indo-West Pacific species. Crustaceana, 75, 1363 - 1374. https: // doi. org / 10.1163 / 156854002321629790" type="journal article" year="2003">Ahyong & Brown 2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B12F4B1FFFB4FFD2FEB5F93F888BF91D" author="Ahyong, S. T." box="[304,461,1699,1725]" pageId="4" pageNumber="265" pagination="1 - 72" refId="ref3528" refString="Ahyong, S. T. (2008) Deepwater crabs from seamounts and chemosynthetic habitats off eastern New Zealand (Crustacea: Decapoda: Brachyura). Zootaxa, 1708 (1), 1 - 72. https: // doi. org / 10.11646 / zootaxa. 1708.1.1" type="journal article" year="2008">Ahyong 2008</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="B12F4B1FFFB4FFD2FE5DF93F8B55F91D" author="Ahyong, S. T." box="[472,531,1699,1725]" pageId="4" pageNumber="265" pagination="384 - 390" refId="ref3579" refString="Ahyong, S. T. (2014) Cymonomid crabs of the MAINBAZA Expedition (Decapoda: Brachyura). Zootaxa, 3821 (3), 384 - 390. https: // doi. org / 10.11646 / zootaxa. 3821.3.7" type="journal article" year="2014">2014</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="B12F4B1FFFB4FFD2FD9BF93F8B1FF91D" author="Ahyong, S. T." box="[542,601,1699,1725]" pageId="4" pageNumber="265" pagination="33 - 69" refId="ref3623" refString="Ahyong, S. T. (2019) The cymonomid crabs of New Zealand and Australia (Crustacea: Brachyura: Cyclodorripoida). Records of the Australian Museum, 71 (2), 33 - 69. https: // doi. org / 10.3853 / j. 2201 - 4349.71.2019.1682" type="journal article" year="2019">2019</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B12F4B1FFFB4FFD2FDE1F93F8A07F91D" author="Ahyong, S. T. & Mitra, S. & Ng, P. K. L." box="[612,833,1699,1725]" pageId="4" pageNumber="265" pagination="62 - 69" refId="ref3732" refString="Ahyong, S. T., Mitra, S. & Ng, P. K. L. (2020). Cymonomid crabs from southwestern Indonesia and redescription of Cymonomus andamanicus Alcock, 1905. Raffles Bulletin of Zoology, 68, 62 - 69." type="journal article" year="2020">
|
||
Ahyong
|
||
<emphasis id="E7CAEAFCFFB4FFD2FD43F93F8BB1F91D" box="[710,759,1699,1725]" italics="true" pageId="4" pageNumber="265">et al</emphasis>
|
||
. 2020
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B12F4B1FFFB4FFD2FCCEF93F8D68F91D" author="Ahyong, S. T. & Ng, P. K. L." box="[843,1070,1699,1725]" pageId="4" pageNumber="265" pagination="233 - 246" refId="ref3781" refString="Ahyong, S. T. & Ng, P. K. L. (2009) The Cymonomidae of the Philippines (Crustacea: Decapoda: Brachyura), with descriptions of four new species. Raffles Bulletin of Zoology, Supplement 20, 233 - 246." type="journal article" year="2009">Ahyong & Ng 2009</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="B12F4B1FFFB4FFD2FBBCF93F8D35F91D" author="Ahyong, S. T. & Ng, P. K. L." box="[1081,1139,1699,1725]" pageId="4" pageNumber="265" pagination="479 - 487" refId="ref3831" refString="Ahyong, S. T. & Ng, P. K. L. (2011) Cyclodorippoid crabs from the Philippines collected by the PANGLAO 2004 - 2005 and AURORA 2007 expeditions. Zoologischer Anzeiger, 250, 479 - 487. https: // doi. org / 10.1016 / j. jcz. 2011.06.001" type="journal article" year="2011">2011</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="B12F4B1FFFB4FFD2FBFBF93F8DFFF91D" author="Ahyong, S. T. & Ng, P. K. L." box="[1150,1209,1699,1725]" pageId="4" pageNumber="265" pagination="1 - 20" refId="ref3889" refString="Ahyong, S. T. & Ng, P. K. L. (2017) East Asian cymonomid crabs (Crustacea: Brachyura). Zoological Studies, 56 (24), 1 - 20. https: // doi. org / 10.6620 / ZS. 2017.56 - 24" type="journal article" year="2017">2017</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B12F4B1FFFB4FFD2FB40F93F8CD1F91D" author="Takeda, M. & Ohtsuchi, N. & Komatsu, H." box="[1221,1431,1699,1725]" pageId="4" pageNumber="265" pagination="65 - 97" refId="ref4534" refString="Takeda, M., Ohtsuchi, N. & Komatsu, H. (2021) Crabs (Crustacea, Decapoda) from the Sea off East and Southeast Asia collected by the RV Hakuho Maru (KH- 72 - 1 Cruise) 1. Sulu Sea and Sibutu Passage. Bulletin of the National Museum of Nature and Science, Series A (Zoology), 47, 65 - 97." type="journal article" year="2021">
|
||
Takeda
|
||
<emphasis id="E7CAEAFCFFB4FFD2FA98F93F8C08F91D" box="[1309,1358,1699,1725]" italics="true" pageId="4" pageNumber="265">et al</emphasis>
|
||
. 2021
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B12F4B1FFFB4FFD2FF12F95B883BF941" author="Ahyong, S. T. & Ng, P. K. L." box="[151,381,1735,1761]" pageId="4" pageNumber="265" pagination="116 - 119" refId="ref3943" refString="Ahyong, S. T. & Ng, P. K. L. (2023) Cymonomus dwi, a new species of deep-water cymonomid crab from Indonesia (Crustacea: Brachyura). Raffles Bulletin of Zoology, 71, 116 - 119. https: // doi. org / 10.26107 / RBZ- 2023 - 0010" type="journal article" year="2023">Ahyong & Ng 2023</bibRefCitation>
|
||
). The Atlanto-Mediterranean/East Pacific species of
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FC48F95B8D11F941" authorityName="A. Milne-Edwards" authorityYear="1880" box="[973,1111,1735,1761]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E7CAEAFCFFB4FFD2FC48F95B8D11F941" box="[973,1111,1735,1761]" italics="true" pageId="4" pageNumber="265">Cymonomus</emphasis>
|
||
</taxonomicName>
|
||
are under revision by one of us (MT) so that further additions are anticipated in the course of this effort, as well as range extensions building on previous work (
|
||
<bibRefCitation id="B12F4B1FFFB4FFD2FEF3F8938B55F889" author="Tavares, M." box="[374,531,1807,1833]" pageId="4" pageNumber="265" pagination="623 - 648" refId="ref4608" refString="Tavares, M. (1991) Especes nouvelles de Cyclodorippoidea Ortmann et remarques sur les genres Tymolus Stimpson et Cyclodorippe A. Milne Edwards (Crustacea, Decapoda, Brachyura). Bulletin du Museum national d'Histoire naturelle, Paris, 4 A, 12 (3 - 4), 623 - 648. [dated 1990, published 1991] https: // doi. org / 10.5962 / p. 289575" type="journal article" year="1991">Tavares 1991</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="B12F4B1FFFB4FFD2FDA6F8938B2CF889" author="Tavares, M." box="[547,618,1807,1833]" pageId="4" pageNumber="265" pagination="137 - 144" refId="ref4683" refString="Tavares, M. (1993 a) Description preliminaire de quatre nouveaux genres et trois nouvelles especes de Cyclodorippoidea Americains (Crustacea, Decapoda Brachyura). Vie et Milieu, 43, 137 - 144." type="journal article" year="1993">1993a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="B12F4B1FFFB4FFD2FDFFF8938BF3F889" author="Tavares, M." box="[634,693,1807,1833]" pageId="4" pageNumber="265" pagination="203 - 208" refId="ref4784" refString="Tavares, M. (1994) Description de Cymonomus leblondi sp. nov. de Guadeloupe, avec une clef des Cymonomus americains (Decapoda, Brachyura, Cymonomidae). Bulletin du Museum national d'Histoire naturelle, Paris, Serie 4, 16, 203 - 208." type="journal article" year="1994">1994</bibRefCitation>
|
||
; Campos Junior 1997). Additional insights into the cymonomid assemblage of the Gulf of Mexico region are, however, constrained by limited appropriate sampling in the region over recent decades, which also limits insight into microhabitats and substrate dependencies of
|
||
<taxonomicName id="12BE4D6DFFB4FFD2FB44F8CB8C0DF8D1" authorityName="A. Milne-Edwards" authorityYear="1880" box="[1217,1355,1879,1905]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E7CAEAFCFFB4FFD2FB44F8CB8C0DF8D1" box="[1217,1355,1879,1905]" italics="true" pageId="4" pageNumber="265">Cymonomus</emphasis>
|
||
</taxonomicName>
|
||
and its confamilials.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |