222 lines
18 KiB
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222 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/nl.43.54916" ID-GBIF-Dataset="50ba320b-8af3-4e5c-8521-0751d34ec29b" ID-GBIF-Taxon="170296335" ID-Pensoft-Pub="2367-5365-43-337" ID-Pensoft-UUID="EB2A020693ED527CA0437F9679DF9098" ID-ZooBank="14033F5AF5E742B9894276B050E36B70" ModsDocID="2367-5365-43-337" checkinTime="1605112368173" checkinUser="pensoft" docAuthor="Wikstroem, Bo, Huemer, Peter, Mutanen, Marko, Tyllinen, Juha & Kaila, Lauri" docDate="2020" docId="BDA8601C15E6525CB246D374EFDA8E7B" docLanguage="en" docName="NotaLepidopt 43: 337-364" docOrigin="Nota Lepidopterologica 43" docSource="http://dx.doi.org/10.3897/nl.43.54916" docTitle="Pyralis princeps Butler 1889" docType="treatment" docVersion="3" id="EB2A020693ED527CA0437F9679DF9098" lastPageNumber="337" masterDocId="EB2A020693ED527CA0437F9679DF9098" masterDocTitle="Pyralis cardinalis, a charismatic new species related to P. regalis [Denis & Schiffermueller], 1775, first recognized in Finland (Lepidoptera, Pyralidae)" masterLastPageNumber="364" masterPageNumber="337" pageId="0" pageNumber="337" updateTime="1643441744806" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Pyralis cardinalis, a charismatic new species related to P. regalis [Denis & Schiffermueller], 1775, first recognized in Finland (Lepidoptera, Pyralidae)</mods:title>
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Wikstroem, Bo</mods:namePart>
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<mods:affiliation>Yli-Haakkointie 13, 03100 Nummela, Finland e-mail: bo. r. wikstrom @ gmail. com</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Huemer, Peter</mods:namePart>
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<mods:affiliation>Tiroler Landesmuseen-Betriebsgesellschaft m. b. H., Sammlungs- und Forschunszentrum, Naturwissenschaftliche Sammlungen, Hall in Tirol, Austria; e-mail: p. huemer @ tiroler-landesmuseen. at</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Mutanen, Marko</mods:namePart>
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<mods:affiliation>Ecology and Genetics Research Unit, PO BOX 3000, FI- 90014 University of Oulu, Finland; e-mail: marko. mutanen @ oulu. fi</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-4464-6308</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tyllinen, Juha</mods:namePart>
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<mods:affiliation>Tammistontie 4 a E 31, FI- 01250 Vantaa; e-mail: jkt-koti @ kolumbus. fi</mods:affiliation>
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</mods:name>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kaila, Lauri</mods:namePart>
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<mods:affiliation>Finnish Museum of Natural History, Zoology Unit, University of Helsinki, Finland; e-mail: Lauri. Kaila @ Helsinki. fi (corresponding author)</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-0277-1872</mods:nameIdentifier>
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<mods:nameIdentifier type="email">lauri.kaila@helsinki.fi</mods:nameIdentifier>
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<mods:title>Nota Lepidopterologica</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>43</mods:number>
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<mods:start>337</mods:start>
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<mods:end>364</mods:end>
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<mods:url>http://dx.doi.org/10.3897/nl.43.54916</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/nl.43.54916</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2367-5365-43-337</mods:identifier>
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<treatment ID-GBIF-Taxon="170296335" LSID="urn:lsid:plazi:treatment:BDA8601C15E6525CB246D374EFDA8E7B" httpUri="http://treatment.plazi.org/id/BDA8601C15E6525CB246D374EFDA8E7B" lastPageNumber="337" pageId="0" pageNumber="337">
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<subSubSection pageId="0" pageNumber="337" type="nomenclature">
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<paragraph pageId="0" pageNumber="337">
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<taxonomicName LSID="BDA8601C-15E6-525C-B246-D374EFDA8E7B" authority="Butler, 1889" authorityName="Butler" authorityYear="1889" class="Insecta" family="Pyralidae" genus="Pyralis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyralis princeps" order="Lepidoptera" pageId="0" pageNumber="337" phylum="Arthropoda" rank="species" species="princeps">Pyralis princeps Butler, 1889</taxonomicName>
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<figureCitation captionStart="Figures 18–21" captionStartId="F6" captionText="Figures 18 - 21. Pyralis spp., habitus. 18. P. princeps Butler, ♂; Nepal, Godawari nr. Kathmandu, leg. exp. Albrecht et al. 19. P. joannisi Leraut, ♂; Vietnam, Bac Kan province, leg. A. Wikstroem. 20, 21. P. perversalis (Herrich-Schaeffer), habitus. 20. ♂, Czech Republic, Bohemia sept., leg. J. Liska; 21. ♀, S. Ural, Cheliabinsk district, leg. T. Nupponen." figureDoi="10.3897/nl.43.54916.figures18-21" httpUri="https://binary.pensoft.net/fig/473651" pageId="0" pageNumber="337">Figs 18</figureCitation>
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<figureCitation captionStart="Figures 25–27" captionStartId="F8" captionText="Figures 25 - 27. ♂ genitalia of Pyralis spp. 25. P. kacheticalis (Christoph), GPBW 7942, Greece, Lesvos, Anaxos. 26. P. princeps Butler, L. Kaila prep. 6296, Nepal, Godawari, 27. P. joannisi Leraut, L. Kaila prep. 6297, Vietnam, Bac Kan Prov." figureDoi="10.3897/nl.43.54916.figures25-27" httpUri="https://binary.pensoft.net/fig/473653" pageId="0" pageNumber="337">, 26</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="337" type="reference_group">
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<paragraph pageId="0" pageNumber="337">
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<taxonomicName authorityName="Butler" authorityYear="1889" class="Insecta" family="Pyralidae" genus="Pyralis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyralis princeps" order="Lepidoptera" pageId="0" pageNumber="337" phylum="Arthropoda" rank="species" species="princeps">Pyralis princeps</taxonomicName>
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Butler, 1889(7): 91, pl. 134, fig. 12.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="337" type="materials_examined">
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<paragraph pageId="0" pageNumber="337">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="337">
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Type material.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="337">Lectotype</emphasis>
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, here designated: India, Dharamshala, Photograph of
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<emphasis bold="true" italics="true" pageId="0" pageNumber="337">lectotype</emphasis>
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of
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<taxonomicName authorityName="Butler" authorityYear="1889" class="Insecta" family="Pyralidae" genus="Pyralis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyralis princeps" order="Lepidoptera" pageId="0" pageNumber="337" phylum="Arthropoda" rank="species" species="princeps">
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<emphasis italics="true" pageId="0" pageNumber="337">Pyralis princeps</emphasis>
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</taxonomicName>
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made available by David Lees, labelled: LECTO-TYPE [blue round label]; MANUSCRIPT LECTOTYPE; Type locality India Dharmsala; Lectotype
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<taxonomicName authorityName="Butler" authorityYear="1889" class="Insecta" family="Pyralidae" genus="Pyralis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyralis princeps" order="Lepidoptera" pageId="0" pageNumber="337" phylum="Arthropoda" rank="species" species="princeps">
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<emphasis italics="true" pageId="0" pageNumber="337">Pyralis princeps</emphasis>
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</taxonomicName>
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det. M. Shaffer 1985[]; stat. n. [abdomen in capsule] (NHMUK).
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</paragraph>
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<paragraph pageId="0" pageNumber="337">
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<emphasis bold="true" pageId="0" pageNumber="337">Other material.</emphasis>
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Nepal. 2♂,
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<geoCoordinate degrees="27" direction="north" minutes="40" orientation="latitude" precision="925" value="27.666666">27°40'N</geoCoordinate>
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,
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<geoCoordinate degrees="85" direction="east" minutes="25" orientation="longitude" precision="925" value="85.416664">85°25'E</geoCoordinate>
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, Godavari, 15 km SE Kathmandu, 1500 m, 8.-9.v. 1996 leg. Exp. A. Albrecht, O.
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<normalizedToken originalValue="Biström">Bistroem</normalizedToken>
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, K. Mikkola & A. Wikberg, L. Kaila prep. 6296, BOLD sample ID: TLMF Lep 05678, BOLD sample ID: TLMF Lep 05679, http://id.luomus.fi/HV.369 (MZH).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="337" type="diagnosis">
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<paragraph pageId="0" pageNumber="337">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="337">
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The separation of this species from other species is explained in the key. In the genitalia, the dense bush of long spines at the posterior end of vesica is characteristic, and the genital characters readily separate it from the externally similar but smaller
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<taxonomicName genus="P." lsidName="P. joannisi" pageId="0" pageNumber="337" rank="species" species="joannisi">
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<emphasis italics="true" pageId="0" pageNumber="337">P. joannisi</emphasis>
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</taxonomicName>
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; see the key for further details.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="337" type="molecular data">
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<paragraph pageId="0" pageNumber="337">Molecular data.</paragraph>
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<paragraph pageId="0" pageNumber="337">
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BIN: replace: BOLD:ABA8505. The intraspecific mean and maximum divergence of the barcode region is 0% (
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<emphasis italics="true" pageId="0" pageNumber="337">N</emphasis>
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= 2). The minimum distance to the nearest neighbour,
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<taxonomicName genus="P." lsidName="P. kacheticalis" pageId="0" pageNumber="337" rank="species" species="kacheticalis">
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<emphasis italics="true" pageId="0" pageNumber="337">P. kacheticalis</emphasis>
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</taxonomicName>
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, is 8.26%.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="337" type="distribution">
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<paragraph pageId="0" pageNumber="337">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="337">India, Nepal.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="337" type="remarks">
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<paragraph pageId="0" pageNumber="337">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="337">
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<taxonomicName authorityName="Butler" authorityYear="1889" class="Insecta" family="Pyralidae" genus="Pyralis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyralis princeps" order="Lepidoptera" pageId="0" pageNumber="337" phylum="Arthropoda" rank="species" species="princeps">
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<emphasis italics="true" pageId="0" pageNumber="337">Pyralis princeps</emphasis>
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</taxonomicName>
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was described from four specimens collected in India (Himachal Pradesh, Dharamshala [Dharmsala]) and two from
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<normalizedToken originalValue="”Yezo”">"Yezo"</normalizedToken>
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[region from Northern Japan to Kamchatka] (Butler, 1889). However, the latter two are misidentifications of
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<taxonomicName genus="P." lsidName="P. cardinalis" pageId="0" pageNumber="337" rank="species" species="cardinalis">
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<emphasis italics="true" pageId="0" pageNumber="337">P. cardinalis</emphasis>
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</taxonomicName>
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, this interpretation also supported by our examination of some Japanese and South Korean specimens. Following
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<bibRefCitation author="ICZN" journalOrPublisher="The International Trust for Zoological Nomenclature, London" pageId="0" pageNumber="337" refId="B10" refString="ICZN, 1999. International Code of Zoological Nomenclature. Fourth edition. The International Trust for Zoological Nomenclature, London" title="International Code of Zoological Nomenclature. Fourth edition." year="1999">ICZN (1999)</bibRefCitation>
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, a lectotype from Dharamshala [Dharmsala], already labelled as such by M. Shaffer in NHMUK, is here designated in order to fix the identity of the species and conserve nomenclatural stability.We have examined photographs of the specimen and its labels which confirm our interpretation. Outside Europe, the name
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<taxonomicName genus="P." lsidName="P. princeps" pageId="0" pageNumber="337" rank="species" species="princeps">
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<emphasis italics="true" pageId="0" pageNumber="337">P. princeps</emphasis>
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</taxonomicName>
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Butler has been somewhat inconsistently used for the East Palearctic populations. E.g.
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<bibRefCitation author="Shibuya, J" journalOrPublisher="Journal of the Faculty of Agriculture, Hokkaido Imperial University" pageId="0" pageNumber="337" pagination="149 - 176" refId="B24" refString="Shibuya, J, 1928. The systematic study on the Japanese Pyralinae (Lepid.). Journal of the Faculty of Agriculture, Hokkaido Imperial University 21 (4): 149 - 176" title="The systematic study on the Japanese Pyralinae (Lepid.)." volume="21" year="1928">Shibuya (1928)</bibRefCitation>
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, in his treatment of Japanese species of
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<taxonomicName class="Insecta" family="Pyralidae" genus="Pyralis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyralis" order="Lepidoptera" pageId="0" pageNumber="337" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="337">Pyralis</emphasis>
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</taxonomicName>
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and related genera, only recognized
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<taxonomicName genus="P." lsidName="P. regalis" pageId="0" pageNumber="337" rank="species" species="regalis">
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<emphasis italics="true" pageId="0" pageNumber="337">P. regalis</emphasis>
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</taxonomicName>
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of the relevant taxa to occur there. Inoue
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<emphasis italics="true" pageId="0" pageNumber="337">et al.</emphasis>
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(1982), plate 46: Figs
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<figureCitation captionStart="Figures 4–11" captionStartId="F4" captionText="Figures 4 - 11. Pyralis sp., habitus. 4 - 7. P. cardinalis sp. nov., 4. ♂, Finland, V, Kemioensaari, leg. J. Tyllinen. 5. ♀, A, Lemland, leg. B. Wikstroem & K. Vaalamo; 6. ♂, Finland, U, Kirkkonummi, leg. J. Junnilainen; 7. ♂, Finland, U, Raasepori, leg. L. Kaila. 8 - 11. P. regalis. 8. ♂, Greece, Lesvos, leg. L. Kaila & J. Kullberg. 9. ♀, Slovenia, Granska Gora, leg. B. Wikstroem; 10. ♂, Italy, Tuscany, Cortona, leg. B. Wikstroem. 11. Bulgaria, 5 km S. Blagoevgrad, ♀, leg. B. A. Bengtsson." figureDoi="10.3897/nl.43.54916.figures4-11" httpUri="https://binary.pensoft.net/fig/473649" pageId="0" pageNumber="337">4</figureCitation>
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,
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<figureCitation captionStart="Figures 4–11" captionStartId="F4" captionText="Figures 4 - 11. Pyralis sp., habitus. 4 - 7. P. cardinalis sp. nov., 4. ♂, Finland, V, Kemioensaari, leg. J. Tyllinen. 5. ♀, A, Lemland, leg. B. Wikstroem & K. Vaalamo; 6. ♂, Finland, U, Kirkkonummi, leg. J. Junnilainen; 7. ♂, Finland, U, Raasepori, leg. L. Kaila. 8 - 11. P. regalis. 8. ♂, Greece, Lesvos, leg. L. Kaila & J. Kullberg. 9. ♀, Slovenia, Granska Gora, leg. B. Wikstroem; 10. ♂, Italy, Tuscany, Cortona, leg. B. Wikstroem. 11. Bulgaria, 5 km S. Blagoevgrad, ♀, leg. B. A. Bengtsson." figureDoi="10.3897/nl.43.54916.figures4-11" httpUri="https://binary.pensoft.net/fig/473649" pageId="0" pageNumber="337">5</figureCitation>
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illustrated Japanese specimens that seem identical to
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<taxonomicName genus="P." lsidName="P. cardinalis" pageId="0" pageNumber="337" rank="species" species="cardinalis">
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<emphasis italics="true" pageId="0" pageNumber="337">P. cardinalis</emphasis>
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</taxonomicName>
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and whose occurrence in Japan is verified in the present study. Neither of these authors mention other species, but they consider
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<taxonomicName genus="P." lsidName="P. princeps" pageId="0" pageNumber="337" rank="species" species="princeps">
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<emphasis italics="true" pageId="0" pageNumber="337">P. princeps</emphasis>
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</taxonomicName>
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a synonym of
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<taxonomicName genus="P." lsidName="P. regalis" pageId="0" pageNumber="337" rank="species" species="regalis">
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<emphasis italics="true" pageId="0" pageNumber="337">P. regalis</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<caption doi="10.3897/nl.43.54916.figures18-21" httpUri="https://binary.pensoft.net/fig/473651" pageId="0" pageNumber="337" start="Figures 18–21" startId="F6">
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<paragraph pageId="0" pageNumber="337">
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<emphasis bold="true" pageId="0" pageNumber="337">Figures 18-21.</emphasis>
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<taxonomicName class="Insecta" family="Pyralidae" genus="Pyralis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyralis" order="Lepidoptera" pageId="0" pageNumber="337" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="337">Pyralis</emphasis>
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</taxonomicName>
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spp., habitus.
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<emphasis bold="true" pageId="0" pageNumber="337">18.</emphasis>
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<taxonomicName genus="P." lsidName="P. princeps" pageId="0" pageNumber="337" rank="species" species="princeps">
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<emphasis italics="true" pageId="0" pageNumber="337">P. princeps</emphasis>
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</taxonomicName>
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Butler, ♂; Nepal, Godawari nr. Kathmandu, leg. exp. Albrecht et al.
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<emphasis bold="true" pageId="0" pageNumber="337">19.</emphasis>
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<taxonomicName genus="P." lsidName="P. joannisi" pageId="0" pageNumber="337" rank="species" species="joannisi">
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<emphasis italics="true" pageId="0" pageNumber="337">P. joannisi</emphasis>
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</taxonomicName>
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Leraut, ♂; Vietnam, Bac Kan province, leg. A.
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<normalizedToken originalValue="Wikström">Wikstroem</normalizedToken>
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.
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<emphasis bold="true" pageId="0" pageNumber="337">20, 21.</emphasis>
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<taxonomicName genus="P." lsidName="P. perversalis" pageId="0" pageNumber="337" rank="species" species="perversalis">
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<emphasis italics="true" pageId="0" pageNumber="337">P. perversalis</emphasis>
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</taxonomicName>
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(
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<normalizedToken originalValue="Herrich-Schäffer">Herrich-Schaeffer</normalizedToken>
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), habitus.
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<emphasis bold="true" pageId="0" pageNumber="337">20.</emphasis>
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♂, Czech Republic, Bohemia sept., leg. J.
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<normalizedToken originalValue="Liška">Liska</normalizedToken>
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;
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<emphasis bold="true" pageId="0" pageNumber="337">21.</emphasis>
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♀, S. Ural, Cheliabinsk district, leg. T. Nupponen.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |