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<document id="E31F3026715803AF96204E575B162F89" ID-DOI="10.11646/zootaxa.5403.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="10561942" ID-ZooBank="127A062D-8597-4DD8-B3EE-1B94A3F0B878" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1706104528303" checkinUser="plazi" docAuthor="Fleck, Günther" docDate="2024" docId="03F387EEFFCEFF8BFF3DFF05FAA87969" docLanguage="en" docName="zootaxa.5403.3.2.pdf" docOrigin="Zootaxa 5403 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5403.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neocaledosynthemis Fleck 2024, gen. nov." docType="treatment" docVersion="1" lastPageNumber="329" masterDocId="FFCAFF96FFC6FF82FFAAFF92FFA97D5C" masterDocTitle="A remarkable new synthemistid from New Caledonia (Odonata: Anisoptera: Synthemistidae s. str.). Taxonomic and phylogenetic note on New Caledonian Synthemistidae and erection of a new genus" masterLastPageNumber="330" masterPageNumber="320" pageNumber="328" updateTime="1706193327035" updateUser="julia" zenodo-license-document="CLOSED">
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<mods:title id="6AAE71D8948501E17174EA9B42F205E0">A remarkable new synthemistid from New Caledonia (Odonata: Anisoptera: Synthemistidae s. str.). Taxonomic and phylogenetic note on New Caledonian Synthemistidae and erection of a new genus</mods:title>
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<treatment id="03F387EEFFCEFF8BFF3DFF05FAA87969" LSID="urn:lsid:plazi:treatment:03F387EEFFCEFF8BFF3DFF05FAA87969" httpUri="http://treatment.plazi.org/id/03F387EEFFCEFF8BFF3DFF05FAA87969" lastPageId="9" lastPageNumber="329" pageId="8" pageNumber="328">
<subSubSection id="C3406573FFCEFF8AFF3DFF05FDAD7DED" box="[151,516,151,178]" pageId="8" pageNumber="328" type="nomenclature">
<paragraph id="8BE536F8FFCEFF8AFF3DFF05FDAD7DED" blockId="8.[151,516,151,178]" box="[151,516,151,178]" pageId="8" pageNumber="328">
<heading id="D0AD8194FFCEFF8AFF3DFF05FDAD7DED" bold="true" box="[151,516,151,178]" fontSize="11" level="1" pageId="8" pageNumber="328" reason="1">
<emphasis id="B92EEAEAFFCEFF8AFF3DFF05FDAD7DED" bold="true" box="[151,516,151,178]" pageId="8" pageNumber="328">
<taxonomicName id="4C5A4D7BFFCEFF8AFF3DFF05FE267DEE" authority="Fleck, 2024" authorityName="Fleck" authorityYear="2024" box="[151,399,151,178]" class="Insecta" family="Synthemistidae" genus="Neocaledosynthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B92EEAEAFFCEFF8AFF3DFF05FE267DEE" bold="true" box="[151,399,151,178]" italics="true" pageId="8" pageNumber="328">Neocaledosynthemis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D5791FFCEFF8AFE3CFF05FDAD7DED" box="[406,516,151,177]" pageId="8" pageNumber="328" rank="genus">
<emphasis id="B92EEAEAFFCEFF8AFE3CFF05FE6B7DED" bold="true" box="[406,450,151,177]" italics="true" pageId="8" pageNumber="328">gen</emphasis>
. nov.
</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3406573FFCEFF8AFF3DFF4DFDCE7DA5" box="[151,615,223,249]" pageId="8" pageNumber="328" type="type_taxon">
<paragraph id="8BE536F8FFCEFF8AFF3DFF4DFDCE7DA5" blockId="8.[151,817,223,322]" box="[151,615,223,249]" pageId="8" pageNumber="328">
<typeStatus id="54E1885AFFCEFF8AFF3DFF4DFF677DA5" box="[151,206,223,249]" pageId="8" pageNumber="328">Type</typeStatus>
species.
<taxonomicName id="4C5A4D7BFFCEFF8AFE9EFF4DFDCE7DA5" authority="Campion" authorityName="Campion" authorityYear="1921" box="[308,615,223,249]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="fenella">
<emphasis id="B92EEAEAFFCEFF8AFE9EFF4DFE517DA5" box="[308,504,223,249]" italics="true" pageId="8" pageNumber="328">Synthemis fenella</emphasis>
Campion
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3406573FFCEFF8AFF6DFE91FCB17C41" box="[199,792,259,286]" pageId="8" pageNumber="328" type="description">
<paragraph id="8BE536F8FFCEFF8AFF6DFE91FCB17C41" blockId="8.[151,817,223,322]" box="[199,792,259,286]" pageId="8" pageNumber="328">
Other included species.
<taxonomicName id="4C5A4D7BFFCEFF8AFE70FE91FCB17C41" authority="Lieftinck" authorityName="Lieftinck" box="[474,792,259,285]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="ariadne">
<emphasis id="B92EEAEAFFCEFF8AFE70FE91FD037C41" box="[474,682,259,285]" italics="true" pageId="8" pageNumber="328">Synthemis ariadne</emphasis>
Lieftinck
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3406573FFCEFF8AFF6DFEB5FC987C1D" box="[199,817,295,322]" pageId="8" pageNumber="328" type="etymology">
<paragraph id="8BE536F8FFCEFF8AFF6DFEB5FC987C1D" blockId="8.[151,817,223,322]" box="[199,817,295,322]" pageId="8" pageNumber="328">
<emphasis id="B92EEAEAFFCEFF8AFF6DFEB5FEEB7C1E" bold="true" box="[199,322,295,322]" italics="true" pageId="8" pageNumber="328">Etymology</emphasis>
. After
<collectingCountry id="F34D7668FFCEFF8AFE3BFEBAFD977C1D" box="[401,574,295,322]" name="New Caledonia" pageId="8" pageNumber="328">New Caledonia</collectingCountry>
and
<taxonomicName id="4C5A4D7BFFCEFF8AFDDEFEB5FC987C1D" authority="Selys." authorityName="Selys." box="[628,817,295,321]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAEAFFCEFF8AFDDEFEB5FD4F7C1D" box="[628,742,295,321]" italics="true" pageId="8" pageNumber="328">Synthemis</emphasis>
Selys.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3406573FFCEFF8BFF3DFE01FAA87969" lastPageId="9" lastPageNumber="329" pageId="8" pageNumber="328" type="diagnosis">
<paragraph id="8BE536F8FFCEFF8AFF3DFE01FDC27CF2" blockId="8.[151,619,403,430]" box="[151,619,403,430]" pageId="8" pageNumber="328">
<heading id="D0AD8194FFCEFF8AFF3DFE01FDC27CF2" bold="true" box="[151,619,403,430]" fontSize="11" level="1" pageId="8" pageNumber="328" reason="1">
<emphasis id="B92EEAEAFFCEFF8AFF3DFE01FDC27CF2" bold="true" box="[151,619,403,430]" italics="true" pageId="8" pageNumber="328">Justification and differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BE536F8FFCEFF8AFF3DFE49FAA67FD9" blockId="8.[151,1437,475,2014]" pageId="8" pageNumber="328">
<emphasis id="B92EEAEAFFCEFF8AFF3DFE49FEC77CA9" box="[151,366,475,501]" italics="true" pageId="8" pageNumber="328">Material examined</emphasis>
: male, female and larva (reared) of all New Caledonian species of synthemistids (female
<taxonomicName id="4C5A4D7BFFCEFF8AFA29FE4EFF477F45" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="ariadne">
<emphasis id="B92EEAEAFFCEFF8AFA29FE4EFF477F45" italics="true" pageId="8" pageNumber="328">C. ariadne</emphasis>
</taxonomicName>
based on photos), except
<taxonomicName id="4C5A4D7BFFCEFF8AFDBAFD92FD4D7F45" authority="Davies" authorityName="Davies" box="[528,740,511,538]" class="Insecta" genus="Calesynthemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="pamelae">
<emphasis id="B92EEAEAFFCEFF8AFDBAFD92FD267F45" box="[528,655,511,538]" italics="true" pageId="8" pageNumber="328">C. pamelae</emphasis>
Davies
</taxonomicName>
and larva
<taxonomicName id="4C5A4D7BFFCEFF8AFCF2FD92FBA17F45" authorityName="Fleck" authorityYear="2024" box="[856,1032,511,538]" class="Insecta" genus="Calesynthemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="jeanlegrandi">
<emphasis id="B92EEAEAFFCEFF8AFCF2FD92FBA17F45" box="[856,1032,511,538]" italics="true" pageId="8" pageNumber="328">C. jeanlegrandi</emphasis>
</taxonomicName>
. In addition to characters accessible in literature for non-New-Caledonian synthemistids male, female and larva of
<taxonomicName id="4C5A4D7BFFCEFF8AFB94FDB1FA317F61" authority="Tillyard" authorityName="Tillyard" authorityYear="1910" box="[1086,1432,547,573]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="tasmanica">
<emphasis id="B92EEAEAFFCEFF8AFB94FDB1FA867F61" box="[1086,1327,547,573]" italics="true" pageId="8" pageNumber="328">Synthemis tasmanica</emphasis>
Tillyard
</taxonomicName>
,
<taxonomicName id="4C5A4D7BFFCEFF8AFF3DFDD5FD947F3D" authority="Tillyard" authorityName="Tillyard" authorityYear="1908" box="[151,573,583,609]" class="Insecta" family="Synthemistidae" genus="Austrosynthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="cyanitincta">
<emphasis id="B92EEAEAFFCEFF8AFF3DFDD5FE7F7F3D" box="[151,470,583,609]" italics="true" pageId="8" pageNumber="328">Austrosynthemis cyanitincta</emphasis>
Tillyard
</taxonomicName>
,
<taxonomicName id="4C5A4D7BFFCEFF8AFDE6FDD5FC427F3D" authority="Tillyard" authorityName="Tillyard" authorityYear="1909" box="[588,1003,583,609]" class="Insecta" family="Synthemistidae" genus="Tonyosynthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="claviculata">
<emphasis id="B92EEAEAFFCEFF8AFDE6FDD5FC2D7F3D" box="[588,900,583,609]" italics="true" pageId="8" pageNumber="328">Tonyosynthemis claviculata</emphasis>
Tillyard
</taxonomicName>
,
<taxonomicName id="4C5A4D7BFFCEFF8AFC50FDD5FA317F3D" authority="Martin" authorityName="Martin" authorityYear="1901" box="[1018,1432,583,609]" class="Insecta" family="Synthemistidae" genus="Choristhemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="flavoterminata">
<emphasis id="B92EEAEAFFCEFF8AFC50FDD5FA967F3D" box="[1018,1343,583,609]" italics="true" pageId="8" pageNumber="328">Choristhemis flavoterminata</emphasis>
Martin
</taxonomicName>
,
<taxonomicName id="4C5A4D7BFFCEFF8AFF3DFDF9FDBF7FD9" authority="Lieftinck" authorityName="Lieftinck" box="[151,534,619,645]" class="Insecta" family="Synthemistidae" genus="Palaeosynthemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="cyrene">
<emphasis id="B92EEAEAFFCEFF8AFF3DFDF9FE0F7FD9" box="[151,422,619,645]" italics="true" pageId="8" pageNumber="328">Palaeosynthemis cyrene</emphasis>
Lieftinck
</taxonomicName>
,
<taxonomicName id="4C5A4D7BFFCEFF8AFD8BFDF9FC8C7FDA" box="[545,805,619,646]" class="Insecta" family="Synthemistidae" genus="Archaeosynthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B92EEAEAFFCEFF8AFD8BFDF9FD447FD9" box="[545,749,619,645]" italics="true" pageId="8" pageNumber="328">Archaeosynthemis</emphasis>
spp.
</taxonomicName>
, and
<taxonomicName id="4C5A4D7BFFCEFF8AFCCAFDF9FB8A7FDA" box="[864,1059,619,646]" class="Insecta" family="Synthemistidae" genus="Eusynthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B92EEAEAFFCEFF8AFCCAFDF9FC477FD9" box="[864,1006,619,645]" italics="true" pageId="8" pageNumber="328">Eusynthemis</emphasis>
spp.
</taxonomicName>
were also examined.
</paragraph>
<paragraph id="8BE536F8FFCEFF8AFF6DFD1DFB3A7E49" blockId="8.[151,1437,475,2014]" pageId="8" pageNumber="328">
<bibRefCitation id="EFCB4B09FFCEFF8AFF6DFD1DFEFE7FF6" author="Carle, F. L." box="[199,343,655,682]" pageId="8" pageNumber="328" pagination="383 - 424" refId="ref6263" refString="Carle, F. L. (1995) Evolution, taxonomy, and biogeography of ancient Gondwanian libelluloides, with comments on anisopteroid evolution and phylogenetic systematic (Anisoptera: Libelluloidea). Odonatologica, 24 (4), 383 - 424." type="journal article" year="1995">Carle (1995)</bibRefCitation>
, on the supposed basis of a pair of male apomorphic characters (see below discussion), erected the genus
<taxonomicName id="4C5A4D7BFFCEFF8AFF74FD21FE287F91" authorityName="sensu Carle" authorityYear="1995" box="[222,385,691,717]" class="Insecta" genus="Calesynthemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAEAFFCEFF8AFF74FD21FE287F91" box="[222,385,691,717]" italics="true" pageId="8" pageNumber="328">Calesynthemis</emphasis>
</taxonomicName>
to accomodate all
<taxonomicName id="4C5A4D7BFFCEFF8AFDE4FD21FCBF7F91" box="[590,790,691,717]" pageId="8" pageNumber="328">
<emphasis id="B92EEAEAFFCEFF8AFDE4FD21FD697F91" box="[590,704,691,717]" italics="true" pageId="8" pageNumber="328">Synthemis</emphasis>
species
</taxonomicName>
from
<collectingCountry id="F34D7668FFCEFF8AFCFDFD26FBAA7F91" box="[855,1027,691,718]" name="New Caledonia" pageId="8" pageNumber="328">New Caledonia</collectingCountry>
ie
<taxonomicName id="4C5A4D7BFFCEFF8AFB8BFD21FB5E7F91" authorityName="Selys" authorityYear="1871" box="[1057,1271,691,717]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="miranda">
<emphasis id="B92EEAEAFFCEFF8AFB8BFD21FB5E7F91" box="[1057,1271,691,717]" italics="true" pageId="8" pageNumber="328">Synthemis miranda</emphasis>
</taxonomicName>
(
<typeStatus id="54E1885AFFCEFF8AFAAEFD21FA9F7F91" box="[1284,1334,691,717]" pageId="8" pageNumber="328">type</typeStatus>
species),
<taxonomicName id="4C5A4D7BFFCEFF8AFF3DFD45FEA27FAD" box="[151,267,727,753]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="ariadne">
<emphasis id="B92EEAEAFFCEFF8AFF3DFD45FEA27FAD" box="[151,267,727,753]" italics="true" pageId="8" pageNumber="328">S. ariadne</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5A4D7BFFCEFF8AFEB3FD45FDA67FAD" authority="Lieftinck" authorityName="Lieftinck" authorityYear="1971" box="[281,527,727,753]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="campioni">
<emphasis id="B92EEAEAFFCEFF8AFEB3FD45FE377FAD" box="[281,414,727,753]" italics="true" pageId="8" pageNumber="328">S. campioni</emphasis>
Lieftinck
</taxonomicName>
,
<taxonomicName id="4C5A4D7BFFCEFF8AFDB0FD45FD2A7FAD" authorityName="Campion" authorityYear="1921" box="[538,643,727,753]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="fenella">
<emphasis id="B92EEAEAFFCEFF8AFDB0FD45FD2A7FAD" box="[538,643,727,753]" italics="true" pageId="8" pageNumber="328">S. fenella</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5A4D7BFFCEFF8AFD3BFD45FC3D7FAD" authority="Campion" authorityName="Campion" authorityYear="1921" box="[657,916,727,753]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="flexicauda">
<emphasis id="B92EEAEAFFCEFF8AFD3BFD45FC8A7FAD" box="[657,803,727,753]" italics="true" pageId="8" pageNumber="328">S. flexicauda</emphasis>
Campion
</taxonomicName>
,
<taxonomicName id="4C5A4D7BFFCEFF8AFC35FD45FB097FAD" authority="Campion" authorityName="Campion" authorityYear="1921" box="[927,1184,727,753]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="montaguei">
<emphasis id="B92EEAEAFFCEFF8AFC35FD45FB987FAD" box="[927,1073,727,753]" italics="true" pageId="8" pageNumber="328">S. montaguei</emphasis>
Campion
</taxonomicName>
and
<taxonomicName id="4C5A4D7BFFCEFF8AFB7DFD45FAD97FAD" box="[1239,1392,727,753]" class="Insecta" genus="Calesynthemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="serendipita">
<emphasis id="B92EEAEAFFCEFF8AFB7DFD45FAD97FAD" box="[1239,1392,727,753]" italics="true" pageId="8" pageNumber="328">S. serendipita</emphasis>
</taxonomicName>
. As these species are all endemic, the genus
<taxonomicName id="4C5A4D7BFFCEFF8AFDF0FD69FD547E49" authorityName="sensu Carle" authorityYear="1995" box="[602,765,763,789]" class="Insecta" genus="Calesynthemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAEAFFCEFF8AFDF0FD69FD547E49" box="[602,765,763,789]" italics="true" pageId="8" pageNumber="328">Calesynthemis</emphasis>
</taxonomicName>
is also restricted to
<collectingCountry id="F34D7668FFCEFF8AFC74FD6EFB267E49" box="[990,1167,763,790]" name="New Caledonia" pageId="8" pageNumber="328">New Caledonia</collectingCountry>
.
</paragraph>
<paragraph id="8BE536F8FFCEFF8AFF6DFC8DFB777EDE" blockId="8.[151,1437,475,2014]" pageId="8" pageNumber="328">
<bibRefCitation id="EFCB4B09FFCEFF8AFF6DFC8DFEC07E65" author="Davies, D. A. L." box="[199,361,799,825]" pageId="8" pageNumber="328" pagination="229 - 251" refId="ref6221" refString="Davies, D. A. L. (2002) The Odonate fauna of New Caledonia, including the descriptions of a new species and a new subspecies. Odonatologic a, 31 (3), 229 - 251." type="journal article" year="2002">Davies (2002)</bibRefCitation>
did not recognize the genus
<taxonomicName id="4C5A4D7BFFCEFF8AFD12FC8DFCF27E65" authorityName="sensu Carle" authorityYear="1995" box="[696,859,799,825]" class="Insecta" genus="Calesynthemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAEAFFCEFF8AFD12FC8DFCF27E65" box="[696,859,799,825]" italics="true" pageId="8" pageNumber="328">Calesynthemis</emphasis>
</taxonomicName>
and described under
<taxonomicName id="4C5A4D7BFFCEFF8AFBFEFC8DFB6F7E65" box="[1108,1222,799,825]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAEAFFCEFF8AFBFEFC8DFB6F7E65" box="[1108,1222,799,825]" italics="true" pageId="8" pageNumber="328">Synthemis</emphasis>
</taxonomicName>
, a new species (
<taxonomicName id="4C5A4D7BFFCEFF8AFA2DFC8DFF5F7E01" class="Insecta" genus="Calesynthemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="pamelae">
<emphasis id="B92EEAEAFFCEFF8AFA2DFC8DFF5F7E01" italics="true" pageId="8" pageNumber="328">S. pamelae</emphasis>
</taxonomicName>
). Given the available characters and the possible close affinity with
<taxonomicName id="4C5A4D7BFFCEFF8AFC42FCD6FBCE7E01" authorityName="Selys" authorityYear="1871" box="[1000,1127,835,862]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="miranda">
<emphasis id="B92EEAEAFFCEFF8AFC42FCD6FBCE7E01" box="[1000,1127,835,862]" italics="true" pageId="8" pageNumber="328">C. miranda</emphasis>
</taxonomicName>
(being “its nearest relative” after Davies),
<taxonomicName id="4C5A4D7BFFCEFF8AFE9DFCF5FE1B7EDD" box="[311,434,871,897]" class="Insecta" genus="Calesynthemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="pamelae">
<emphasis id="B92EEAEAFFCEFF8AFE9DFCF5FE1B7EDD" box="[311,434,871,897]" italics="true" pageId="8" pageNumber="328">S. pamelae</emphasis>
</taxonomicName>
should have probably been placed in
<taxonomicName id="4C5A4D7BFFCEFF8AFCF3FCF5FB727EDE" authority="sensu Carle (1995)" authorityName="sensu Carle" authorityYear="1995" box="[857,1243,871,898]" class="Insecta" genus="Calesynthemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAEAFFCEFF8AFCF3FCF5FC557EDD" box="[857,1020,871,897]" italics="true" pageId="8" pageNumber="328">Calesynthemis</emphasis>
sensu
<bibRefCitation id="EFCB4B09FFCEFF8AFBE3FCF5FB727EDE" author="Carle, F. L." box="[1097,1243,871,898]" pageId="8" pageNumber="328" pagination="383 - 424" refId="ref6263" refString="Carle, F. L. (1995) Evolution, taxonomy, and biogeography of ancient Gondwanian libelluloides, with comments on anisopteroid evolution and phylogenetic systematic (Anisoptera: Libelluloidea). Odonatologica, 24 (4), 383 - 424." type="journal article" year="1995">Carle (1995)</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE536F8FFCEFF8AFF6DFC19FE457905" blockId="8.[151,1437,475,2014]" pageId="8" pageNumber="328">
The New Caledonian synthemistids, even if probably rather closely related to true
<taxonomicName id="4C5A4D7BFFCEFF8AFBC4FC19FB497EF9" box="[1134,1248,907,933]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAEAFFCEFF8AFBC4FC19FB497EF9" box="[1134,1248,907,933]" italics="true" pageId="8" pageNumber="328">Synthemis</emphasis>
</taxonomicName>
(ie
<taxonomicName id="4C5A4D7BFFCEFF8AFAA6FC19FEBA7E95" authority="Burmeister" authorityName="Burmeister" authorityYear="1839" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="eustalacta">
<emphasis id="B92EEAEAFFCEFF8AFAA6FC19FA357EF9" box="[1292,1436,907,933]" italics="true" pageId="8" pageNumber="328">S. eustalacta</emphasis>
Burmeister
</taxonomicName>
from South East
<collectingCountry id="F34D7668FFCEFF8AFE4CFC3DFDE47E95" box="[486,589,943,969]" name="Australia" pageId="8" pageNumber="328">Australia</collectingCountry>
and
<taxonomicName id="4C5A4D7BFFCEFF8AFD21FC3DFCB77E95" authorityName="Tillyard" authorityYear="1910" box="[651,798,943,969]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="tasmanica">
<emphasis id="B92EEAEAFFCEFF8AFD21FC3DFCB77E95" box="[651,798,943,969]" italics="true" pageId="8" pageNumber="328">S. tasmanica</emphasis>
</taxonomicName>
, from
<collectingRegion id="499EF81AFFCEFF8AFCDBFC3DFC777E95" box="[881,990,943,969]" country="Australia" name="Tasmania" pageId="8" pageNumber="328">Tasmania</collectingRegion>
) are nevertheless sufficiently different to be removed from this genus. Indeed, the true
<taxonomicName id="4C5A4D7BFFCEFF8AFD62FC41FC937EB1" box="[712,826,979,1005]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAEAFFCEFF8AFD62FC41FC937EB1" box="[712,826,979,1005]" italics="true" pageId="8" pageNumber="328">Synthemis</emphasis>
</taxonomicName>
have, among others, distinctly different male hamuli posteriores, male
<taxonomicName id="4C5A4D7BFFCEFF8AFECBFC65FDB2794D" box="[353,539,1015,1041]" class="Insecta" family="Pneumoridae" genus="Vesica" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="327" phylum="Arthropoda" rank="species" species="spermalis">vesica spermalis</taxonomicName>
<collectionCode id="ED4BAE3DFFCEFF8AFD8EFC65FD94794D" box="[548,573,1015,1041]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="328" type="Museum">V</collectionCode>
1, male organisation of S2 ventral tergum, male epiproct, female ovipositor, FW relative position of arculus, HW anal loop construction, larval frontal plate, larval premental distal margin, and larval mask setal organisation.
</paragraph>
<paragraph id="8BE536F8FFCEFF8AFF6DFBF1FC4B7825" blockId="8.[151,1437,475,2014]" pageId="8" pageNumber="328">
<bibRefCitation id="EFCB4B09FFCEFF8AFF6DFBF1FEF87922" author="Fleck, G." box="[199,337,1123,1150]" pageId="8" pageNumber="328" pagination="165 - 166" refId="ref6308" refString="Fleck, G. (2005) Contribution a la connaissance des Odonates de Nouvelle Caledonie. Notes sur la larve de Synthemis ariadne Lieftinck, 1975 (Anisoptera, Synthemistidae). Bulletin de la Societe Entomologique de France, 110 (2), 165 - 166. https: // doi. org / 10.3406 / bsef. 2005.16214" type="journal article" year="2005">Fleck (2005</bibRefCitation>
, in
<bibRefCitation id="EFCB4B09FFCEFF8AFED0FBF1FD907921" author="Fleck, G. &amp; De Marmels, J. &amp; Theischinger, G." box="[378,569,1123,1149]" pageId="8" pageNumber="328" pagination="263 - 319" refId="ref6370" refString="Fleck, G., De Marmels, J. &amp; Theischinger, G. (2020) Chapter 12. Ordre des Odonata (Odonates ou Demoiselles et Libellules). In: Aberlenc, H. - P. (Ed.), Les Insectes du Monde. Biodiversite. Classification. Cles de determination des familles. Vols. 1 &amp; 2. Versailles, Montpellier &amp; Plaissan, Quae &amp; Museo, pp. 263 - 319 (text) &amp; pp. 102 - 145 (plates). [in French]" type="book chapter" year="2020">
Fleck
<emphasis id="B92EEAEAFFCEFF8AFE15FBF6FE517921" box="[447,504,1123,1149]" italics="true" pageId="8" pageNumber="328">et al.</emphasis>
2020
</bibRefCitation>
) considered New Caledonian synthemistids as
<emphasis id="B92EEAEAFFCEFF8AFBE1FBF1FA9F7921" box="[1099,1334,1123,1149]" italics="true" pageId="8" pageNumber="328">
<taxonomicName id="4C5A4D7BFFCEFF8AFBE1FBF1FB147921" box="[1099,1213,1123,1149]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="genus" sensu="lato">Synthemis</taxonomicName>
<taxonomicNameLabel id="A21D5791FFCEFF8AFB6EFBF6FA9F7921" box="[1220,1334,1123,1149]" pageId="8" pageNumber="328" sensu="lato">sensu lato</taxonomicNameLabel>
</emphasis>
, because the species
<taxonomicName id="4C5A4D7BFFCEFF8AFEB5FB15FE3F79FD" box="[287,406,1159,1185]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="ariadne">
<emphasis id="B92EEAEAFFCEFF8AFEB5FB15FE3F79FD" box="[287,406,1159,1185]" italics="true" pageId="8" pageNumber="328">S. ariadne</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5A4D7BFFCEFF8AFE79FB15FD9679FD" authorityName="Campion" authorityYear="1921" box="[467,575,1159,1185]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="fenella">
<emphasis id="B92EEAEAFFCEFF8AFE79FB15FD9679FD" box="[467,575,1159,1185]" italics="true" pageId="8" pageNumber="328">S. fenella</emphasis>
</taxonomicName>
were not considered belonging to
<taxonomicName id="4C5A4D7BFFCEFF8AFC7DFB15FBD379FD" authorityName="sensu Carle" authorityYear="1995" box="[983,1146,1159,1185]" class="Insecta" genus="Calesynthemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAEAFFCEFF8AFC7DFB15FBD379FD" box="[983,1146,1159,1185]" italics="true" pageId="8" pageNumber="328">Calesynthemis</emphasis>
</taxonomicName>
, and
<bibRefCitation id="EFCB4B09FFCEFF8AFB15FB15FF4B7999" author="Fleck, G. &amp; El Adouzi, M." pageId="8" pageNumber="328" pagination="589 - 594" refId="ref6469" refString="Fleck, G. &amp; El Adouzi, M. (2013) The larva of the genus Palaeosynthemis Forster, 1903 (Odonata: Anisoptera: Synthemistidae) and a generic key to the larvae of non-New Caledonian Synthemistidae. Zootaxa, 3619 (5), 589 - 594. https: // doi. org / 10.11646 / zootaxa. 3619.5.8" type="journal article" year="2013">Fleck &amp; El Adouzi (2013)</bibRefCitation>
did not treat New Caledonian synthemistids and mentioned “New Caledonian synthemistids are in need of a revision and probably to be split into two genera”. In the matter of fact, the species
<taxonomicName id="4C5A4D7BFFCEFF8AFBF4FB5DFB7879B5" box="[1118,1233,1231,1257]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="ariadne">
<emphasis id="B92EEAEAFFCEFF8AFBF4FB5DFB7879B5" box="[1118,1233,1231,1257]" italics="true" pageId="8" pageNumber="328">S. ariadne</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5A4D7BFFCEFF8AFAA3FB5DFADB79B5" authorityName="Campion" authorityYear="1921" box="[1289,1394,1231,1257]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="fenella">
<emphasis id="B92EEAEAFFCEFF8AFAA3FB5DFADB79B5" box="[1289,1394,1231,1257]" italics="true" pageId="8" pageNumber="328">S. fenella</emphasis>
</taxonomicName>
are distinctly different from other
<taxonomicName id="4C5A4D7BFFCEFF8AFE45FB61FD3B7851" authorityName="sensu Carle" authorityYear="1995" box="[495,658,1267,1293]" class="Insecta" genus="Calesynthemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAEAFFCEFF8AFE45FB61FD3B7851" box="[495,658,1267,1293]" italics="true" pageId="8" pageNumber="328">Calesynthemis</emphasis>
</taxonomicName>
, not only by virtue of their smaller size, but for example their larvae are remarkable and unique among synthemistids by the absence of long body setae giving them a strong glabrous and finely granulose aspect, and by the absence of dentition on the labial palps. The following characters permit the separation of
<taxonomicName id="4C5A4D7BFFCEFF8AFE9AFACDFE317825" authorityName="Campion" authorityYear="1921" box="[304,408,1375,1401]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="fenella">
<emphasis id="B92EEAEAFFCEFF8AFE9AFACDFE317825" box="[304,408,1375,1401]" italics="true" pageId="8" pageNumber="328">S. fenella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5A4D7BFFCEFF8AFE64FACDFDE87825" box="[462,577,1375,1401]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="ariadne">
<emphasis id="B92EEAEAFFCEFF8AFE64FACDFDE87825" box="[462,577,1375,1401]" italics="true" pageId="8" pageNumber="328">S. ariadne</emphasis>
</taxonomicName>
from other New Caledonian species:
</paragraph>
<paragraph id="8BE536F8FFCEFF8AFF6DFA16FAC278B9" blockId="8.[151,1437,475,2014]" pageId="8" pageNumber="328">
- All discoidal triangles free. They are generally free in
<taxonomicName id="4C5A4D7BFFCEFF8AFC83FA11FC3978C1" authorityName="Campion" authorityYear="1921" box="[809,912,1411,1437]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="fenella">
<emphasis id="B92EEAEAFFCEFF8AFC83FA11FC3978C1" box="[809,912,1411,1437]" italics="true" pageId="8" pageNumber="328">S. fenella</emphasis>
</taxonomicName>
, in
<taxonomicName id="4C5A4D7BFFCEFF8AFC12FA11FB8378C1" box="[952,1066,1411,1437]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="ariadne">
<emphasis id="B92EEAEAFFCEFF8AFC12FA11FB8378C1" box="[952,1066,1411,1437]" italics="true" pageId="8" pageNumber="328">S. ariadne</emphasis>
</taxonomicName>
rarely crossed in one or even two wings. In other New Caledonian representatives discoidal triangles are crossed at least in one pair of wings (usually on both pair of wings). HW discoidal triangle is generally free in male
<taxonomicName id="4C5A4D7BFFCEFF8AFC1BFA5EFB9878B9" authorityName="Selys" authorityYear="1871" box="[945,1073,1483,1510]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="miranda">
<emphasis id="B92EEAEAFFCEFF8AFC1BFA5EFB9878B9" box="[945,1073,1483,1510]" italics="true" pageId="8" pageNumber="328">C. miranda</emphasis>
</taxonomicName>
and occasionally in female.
</paragraph>
<paragraph id="8BE536F8FFCEFF8AFF6DFA62FDB57BC5" blockId="8.[151,1437,475,2014]" pageId="8" pageNumber="328">
- Arculus not strongly shifted distal to Ax2. In
<taxonomicName id="4C5A4D7BFFCEFF8AFD40FA7DFCFD7B55" authorityName="Campion" authorityYear="1921" box="[746,852,1519,1545]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="fenella">
<emphasis id="B92EEAEAFFCEFF8AFD40FA7DFCFD7B55" box="[746,852,1519,1545]" italics="true" pageId="8" pageNumber="328">S. fenella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5A4D7BFFCEFF8AFC3AFA7DFBAF7B55" box="[912,1030,1519,1545]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="ariadne">
<emphasis id="B92EEAEAFFCEFF8AFC3AFA7DFBAF7B55" box="[912,1030,1519,1545]" italics="true" pageId="8" pageNumber="328">S. ariadne</emphasis>
</taxonomicName>
the arculus is often placed slightly proximal to Ax2 or at Ax2, it is regularly shifted slightly distally (rarely distinctly in HW). In most other species, except
<taxonomicName id="4C5A4D7BFFCEFF8AFF4CF9AAFED37B0D" authorityName="Campion" authorityYear="1921" box="[230,378,1591,1618]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="flexicauda">
<emphasis id="B92EEAEAFFCEFF8AFF4CF9AAFED37B0D" box="[230,378,1591,1618]" italics="true" pageId="8" pageNumber="328">C. flexicauda</emphasis>
</taxonomicName>
, the origin of arculus is distinctly shifted distal to Ax2, most of the time in both pair of wings. In
<taxonomicName id="4C5A4D7BFFCEFF8AFF3DF9CEFE857B29" authorityName="Campion" authorityYear="1921" box="[151,300,1627,1654]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="flexicauda">
<emphasis id="B92EEAEAFFCEFF8AFF3DF9CEFE857B29" box="[151,300,1627,1654]" italics="true" pageId="8" pageNumber="328">C. flexicauda</emphasis>
</taxonomicName>
it is placed at Ax2 or slightly distally shifted. Note that in some
<taxonomicName id="4C5A4D7BFFCEFF8AFC45F9C9FB7B7B2A" authority="Forster" authorityName="Forster" authorityYear="1903" box="[1007,1234,1627,1654]" class="Insecta" family="Synthemistidae" genus="Eusynthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAEAFFCEFF8AFC45F9C9FBD47B29" box="[1007,1149,1627,1653]" italics="true" pageId="8" pageNumber="328">Eusynthemis</emphasis>
Förster
</taxonomicName>
and allied genera, the arculus is shifted distal to Ax2.
</paragraph>
<paragraph id="8BE536F8FFCEFF8AFF6DF936FB277BBE" blockId="8.[151,1437,475,2014]" pageId="8" pageNumber="328">
- Male cerci not longer than S9+10. In
<taxonomicName id="4C5A4D7BFFCEFF8AFD20F931FD5A7BE1" authorityName="Campion" authorityYear="1921" box="[650,755,1699,1725]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="fenella">
<emphasis id="B92EEAEAFFCEFF8AFD20F931FD5A7BE1" box="[650,755,1699,1725]" italics="true" pageId="8" pageNumber="328">S. fenella</emphasis>
</taxonomicName>
they are about as long as S9+10 and in
<taxonomicName id="4C5A4D7BFFCEFF8AFB6EF931FA917BE1" box="[1220,1336,1699,1725]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="ariadne">
<emphasis id="B92EEAEAFFCEFF8AFB6EF931FA917BE1" box="[1220,1336,1699,1725]" italics="true" pageId="8" pageNumber="328">S. ariadne</emphasis>
</taxonomicName>
they are slightly shorter than S9+10. In other species they are longer, most of the time distinctly so.
</paragraph>
<paragraph id="8BE536F8FFCEFF8AFF6DF97EFB0C7A76" blockId="8.[151,1437,475,2014]" pageId="8" pageNumber="328">
- Male penis dh1 and dh2 placed on a common stem (
<figureCitation id="13612A7DFFCEFF8AFC90F979FC267A59" box="[826,911,1771,1797]" captionStart="FIGURES 2224. 22" captionStartId="7.[152,270,1259,1283]" captionTargetBox="[162,1436,777,1236]" captionTargetId="figure-298@7.[151,1436,777,1236]" captionTargetPageId="7" captionText="FIGURES 2224. 22. Comparison of paratype female Calesynthemis jeanlegrandi sp. nov. (left) and female Calesynthemis serendipita (right). 23. Neocaledosynthemis gen. nov. N. fenella. Vesica spermalis (or penis) in right lateral view.Arrow indicates common stem of dh1 and dh2. 24. Synthemis tasmanica. Vesica spermalis (or penis). V3 and V4 in right lateral view. Note the remarkable similarity with V3 and V4 of Neocaledosynthemis fenella gen. nov." figureDoi="http://doi.org/10.5281/zenodo.10561958" httpUri="https://zenodo.org/record/10561958/files/figure.png" pageId="8" pageNumber="328">Fig. 23</figureCitation>
, see arrow, compare with
<figureCitation id="13612A7DFFCEFF8AFB15F979FAAC7A59" box="[1215,1285,1771,1797]" captionStart="FIGURES 610" captionStartId="3.[152,270,1548,1572]" captionTargetBox="[151,1435,489,1525]" captionTargetId="figure-182@3.[151,1435,489,1525]" captionTargetPageId="3" captionText="FIGURES 610. Calesynthemis jeanlegrandi sp. nov., male holotype. Vesica spermalis (or penis). 6. Left general view. 7. Ventral general view. 8. Detail of V2V4 in right lateral view. 9. Detail of V3 in ventral view. 10. Detail of V3V4 in anteroventral view. Note the strongly divergent branches of dh1." figureDoi="http://doi.org/10.5281/zenodo.10561950" httpUri="https://zenodo.org/record/10561950/files/figure.png" pageId="8" pageNumber="328">Fig. 8</figureCitation>
). In all other species they are distinctly separated or approximate on
<collectionCode id="ED4BAE3DFFCEFF8AFCA9F89DFCB07A75" box="[771,793,1807,1833]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="328" type="Museum">V</collectionCode>
4, never placed on a common stem.
</paragraph>
<paragraph id="8BE536F8FFCEFF8BFF6DF8A6FE997CD5" blockId="8.[151,1437,475,2014]" lastBlockId="9.[151,1437,151,1077]" lastPageId="9" lastPageNumber="329" pageId="8" pageNumber="328">
- Female
<collectionCode id="ED4BAE3DFFCEFF8AFE95F8A1FEFE7A11" box="[319,343,1843,1869]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="328" type="Museum">V</collectionCode>
1 (vulvar lamina) short and stout with U-shaped medial notch. In New Caledonian female synthemistids the
<collectionCode id="ED4BAE3DFFCEFF8AFEF5F8C5FEDE7A2D" box="[351,375,1879,1905]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="328" type="Museum">V</collectionCode>
1 are in their proximal part parallel and contiguous, in their distal part divergent, the notch being V-shaped or U-shaped; the base of the notch presents additionally a pair of denticles. In
<taxonomicName id="4C5A4D7BFFCEFF8AFB2FF8E9FB467AC9" authorityName="Campion" authorityYear="1921" box="[1157,1263,1915,1941]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="fenella">
<emphasis id="B92EEAEAFFCEFF8AFB2FF8E9FB467AC9" box="[1157,1263,1915,1941]" italics="true" pageId="8" pageNumber="328">S. fenella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5A4D7BFFCEFF8AFA82F8E9FA357AC9" box="[1320,1436,1915,1941]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="ariadne">
<emphasis id="B92EEAEAFFCEFF8AFA82F8E9FA357AC9" box="[1320,1436,1915,1941]" italics="true" pageId="8" pageNumber="328">S. ariadne</emphasis>
</taxonomicName>
<collectionCode id="ED4BAE3DFFCEFF8AFF3DF80DFF067AE5" box="[151,175,1951,1977]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="328" type="Museum">V</collectionCode>
1 is restricted to the basal 1/3 to basal 1/4 of S9 sternum (contra
<bibRefCitation id="EFCB4B09FFCEFF8AFCD3F80DFC557AE6" author="Carle, F. L." box="[889,1020,1951,1978]" pageId="8" pageNumber="328" pagination="383 - 424" refId="ref6263" refString="Carle, F. L. (1995) Evolution, taxonomy, and biogeography of ancient Gondwanian libelluloides, with comments on anisopteroid evolution and phylogenetic systematic (Anisoptera: Libelluloidea). Odonatologica, 24 (4), 383 - 424." type="journal article" year="1995">Carle, 1995</bibRefCitation>
) (and also shorter than 1/2 the length of S10, contra
<bibRefCitation id="EFCB4B09FFCEFF8AFEE8F851FE6E7A82" author="Carle, F. L." box="[322,455,1987,2014]" pageId="8" pageNumber="328" pagination="383 - 424" refId="ref6263" refString="Carle, F. L. (1995) Evolution, taxonomy, and biogeography of ancient Gondwanian libelluloides, with comments on anisopteroid evolution and phylogenetic systematic (Anisoptera: Libelluloidea). Odonatologica, 24 (4), 383 - 424." type="journal article" year="1995">Carle, 1995</bibRefCitation>
) with proximal part strongly reduced, and notch U-shaped in
<taxonomicName id="4C5A4D7BFFCEFF8AFB27F851FB5E7A81" authorityName="Campion" authorityYear="1921" box="[1165,1271,1987,2013]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="fenella">
<emphasis id="B92EEAEAFFCEFF8AFB27F851FB5E7A81" box="[1165,1271,1987,2013]" italics="true" pageId="8" pageNumber="328">S. fenella</emphasis>
</taxonomicName>
, and probably derived from the
<taxonomicName id="4C5A4D7BFFCFFF8BFEF7FF05FE6E7DED" authorityName="Campion" authorityYear="1921" box="[349,455,151,177]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="fenella">
<emphasis id="B92EEAEAFFCFFF8BFEF7FF05FE6E7DED" box="[349,455,151,177]" italics="true" pageId="9" pageNumber="329">S. fenella</emphasis>
</taxonomicName>
<typeStatus id="54E1885AFFCFFF8BFE7AFF05FDA97DED" box="[464,512,151,177]" pageId="9" pageNumber="329">type</typeStatus>
in
<taxonomicName id="4C5A4D7BFFCFFF8BFD8DFF05FD327DED" box="[551,667,151,177]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="ariadne">
<emphasis id="B92EEAEAFFCFFF8BFD8DFF05FD327DED" box="[551,667,151,177]" italics="true" pageId="9" pageNumber="329">S. ariadne</emphasis>
</taxonomicName>
since distal part is similar to that of
<taxonomicName id="4C5A4D7BFFCFFF8BFBEBFF05FB027DED" authorityName="Campion" authorityYear="1921" box="[1089,1195,151,177]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="fenella">
<emphasis id="B92EEAEAFFCFFF8BFBEBFF05FB027DED" box="[1089,1195,151,177]" italics="true" pageId="9" pageNumber="329">S. fenella</emphasis>
</taxonomicName>
but basal part shows shortly divergent inner margins (apparently unique within New Caledonian synthemistids). In other species, except
<taxonomicName id="4C5A4D7BFFCFFF8BFF3DFF4DFE9F7DA5" box="[151,310,223,249]" class="Insecta" genus="Calesynthemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="serendipita">
<emphasis id="B92EEAEAFFCFFF8BFF3DFF4DFE9F7DA5" box="[151,310,223,249]" italics="true" pageId="9" pageNumber="329">C. serendipita</emphasis>
</taxonomicName>
, the
<collectionCode id="ED4BAE3DFFCFFF8BFEDAFF4DFE217DA5" box="[368,392,223,249]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="9" pageNumber="329" type="Museum">V</collectionCode>
1 are overlapping about 1/2 of S9 sternum, their proximal part is rather long and their notch V-shaped. In
<taxonomicName id="4C5A4D7BFFCFFF8BFE9BFE91FE787C41" box="[305,465,259,285]" class="Insecta" genus="Calesynthemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="serendipita">
<emphasis id="B92EEAEAFFCFFF8BFE9BFE91FE787C41" box="[305,465,259,285]" italics="true" pageId="9" pageNumber="329">C. serendipita</emphasis>
</taxonomicName>
the ovipositor is short, hardly longer than 1/3 of S9 (contra
<bibRefCitation id="EFCB4B09FFCFFF8BFB26FE91FAA57C41" author="Carle, F. L." box="[1164,1292,258,285]" pageId="9" pageNumber="329" pagination="383 - 424" refId="ref6263" refString="Carle, F. L. (1995) Evolution, taxonomy, and biogeography of ancient Gondwanian libelluloides, with comments on anisopteroid evolution and phylogenetic systematic (Anisoptera: Libelluloidea). Odonatologica, 24 (4), 383 - 424." type="journal article" year="1995">Carle 1995</bibRefCitation>
), with notch somewhat U-shaped, but with basal part having valves contiguous over a relatively long distance (considering the distal part only, probably a convergence with
<taxonomicName id="4C5A4D7BFFCFFF8BFD3AFED9FD517C39" authorityName="Campion" authorityYear="1921" box="[656,760,331,357]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="fenella">
<emphasis id="B92EEAEAFFCFFF8BFD3AFED9FD517C39" box="[656,760,331,357]" italics="true" pageId="9" pageNumber="329">S. fenella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5A4D7BFFCFFF8BFC87FED9FC367C39" box="[813,927,331,357]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="ariadne">
<emphasis id="B92EEAEAFFCFFF8BFC87FED9FC367C39" box="[813,927,331,357]" italics="true" pageId="9" pageNumber="329">S. ariadne</emphasis>
</taxonomicName>
given the close affinity of
<taxonomicName id="4C5A4D7BFFCFFF8BFB6DFED9FACD7C39" box="[1223,1380,331,357]" class="Insecta" genus="Calesynthemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="serendipita">
<emphasis id="B92EEAEAFFCFFF8BFB6DFED9FACD7C39" box="[1223,1380,331,357]" italics="true" pageId="9" pageNumber="329">C. serendipita</emphasis>
</taxonomicName>
with
<taxonomicName id="4C5A4D7BFFCFFF8BFF3DFEFDFE897CD4" authorityName="Lieftinck" authorityYear="1971" box="[151,288,367,393]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="campioni">
<emphasis id="B92EEAEAFFCFFF8BFF3DFEFDFE897CD4" box="[151,288,367,393]" italics="true" pageId="9" pageNumber="329">C. campioni</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BE536F8FFCFFF8BFF6DFE01FDBC7CA9" blockId="9.[151,1437,151,1077]" pageId="9" pageNumber="329">
- Female cerci not longer than S9+10. In
<taxonomicName id="4C5A4D7BFFCFFF8BFD31FE01FCAD7CF1" authorityName="Campion" authorityYear="1921" box="[667,772,403,429]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="fenella">
<emphasis id="B92EEAEAFFCFFF8BFD31FE01FCAD7CF1" box="[667,772,403,429]" italics="true" pageId="9" pageNumber="329">S. fenella</emphasis>
</taxonomicName>
they are about as long as S9, in
<taxonomicName id="4C5A4D7BFFCFFF8BFBDDFE01FB427CF1" box="[1143,1259,403,429]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="ariadne">
<emphasis id="B92EEAEAFFCFFF8BFBDDFE01FB427CF1" box="[1143,1259,403,429]" italics="true" pageId="9" pageNumber="329">S. ariadne</emphasis>
</taxonomicName>
slightly shorter than S9+10. In other species they are generally distinctly longer than S9+10 except
<taxonomicName id="4C5A4D7BFFCFFF8BFBEDFE25FB6E7C8D" authorityName="Selys" authorityYear="1871" box="[1095,1223,439,465]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="miranda">
<emphasis id="B92EEAEAFFCFFF8BFBEDFE25FB6E7C8D" box="[1095,1223,439,465]" italics="true" pageId="9" pageNumber="329">C. miranda</emphasis>
</taxonomicName>
where they appear up turned and shorter than S9+10.
</paragraph>
<paragraph id="8BE536F8FFCFFF8BFF6DFE6DFB037F3D" blockId="9.[151,1437,151,1077]" pageId="9" pageNumber="329">- Larva glabrous, deprived of long setae and long hair-like setae and with fine granulose appearance (unique character within synthemistids). All other larvae present long setae and/or long hair-like setae at least on frontal plate, occipital lateral margins, pronotal lateral lobes, lateral margins of thorax and abdomen.</paragraph>
<paragraph id="8BE536F8FFCFFF8BFF6DFDF9FD8D7FF5" blockId="9.[151,1437,151,1077]" pageId="9" pageNumber="329">- Larva with distal margin of labial palp lacking clear dentition (a unique character within synthemistids). All other larvae have distinct dentition.</paragraph>
<paragraph id="8BE536F8FFCFFF8BFF6DFD21FC637EF9" blockId="9.[151,1437,151,1077]" pageId="9" pageNumber="329">
- Larva with only 6 or fewer antennomeres (apparently unique within synthemistids). Contrary to the claim of
<bibRefCitation id="EFCB4B09FFCFFF8BFF3DFD45FEFA7FAD" author="Lieftinck, M. A." box="[151,339,727,753]" pageId="9" pageNumber="329" pagination="165 - 200" refId="ref6673" refString="Lieftinck, M. A. (1976) The dragonflies odonata of New Caledonia and the Loyalty Islands: Part 2. Immature stages. Cahiers Office de la Recherche Scientifique et Technique Outre-Mer (ORSTOM), Serie Hydrobiologie, 10 (3), 165 - 200." type="journal article" year="1976">Lieftinck (1976)</bibRefCitation>
, from 12 F-0 and 2 F-1 checked
<taxonomicName id="4C5A4D7BFFCFFF8BFD63FD45FC9B7FAD" authorityName="Campion" authorityYear="1921" box="[713,818,727,753]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="fenella">
<emphasis id="B92EEAEAFFCFFF8BFD63FD45FC9B7FAD" box="[713,818,727,753]" italics="true" pageId="9" pageNumber="329">S. fenella</emphasis>
</taxonomicName>
larvae from different localities, only one F-0 seems to have 7 antennomeres on one antenna (moreover the character is not clearly cut since the interantennomeral joint is not clearly marked!);
<specimenCount id="9D5CFD71FFCFFF8BFE1AFC8DFDCE7E65" box="[432,615,799,825]" count="3" pageId="9" pageNumber="329" type="generic">three specimens</specimenCount>
exhibit 5 antennomeres in a least one antenna. All
<taxonomicName id="4C5A4D7BFFCFFF8BFB6AFC8DFA9F7E65" box="[1216,1334,799,825]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="ariadne">
<emphasis id="B92EEAEAFFCFFF8BFB6AFC8DFA9F7E65" box="[1216,1334,799,825]" italics="true" pageId="9" pageNumber="329">S. ariadne</emphasis>
</taxonomicName>
checked larvae (6 F-0 and 2 F-1) have 6 antennomeres. Thus,
<taxonomicName id="4C5A4D7BFFCFFF8BFD5CFCD1FCF67E01" authorityName="Campion" authorityYear="1921" box="[758,863,835,861]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="fenella">
<emphasis id="B92EEAEAFFCFFF8BFD5CFCD1FCF67E01" box="[758,863,835,861]" italics="true" pageId="9" pageNumber="329">S. fenella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5A4D7BFFCFFF8BFC32FCD1FBA57E01" box="[920,1036,835,861]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="ariadne">
<emphasis id="B92EEAEAFFCFFF8BFC32FCD1FBA57E01" box="[920,1036,835,861]" italics="true" pageId="9" pageNumber="329">S. ariadne</emphasis>
</taxonomicName>
larvae can be considered to have 6 antennomeres (sometimes with one less, very exceptionally, and aberrantly, with one more). All other species have 7 antennomeres, and antennae, comparatively to body dimension, longer.
</paragraph>
<paragraph id="8BE536F8FFCFFF8BFF6DFC3DFCEE7EB1" blockId="9.[151,1437,151,1077]" pageId="9" pageNumber="329">- Larva with only 2 long premental setae and 2 or 3 palpal setae (apparently unique within synthemistids). Other larvae have at least 3 long premental setae and 4 palpal setae.</paragraph>
<paragraph id="8BE536F8FFCFFF8BFF78FC65FAA87969" blockId="9.[151,1437,151,1077]" pageId="9" pageNumber="329">
Considering all the above-mentioned characters, any of them unique among synthemistids and considered synapomorphic,
<taxonomicName id="4C5A4D7BFFCFFF8BFEF8FB89FE137969" authorityName="Campion" authorityYear="1921" box="[338,442,1051,1077]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="fenella">
<emphasis id="B92EEAEAFFCFFF8BFEF8FB89FE137969" box="[338,442,1051,1077]" italics="true" pageId="9" pageNumber="329">S. fenella</emphasis>
</taxonomicName>
(
<typeStatus id="54E1885AFFCFFF8BFE63FB89FE527969" box="[457,507,1051,1077]" pageId="9" pageNumber="329">type</typeStatus>
species) and
<taxonomicName id="4C5A4D7BFFCFFF8BFD38FB89FCAC7969" box="[658,773,1051,1077]" class="Insecta" family="Synthemistidae" genus="Synthemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="ariadne">
<emphasis id="B92EEAEAFFCFFF8BFD38FB89FCAC7969" box="[658,773,1051,1077]" italics="true" pageId="9" pageNumber="329">S. ariadne</emphasis>
</taxonomicName>
are placed in the genus
<taxonomicName id="4C5A4D7BFFCFFF8BFBBEFB89FB537969" authorityName="Fleck" authorityYear="2024" box="[1044,1274,1051,1077]" class="Insecta" family="Synthemistidae" genus="Neocaledosynthemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="329" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAEAFFCFFF8BFBBEFB89FB537969" box="[1044,1274,1051,1077]" italics="true" pageId="9" pageNumber="329">Neocaledosynthemis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>