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<mods:title id="1431A8FC4D03C2899DADC1574C8B07F5">On the subgenus Eurysunius Reitter of Turkey IX. Three new species from central northern Anatolia, with an identification key to Turkish species (Coleoptera: Staphylinidae: Paederinae, Astenus)</mods:title>
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<mods:date id="53FA69E394BD023187B117BB67AB3629">2024</mods:date>
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<subSubSection id="C3406547FF8A356CFF6CFAE9FCDC1948" box="[151,843,1329,1356]" pageId="3" pageNumber="71" type="nomenclature">
<paragraph id="8BE536CCFF8A356CFF6CFAE9FCDC1948" blockId="3.[151,843,1329,1356]" box="[151,843,1329,1356]" pageId="3" pageNumber="71">
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<taxonomicName id="4C5A4D4FFF8A356CFF6CFAE9FD9F194F" authorityName="Anlaş" authorityYear="2024" box="[151,520,1329,1356]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="71" phylum="Arthropoda" rank="species" species="hititicus" status="sp. nov." subGenus="Eurysunius">
<emphasis id="B92EEADEFF8A356CFF6CFAE9FF60194F" bold="true" box="[151,247,1330,1356]" italics="true" pageId="3" pageNumber="71">Astenus</emphasis>
(
<emphasis id="B92EEADEFF8A356CFEF3FAEAFE031948" bold="true" box="[264,404,1329,1355]" italics="true" pageId="3" pageNumber="71">Eurysunius</emphasis>
)
<emphasis id="B92EEADEFF8A356CFE5DFAEAFD9F194F" bold="true" box="[422,520,1329,1356]" italics="true" pageId="3" pageNumber="71">hititicus</emphasis>
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<taxonomicNameLabel id="A21D57A5FF8A356CFDF4FAE9FDC6194F" box="[527,593,1330,1356]" pageId="3" pageNumber="71" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="13612A49FF8A356CFD9AFAEAFD731948" box="[609,740,1329,1355]" captionStart="FIGURES 916" captionStartId="3.[151,264,1151,1175]" captionTargetBox="[427,1165,185,1121]" captionTargetId="figure-41@3.[422,1166,181,1127]" captionTargetPageId="3" captionText="FIGURES 916. Astenus (Eurysunius) hititicus sp. n. 9—habitus; 10—forebody; 11—Male sternite VII; 12—Male sternite VIII; 13, 14—Aedeagus in lateral view; 15, 16—Aedeagus in ventral view. Scale bars: 0.5 mm (Figs. 910), 0.2 mm (Fig. 1116)." figureDoi="http://doi.org/10.5281/zenodo.12526989" httpUri="https://zenodo.org/record/12526989/files/figure.png" pageId="3" pageNumber="71">Figs. 916</figureCitation>
;
<figureCitation id="13612A49FF8A356CFD0AFAE9FCD61948" box="[753,833,1329,1356]" captionStart="MAP 1" captionStartId="6.[151,207,1425,1449]" captionTargetBox="[151,1436,489,1401]" captionTargetId="figure-135@6.[151,1436,489,1401]" captionText="MAP 1. Distribution of Astenus (Eurysunius) species in Turkey (28 species)" figureDoi="http://doi.org/10.5281/zenodo.12547399" httpUri="https://zenodo.org/record/12547399/files/figure.png" pageId="3" pageNumber="71">Map 1</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3406547FF8A356CFF6CFAA1FCF11A6F" pageId="3" pageNumber="71" type="materials_examined">
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<emphasis id="B92EEADEFF8A356CFF6CFAA1FED21990" bold="true" box="[151,325,1401,1428]" pageId="3" pageNumber="71">Type material.</emphasis>
<materialsCitation id="3B323C91FF8A356CFEB7FAA1FEEE19BB" ID-GBIF-Occurrence="4899575311" collectingDate="2022-05-09" collectorName="Kacar &amp; Celik." country="Turkey" elevation="1112" latitude="40.371666" location="Cagsak" longLatPrecision="19" longitude="34.52861" pageId="3" pageNumber="71" specimenCount="1" specimenCount-male="1" stateProvince="Corum" typeStatus="holotype">
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<typeStatus id="54E1886EFF8A356CFEB7FAA1FE2A1997" box="[332,445,1402,1428]" pageId="3" pageNumber="71" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F34D765CFF8A356CFE33FAA1FDD51997" box="[456,578,1402,1428]" name="Turkey" pageId="3" pageNumber="71">TURKEY</collectingCountry>
:
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,
<collectingRegion id="499EF82EFF8A356CFD80FAA1FD461997" box="[635,721,1402,1428]" country="Turkey" name="Corum" pageId="3" pageNumber="71">Çorum</collectingRegion>
,
<location id="8E856017FF8A356CFD26FAA1FCB81997" LSID="urn:lsid:plazi:treatment:03F387DAFF8A356BFF6CFAE9FF201956:8E856017FF8A356CFD26FAA1FCB81997" box="[733,815,1402,1428]" country="Turkey" latitude="40.371666" longLatPrecision="19" longitude="34.52861" name="Cagsak" pageId="3" pageNumber="71" stateProvince="Corum">Çağşak</location>
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SE,
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,
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,
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,
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<collectingDate id="EFA0E9E4FF8A356CFAD8FAA1FA0F1997" box="[1315,1432,1402,1428]" pageId="3" pageNumber="71" value="2022-05-09">09.V.2022</collectingDate>
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, leg.
<collectorName id="26AF531AFF8A356CFF32FA45FE9B19BB" box="[201,268,1438,1464]" pageId="3" pageNumber="71">Kacar</collectorName>
&amp;
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/
<materialsCitation id="3B323C91FF8A356CFE69FA45FE9219DF" ID-GBIF-Occurrence="4899575306" collectionCode="AZMM" country="Turkey" determinerName="S. Anlas" pageId="3" pageNumber="71" specimenCount="1" specimenCount-male="1" stateProvince="Corum" typeStatus="holotype">
<typeStatus id="54E1886EFF8A356CFE69FA45FD9019BB" box="[402,519,1438,1464]" pageId="3" pageNumber="71" type="holotype">Holotypus</typeStatus>
<specimenCount id="9D5CFD45FF8A356CFDEBFA45FDB019BB" box="[528,551,1438,1464]" count="1" pageId="3" pageNumber="71" type="male"></specimenCount>
,
<taxonomicName id="4C5A4D4FFF8A356CFDCEFA45FC1F19BB" authorityName="Anlaş" authorityYear="2024" box="[565,904,1438,1464]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="71" phylum="Arthropoda" rank="species" species="hititicus" status="sp. nov." subGenus="Eurysunius">
<emphasis id="B92EEADEFF8A356CFDCEFA45FD1B19BB" box="[565,652,1438,1464]" italics="true" pageId="3" pageNumber="71">Astenus</emphasis>
(
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)
<emphasis id="B92EEADEFF8A356CFCD5FA45FC1F19BB" box="[814,904,1438,1464]" italics="true" pageId="3" pageNumber="71">hititicus</emphasis>
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<taxonomicNameLabel id="A21D57A5FF8A356CFC69FA45FC4719BB" box="[914,976,1438,1464]" pageId="3" pageNumber="71" rank="species">sp. n.</taxonomicNameLabel>
det.
<determinerName id="7E39DA08FF8A356CFBF0FA45FBFD19BB" box="[1035,1130,1438,1464]" pageId="3" pageNumber="71">S. Anlaş</determinerName>
2024” &lt;red printed label&gt; (
<collectionCode id="ED4BAE09FF8A356CFF64FA19FF6A19DF" box="[159,253,1474,1500]" pageId="3" pageNumber="71">AZMM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B323C91FF8A356CFEEAFA19FC9919DF" ID-GBIF-Occurrence="4899575316" box="[273,782,1474,1500]" collectionCode="AZMM" country="Turkey" determinerName="S. Anlas" pageId="3" pageNumber="71" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Corum" typeStatus="paratype">
<typeStatus id="54E1886EFF8A356CFEEAFA19FE1519DF" box="[273,386,1474,1500]" pageId="3" pageNumber="71" type="paratype">Paratypes</typeStatus>
:
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,
<specimenCount id="9D5CFD45FF8A356CFE38FA19FE6B19DF" box="[451,508,1474,1500]" count="3" pageId="3" pageNumber="71" type="female">3♀♀</specimenCount>
, same data as holotype
</materialsCitation>
.
<materialsCitation id="3B323C91FF8A356CFCE1FA19FD231A03" ID-GBIF-Occurrence="4899575309" collectingDate="2022-04-03" collectorName="Orgel &amp; Kacar &amp; Celik." country="Turkey" elevation="1100" latitude="40.35611" location="Cagsak" longLatPrecision="19" longitude="34.563053" pageId="3" pageNumber="71" specimenCount="1" specimenCount-female="1" stateProvince="Corum" typeStatus="paratype">
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,
<collectingRegion id="499EF82EFF8A356CFCB5FA19FC0819DF" box="[846,927,1474,1500]" country="Turkey" name="Corum" pageId="3" pageNumber="71">Çorum</collectingRegion>
,
<location id="8E856017FF8A356CFC57FA19FC6919DF" LSID="urn:lsid:plazi:treatment:03F387DAFF8A356BFF6CFAE9FF201956:8E856017FF8A356CFC57FA19FC6919DF" box="[940,1022,1474,1500]" country="Turkey" latitude="40.35611" longLatPrecision="19" longitude="34.563053" name="Cagsak" pageId="3" pageNumber="71" stateProvince="Corum">Çağşak</location>
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NE,
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,
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,
<quantity id="4CA29B29FF8A356CFF6CFA3DFF791A03" box="[151,238,1510,1536]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="71" unit="m" value="1100.0">
<elevation id="0077D1FFFF8A356CFF6CFA3DFF791A03" box="[151,238,1510,1536]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="71" unit="m" value="1100.0">1100 m</elevation>
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,
<date id="FFE4100CFF8A356CFF00FA3DFEED1A03" box="[251,378,1510,1536]" pageId="3" pageNumber="71" value="2022-04-03">
<collectingDate id="EFA0E9E4FF8A356CFF00FA3DFEED1A03" box="[251,378,1510,1536]" pageId="3" pageNumber="71" value="2022-04-03">03.IV.2022</collectingDate>
</date>
, leg.
<collectorName id="26AF531AFF8A356CFE43FA3CFE6D1A01" box="[440,506,1511,1538]" pageId="3" pageNumber="71">Örgel</collectorName>
,
<collectorName id="26AF531AFF8A356CFDFCFA3DFDDD1A03" box="[519,586,1510,1536]" pageId="3" pageNumber="71">Kacar</collectorName>
&amp;
<collectorName id="26AF531AFF8A356CFD8BFA3DFD231A03" box="[624,692,1510,1536]" pageId="3" pageNumber="71">Çelik.</collectorName>
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<materialsCitation id="3B323C91FF8A356CFD47FA3DFDCD1A27" ID-GBIF-Occurrence="4899575304" collectingDate="2022-04-03" collectorName="Orgel &amp; Kacar &amp; Celik." country="Turkey" elevation="1095" latitude="40.35694" location="Cagsak" longLatPrecision="19" longitude="34.59778" pageId="3" pageNumber="71" specimenCount="1" specimenCount-female="1" stateProvince="Corum" typeStatus="paratype">
<specimenCount id="9D5CFD45FF8A356CFD47FA3DFD481A03" box="[700,735,1510,1536]" count="1" pageId="3" pageNumber="71" type="female">1♀</specimenCount>
,
<collectingRegion id="499EF82EFF8A356CFD15FA3DFCA81A03" box="[750,831,1510,1536]" country="Turkey" name="Corum" pageId="3" pageNumber="71">Çorum</collectingRegion>
,
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SW,
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,
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,
<quantity id="4CA29B29FF8A356CFAC4FA3EFA001A03" box="[1343,1431,1509,1536]" metricMagnitude="3" metricUnit="m" metricValue="1.095" pageId="3" pageNumber="71" unit="m" value="1095.0">
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,
<date id="FFE4100CFF8A356CFF6CF9D1FE821A27" box="[151,277,1546,1572]" pageId="3" pageNumber="71" value="2022-04-03">
<collectingDate id="EFA0E9E4FF8A356CFF6CF9D1FE821A27" box="[151,277,1546,1572]" pageId="3" pageNumber="71" value="2022-04-03">03.IV.2022</collectingDate>
</date>
, leg.
<collectorName id="26AF531AFF8A356CFEA3F9D0FE0D1A25" box="[344,410,1547,1574]" pageId="3" pageNumber="71">Örgel</collectorName>
,
<collectorName id="26AF531AFF8A356CFE52F9D1FE7B1A27" box="[425,492,1546,1572]" pageId="3" pageNumber="71">Kacar</collectorName>
&amp;
<collectorName id="26AF531AFF8A356CFDEDF9D1FDCD1A27" box="[534,602,1546,1572]" pageId="3" pageNumber="71">Çelik.</collectorName>
</materialsCitation>
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,
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,
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,
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NE,
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,
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,
<quantity id="4CA29B29FF8A356CFAC4F9D1FA001A27" box="[1343,1431,1546,1572]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="3" pageNumber="71" unit="m" value="1110.0">
<elevation id="0077D1FFFF8A356CFAC4F9D1FA001A27" box="[1343,1431,1546,1572]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="3" pageNumber="71" unit="m" value="1110.0">1110 m</elevation>
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,
<date id="FFE4100CFF8A356CFF6CF9F5FE811A4B" box="[151,278,1582,1608]" pageId="3" pageNumber="71" value="2022-04-14">
<collectingDate id="EFA0E9E4FF8A356CFF6CF9F5FE811A4B" box="[151,278,1582,1608]" pageId="3" pageNumber="71" value="2022-04-14">14.IV.2022</collectingDate>
</date>
, leg.
<collectorName id="26AF531AFF8A356CFEB4F9F5FE051A4B" box="[335,402,1582,1608]" pageId="3" pageNumber="71">Kacar</collectorName>
&amp;
<collectorName id="26AF531AFF8A356CFE48F9F5FE601A4B" box="[435,503,1582,1608]" pageId="3" pageNumber="71">Çelik.</collectorName>
</materialsCitation>
<materialsCitation id="3B323C91FF8A356CFE07F9F5FED51A6F" ID-GBIF-Occurrence="4899575302" collectingDate="2022-04-15" collectorName="Kacar &amp; Celik." country="Turkey" elevation="1030" latitude="40.396114" location="Laloglu" longLatPrecision="19" longitude="34.545834" pageId="3" pageNumber="71" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Corum" typeStatus="paratype">
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,
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,
<collectingRegion id="499EF82EFF8A356CFDA7F9F5FD3A1A4B" box="[604,685,1582,1608]" country="Turkey" name="Corum" pageId="3" pageNumber="71">Çorum</collectingRegion>
,
<location id="8E856017FF8A356CFD4DF9F5FC9A1A4B" LSID="urn:lsid:plazi:treatment:03F387DAFF8A356BFF6CFAE9FF201956:8E856017FF8A356CFD4DF9F5FC9A1A4B" box="[694,781,1582,1608]" country="Turkey" latitude="40.396114" longLatPrecision="19" longitude="34.545834" name="Laloglu" pageId="3" pageNumber="71" stateProvince="Corum">Laloğlu</location>
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E,
<geoCoordinate id="EE6E500BFF8A356CFC94F9F6FC641A44" box="[879,1011,1581,1607]" degrees="40" direction="north" minutes="23" orientation="latitude" pageId="3" pageNumber="71" precision="15" seconds="46" value="40.396114">40°23'46''N</geoCoordinate>
,
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,
<quantity id="4CA29B29FF8A356CFB71F9F5FB771A4B" box="[1162,1248,1582,1608]" metricMagnitude="3" metricUnit="m" metricValue="1.03" pageId="3" pageNumber="71" unit="m" value="1030.0">
<elevation id="0077D1FFFF8A356CFB71F9F5FB771A4B" box="[1162,1248,1582,1608]" metricMagnitude="3" metricUnit="m" metricValue="1.03" pageId="3" pageNumber="71" unit="m" value="1030.0">1030 m</elevation>
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,
<date id="FFE4100CFF8A356CFB11F9F6FAFE1A4B" box="[1258,1385,1581,1608]" pageId="3" pageNumber="71" value="2022-04-15">
<collectingDate id="EFA0E9E4FF8A356CFB11F9F6FAFE1A4B" box="[1258,1385,1581,1608]" pageId="3" pageNumber="71" value="2022-04-15">15.IV.2022</collectingDate>
</date>
, leg.
<collectorName id="26AF531AFF8A356CFF6CF989FF4D1A6F" box="[151,218,1618,1644]" pageId="3" pageNumber="71">Kacar</collectorName>
&amp;
<collectorName id="26AF531AFF8A356CFF05F989FED51A6F" box="[254,322,1618,1644]" pageId="3" pageNumber="71">Çelik.</collectorName>
</materialsCitation>
&lt;all
<typeStatus id="54E1886EFF8A356CFE80F989FE711A6F" box="[379,486,1618,1644]" pageId="3" pageNumber="71" type="paratype">paratypes</typeStatus>
with red printed label&gt; (
<collectionCode id="ED4BAE09FF8A356CFD07F989FCCD1A6F" box="[764,858,1618,1644]" pageId="3" pageNumber="71">AZMM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3406547FF8A356BFF3CF9AEFE45187E" lastPageId="4" lastPageNumber="72" pageId="3" pageNumber="71" type="description">
<paragraph id="8BE536CCFF8A356CFF3CF9AEFCA81AFF" blockId="3.[151,1437,1401,2040]" pageId="3" pageNumber="71">
<emphasis id="B92EEADEFF8A356CFF3CF9AEFECE1A8C" bold="true" box="[199,345,1653,1679]" pageId="3" pageNumber="71">Description.</emphasis>
Habitus as in
<figureCitation id="13612A49FF8A356CFE0CF9ADFDAC1A93" box="[503,571,1654,1680]" captionStart="FIGURES 916" captionStartId="3.[151,264,1151,1175]" captionTargetBox="[427,1165,185,1121]" captionTargetId="figure-41@3.[422,1166,181,1127]" captionTargetPageId="3" captionText="FIGURES 916. Astenus (Eurysunius) hititicus sp. n. 9—habitus; 10—forebody; 11—Male sternite VII; 12—Male sternite VIII; 13, 14—Aedeagus in lateral view; 15, 16—Aedeagus in ventral view. Scale bars: 0.5 mm (Figs. 910), 0.2 mm (Fig. 1116)." figureDoi="http://doi.org/10.5281/zenodo.12526989" httpUri="https://zenodo.org/record/12526989/files/figure.png" pageId="3" pageNumber="71">Fig. 9</figureCitation>
. Body
<quantity id="4CA29B29FF8A356CFD72F9AEFC871A93" box="[649,784,1653,1680]" metricMagnitude="-3" metricUnit="m" metricValue="4.449999999999999" metricValueMax="4.6" metricValueMin="4.3" pageId="3" pageNumber="71" unit="mm" value="4.449999999999999" valueMax="4.6" valueMin="4.3">4.34.6 mm</quantity>
long. Coloration: head and pronotum blackish, sometimes pronotum more or less dark brown, posterior half of elytra reddish, with anterior area brown, abdomen black or dark brown with the posterior margins of segments VIVIII reddish brown, antennae reddish and legs with the femora dark-brown, the tibiae brown, and the tarsi yellowish brown.
</paragraph>
<paragraph id="8BE536CCFF8A356CFF3CF8DDFED91BB3" blockId="3.[151,1437,1401,2040]" pageId="3" pageNumber="71">
Head transverse, 1.201.25 times as wide as long (
<figureCitation id="13612A49FF8A356CFD02F8DDFCF91B23" box="[761,878,1798,1824]" captionStart="FIGURES 916" captionStartId="3.[151,264,1151,1175]" captionTargetBox="[427,1165,185,1121]" captionTargetId="figure-41@3.[422,1166,181,1127]" captionTargetPageId="3" captionText="FIGURES 916. Astenus (Eurysunius) hititicus sp. n. 9—habitus; 10—forebody; 11—Male sternite VII; 12—Male sternite VIII; 13, 14—Aedeagus in lateral view; 15, 16—Aedeagus in ventral view. Scale bars: 0.5 mm (Figs. 910), 0.2 mm (Fig. 1116)." figureDoi="http://doi.org/10.5281/zenodo.12526989" httpUri="https://zenodo.org/record/12526989/files/figure.png" pageId="3" pageNumber="71">Figs 910</figureCitation>
); posterior margin concave, punctation umbilicate and very dense, but not very deep; microsculpture indistinct; pubescence yellowish, short, sparse and decumbent. Eyes relatively small in size with slightly protruding, shorter than postocular region in dorsal view. Antennae moderately slender,
<quantity id="4CA29B29FF8A356CFE82F8A9FD8C1B8F" box="[377,539,1906,1932]" metricMagnitude="-3" metricUnit="m" metricValue="1.16" metricValueMax="1.18" metricValueMin="1.14" pageId="3" pageNumber="71" unit="mm" value="1.16" valueMax="1.18" valueMin="1.14">1.141.18 mm</quantity>
long, all antennomeres oblong, antennomere III approximately 2.2 times as long as wide (
<figureCitation id="13612A49FF8A356CFF00F84DFED71BB3" box="[251,320,1942,1968]" captionStart="FIGURES 916" captionStartId="3.[151,264,1151,1175]" captionTargetBox="[427,1165,185,1121]" captionTargetId="figure-41@3.[422,1166,181,1127]" captionTargetPageId="3" captionText="FIGURES 916. Astenus (Eurysunius) hititicus sp. n. 9—habitus; 10—forebody; 11—Male sternite VII; 12—Male sternite VIII; 13, 14—Aedeagus in lateral view; 15, 16—Aedeagus in ventral view. Scale bars: 0.5 mm (Figs. 910), 0.2 mm (Fig. 1116)." figureDoi="http://doi.org/10.5281/zenodo.12526989" httpUri="https://zenodo.org/record/12526989/files/figure.png" pageId="3" pageNumber="71">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="8BE536CCFF8A356BFF3CF861FECB1D42" blockId="3.[151,1437,1401,2040]" lastBlockId="4.[151,1437,151,1365]" lastPageId="4" lastPageNumber="72" pageId="3" pageNumber="71">
Pronotum transverse and octagonal shape, 1.181.22 times as wide as long (
<figureCitation id="13612A49FF8A356CFBEBF861FB131BD7" box="[1040,1156,1978,2004]" captionStart="FIGURES 916" captionStartId="3.[151,264,1151,1175]" captionTargetBox="[427,1165,185,1121]" captionTargetId="figure-41@3.[422,1166,181,1127]" captionTargetPageId="3" captionText="FIGURES 916. Astenus (Eurysunius) hititicus sp. n. 9—habitus; 10—forebody; 11—Male sternite VII; 12—Male sternite VIII; 13, 14—Aedeagus in lateral view; 15, 16—Aedeagus in ventral view. Scale bars: 0.5 mm (Figs. 910), 0.2 mm (Fig. 1116)." figureDoi="http://doi.org/10.5281/zenodo.12526989" httpUri="https://zenodo.org/record/12526989/files/figure.png" pageId="3" pageNumber="71">Figs 910</figureCitation>
); approximately as broad as head or slightly narrower than head, slightly widest at anterior angles and narrowed posteriorly; posteriorly with median impression; punctation somewhat coarser, non-umbilicate and sparser than that of head; interstices on average about as wide as diameter of punctures, with shallow microsculpture; pubescence short, decumbent and yellowish, more distinct in middle area of the posterior margin; lateral margins each with three long black setae; one in anterior angle (on avarege
<quantity id="4CA29B29FF8D356BFDF7FED9FDF11D1E" box="[524,614,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="72" unit="mm" value="2.5">2.5 mm</quantity>
), one at anterior third (on avarege
<quantity id="4CA29B29FF8D356BFC1FFED8FBAA1D1E" box="[996,1085,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="4" pageNumber="72" unit="mm" value="2.2">2.2 mm</quantity>
), and one in posterior angle (on avarege
<quantity id="4CA29B29FF8D356BFF0EFEFDFED81D42" box="[245,335,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="72" unit="mm" value="2.5">2.5 mm</quantity>
).
</paragraph>
<paragraph id="8BE536CCFF8D356BFF3CFE90FAFE1DAE" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="72">
Elytra distinctly transverse, 1.701.80 times as long, 1.151.20 times as wide as pronotum (
<figureCitation id="13612A49FF8D356BFB24FE90FAC11D66" box="[1247,1366,331,357]" captionStart="FIGURES 916" captionStartId="3.[151,264,1151,1175]" captionTargetBox="[427,1165,185,1121]" captionTargetId="figure-41@3.[422,1166,181,1127]" captionTargetPageId="3" captionText="FIGURES 916. Astenus (Eurysunius) hititicus sp. n. 9—habitus; 10—forebody; 11—Male sternite VII; 12—Male sternite VIII; 13, 14—Aedeagus in lateral view; 15, 16—Aedeagus in ventral view. Scale bars: 0.5 mm (Figs. 910), 0.2 mm (Fig. 1116)." figureDoi="http://doi.org/10.5281/zenodo.12526989" httpUri="https://zenodo.org/record/12526989/files/figure.png" pageId="4" pageNumber="72">Figs 910</figureCitation>
); 0.65 times as long as pronotum; microsculpture absent, punctation dense and granulose, non-umbilicate; pubescence yellowish brown, posterior margin of each elytron with 57 long black setae. Hind wings completely reduced.
</paragraph>
<paragraph id="8BE536CCFF8D356BFF3CFE6CFB931E1A" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="72">
Abdomen narower than elytra, 0.9 times as broad as elytra; widest at segment V, segments IIIVI distinctly transverse (
<figureCitation id="13612A49FF8D356BFEE6FE00FEF01DF6" box="[285,359,475,501]" captionStart="FIGURES 916" captionStartId="3.[151,264,1151,1175]" captionTargetBox="[427,1165,185,1121]" captionTargetId="figure-41@3.[422,1166,181,1127]" captionTargetPageId="3" captionText="FIGURES 916. Astenus (Eurysunius) hititicus sp. n. 9—habitus; 10—forebody; 11—Male sternite VII; 12—Male sternite VIII; 13, 14—Aedeagus in lateral view; 15, 16—Aedeagus in ventral view. Scale bars: 0.5 mm (Figs. 910), 0.2 mm (Fig. 1116)." figureDoi="http://doi.org/10.5281/zenodo.12526989" httpUri="https://zenodo.org/record/12526989/files/figure.png" pageId="4" pageNumber="72">Fig. 9</figureCitation>
), punctation moderately sparse; interstices with microsculpture; pubescence reddish brown; posterior margin of tergite VII with very narrow rudiment of a palisade fringe.
</paragraph>
<paragraph id="8BE536CCFF8D356BFF3CFDF8FE051EAA" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="72">
<specimenCount id="9D5CFD45FF8D356BFF3CFDF8FF481E3E" box="[199,223,547,573]" count="1" pageId="4" pageNumber="72" type="male"></specimenCount>
: sternite VII in posterior median area undepressed, with some modified dark and short setae, posterior margin not concave (
<figureCitation id="13612A49FF8D356BFE70FD9CFE481E62" box="[395,479,583,609]" captionStart="FIGURES 916" captionStartId="3.[151,264,1151,1175]" captionTargetBox="[427,1165,185,1121]" captionTargetId="figure-41@3.[422,1166,181,1127]" captionTargetPageId="3" captionText="FIGURES 916. Astenus (Eurysunius) hititicus sp. n. 9—habitus; 10—forebody; 11—Male sternite VII; 12—Male sternite VIII; 13, 14—Aedeagus in lateral view; 15, 16—Aedeagus in ventral view. Scale bars: 0.5 mm (Figs. 910), 0.2 mm (Fig. 1116)." figureDoi="http://doi.org/10.5281/zenodo.12526989" httpUri="https://zenodo.org/record/12526989/files/figure.png" pageId="4" pageNumber="72">Fig. 11</figureCitation>
); sternite VIII deeply and acutely incised posteriorly, pubescence unmodified (
<figureCitation id="13612A49FF8D356BFA94FD9CFF211E86" captionStart="FIGURES 916" captionStartId="3.[151,264,1151,1175]" captionTargetBox="[427,1165,185,1121]" captionTargetId="figure-41@3.[422,1166,181,1127]" captionTargetPageId="3" captionText="FIGURES 916. Astenus (Eurysunius) hititicus sp. n. 9—habitus; 10—forebody; 11—Male sternite VII; 12—Male sternite VIII; 13, 14—Aedeagus in lateral view; 15, 16—Aedeagus in ventral view. Scale bars: 0.5 mm (Figs. 910), 0.2 mm (Fig. 1116)." figureDoi="http://doi.org/10.5281/zenodo.12526989" httpUri="https://zenodo.org/record/12526989/files/figure.png" pageId="4" pageNumber="72">Fig. 12</figureCitation>
); aedeagus approximately
<quantity id="4CA29B29FF8D356BFE1AFDB1FDD21E86" box="[481,581,618,645]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="4" pageNumber="72" unit="mm" value="0.65">0.65 mm</quantity>
long, as in
<figureCitation id="13612A49FF8D356BFD3CFDB0FCFE1E86" box="[711,873,619,645]" captionStart="FIGURES 916" captionStartId="3.[151,264,1151,1175]" captionTargetBox="[427,1165,185,1121]" captionTargetId="figure-41@3.[422,1166,181,1127]" captionTargetPageId="3" captionText="FIGURES 916. Astenus (Eurysunius) hititicus sp. n. 9—habitus; 10—forebody; 11—Male sternite VII; 12—Male sternite VIII; 13, 14—Aedeagus in lateral view; 15, 16—Aedeagus in ventral view. Scale bars: 0.5 mm (Figs. 910), 0.2 mm (Fig. 1116)." figureDoi="http://doi.org/10.5281/zenodo.12526989" httpUri="https://zenodo.org/record/12526989/files/figure.png" pageId="4" pageNumber="72">Figures 1316</figureCitation>
, and apical portion of ventral process with curled-shaped in lateral view.
</paragraph>
<paragraph id="8BE536CCFF8D356BFF3CFD69FB111836" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="72">
<emphasis id="B92EEADEFF8D356BFF3CFD69FE261ECE" bold="true" box="[199,433,690,717]" pageId="4" pageNumber="72">Comparative notes.</emphasis>
The species is distinguished from all its consubgeners except
<taxonomicName id="4C5A4D4FFF8D356BFB95FD68FA0B1ECE" authority="Assing, 2002" authorityName="Assing" authorityYear="2002" box="[1134,1436,691,717]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="sexsetosus">
<emphasis id="B92EEADEFF8D356BFB95FD68FA951ECF" box="[1134,1282,691,717]" italics="true" pageId="4" pageNumber="72">A. sexsetosus</emphasis>
<bibRefCitation id="EFCB4B3DFF8D356BFAF3FD68FA0B1ECE" author="Assing, V." box="[1288,1436,691,717]" pageId="4" pageNumber="72" pagination="265 - 274" refId="ref8983" refString="Assing, V. (2002) On the Turkish and Caucasian species of Eurysunius, subgenus of Astenus Dejean, with an appendix on A. breuili Jarrige (Coleoptera: Staphylinidae, Paederinae). Linzer Biologische Beitrage, 34, 265 - 274." type="journal article" year="2002">Assing, 2002</bibRefCitation>
</taxonomicName>
(
<collectingRegion id="499EF82EFF8D356BFF5AFD0CFF6B1EF2" box="[161,252,727,753]" country="Turkey" name="Kayseri" pageId="4" pageNumber="72">Kayseri</collectingRegion>
),
<taxonomicName id="4C5A4D4FFF8D356BFEF6FD0CFD851EF2" authority="Assing, 2015" authorityName="Assing" authorityYear="2015" box="[269,530,726,753]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="kociani">
<emphasis id="B92EEADEFF8D356BFEF6FD0CFEEC1EF2" box="[269,379,727,753]" italics="true" pageId="4" pageNumber="72">A. kociani</emphasis>
<bibRefCitation id="EFCB4B3DFF8D356BFE7AFD0CFD851EF2" author="Assing, V." box="[385,530,726,753]" pageId="4" pageNumber="72" pagination="1113 - 1118" refId="ref9178" refString="Assing, V. (2015) A new myrmecophilous species of Eurysunius from Turkey (Coleoptera: Staphylinidae: Paederinae). Linzer Biologische Beitrage, 47, 1113 - 1118." type="journal article" year="2015">Assing, 2015</bibRefCitation>
</taxonomicName>
(
<collectingRegion id="499EF82EFF8D356BFDE5FD0CFD1E1EF2" box="[542,649,727,753]" country="Turkey" name="Nevsehir" pageId="4" pageNumber="72">Nevşehir</collectingRegion>
),
<taxonomicName id="4C5A4D4FFF8D356BFD61FD0CFC1A1EF2" authority="Anlas, 2021" authorityName="Anlas" authorityYear="2021" box="[666,909,727,753]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="veyseli">
<emphasis id="B92EEADEFF8D356BFD61FD0CFC941EF2" box="[666,771,727,753]" italics="true" pageId="4" pageNumber="72">A. veyseli</emphasis>
<bibRefCitation id="EFCB4B3DFF8D356BFCF3FD0CFC1A1EF2" author="Anlas, S." box="[776,909,727,753]" pageId="4" pageNumber="72" pagination="343 - 352" refId="ref8873" refString="Anlas, S. (2021) On the subgenus Eurysunius Reitter of Turkey VII. Four new species from central Anatolia, and additional records (Coleoptera: Staphylinidae: Paederinae, Astenus). Zootaxa, 4991 (1), 343 - 352. https: // doi. org / 10.11646 / zootaxa. 4991.2.7" type="journal article" year="2021">Anlaş, 2021</bibRefCitation>
</taxonomicName>
(
<collectingRegion id="499EF82EFF8D356BFC61FD0CFC4E1EF2" box="[922,985,727,753]" country="Turkey" name="Sivas" pageId="4" pageNumber="72">Sivas</collectingRegion>
) and
<taxonomicName id="4C5A4D4FFF8D356BFBE8FD0CFA8C1EF2" authority="Anlas, 2021" authorityName="Anlas" authorityYear="2021" box="[1043,1307,727,753]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="gultekini">
<emphasis id="B92EEADEFF8D356BFBE8FD0CFB061EF2" box="[1043,1169,727,753]" italics="true" pageId="4" pageNumber="72">A. gultekini</emphasis>
<bibRefCitation id="EFCB4B3DFF8D356BFB6DFD0CFA8C1EF2" author="Anlas, S." box="[1174,1307,727,753]" pageId="4" pageNumber="72" pagination="343 - 352" refId="ref8873" refString="Anlas, S. (2021) On the subgenus Eurysunius Reitter of Turkey VII. Four new species from central Anatolia, and additional records (Coleoptera: Staphylinidae: Paederinae, Astenus). Zootaxa, 4991 (1), 343 - 352. https: // doi. org / 10.11646 / zootaxa. 4991.2.7" type="journal article" year="2021">Anlaş, 2021</bibRefCitation>
</taxonomicName>
(
<collectingRegion id="499EF82EFF8D356BFADCFD0CFA031EF2" box="[1319,1428,727,753]" country="Turkey" name="Kirikkale" pageId="4" pageNumber="72">Kırıkkale</collectingRegion>
) by the presence of three (rather than two or one) long setae on either of the lateral margins of the pronotum. It differs from
<emphasis id="B92EEADEFF8D356BFF2EFCC4FE7D1F3A" box="[213,490,799,825]" italics="true" pageId="4" pageNumber="72">
<taxonomicName id="4C5A4D4FFF8D356BFF2EFCC4FEFB1F3B" authorityName="Assing" authorityYear="2002" box="[213,364,799,825]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="sexsetosus">A. sexsetosus</taxonomicName>
,
<taxonomicName id="4C5A4D4FFF8D356BFE83FCC4FE7D1F3A" authorityName="Assing" authorityYear="2015" box="[376,490,799,825]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="kociani">A. kociani</taxonomicName>
</emphasis>
,
<taxonomicName id="4C5A4D4FFF8D356BFE02FCC4FDF21F3A" authorityName="Anlas" authorityYear="2021" box="[505,613,799,825]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="veyseli">
<emphasis id="B92EEADEFF8D356BFE02FCC4FDF21F3A" box="[505,613,799,825]" italics="true" pageId="4" pageNumber="72">A. veyseli</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5A4D4FFF8D356BFD65FCC4FCB71F3A" authorityName="Anlas" authorityYear="2021" box="[670,800,799,825]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="gultekini">
<emphasis id="B92EEADEFF8D356BFD65FCC4FCB71F3A" box="[670,800,799,825]" italics="true" pageId="4" pageNumber="72">A. gultekini</emphasis>
</taxonomicName>
by the different morphology of the aedeagus, especially apical portion of ventral process with curled-shaped in lateral view, it is additionally separated by the different coloration of the body (
<taxonomicName id="4C5A4D4FFF8D356BFE58FCBCFDAF1F83" authorityName="Assing" authorityYear="2002" box="[419,568,871,897]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="sexsetosus">
<emphasis id="B92EEADEFF8D356BFE58FCBCFDAF1F83" box="[419,568,871,897]" italics="true" pageId="4" pageNumber="72">A. sexsetosus</emphasis>
</taxonomicName>
: Head, pronotum and abdomen blackish brown, elytra yellowish, sometimes with a more or less infuscate portion near the scutellum.
<taxonomicName id="4C5A4D4FFF8D356BFCC0FC50FC271FA6" authorityName="Assing" authorityYear="2015" box="[827,944,907,933]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="kociani">
<emphasis id="B92EEADEFF8D356BFCC0FC50FC271FA6" box="[827,944,907,933]" italics="true" pageId="4" pageNumber="72">A. kociani</emphasis>
</taxonomicName>
: head and pronotum blackish; elytra dark yellowish-brown, with the region near the scutellum infuscate.
<taxonomicName id="4C5A4D4FFF8D356BFCA1FC74FC521FCA" authorityName="Anlas" authorityYear="2021" box="[858,965,943,969]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="veyseli">
<emphasis id="B92EEADEFF8D356BFCA1FC74FC521FCA" box="[858,965,943,969]" italics="true" pageId="4" pageNumber="72">A. veyseli</emphasis>
</taxonomicName>
: head and pronotum black or dark brown, more than anterior half of elytra black or dark brown, with posterior area yellowish brown, abdomen black or dark brown with the posterior margins of segments VIVIII reddish brown.
<taxonomicName id="4C5A4D4FFF8D356BFC40FC2CFBAA1812" authorityName="Anlas" authorityYear="2021" box="[955,1085,1015,1041]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="gultekini">
<emphasis id="B92EEADEFF8D356BFC40FC2CFBAA1812" box="[955,1085,1015,1041]" italics="true" pageId="4" pageNumber="72">A. gultekini</emphasis>
</taxonomicName>
: head, pronotum and abdomen blackish; elytra yellowish brown, with area near scutellum and anterior margin infuscate).
</paragraph>
<paragraph id="8BE536CCFF8D356BFF3CFBE4FE45187E" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="72">
For descriptions and illustrations of these species see
<bibRefCitation id="EFCB4B3DFF8D356BFCE2FBE4FC26185A" author="Assing, V." box="[793,945,1087,1113]" pageId="4" pageNumber="72" pagination="265 - 274" refId="ref8983" refString="Assing, V. (2002) On the Turkish and Caucasian species of Eurysunius, subgenus of Astenus Dejean, with an appendix on A. breuili Jarrige (Coleoptera: Staphylinidae, Paederinae). Linzer Biologische Beitrage, 34, 265 - 274." type="journal article" year="2002">Assing (2002</bibRefCitation>
,
<bibRefCitation id="EFCB4B3DFF8D356BFC47FBE5FC6F185A" author="Assing, V." box="[956,1016,1086,1113]" pageId="4" pageNumber="72" pagination="1113 - 1118" refId="ref9178" refString="Assing, V. (2015) A new myrmecophilous species of Eurysunius from Turkey (Coleoptera: Staphylinidae: Paederinae). Linzer Biologische Beitrage, 47, 1113 - 1118." type="journal article" year="2015">2015</bibRefCitation>
),
<bibRefCitation id="EFCB4B3DFF8D356BFBF1FBE4FB0B185A" author="Anlas, S." box="[1034,1180,1087,1113]" pageId="4" pageNumber="72" pagination="343 - 352" refId="ref8873" refString="Anlas, S. (2021) On the subgenus Eurysunius Reitter of Turkey VII. Four new species from central Anatolia, and additional records (Coleoptera: Staphylinidae: Paederinae, Astenus). Zootaxa, 4991 (1), 343 - 352. https: // doi. org / 10.11646 / zootaxa. 4991.2.7" type="journal article" year="2021">Anlaş (2021)</bibRefCitation>
and also see key to the Turkish
<taxonomicName id="4C5A4D4FFF8D356BFF0FFBB8FEE5187F" box="[244,370,1123,1148]" pageId="4" pageNumber="76" rank="subGenus" subGenus="Eurysunius">
<emphasis id="B92EEADEFF8D356BFF0FFBB8FEE5187F" box="[244,370,1123,1148]" italics="true" pageId="4" pageNumber="72">Eurysunius</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
<subSubSection id="C3406547FF8D356BFF3CFB5CFC4018EA" pageId="4" pageNumber="72" type="etymology">
<paragraph id="8BE536CCFF8D356BFF3CFB5CFC4018EA" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="72">
<emphasis id="B92EEADEFF8D356BFF3CFB5CFED918A2" bold="true" box="[199,334,1159,1185]" pageId="4" pageNumber="72">Etymology.</emphasis>
<taxonomicName id="4C5A4D4FFF8D356BFEACFB5CFD8618A2" authorityName="Anlaş" authorityYear="2024" box="[343,529,1159,1185]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="hititicus">
<emphasis id="B92EEADEFF8D356BFEACFB5CFD8618A2" box="[343,529,1159,1185]" italics="true" pageId="4" pageNumber="72">Astenus hititicus</emphasis>
</taxonomicName>
is named after the Hittites (Turkish: Hititler), an Anatolian ancient people who established an empire from the 17
<superScript id="7C2F9B84FF8D356BFDEDFB71FDB418BB" attach="left" box="[534,547,1194,1208]" fontSize="6" pageId="4" pageNumber="72">th</superScript>
century BC to the 12th century BC at Hattusa. The Hittite Capital is located in
<collectingRegion id="499EF82EFF8D356BFF6CFB14FEC618EA" box="[151,337,1231,1257]" country="Turkey" name="Corum" pageId="4" pageNumber="72">Çorum province</collectingRegion>
, in the territory in which the new species was discovered.
</paragraph>
</subSubSection>
<subSubSection id="C3406547FF8D356BFF3CFB29FF201956" pageId="4" pageNumber="72" type="distribution">
<paragraph id="8BE536CCFF8D356BFF3CFB29FF201956" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="72">
<emphasis id="B92EEADEFF8D356BFF3CFB29FD82190F" bold="true" box="[199,533,1266,1293]" pageId="4" pageNumber="72">Distribution and bionomics.</emphasis>
The species was collected from five localities from
<collectingRegion id="499EF82EFF8D356BFBA4FB28FA81190E" box="[1119,1302,1267,1293]" country="Turkey" name="Corum" pageId="4" pageNumber="72">Çorum province</collectingRegion>
(
<figureCitation id="13612A49FF8D356BFADEFB28FAF9190E" box="[1317,1390,1267,1293]" captionStart="MAP 1" captionStartId="6.[151,207,1425,1449]" captionTargetBox="[151,1436,489,1401]" captionTargetId="figure-135@6.[151,1436,489,1401]" captionText="MAP 1. Distribution of Astenus (Eurysunius) species in Turkey (28 species)" figureDoi="http://doi.org/10.5281/zenodo.12547399" httpUri="https://zenodo.org/record/12547399/files/figure.png" pageId="4" pageNumber="72">Map 1</figureCitation>
), in various grassland areas at altitudes of
<quantity id="4CA29B29FF8D356BFDC7FACCFD401932" box="[572,727,1303,1329]" metricMagnitude="3" metricUnit="m" metricValue="1.0699999999999998" metricValueMax="1.1099999999999999" metricValueMin="1.03" pageId="4" pageNumber="72" unit="m" value="1070.0" valueMax="1110.0" valueMin="1030.0">10301110 m</quantity>
. The
<typeStatus id="54E1886EFF8D356BFCE9FACCFCD51932" box="[786,834,1303,1329]" pageId="4" pageNumber="72">type</typeStatus>
specimens were collected in the nests of
<taxonomicName id="4C5A4D4FFF8D356BFAF0FACCFF201956" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B92EEADEFF8D356BFAF0FACCFA0B1933" box="[1291,1436,1303,1328]" italics="true" pageId="4" pageNumber="72">Tetramorium</emphasis>
sp.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>