258 lines
30 KiB
XML
258 lines
30 KiB
XML
<document id="5C16C615B4706B2EB27F97E79E20FD91" ID-CLB-Dataset="8143" ID-DOI="10.2478/aemnp-2019-0020" ID-GBIF-Dataset="a03ecce2-b526-4f94-aa65-620016956099" ID-ISSN="1804-6487" ID-Zenodo-Dep="4488918" ID-ZooBank="9F309FCC-A2ED-47B9-BC37-D0C4A3B482E5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612219281920" checkinUser="felipe" docAuthor="Fikáček, Martin & Liu, Hsing-Che" docDate="2019" docId="03F387C57A20FFB1FC5A07C035C9B19F" docLanguage="en" docName="ActaEntMusNatPra.59.1.229-252.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 59 (1)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Thysanarthria chui Fikáček & Liu 2019, sp. nov." docType="treatment" docVersion="9" lastPageNumber="244" masterDocId="FFCAFFBD7A2EFFBEFF8901283431B709" masterDocTitle="A review of Thysanarthria with description of seven new species and comments on its relationship to Chaetarthria (Hydrophilidae: Chaetarthriini)" masterLastPageNumber="252" masterPageNumber="229" pageNumber="243" updateTime="1698926077238" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="0CA24AA3C32C7925DB18A12A9FCC1034">A review of Thysanarthria with description of seven new species and comments on its relationship to Chaetarthria (Hydrophilidae: Chaetarthriini)</mods:title>
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<mods:name id="B68044AA83772CF3AEE69E1FDEE7E027" type="personal">
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<mods:namePart id="9CC664EF84437B86D121A845B8DE52F8">Fikáček, Martin</mods:namePart>
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<mods:affiliation id="E7B4EAF1A0FEE9A176549B430C326546">Department of Entomology, National Museum, Cirkusová 1740, CZ- 19100 Praha 9 – Horní Počernice, Czech Republic & Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 12844 Praha 2, Czech Republic</mods:affiliation>
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<mods:nameIdentifier id="81551FC35670822336D676BF347B4809" type="email">mfikacek@gmail.com</mods:nameIdentifier>
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<mods:namePart id="8AE8CBD60491B9784FB357CC85B3320E">Liu, Hsing-Che</mods:namePart>
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<mods:affiliation id="5D580840EE2E86B21E45227AABAB1149">Department of Environmental Engineering and Management, Chaoyang University of Technology, No. 168, Jifeng E. Rd., Taichung City 413, Taiwan</mods:affiliation>
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<mods:nameIdentifier id="ED2961FAFAE113326BB54393B20B4978" type="email">td965771@gmail.com</mods:nameIdentifier>
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<mods:title id="425F01D673DE84840796F511507D7589">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:part id="31F95E5EF9A9517CAC1029DA05658805">
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<mods:date id="A9101617B6ABE7195D8CE88096E1914B">2019</mods:date>
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<mods:number id="DE2674710DA1892D3667CE94F3EDA29F">2019-06-20</mods:number>
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<mods:number id="BFEA658FCE030EDBB21581A27078B28A">59</mods:number>
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<mods:identifier id="F873C6FC89228D53C832F791B531184D" type="DOI">10.2478/aemnp-2019-0020</mods:identifier>
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<treatment id="03F387C57A20FFB1FC5A07C035C9B19F" ID-DOI="http://doi.org/10.5281/zenodo.4548835" ID-GBIF-Taxon="177423427" ID-Zenodo-Dep="4548835" LSID="urn:lsid:plazi:treatment:03F387C57A20FFB1FC5A07C035C9B19F" httpUri="http://treatment.plazi.org/id/03F387C57A20FFB1FC5A07C035C9B19F" lastPageId="15" lastPageNumber="244" pageId="14" pageNumber="243">
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<subSubSection id="C34065587A20FFB0FC5A07C03134B1F6" box="[979,1285,1768,1791]" pageId="14" pageNumber="243" type="nomenclature">
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<paragraph id="8BE536D37A20FFB0FC5A07C03134B1F6" blockId="14.[979,1285,1768,1791]" box="[979,1285,1768,1791]" pageId="14" pageNumber="243">
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<heading id="D0AD81BF7A20FFB0FC5A07C03134B1F6" box="[979,1285,1768,1791]" centered="true" fontSize="10" level="2" pageId="14" pageNumber="243" reason="3">
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<emphasis id="B92EEAC17A20FFB0FC5A07C03134B1F6" bold="true" box="[979,1285,1768,1791]" pageId="14" pageNumber="243">
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<taxonomicName id="4C5A4D507A20FFB0FC5A07C03098B1F6" ID-CoL="8YBVT" authority="Fikáček & Liu, 2019" authorityName="Fikáček & Liu" authorityYear="2019" box="[979,1193,1768,1791]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="243" phylum="Arthropoda" rank="species" species="chui" status="sp. nov.">
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<emphasis id="B92EEAC17A20FFB0FC5A07C03098B1F6" bold="true" box="[979,1193,1768,1791]" italics="true" pageId="14" pageNumber="243">Thysanarthria chui</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21D57BA7A20FFB0FB3907C03134B1F6" box="[1200,1285,1768,1791]" pageId="14" pageNumber="243" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34065587A20FFB0FBAE062E3080B010" box="[1063,1201,1798,1817]" pageId="14" pageNumber="243" type="description">
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<paragraph id="8BE536D37A20FFB0FBAE062E3080B010" blockId="14.[828,1438,1798,2072]" box="[1063,1201,1798,1817]" pageId="14" pageNumber="243">
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(
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<figureCitation id="13612A567A20FFB0FBA4062E3050B010" box="[1069,1121,1798,1817]" captionStart="Fig" captionStartId="16.[151,182,1976,1995]" captionTargetBox="[204,1420,202,1944]" captionTargetId="figure-108@16.[162,1439,189,1956]" captionTargetPageId="16" captionText="Fig. 8. Male genitalia of Thysanarthria species, holotypes.A–E – T. chui sp. nov.(A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. hongsonensis Hebauer, 2001 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view). K–O – T. madurensis Hebauer, 2001 (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – details of median lobe and parameres in dorsal and ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4488940" httpUri="https://zenodo.org/record/4488940/files/figure.png" pageId="14" pageNumber="243">Figs 8</figureCitation>
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A–E,
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<figureCitation id="13612A567A20FFB0FB1C062E309BB010" box="[1173,1194,1798,1817]" captionStart="Fig" captionStartId="20.[151,182,1949,1968]" captionTargetBox="[151,1436,401,1931]" captionTargetId="figure-13@20.[151,1437,401,1931]" captionTargetPageId="20" captionText="Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu." figureDoi="http://doi.org/10.5281/zenodo.4488946" httpUri="https://zenodo.org/record/4488946/files/figure.png" pageId="14" pageNumber="243">11</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34065587A20FFB1FCB50618362BB60F" lastPageId="15" lastPageNumber="244" pageId="14" pageNumber="243" type="materials_examined">
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<paragraph id="8BE536D37A20FFB0FCB50618303CB0EA" blockId="14.[828,1438,1798,2072]" pageId="14" pageNumber="243">
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<emphasis id="B92EEAC17A20FFB0FCB5061837F1B04A" bold="true" box="[828,960,1840,1859]" pageId="14" pageNumber="243">Type material.</emphasis>
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<materialsCitation id="3B323C8E7A20FFB0FC4D06183746B071" ID-GBIF-Occurrence="3756360302" collectingDate="2003-04-22" collectingDateMax="2003-04-23" collectingDateMin="2003-04-22" collectingMethod="at light" collectionCode="NMNS" collectorName="C. S. Lin" country="Taiwan" location="Shan Ping Forest Ecological Garden" municipality="Shanpin" pageId="14" pageNumber="243" specimenCount="1" specimenCount-male="1" stateProvince="Kaohsiung" typeStatus="holotype">
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<typeStatus id="54E188717A20FFB0FC4D06183029B048" box="[964,1048,1840,1859]" pageId="14" pageNumber="243" type="holotype">HOLOTYPE</typeStatus>
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:
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<specimenCount id="9D5CFD5A7A20FFB0FBAA06023002B04A" box="[1059,1075,1834,1859]" pageId="14" pageNumber="243" type="male">♂</specimenCount>
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(
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<collectionCode id="ED4BAE167A20FFB0FBB40618304EB04A" box="[1085,1151,1840,1859]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="14" pageNumber="243">NMNS</collectionCode>
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),
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<emphasis id="B92EEAC17A20FFB0FB04061830DAB04A" bold="true" box="[1165,1259,1840,1859]" pageId="14" pageNumber="243">
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<collectingCountry id="F34D76437A20FFB0FB04061830D6B04A" box="[1165,1255,1840,1859]" name="Taiwan" pageId="14" pageNumber="243">TAIWAN</collectingCountry>
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:
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</emphasis>
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<collectingRegion id="499EF8317A20FFB0FB790618317EB04A" box="[1264,1359,1840,1859]" country="Taiwan" name="Kaohsiung" pageId="14" pageNumber="243">Kaohsiung</collectingRegion>
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,
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<collectingMunicipality id="6B81ACA97A20FFB0FADF061831ADB04A" box="[1366,1436,1840,1859]" country="Taiwan" name="Shanpin" pageId="14" pageNumber="243" stateProvince="Kaohsiung">Shanpin</collectingMunicipality>
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[=
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<location id="8E8560087A20FFB0FCDD066330BFB057" LSID="urn:lsid:plazi:treatment:03F387C57A20FFB1FC5A07C035C9B19F:8E8560087A20FFB0FCDD066330BFB057" box="[852,1166,1867,1886]" country="Taiwan" municipality="Shanpin" name="Shan Ping Forest Ecological Garden" pageId="14" pageNumber="243" stateProvince="Kaohsiung">Shan Ping Forest Ecological Garden</location>
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],
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<collectingMethod id="521B4EC47A20FFB0FB12066330E8B057" box="[1179,1241,1867,1886]" pageId="14" pageNumber="243">at light</collectingMethod>
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,
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<date id="FFE410137A20FFB0FB680663316EB057" box="[1249,1375,1867,1886]" pageId="14" pageNumber="243" value="2003-04-22" valueMax="2003-04-23" valueMin="2003-04-22">
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<collectingDate id="EFA0E9FB7A20FFB0FB680663316EB057" box="[1249,1375,1867,1886]" pageId="14" pageNumber="243" value="2003-04-22" valueMax="2003-04-23" valueMin="2003-04-22">22.-23.iv.2003</collectingDate>
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</date>
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, lgt.
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<collectorName id="26AF53057A20FFB0FA0006633742B071" pageId="14" pageNumber="243">C. S. Lin</collectorName>
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||
.
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</materialsCitation>
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<materialsCitation id="3B323C8E7A20FFB0FCF5064D303CB0EA" ID-GBIF-Occurrence="3756360335" latitude="24.054983" longitude="120.75543" pageId="14" pageNumber="243" typeStatus="paratype">
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<typeStatus id="54E188717A20FFB0FCF5064D37E5B07E" box="[892,980,1893,1912]" pageId="14" pageNumber="243" type="paratype">PARATYPES</typeStatus>
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:
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<specimenCount id="9D5CFD5A7A20FFB0FC69064D3019B071" box="[992,1064,1893,1912]" pageId="14" pageNumber="243" type="generic">20 spec.</specimenCount>
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(
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<collectionCode id="ED4BAE167A20FFB0FBBB064D3042B071" box="[1074,1139,1893,1912]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="14" pageNumber="243">NMNS</collectionCode>
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,
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<collectionCode id="ED4BAE167A20FFB0FBF5064D309FB071" box="[1148,1198,1893,1912]" country="Taiwan" httpUri="http://grbio.org/cool/a13e-ta1z" name="Taiwan Agricultural Research Institute" pageId="14" pageNumber="243">TARI</collectionCode>
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,
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<collectionCode id="ED4BAE167A20FFB0FB31064D30C6B071" box="[1208,1271,1893,1912]" country="Czech Republic" name="National Museum Prague" pageId="14" pageNumber="243">NMPC</collectionCode>
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,
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<collectionCode id="ED4BAE167A20FFB0FA89064D3177B071" box="[1280,1350,1893,1912]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="14" pageNumber="243" type="Museum">NHMW</collectionCode>
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,
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<collectionCode id="ED4BAE167A20FFB0FAC6064D31A2B071" box="[1359,1427,1893,1912]" pageId="14" pageNumber="243">BMNH</collectionCode>
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):
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<emphasis id="B92EEAC17A20FFB0FCB506A837A9B09A" bold="true" box="[828,920,1920,1939]" pageId="14" pageNumber="243">
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<collectingCountry id="F34D76437A20FFB0FCB506A837A5B09A" box="[828,916,1920,1939]" name="Taiwan" pageId="14" pageNumber="243">TAIWAN</collectingCountry>
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:
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</emphasis>
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<collectingRegion id="499EF8317A20FFB0FC1206A83020B09A" box="[923,1041,1920,1939]" country="Taiwan" name="Taichung" pageId="14" pageNumber="243">Taichung City</collectingRegion>
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,Wufeng district,Zhongkeng Industry Road,
|
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<quantity id="4CA29B367A20FFB0FA0B06A83766B0A7" metricMagnitude="3" metricUnit="m" metricValue="4.3" pageId="14" pageNumber="243" unit="km" value="4.3">4.3 km</quantity>
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SEE of Chaoyang Univ. of Technology,
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<geoCoordinate id="EE6E50147A20FFB0FB3C06B3312EB0A7" box="[1205,1311,1947,1966]" degrees="24.054983" direction="north" orientation="latitude" pageId="14" pageNumber="243" precision="1" value="24.054983">24.054983N</geoCoordinate>
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<geoCoordinate id="EE6E50147A20FFB0FAAD06B331A8B0A7" box="[1316,1433,1947,1966]" degrees="120.755433" direction="east" orientation="longitude" pageId="14" pageNumber="243" precision="1" value="120.75543">120.755433E</geoCoordinate>
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,
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<quantity id="4CA29B367A20FFB0FCB5069D3740B0C1" box="[828,881,1973,1992]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="14" pageNumber="243" unit="m" value="180.0">180 m</quantity>
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,
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<date id="FFE410137A20FFB0FCF0069D3037B0C1" box="[889,1030,1973,1992]" pageId="14" pageNumber="243" value="2019-04-25" valueMax="2019-05-14" valueMin="2019-04-25">25.iv.–14.v.2019</date>
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, H.-C. Liu lgt., on sandy banks of of small slowly running lowland stream.
|
||
</materialsCitation>
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</paragraph>
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||
<paragraph id="8BE536D37A20FFB1FCB506C3362BB60F" blockId="14.[828,1438,1798,2072]" lastBlockId="15.[151,761,190,262]" lastPageId="15" lastPageNumber="244" pageId="14" pageNumber="243">
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||
<emphasis id="B92EEAC17A20FFB0FCB506C3307EB0F7" bold="true" box="[828,1103,2027,2046]" pageId="14" pageNumber="243">Additional material examined.</emphasis>
|
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<materialsCitation id="3B323C8E7A20FFB0FBDB06C3305FBF11" ID-GBIF-Occurrence="3756360303" pageId="14" pageNumber="243">
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<emphasis id="B92EEAC17A20FFB0FBDB06C33103B0F7" bold="true" box="[1106,1330,2027,2046]" pageId="14" pageNumber="243">
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<collectingCountry id="F34D76437A20FFB0FBDB06C3309AB0F7" box="[1106,1195,2027,2046]" name="Taiwan" pageId="14" pageNumber="243">TAIWAN</collectingCountry>
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||
: LANYU ISLAND:
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||
</emphasis>
|
||
<specimenCount id="9D5CFD5A7A20FFB0FABF06C33165B0F7" box="[1334,1364,2023,2046]" pageId="14" pageNumber="243" type="female">1 ♀</specimenCount>
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(
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||
<collectionCode id="ED4BAE167A20FFB0FAD706C331A5B0F7" box="[1374,1428,2027,2046]" country="Taiwan" httpUri="http://grbio.org/cool/a13e-ta1z" name="Taiwan Agricultural Research Institute" pageId="14" pageNumber="243">TARI</collectionCode>
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): Lanyu,
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||
<date id="FFE410137A20FFB0FCF5092D37D1BF11" box="[892,992,2053,2072]" pageId="14" pageNumber="243" value="2015-07-27">27.vii.2015</date>
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||
, lgt.Y.-T. Wang.
|
||
</materialsCitation>
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The specimen corresponds with the
|
||
<typeStatus id="54E188717A21FFB1FF1E019634D1B7D8" box="[151,224,190,209]" pageId="15" pageNumber="244" type="holotype">holotype</typeStatus>
|
||
in all extrernal characters and we are hence assigning it to
|
||
<emphasis id="B92EEAC17A21FFB1FD32019636C8B7D8" box="[699,761,190,209]" italics="true" pageId="15" pageNumber="244">
|
||
<taxonomicName id="4C5A4D507A21FFB1FD32019636C4B7D8" authorityName="Fikáček & Liu" authorityYear="2019" box="[699,757,190,209]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="244" phylum="Arthropoda" rank="species" species="chui">T.chui</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
However, since male genitalia are needed for reliable identification, I am not including this specimen among the types.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34065587A21FFB1FF1E003735AAB4FF" pageId="15" pageNumber="244" type="description">
|
||
<paragraph id="8BE536D37A21FFB1FF1E003735AAB4FF" blockId="15.[151,761,287,1686]" pageId="15" pageNumber="244">
|
||
<emphasis id="B92EEAC17A21FFB1FF1E0037356FB63F" bold="true" box="[151,350,287,310]" pageId="15" pageNumber="244">
|
||
Description.
|
||
<emphasis id="B92EEAC17A21FFB1FEAF0037356FB63F" bold="true" box="[294,350,287,310]" italics="true" pageId="15" pageNumber="244">Body</emphasis>
|
||
</emphasis>
|
||
length
|
||
<quantity id="4CA29B367A21FFB1FE2200373619B63F" box="[427,552,287,310]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="15" pageNumber="244" unit="mm" value="1.45" valueMax="1.6" valueMin="1.3">1.3–1.6 mm</quantity>
|
||
(
|
||
<typeStatus id="54E188717A21FFB1FDBC003736A5B63F" box="[565,660,287,310]" pageId="15" pageNumber="244" type="holotype">holotype</typeStatus>
|
||
<quantity id="4CA29B367A21FFB1FD10003736DDB63F" box="[665,748,287,310]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="15" pageNumber="244" unit="mm" value="1.6">1.6 mm</quantity>
|
||
), maximum body width 0.9–1.0 mm (
|
||
<typeStatus id="54E188717A21FFB1FDA0001736B9B65F" box="[553,648,319,342]" pageId="15" pageNumber="244" type="holotype">holotype</typeStatus>
|
||
1.0 mm). Head and labrum black; pronotum brown in the middle, becoming weakly paler towards margins; elytra uniformly dark brown; legs brown.
|
||
<emphasis id="B92EEAC17A21FFB1FE1600B735EBB6BF" bold="true" box="[415,474,415,438]" italics="true" pageId="15" pageNumber="244">Head</emphasis>
|
||
without microsculpture on interstices; punctation sparse, each puncture bearing pointed seta. Eyes separated by 3.8× the width of one eye in dorsal view.
|
||
<emphasis id="B92EEAC17A21FFB1FEA900D735BEB51F" bold="true" box="[288,399,511,534]" italics="true" pageId="15" pageNumber="244">Pronotum</emphasis>
|
||
with sparse setiferous punctation similar to that on head; interstices without microsculpture.
|
||
<emphasis id="B92EEAC17A21FFB1FF1E031734EDB55F" bold="true" box="[151,220,575,598]" italics="true" pageId="15" pageNumber="244">Elytra</emphasis>
|
||
with 10 striae sharply impressed except anteromedially (near scutellar shield) where neither striae nor serial punctures are visible; intervals weakly convex at midlength and near apex; interval punctation sparse, setiferous; interstices without microsculpture.
|
||
<emphasis id="B92EEAC17A21FFB1FDDA0397368CB5DF" bold="true" box="[595,701,703,726]" italics="true" pageId="15" pageNumber="244">Aedeagus</emphasis>
|
||
(
|
||
<figureCitation id="13612A567A21FFB1FD4003973493B5FF" captionStart="Fig" captionStartId="16.[151,182,1976,1995]" captionTargetBox="[204,1420,202,1944]" captionTargetId="figure-108@16.[162,1439,189,1956]" captionTargetPageId="16" captionText="Fig. 8. Male genitalia of Thysanarthria species, holotypes.A–E – T. chui sp. nov.(A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. hongsonensis Hebauer, 2001 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view). K–O – T. madurensis Hebauer, 2001 (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – details of median lobe and parameres in dorsal and ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4488940" httpUri="https://zenodo.org/record/4488940/files/figure.png" pageId="15" pageNumber="244">Figs 8</figureCitation>
|
||
A–E)
|
||
<quantity id="4CA29B367A21FFB1FF6D03F73505B5FF" box="[228,308,735,758]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="244" unit="mm" value="0.5">0.5 mm</quantity>
|
||
long. Phallobase indistinctly wider at base of parameres, only very weakly constricted more basally, arcuate in lateral view. Parameres wide basally, gradually narrowing towards apex, outer face subrectangular in basal two thirds, apex projecting into narrow rounded lobe. Median lobe narrow apically, rounded and membranous at apex, reaching to the level of parameral apices, pair of subapical projections missing; gonopore transversely oval, situated in apical fourth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34065587A21FFB1FF1E02D73663B21F" pageId="15" pageNumber="244" type="diagnosis">
|
||
<paragraph id="8BE536D37A21FFB1FF1E02D73663B21F" blockId="15.[151,761,287,1686]" pageId="15" pageNumber="244">
|
||
<emphasis id="B92EEAC17A21FFB1FF1E02D735ACB31F" bold="true" box="[151,413,1023,1046]" pageId="15" pageNumber="244">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C5A4D507A21FFB1FE2102D7364EB31F" authorityName="Fikáček & Liu" authorityYear="2019" box="[424,639,1023,1046]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="244" phylum="Arthropoda" rank="species" species="chui">
|
||
<emphasis id="B92EEAC17A21FFB1FE2102D7364EB31F" box="[424,639,1023,1046]" italics="true" pageId="15" pageNumber="244">Thysanarthria chui</emphasis>
|
||
</taxonomicName>
|
||
externally differs from part of the remaining species in the total absence of dorsal microsculpture and rather dark (brown rather than yellowish) coloration. In the genital morphology it resembles
|
||
<taxonomicName id="4C5A4D507A21FFB1FEDC055735D1B39F" authorityName="Knisch" authorityYear="1924" box="[341,480,1151,1174]" class="Insecta" family="Hydrophilidae" genus="Chaetarthria" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="244" phylum="Arthropoda" rank="species" species="championi">
|
||
<emphasis id="B92EEAC17A21FFB1FEDC055735D1B39F" box="[341,480,1151,1174]" italics="true" pageId="15" pageNumber="244">T. championi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5A4D507A21FFB1FE6705573674B39F" authorityName="Fikáček & Liu" authorityYear="2019" box="[494,581,1151,1174]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="244" phylum="Arthropoda" rank="species" species="bifida">
|
||
<emphasis id="B92EEAC17A21FFB1FE6705573674B39F" box="[494,581,1151,1174]" italics="true" pageId="15" pageNumber="244">T. bifida</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C5A4D507A21FFB1FD09055736EDB39F" authorityName="Fikáček & Liu" authorityYear="2019" box="[640,732,1151,1174]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="244" phylum="Arthropoda" rank="species" species="trifida">
|
||
<emphasis id="B92EEAC17A21FFB1FD09055736EDB39F" box="[640,732,1151,1174]" italics="true" pageId="15" pageNumber="244">T. trifida</emphasis>
|
||
</taxonomicName>
|
||
in the form of the parameres;
|
||
<taxonomicName id="4C5A4D507A21FFB1FE3E05B7363FB3BF" authorityName="Fikáček & Liu" authorityYear="2019" box="[439,526,1183,1206]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="244" phylum="Arthropoda" rank="species" species="bifida">
|
||
<emphasis id="B92EEAC17A21FFB1FE3E05B7363FB3BF" box="[439,526,1183,1206]" italics="true" pageId="15" pageNumber="244">T. bifida</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5A4D507A21FFB1FDC905B736ADB3BF" authorityName="Fikáček & Liu" authorityYear="2019" box="[576,668,1183,1206]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="244" phylum="Arthropoda" rank="species" species="trifida">
|
||
<emphasis id="B92EEAC17A21FFB1FDC905B736ADB3BF" box="[576,668,1183,1206]" italics="true" pageId="15" pageNumber="244">T. trifida</emphasis>
|
||
</taxonomicName>
|
||
differ in the presence of a pair of subapically situated membranous lobes (absent in
|
||
<taxonomicName id="4C5A4D507A21FFB1FEC805F735B9B3FF" authorityName="Fikáček & Liu" authorityYear="2019" box="[321,392,1247,1270]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="244" phylum="Arthropoda" rank="species" species="chui">
|
||
<emphasis id="B92EEAC17A21FFB1FEC805F735B9B3FF" box="[321,392,1247,1270]" italics="true" pageId="15" pageNumber="244">T. chui</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5A4D507A21FFB1FE3005F73672B3FF" authorityName="Knisch" authorityYear="1924" box="[441,579,1247,1270]" class="Insecta" family="Hydrophilidae" genus="Chaetarthria" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="244" phylum="Arthropoda" rank="species" species="championi">
|
||
<emphasis id="B92EEAC17A21FFB1FE3005F73672B3FF" box="[441,579,1247,1270]" italics="true" pageId="15" pageNumber="244">T. championi</emphasis>
|
||
</taxonomicName>
|
||
). For differences from
|
||
<taxonomicName id="4C5A4D507A21FFB1FF5B05D7356CB21F" authorityName="Knisch" authorityYear="1924" box="[210,349,1279,1302]" class="Insecta" family="Hydrophilidae" genus="Chaetarthria" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="244" phylum="Arthropoda" rank="species" species="championi">
|
||
<emphasis id="B92EEAC17A21FFB1FF5B05D7356CB21F" box="[210,349,1279,1302]" italics="true" pageId="15" pageNumber="244">T. championi</emphasis>
|
||
</taxonomicName>
|
||
see under that species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34065587A21FFB1FF1E043736C8B29F" pageId="15" pageNumber="244" type="etymology">
|
||
<paragraph id="8BE536D37A21FFB1FF1E043736C8B29F" blockId="15.[151,761,287,1686]" pageId="15" pageNumber="244">
|
||
<emphasis id="B92EEAC17A21FFB1FF1E04373529B23F" bold="true" box="[151,280,1311,1334]" pageId="15" pageNumber="244">Etymology.</emphasis>
|
||
The first author dedicates this new species to Isaac Chu as thanks for introducing him to
|
||
<collectingCountry id="F34D76437A21FFB1FDD70417369DB25F" box="[606,684,1343,1366]" name="Taiwan" pageId="15" pageNumber="244">Taiwan</collectingCountry>
|
||
and its culture. The first known specimen of the new species was moreover collected in Mr. Chu’s home city of
|
||
<collectingRegion id="499EF8317A21FFB1FDF4045736C5B29F" box="[637,756,1407,1430]" country="Taiwan" name="Kaohsiung" pageId="15" pageNumber="244">Kaohsiung</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34065587A21FFB1FF1E04B73642B11F" pageId="15" pageNumber="244" type="biology_ecology">
|
||
<paragraph id="8BE536D37A21FFB1FF1E04B73642B11F" blockId="15.[151,761,287,1686]" pageId="15" pageNumber="244">
|
||
<emphasis id="B92EEAC17A21FFB1FF1E04B734C2B2BF" bold="true" box="[151,243,1439,1462]" pageId="15" pageNumber="244">Biology.</emphasis>
|
||
<typeStatus id="54E188717A21FFB1FF7204B73552B2BF" box="[251,355,1439,1462]" pageId="15" pageNumber="244" type="paratype">Paratypes</typeStatus>
|
||
were collected on a sandy bank of a slowly running lowland stream; they were actively crawling on the wet sand out of the water during the day (
|
||
<figureCitation id="13612A567A21FFB1FD4104F7349EB11F" captionStart="Fig" captionStartId="20.[151,182,1949,1968]" captionTargetBox="[151,1436,401,1931]" captionTargetId="figure-13@20.[151,1437,401,1931]" captionTargetPageId="20" captionText="Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu." figureDoi="http://doi.org/10.5281/zenodo.4488946" httpUri="https://zenodo.org/record/4488946/files/figure.png" pageId="15" pageNumber="244">Figs 11</figureCitation>
|
||
F–G). The
|
||
<typeStatus id="54E188717A21FFB1FEAF04D735B2B11F" box="[294,387,1535,1558]" pageId="15" pageNumber="244" type="holotype">holotype</typeStatus>
|
||
was collected at light.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34065587A21FFB1FF1E073735C9B19F" pageId="15" pageNumber="244" type="distribution">
|
||
<paragraph id="8BE536D37A21FFB1FF1E073735C9B19F" blockId="15.[151,761,287,1686]" pageId="15" pageNumber="244">
|
||
<emphasis id="B92EEAC17A21FFB1FF1E0737351CB13F" bold="true" box="[151,301,1567,1590]" pageId="15" pageNumber="244">Distribution.</emphasis>
|
||
The species is known from the lowland localities in western and southern
|
||
<collectingCountry id="F34D76437A21FFB1FE720717367CB15F" box="[507,589,1599,1622]" name="Taiwan" pageId="15" pageNumber="244">Taiwan</collectingCountry>
|
||
. The occurrence on Lanyu (= Orchid Island) needs to be confirmed by examination of a male specimen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |