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<document ID-DOI="http://doi.org/10.5281/zenodo.3803500" ID-GBIF-Dataset="fe76f860-39b1-4520-a13c-d01a47dfc130" ID-GBIF-Taxon="163738931" ID-ISSN="1175-5334" ID-Zenodo-Dep="3803500" ID-ZooBank="urn:lsid:zoobank.org:pub:BA570D47-FE77-446C-B000-9208C9A6E957" checkinTime="1587753273724" checkinUser="carolina" docAuthor="Ríos-Tamayo, Duniesky &amp; Lyle, Robin" docDate="2020" docId="03F11B20FFD6FF838EE83937FC5CD749" docLanguage="en" docName="zootaxa.4766.2.2.pdf.imf" docOrigin="Zootaxa 4766 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepthercus confusus Ríos-Tamayo &amp; Lyle 2020, sp. nov." docType="treatment" docVersion="5" lastPageId="11" lastPageNumber="272" masterDocId="FFC86358FFDEFF888E7F3E26FFD2D232" masterDocTitle="The South African genus Lepthercus Purcell, 1902 (Araneae: Mygalomorphae) phylogeny and taxonomy" masterLastPageNumber="305" masterPageNumber="261" pageId="8" pageNumber="269" updateTime="1636538462881" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The South African genus Lepthercus Purcell, 1902 (Araneae: Mygalomorphae) phylogeny and taxonomy</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ríos-Tamayo, Duniesky</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lyle, Robin</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-04-20</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>4766</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>261</mods:start>
<mods:end>305</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.4766.2.2</mods:identifier>
<mods:identifier type="GBIF-Dataset">fe76f860-39b1-4520-a13c-d01a47dfc130</mods:identifier>
<mods:identifier type="ISSN">1175-5334</mods:identifier>
<mods:identifier type="Zenodo-Dep">3765467</mods:identifier>
<mods:identifier type="ZooBank">urn:lsid:zoobank.org:pub:BA570D47-FE77-446C-B000-9208C9A6E957</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3803500" ID-GBIF-Taxon="163738931" ID-Zenodo-Dep="3803500" LSID="urn:lsid:plazi:treatment:03F11B20FFD6FF838EE83937FC5CD749" httpUri="http://treatment.plazi.org/id/03F11B20FFD6FF838EE83937FC5CD749" lastPageId="11" lastPageNumber="272" pageId="8" pageNumber="269">
<subSubSection box="[151,504,1809,1836]" pageId="8" pageNumber="269" type="nomenclature">
<paragraph blockId="8.[151,1110,1809,1906]" box="[151,504,1809,1836]" pageId="8" pageNumber="269">
<heading bold="true" box="[151,504,1809,1836]" fontSize="10" level="2" pageId="8" pageNumber="269" reason="2">
<emphasis bold="true" box="[151,504,1809,1836]" pageId="8" pageNumber="269">
<taxonomicName authority="Ríos-Tamayo &amp; Lyle, 2020" authorityName="Ríos-Tamayo &amp; Lyle" authorityYear="2020" box="[151,403,1809,1836]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="269" phylum="Arthropoda" rank="species" species="confusus" status="sp. nov.">
<emphasis bold="true" box="[151,403,1809,1836]" italics="true" pageId="8" pageNumber="269">Lepthercus confusus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[410,504,1809,1835]" pageId="8" pageNumber="269" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="269" type="description">
<paragraph blockId="8.[151,1110,1809,1906]" box="[151,411,1845,1871]" pageId="8" pageNumber="269">
(
<figureCitation box="[159,235,1845,1871]" captionStart="FIGURE" captionStartId="9.[151,251,1321,1346]" captionTargetBox="[151,1436,309,1296]" captionTargetId="figure@9.[151,1436,309,1296]" captionText="FIGURE. 3. Lepthercus confusus sp. nov. (male holotype, NCA 2011/318). A cephalothorax and abdomen, dorsal view; B sternum, ventral view; C chelicerae, prolateral view, arrow indicates the intercheliceral tumesence; D right pedipalp, arrow indicates the small dorsal setae, and a normal anterior part of the retrolateral side of ventral excavation, retrolateral view; E right pedipalp, prolateral view, arrow indicates the abundant proventral rigid setae; FH copulatory bulb; F retrolateral view, G ventral view, H prolateral view; F, H arrow indicates the longitudinal keels along the embolus, G arrow indicates the small lateral keel; I Tibia I, arrow indicates the small cuticular spur." figureDoi="http://doi.org/10.5281/zenodo.3765473" httpUri="https://zenodo.org/record/3765473/files/figure.png" pageId="8" pageNumber="269">Figs. 3</figureCitation>
AI, 4AE, 24)
</paragraph>
<paragraph blockId="8.[151,1110,1809,1906]" box="[151,1110,1882,1906]" pageId="8" pageNumber="269">
http://zoobank.org/
<uri box="[359,1110,1882,1906]" pageId="8" pageNumber="269">
urn:lsid:zoobank.org:act:
<uuid box="[617,1110,1882,1906]" pageId="8" pageNumber="269">19380CC0-30E8-412C-8509-F69267536BC6</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="270" pageId="8" pageNumber="269" type="materials_examined">
<paragraph blockId="8.[151,1437,1952,2015]" pageId="8" pageNumber="269">
<emphasis bold="true" box="[151,567,1952,1979]" pageId="8" pageNumber="269">Material examined. Type material</emphasis>
:
<materialsCitation ID-GBIF-Occurrence="2608670264" collectingDate="2009-06-22" collectionCode="UP" country="South Africa" elevation="2700" latitude="-29.583334" location="Sani Pass" longLatPrecision="1226" longitude="29.283333" pageId="8" pageNumber="269" specimenCode="NCA 2011" specimenCount="1" specimenCount-male="1" stateProvince="KwaZulu-Natal Province" typeStatus="holotype">
<typeStatus box="[585,689,1953,1979]" pageId="8" pageNumber="269">Holotype</typeStatus>
<specimenCount box="[699,720,1953,1979]" pageId="8" pageNumber="269" type="male"></specimenCount>
.
<collectingCountry box="[737,894,1953,1979]" name="South Africa" pageId="8" pageNumber="269">
<subScript attach="left" box="[753,807,1960,1977]" fontSize="7" pageId="8" pageNumber="269">SOUTH</subScript>
<subScript attach="right" box="[836,894,1960,1977]" fontSize="7" pageId="8" pageNumber="269">AFRICA</subScript>
</collectingCountry>
:
<collectingRegion box="[911,1204,1953,1979]" country="South Africa" name="KwaZulu-Natal" pageId="8" pageNumber="269">KwaZuluNatal Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:03F11B20FFD6FF838EE83937FC5CD749:8E87FCEDFFD6FF808ABD3987FAE2D589" box="[1218,1328,1953,1979]" country="South Africa" latitude="-29.583334" longLatPrecision="1226" longitude="29.283333" name="Sani Pass" pageId="8" pageNumber="269" stateProvince="KwaZulu-Natal Province">Sani Pass</location>
(
<geoCoordinate box="[1346,1437,1952,1979]" degrees="29" direction="south" minutes="35" orientation="latitude" pageId="8" pageNumber="269" precision="925" value="-29.583334">29°35S</geoCoordinate>
<geoCoordinate box="[151,244,1988,2015]" degrees="29" direction="east" minutes="17" orientation="longitude" pageId="8" pageNumber="269" precision="925" value="29.283333">29°17E</geoCoordinate>
,
<quantity box="[261,343,1989,2015]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="8" pageNumber="269" unit="m" value="2700.0">
<elevation box="[261,343,1989,2015]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="8" pageNumber="269" unit="m" value="2700.0">2700m</elevation>
</quantity>
),
<date box="[368,490,1989,2015]" pageId="8" pageNumber="269" value="2009-06-22">
<collectingDate box="[368,490,1989,2015]" pageId="8" pageNumber="269" value="2009-06-22">22.vi.2009</collectingDate>
</date>
,
<collectionCode box="[507,542,1989,2015]" country="Papua New Guinea" name="University of Papua and New Guinea" pageId="8" pageNumber="269">UP</collectionCode>
students
<emphasis box="[659,700,1989,2015]" italics="true" pageId="8" pageNumber="269">leg.</emphasis>
, (
<specimenCode box="[727,855,1989,2015]" collectionCode="NCA" pageId="8" pageNumber="269">NCA 2011</specimenCode>
/318)
</materialsCitation>
.—
<typeStatus box="[947,1061,1989,2015]" pageId="8" pageNumber="269">Paratypes</typeStatus>
.
<collectingCountry box="[1077,1236,1989,2015]" name="South Africa" pageId="8" pageNumber="269">
<subScript attach="left" box="[1092,1146,1996,2013]" fontSize="7" pageId="8" pageNumber="269">SOUTH</subScript>
<subScript attach="left" box="[1178,1236,1996,2013]" fontSize="7" pageId="8" pageNumber="269">AFRICA</subScript>
</collectingCountry>
:
<collectingRegion box="[1256,1436,1989,2015]" country="South Africa" name="KwaZulu-Natal" pageId="8" pageNumber="269">KwaZuluNatal</collectingRegion>
</paragraph>
<paragraph blockId="9.[151,1437,152,287]" pageId="9" pageNumber="270">
Province: Weza, Ngeli Forest [
<geoCoordinate box="[509,603,153,179]" degrees="30" direction="south" minutes="34" orientation="latitude" pageId="9" pageNumber="270" precision="925" value="-30.566668">30°34S</geoCoordinate>
<geoCoordinate box="[614,712,152,179]" degrees="29" direction="east" minutes="42" orientation="longitude" pageId="9" pageNumber="270" precision="925" value="29.7">29°42E</geoCoordinate>
],
<date box="[735,814,153,179]" pageId="9" pageNumber="270" value="1962-10">x.1962</date>
, Lawrence
<collectionCode box="[951,972,153,179]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="270">R</collectionCode>
.F.
<emphasis box="[1008,1049,153,179]" italics="true" pageId="9" pageNumber="270">leg.</emphasis>
,
<specimenCount box="[1067,1133,153,179]" pageId="9" pageNumber="270" type="female">3 ♀♀</specimenCount>
(NM8787); Same data as
<typeStatus box="[151,253,189,215]" pageId="9" pageNumber="270">holotype</typeStatus>
:
<date box="[264,373,189,215]" pageId="9" pageNumber="270" value="2010-01-22">22.i.2010</date>
,
<specimenCount box="[385,427,189,215]" pageId="9" pageNumber="270" type="male">1 ♂</specimenCount>
(
<collectionCode box="[442,502,189,215]" pageId="9" pageNumber="270">NCA</collectionCode>
2010/4443);
<date box="[654,761,189,215]" pageId="9" pageNumber="270" value="2008-01-20">20.i.2008</date>
,
<specimenCount box="[774,816,189,215]" pageId="9" pageNumber="270" type="male">1 ♂</specimenCount>
(
<collectionCode box="[831,891,189,215]" pageId="9" pageNumber="270">NCA</collectionCode>
2010/3642); same data as
<typeStatus box="[1193,1291,189,215]" pageId="9" pageNumber="270">holotype</typeStatus>
<specimenCount box="[1298,1362,189,215]" pageId="9" pageNumber="270" type="male">3 ♂♂</specimenCount>
(
<collectionCode box="[1376,1436,189,215]" pageId="9" pageNumber="270">NCA</collectionCode>
2010/3640); same data as
<typeStatus box="[454,552,225,251]" pageId="9" pageNumber="270">holotype</typeStatus>
<date box="[562,670,225,251]" pageId="9" pageNumber="270" value="2009-01-22">22.i.2009</date>
,
<specimenCount box="[684,750,225,251]" pageId="9" pageNumber="270" type="male">2 ♂♂</specimenCount>
(
<collectionCode box="[766,826,225,251]" pageId="9" pageNumber="270">NCA</collectionCode>
2011/379); same data as
<typeStatus box="[1123,1221,225,251]" pageId="9" pageNumber="270">holotype</typeStatus>
(
<geoCoordinate box="[1239,1329,224,251]" degrees="29" direction="south" minutes="36" orientation="latitude" pageId="9" pageNumber="270" precision="925" value="-29.6">29°36S</geoCoordinate>
<geoCoordinate box="[1339,1432,224,251]" degrees="29" direction="east" minutes="18" orientation="longitude" pageId="9" pageNumber="270" precision="925" value="29.3">29°18E</geoCoordinate>
,
<quantity box="[151,233,261,287]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="9" pageNumber="270" unit="m" value="2400.0">2400m</quantity>
),
<date box="[252,361,261,287]" pageId="9" pageNumber="270" value="2006-01-24">24.i.2006</date>
,
<specimenCount box="[372,435,261,287]" pageId="9" pageNumber="270" type="male">2 ♂♂</specimenCount>
(
<collectionCode box="[449,509,261,287]" pageId="9" pageNumber="270">NCA</collectionCode>
2010/3735).
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3765473" ID-Zenodo-Dep="3765473" httpUri="https://zenodo.org/record/3765473/files/figure.png" pageId="9" pageNumber="270" startId="9.[151,251,1321,1346]" targetBox="[151,1436,309,1296]" targetPageId="9">
<paragraph blockId="9.[151,1437,1321,1526]" pageId="9" pageNumber="270">
<emphasis bold="true" box="[151,280,1321,1346]" pageId="9" pageNumber="270">FIGURE. 3.</emphasis>
<taxonomicName authority="Ríos-Tamayo &amp; Lyle, 2020" authorityName="Ríos-Tamayo &amp; Lyle" authorityYear="2020" box="[286,492,1322,1346]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="confusus" status="sp. nov.">
<emphasis box="[286,492,1322,1346]" italics="true" pageId="9" pageNumber="270">Lepthercus confusus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[498,577,1322,1346]" pageId="9" pageNumber="270">
<taxonomicNameLabel box="[498,577,1322,1346]" pageId="9" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(male
<typeStatus box="[647,738,1321,1346]" pageId="9" pageNumber="270">holotype</typeStatus>
,
<collectionCode box="[747,799,1322,1346]" pageId="9" pageNumber="270">NCA</collectionCode>
2011/318).
<emphasis bold="true" box="[920,938,1321,1346]" pageId="9" pageNumber="270">A</emphasis>
cephalothorax and abdomen, dorsal view;
<emphasis bold="true" box="[1369,1386,1322,1346]" pageId="9" pageNumber="270">B</emphasis>
sternum, ventral view;
<emphasis bold="true" box="[344,362,1357,1382]" pageId="9" pageNumber="270">C</emphasis>
chelicerae, prolateral view, arrow indicates the intercheliceral tumesence;
<emphasis bold="true" box="[1107,1125,1358,1381]" pageId="9" pageNumber="270">D</emphasis>
right pedipalp, arrow indicates the small dorsal setae, and a normal anterior part of the retrolateral side of ventral excavation, retrolateral view;
<emphasis bold="true" box="[1267,1284,1394,1418]" pageId="9" pageNumber="270">E</emphasis>
right pedipalp, prolateral view, arrow indicates the abundant proventral rigid setae;
<emphasis bold="true" box="[834,882,1430,1454]" pageId="9" pageNumber="270">FH</emphasis>
copulatory bulb;
<emphasis bold="true" box="[1059,1074,1430,1454]" pageId="9" pageNumber="270">F</emphasis>
retrolateral view,
<emphasis bold="true" box="[1256,1276,1429,1454]" pageId="9" pageNumber="270">G</emphasis>
ventral view,
<emphasis bold="true" box="[1417,1437,1430,1454]" pageId="9" pageNumber="270">H</emphasis>
prolateral view;
<emphasis bold="true" box="[317,332,1466,1490]" pageId="9" pageNumber="270">F</emphasis>
,
<emphasis bold="true" box="[345,365,1466,1490]" pageId="9" pageNumber="270">H</emphasis>
arrow indicates the longitudinal keels along the embolus,
<emphasis bold="true" box="[956,976,1465,1490]" pageId="9" pageNumber="270">G</emphasis>
arrow indicates the small lateral keel;
<emphasis bold="true" box="[1368,1378,1466,1490]" pageId="9" pageNumber="270">I</emphasis>
Tibia I, arrow indicates the small cuticular spur.
</paragraph>
</caption>
<subSubSection pageId="9" pageNumber="270" type="etymology">
<paragraph blockId="9.[151,1437,1572,2031]" pageId="9" pageNumber="270">
<emphasis bold="true" box="[199,334,1572,1598]" pageId="9" pageNumber="270">Etymology.</emphasis>
The specific epithet refers to the great confusion felt in the beginning stages of this work about the placement of this species.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="270" type="diagnosis">
<paragraph blockId="9.[151,1437,1572,2031]" pageId="9" pageNumber="270">
<emphasis bold="true" box="[199,322,1643,1670]" pageId="9" pageNumber="270">Diagnosis.</emphasis>
Males of
<taxonomicName authority="Ríos-Tamayo &amp; Lyle, 2020" authorityName="Ríos-Tamayo &amp; Lyle" authorityYear="2020" box="[437,565,1644,1670]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="confusus" status="sp. nov.">
<emphasis box="[437,565,1644,1670]" italics="true" pageId="9" pageNumber="270">L. confusus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[573,664,1644,1670]" pageId="9" pageNumber="270">
<taxonomicNameLabel box="[573,664,1644,1670]" pageId="9" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, can be distinguished from males of “Group dregei” by a short bulb with a tapering embolus with longitudinal keels (
<figureCitation box="[693,805,1680,1707]" captionStart="FIGURE" captionStartId="9.[151,251,1321,1346]" captionTargetBox="[151,1436,309,1296]" captionTargetId="figure@9.[151,1436,309,1296]" captionText="FIGURE. 3. Lepthercus confusus sp. nov. (male holotype, NCA 2011/318). A cephalothorax and abdomen, dorsal view; B sternum, ventral view; C chelicerae, prolateral view, arrow indicates the intercheliceral tumesence; D right pedipalp, arrow indicates the small dorsal setae, and a normal anterior part of the retrolateral side of ventral excavation, retrolateral view; E right pedipalp, prolateral view, arrow indicates the abundant proventral rigid setae; FH copulatory bulb; F retrolateral view, G ventral view, H prolateral view; F, H arrow indicates the longitudinal keels along the embolus, G arrow indicates the small lateral keel; I Tibia I, arrow indicates the small cuticular spur." figureDoi="http://doi.org/10.5281/zenodo.3765473" httpUri="https://zenodo.org/record/3765473/files/figure.png" pageId="9" pageNumber="270">Fig. 3F, H</figureCitation>
) similar to
<taxonomicName authorityName="Purcell" authorityYear="1902" box="[931,1031,1680,1706]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="dregei">
<emphasis box="[931,1031,1680,1706]" italics="true" pageId="9" pageNumber="270">L. dregei</emphasis>
</taxonomicName>
but differs by their slender embolus, with a small lateral keel (
<figureCitation box="[436,555,1716,1742]" captionStart="FIGURE" captionStartId="9.[151,251,1321,1346]" captionTargetBox="[151,1436,309,1296]" captionTargetId="figure@9.[151,1436,309,1296]" captionText="FIGURE. 3. Lepthercus confusus sp. nov. (male holotype, NCA 2011/318). A cephalothorax and abdomen, dorsal view; B sternum, ventral view; C chelicerae, prolateral view, arrow indicates the intercheliceral tumesence; D right pedipalp, arrow indicates the small dorsal setae, and a normal anterior part of the retrolateral side of ventral excavation, retrolateral view; E right pedipalp, prolateral view, arrow indicates the abundant proventral rigid setae; FH copulatory bulb; F retrolateral view, G ventral view, H prolateral view; F, H arrow indicates the longitudinal keels along the embolus, G arrow indicates the small lateral keel; I Tibia I, arrow indicates the small cuticular spur." figureDoi="http://doi.org/10.5281/zenodo.3765473" httpUri="https://zenodo.org/record/3765473/files/figure.png" pageId="9" pageNumber="270">Fig. 3F, G</figureCitation>
). Further it is differentiated by the presence of a small cuticular spur on tibia I (
<figureCitation box="[160,236,1752,1778]" captionStart="FIGURE" captionStartId="9.[151,251,1321,1346]" captionTargetBox="[151,1436,309,1296]" captionTargetId="figure@9.[151,1436,309,1296]" captionText="FIGURE. 3. Lepthercus confusus sp. nov. (male holotype, NCA 2011/318). A cephalothorax and abdomen, dorsal view; B sternum, ventral view; C chelicerae, prolateral view, arrow indicates the intercheliceral tumesence; D right pedipalp, arrow indicates the small dorsal setae, and a normal anterior part of the retrolateral side of ventral excavation, retrolateral view; E right pedipalp, prolateral view, arrow indicates the abundant proventral rigid setae; FH copulatory bulb; F retrolateral view, G ventral view, H prolateral view; F, H arrow indicates the longitudinal keels along the embolus, G arrow indicates the small lateral keel; I Tibia I, arrow indicates the small cuticular spur." figureDoi="http://doi.org/10.5281/zenodo.3765473" httpUri="https://zenodo.org/record/3765473/files/figure.png" pageId="9" pageNumber="270">Fig. 3I</figureCitation>
, longer in others). Females can be distinguished by the shape of their spermathecae with a base with a slight transition to the stalk, straight stalks twisted and oval receptacles inwardly curved (
<figureCitation box="[1083,1170,1788,1814]" captionStart="FIGURE" captionStartId="10.[151,251,1604,1629]" captionTargetBox="[151,1436,417,1579]" captionTargetId="figure@10.[151,1436,417,1579]" captionText="FIGURE. 4. Lepthercus confusus sp. nov. (female paratype, NM 8787). A cephalothorax, dorsal view; B sternum, arrow indicates the anterior maxillary edge slightly concave, ventral view; C abdomen, dorsal view; D spermathecae." figureDoi="http://doi.org/10.5281/zenodo.3765475" httpUri="https://zenodo.org/record/3765475/files/figure.png" pageId="9" pageNumber="270">Fig. 4E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="11" lastPageNumber="272" pageId="9" pageNumber="270" type="description">
<paragraph blockId="9.[151,1437,1572,2031]" lastBlockId="10.[151,1437,153,395]" lastPageId="10" lastPageNumber="271" pageId="9" pageNumber="270">
<emphasis bold="true" box="[199,345,1823,1850]" pageId="9" pageNumber="270">Description.</emphasis>
Male
<typeStatus box="[417,515,1824,1850]" pageId="9" pageNumber="270">holotype</typeStatus>
(NCA 2011/318,
<figureCitation box="[715,780,1824,1850]" captionStart="FIGURE" captionStartId="9.[151,251,1321,1346]" captionTargetBox="[151,1436,309,1296]" captionTargetId="figure@9.[151,1436,309,1296]" captionText="FIGURE. 3. Lepthercus confusus sp. nov. (male holotype, NCA 2011/318). A cephalothorax and abdomen, dorsal view; B sternum, ventral view; C chelicerae, prolateral view, arrow indicates the intercheliceral tumesence; D right pedipalp, arrow indicates the small dorsal setae, and a normal anterior part of the retrolateral side of ventral excavation, retrolateral view; E right pedipalp, prolateral view, arrow indicates the abundant proventral rigid setae; FH copulatory bulb; F retrolateral view, G ventral view, H prolateral view; F, H arrow indicates the longitudinal keels along the embolus, G arrow indicates the small lateral keel; I Tibia I, arrow indicates the small cuticular spur." figureDoi="http://doi.org/10.5281/zenodo.3765473" httpUri="https://zenodo.org/record/3765473/files/figure.png" pageId="9" pageNumber="270">Fig. 3</figureCitation>
AI). Total length 11.40.
<emphasis box="[1066,1176,1825,1851]" italics="true" pageId="9" pageNumber="270">Carapace</emphasis>
(
<figureCitation box="[1192,1283,1824,1850]" captionStart="FIGURE" captionStartId="9.[151,251,1321,1346]" captionTargetBox="[151,1436,309,1296]" captionTargetId="figure@9.[151,1436,309,1296]" captionText="FIGURE. 3. Lepthercus confusus sp. nov. (male holotype, NCA 2011/318). A cephalothorax and abdomen, dorsal view; B sternum, ventral view; C chelicerae, prolateral view, arrow indicates the intercheliceral tumesence; D right pedipalp, arrow indicates the small dorsal setae, and a normal anterior part of the retrolateral side of ventral excavation, retrolateral view; E right pedipalp, prolateral view, arrow indicates the abundant proventral rigid setae; FH copulatory bulb; F retrolateral view, G ventral view, H prolateral view; F, H arrow indicates the longitudinal keels along the embolus, G arrow indicates the small lateral keel; I Tibia I, arrow indicates the small cuticular spur." figureDoi="http://doi.org/10.5281/zenodo.3765473" httpUri="https://zenodo.org/record/3765473/files/figure.png" pageId="9" pageNumber="270">Fig. 3A</figureCitation>
): length 5.21, width 4.17, lateral marginal bristles present, with sparse pubescence.
<emphasis box="[943,1125,1860,1886]" italics="true" pageId="9" pageNumber="270">Cephalic region</emphasis>
: length 3.28; clypeus short with 5 bristles on the edge, numerous long bristles in front of the OQ and between the PME, dorsal line of bristles until the fovea.
<emphasis box="[332,411,1933,1958]" italics="true" pageId="9" pageNumber="270">Fovea:</emphasis>
width 0.45, short and almost straight.
<emphasis box="[854,1101,1932,1958]" italics="true" pageId="9" pageNumber="270">Ocular measurements</emphasis>
: AME 0.14, ALE 0.21, PME 0.17, PLE 0.19, OQ length 0.48, 0.78 width; AMEALE 0.05, PMEPLE 0.05, AMEPME 0.07, ALEPLE 0.08, AMEAME 0.11, OMPOMP 0.31.
<emphasis box="[574,697,2004,2030]" italics="true" pageId="9" pageNumber="270">Chelicerae</emphasis>
: length 2.72, width 1.53, dorsal-anterior bristles present, interc- heliceral tumescence small, well-marked and with setae (
<figureCitation box="[811,902,153,179]" captionStart="FIGURE" captionStartId="9.[151,251,1321,1346]" captionTargetBox="[151,1436,309,1296]" captionTargetId="figure@9.[151,1436,309,1296]" captionText="FIGURE. 3. Lepthercus confusus sp. nov. (male holotype, NCA 2011/318). A cephalothorax and abdomen, dorsal view; B sternum, ventral view; C chelicerae, prolateral view, arrow indicates the intercheliceral tumesence; D right pedipalp, arrow indicates the small dorsal setae, and a normal anterior part of the retrolateral side of ventral excavation, retrolateral view; E right pedipalp, prolateral view, arrow indicates the abundant proventral rigid setae; FH copulatory bulb; F retrolateral view, G ventral view, H prolateral view; F, H arrow indicates the longitudinal keels along the embolus, G arrow indicates the small lateral keel; I Tibia I, arrow indicates the small cuticular spur." figureDoi="http://doi.org/10.5281/zenodo.3765473" httpUri="https://zenodo.org/record/3765473/files/figure.png" pageId="10" pageNumber="271">Fig. 3C</figureCitation>
). Fang groove with 6 promarginal teeth and 7 basomedial denticles.
<emphasis box="[402,488,189,215]" italics="true" pageId="10" pageNumber="271">Labium</emphasis>
: length 0.35, width 0.80, without cuspules.
<emphasis box="[991,1091,189,215]" italics="true" pageId="10" pageNumber="271">Maxillae</emphasis>
: length 1.60, width 0.72, with about 65 minuscule cuspules on inner corner, prolateral face slightly curved, with long uniformly distributed hairs.
<emphasis box="[151,244,261,287]" italics="true" pageId="10" pageNumber="271">Sternum</emphasis>
(
<figureCitation box="[262,353,261,287]" captionStart="FIGURE" captionStartId="9.[151,251,1321,1346]" captionTargetBox="[151,1436,309,1296]" captionTargetId="figure@9.[151,1436,309,1296]" captionText="FIGURE. 3. Lepthercus confusus sp. nov. (male holotype, NCA 2011/318). A cephalothorax and abdomen, dorsal view; B sternum, ventral view; C chelicerae, prolateral view, arrow indicates the intercheliceral tumesence; D right pedipalp, arrow indicates the small dorsal setae, and a normal anterior part of the retrolateral side of ventral excavation, retrolateral view; E right pedipalp, prolateral view, arrow indicates the abundant proventral rigid setae; FH copulatory bulb; F retrolateral view, G ventral view, H prolateral view; F, H arrow indicates the longitudinal keels along the embolus, G arrow indicates the small lateral keel; I Tibia I, arrow indicates the small cuticular spur." figureDoi="http://doi.org/10.5281/zenodo.3765473" httpUri="https://zenodo.org/record/3765473/files/figure.png" pageId="10" pageNumber="271">Fig. 3B</figureCitation>
): length 2.71, maximum width 2.14.
<emphasis box="[782,888,261,287]" italics="true" pageId="10" pageNumber="271">Abdomen</emphasis>
: length 4.42. PMS: length 0.57; PLS: length of basal:medial:apical segments 0.80:0.75:1.11; total length 2.66.
<emphasis box="[869,1152,297,323]" italics="true" pageId="10" pageNumber="271">Lengths of legs and palp</emphasis>
: I 4.16, 2.24, 3.06, 3.13, 2.43, 15.02. II 3.95, 2.35, 2.67, 2.93, 2.21, 14.11. III 3.54, 1.89, 2.39, 3.39, 2.01, 13.22. IV 4.41, 2.15, 3.38, 4.77, 2.24, 16.95. Palp 2.23, 1.26, 1.39, —, 0.76, 5.64.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3765475" ID-Zenodo-Dep="3765475" httpUri="https://zenodo.org/record/3765475/files/figure.png" pageId="10" pageNumber="271" startId="10.[151,251,1604,1629]" targetBox="[151,1436,417,1579]" targetPageId="10">
<paragraph blockId="10.[151,1436,1604,1665]" pageId="10" pageNumber="271">
<emphasis bold="true" box="[151,281,1604,1629]" pageId="10" pageNumber="271">FIGURE. 4.</emphasis>
<taxonomicName authority="Ríos-Tamayo &amp; Lyle, 2020" authorityName="Ríos-Tamayo &amp; Lyle" authorityYear="2020" box="[288,495,1604,1628]" class="Arachnida" family="Nemesiidae" genus="Lepthercus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="confusus" status="sp. nov.">
<emphasis box="[288,495,1604,1628]" italics="true" pageId="10" pageNumber="271">Lepthercus confusus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[501,582,1604,1628]" pageId="10" pageNumber="271">
<taxonomicNameLabel box="[501,582,1604,1628]" pageId="10" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(female
<typeStatus box="[673,763,1605,1629]" pageId="10" pageNumber="271">paratype</typeStatus>
, NM 8787).
<emphasis bold="true" box="[892,910,1604,1629]" pageId="10" pageNumber="271">A</emphasis>
cephalothorax, dorsal view;
<emphasis bold="true" box="[1205,1222,1604,1628]" pageId="10" pageNumber="271">B</emphasis>
sternum, arrow indicates the anterior maxillary edge slightly concave, ventral view;
<emphasis bold="true" box="[805,823,1640,1665]" pageId="10" pageNumber="271">C</emphasis>
abdomen, dorsal view;
<emphasis bold="true" box="[1066,1084,1640,1664]" pageId="10" pageNumber="271">D</emphasis>
spermathecae.
</paragraph>
</caption>
<paragraph blockId="10.[151,1437,1711,2025]" pageId="10" pageNumber="271">
<emphasis box="[199,327,1711,1737]" italics="true" pageId="10" pageNumber="271">Chaetotaxy</emphasis>
: Leg I: femur, 1-1-1-1-1-1-1-1-1 d (centrals), 1-1-1-2 P SUP, 1-0-1-1 R SUP; patella, 1 P A; tibia, 1-1-2 P, 1-1 R, 1-2-1-2-3-2 V, a small spur, with an apical long and curved spine (longer than the spur,
<figureCitation box="[1342,1421,1747,1773]" captionStart="FIGURE" captionStartId="9.[151,251,1321,1346]" captionTargetBox="[151,1436,309,1296]" captionTargetId="figure@9.[151,1436,309,1296]" captionText="FIGURE. 3. Lepthercus confusus sp. nov. (male holotype, NCA 2011/318). A cephalothorax and abdomen, dorsal view; B sternum, ventral view; C chelicerae, prolateral view, arrow indicates the intercheliceral tumesence; D right pedipalp, arrow indicates the small dorsal setae, and a normal anterior part of the retrolateral side of ventral excavation, retrolateral view; E right pedipalp, prolateral view, arrow indicates the abundant proventral rigid setae; FH copulatory bulb; F retrolateral view, G ventral view, H prolateral view; F, H arrow indicates the longitudinal keels along the embolus, G arrow indicates the small lateral keel; I Tibia I, arrow indicates the small cuticular spur." figureDoi="http://doi.org/10.5281/zenodo.3765473" httpUri="https://zenodo.org/record/3765473/files/figure.png" pageId="10" pageNumber="271">Fig. 3I</figureCitation>
); metatarsus, 1-1-0-1 V POST, straight without protuberance; tarsus, 0. Leg II: femur, 1-1-1-1-1-1 d, 1-0-1-1 P SUP, 1-1-1 R SUP; patella, 1 P A; tibia, 1-1 P, 2-2-1-4 V; metatarsus, 1 D M, 2-2-2 V; tarsus, 0. Leg III: femur, 1-1-1-1 d, 1-1-1-1 P SUP, 1-1-1-1 R SUP; patella, 1-1-2 P, 1 R; tibia, 1-1 P, 1-1 D, 1-1 R, 2-1-2-3 V; metatarsus, 1-1-1 P, 2-1-2-2 D, 2-1-1-3 V; tarsus, 0. Leg IV: femur, 1-1-1-1-1 d, 1-1-1-1 P SUP, 1-1-1-1 R SUP; patella, 1 R; tibia, 1 P, 1-1-1 R, 2-2-3 V; metatarsus, 1-1 P, 1-1-2-1-2 D, 2-2-1-3 V; tarsus, 0. Palp (
<figureCitation box="[995,1116,1927,1953]" captionStart="FIGURE" captionStartId="7.[151,251,1647,1672]" captionTargetBox="[151,1436,381,1622]" captionTargetId="figure@7.[151,1436,381,1622]" captionTargetPageId="7" captionText="FIGURE. 2. Preening comb. A Lepthercus haddadi sp. nov. (female paratype, NCA 2007/3847). Maxillary cuspules (males). B L. dippenaarae. C L. kwazuluensis. Serrula. D L. mandelai sp. nov. (female, NCA 2014/415A); E L. filmeri sp. nov. (female, NCA 91/1437); F L. haddadi sp. nov. (male, NCA 2007/1486); G L. haddadi sp. nov. (female, NCA 2007/3847); H L. engelbrechti sp. nov. (female, NCA 2018/385); I L. engelbrechti sp. nov. (male, NCA 2018/384); J L. dregei Purcell, 1902 (female, NCA 2012/2333); K L. sofiae sp. nov. (male, NCA 97/395)." figureDoi="http://doi.org/10.5281/zenodo.3765471" httpUri="https://zenodo.org/record/3765471/files/figure.png" pageId="10" pageNumber="271">Fig. 2D, E</figureCitation>
): femur, 1-1-1-3 d; patella, 1 p; tibia, 1-2 P, 1 RA, 1 v, (width/length: 0.56); tarsus, 0.
</paragraph>
<paragraph blockId="10.[151,1437,1711,2025]" lastBlockId="11.[151,1437,153,1403]" lastPageId="11" lastPageNumber="272" pageId="10" pageNumber="271">
<emphasis box="[199,302,1999,2025]" italics="true" pageId="10" pageNumber="271">Scopulae</emphasis>
: Metatarsi: III, dense, uniformly distributed throughout the segment; III, sparse (1:3A), divided by narrow band of setae; IV, sparse (A), divided by a wide band of setae. Tarsi: I-II, dense, uniformly distributed throughout the segment; III, sparse, uniformly distributed and divided by narrow band of setae; IV, sparse, uniformly distributed and divided by a narrow band of setae.
<emphasis box="[821,977,225,251]" italics="true" pageId="11" pageNumber="272">Trichobothria</emphasis>
: Tibiae: I-II 10-11; III 10-10; IV 12-12. Metatarsi: I (4)1(1)1(2)1; II (5)1(3)1(5)1; III (6)1(2)1(6)1; IV (4)1(3)1(4)1(6)1. Tarsi: I-II 13; III-IV 12.
<emphasis italics="true" pageId="11" pageNumber="272">Preening combs</emphasis>
: III 2 R V2 P V, IV 3/2 R V.
<emphasis box="[567,752,297,323]" italics="true" pageId="11" pageNumber="272">Copulatory bulb</emphasis>
: small, with a short and acuminate embolus with basal keels and the tip curved (
<figureCitation box="[369,434,333,359]" captionStart="FIGURE" captionStartId="9.[151,251,1321,1346]" captionTargetBox="[151,1436,309,1296]" captionTargetId="figure@9.[151,1436,309,1296]" captionText="FIGURE. 3. Lepthercus confusus sp. nov. (male holotype, NCA 2011/318). A cephalothorax and abdomen, dorsal view; B sternum, ventral view; C chelicerae, prolateral view, arrow indicates the intercheliceral tumesence; D right pedipalp, arrow indicates the small dorsal setae, and a normal anterior part of the retrolateral side of ventral excavation, retrolateral view; E right pedipalp, prolateral view, arrow indicates the abundant proventral rigid setae; FH copulatory bulb; F retrolateral view, G ventral view, H prolateral view; F, H arrow indicates the longitudinal keels along the embolus, G arrow indicates the small lateral keel; I Tibia I, arrow indicates the small cuticular spur." figureDoi="http://doi.org/10.5281/zenodo.3765473" httpUri="https://zenodo.org/record/3765473/files/figure.png" pageId="11" pageNumber="272">Fig. 3</figureCitation>
FH), spermatic duct slightly curved.
<emphasis box="[864,929,333,359]" italics="true" pageId="11" pageNumber="272">Color</emphasis>
: Carapace and legs brown-orange. Abdomen dorsally dark without bands, covered with short and long dark hairs; sternum and ventral abdomen lighter; spinnerets yellowish with dark spots.
</paragraph>
<paragraph blockId="11.[151,1437,153,1403]" pageId="11" pageNumber="272">
Female
<typeStatus box="[289,385,441,467]" pageId="11" pageNumber="272">paratype</typeStatus>
(NM8787,
<figureCitation box="[517,582,441,467]" captionStart="FIGURE" captionStartId="10.[151,251,1604,1629]" captionTargetBox="[151,1436,417,1579]" captionTargetId="figure@10.[151,1436,417,1579]" captionText="FIGURE. 4. Lepthercus confusus sp. nov. (female paratype, NM 8787). A cephalothorax, dorsal view; B sternum, arrow indicates the anterior maxillary edge slightly concave, ventral view; C abdomen, dorsal view; D spermathecae." figureDoi="http://doi.org/10.5281/zenodo.3765475" httpUri="https://zenodo.org/record/3765475/files/figure.png" pageId="11" pageNumber="272">Fig. 4</figureCitation>
AE). Total length 12.07.
<emphasis box="[875,985,441,467]" italics="true" pageId="11" pageNumber="272">Carapace</emphasis>
(
<figureCitation box="[1001,1092,441,467]" captionStart="FIGURE" captionStartId="10.[151,251,1604,1629]" captionTargetBox="[151,1436,417,1579]" captionTargetId="figure@10.[151,1436,417,1579]" captionText="FIGURE. 4. Lepthercus confusus sp. nov. (female paratype, NM 8787). A cephalothorax, dorsal view; B sternum, arrow indicates the anterior maxillary edge slightly concave, ventral view; C abdomen, dorsal view; D spermathecae." figureDoi="http://doi.org/10.5281/zenodo.3765475" httpUri="https://zenodo.org/record/3765475/files/figure.png" pageId="11" pageNumber="272">Fig. 4A</figureCitation>
): length 4.06, width 2.74, with small marginal bristles and sparse pubescence.
<emphasis box="[685,866,477,503]" italics="true" pageId="11" pageNumber="272">Cephalic region</emphasis>
: length 2.61, clypeus small (almost absent) with 3 marginal bristles; 4 bristles in front of the OQ and 3 between the PME, line of small bristles until the fovea.
<emphasis box="[1357,1436,514,539]" italics="true" pageId="11" pageNumber="272">Fovea:</emphasis>
width 0.34, almost straight (slightly procurved).
<emphasis box="[690,934,549,575]" italics="true" pageId="11" pageNumber="272">Ocular measurements</emphasis>
: AME 0.10, ALE 0.23, PME 0.13, PLE 0.17, OQ length 0.36, width 0.72; AMEALE 0.04, PMEPLE 0.03, AMEPME 0.06, ALEPLE 0.02, AMEAME 0.09, PMEPME 0.28.
<emphasis box="[351,474,621,647]" italics="true" pageId="11" pageNumber="272">Chelicerae</emphasis>
(
<figureCitation box="[490,581,621,647]" captionStart="FIGURE" captionStartId="10.[151,251,1604,1629]" captionTargetBox="[151,1436,417,1579]" captionTargetId="figure@10.[151,1436,417,1579]" captionText="FIGURE. 4. Lepthercus confusus sp. nov. (female paratype, NM 8787). A cephalothorax, dorsal view; B sternum, arrow indicates the anterior maxillary edge slightly concave, ventral view; C abdomen, dorsal view; D spermathecae." figureDoi="http://doi.org/10.5281/zenodo.3765475" httpUri="https://zenodo.org/record/3765475/files/figure.png" pageId="11" pageNumber="272">Fig. 4D</figureCitation>
): length 2.30, width 1.46; with dorsal-retrolateral dark bristles. Fang groove with 8 promarginal teeth and 18 mesobasal denticles.
<emphasis box="[769,855,657,683]" italics="true" pageId="11" pageNumber="272">Labium</emphasis>
: length 0.27, width 0.67, dorsally slightly concave without cuspules.
<emphasis box="[353,453,693,719]" italics="true" pageId="11" pageNumber="272">Maxillae</emphasis>
: length 1.43, width 0.69, with 85 cuspules on posterior inner corner, prolateral face concave, soft, with long uniformly distributed hairs, serrula present on anterior inner corner.
<emphasis box="[1149,1242,729,755]" italics="true" pageId="11" pageNumber="272">Sternum</emphasis>
(
<figureCitation box="[1257,1346,729,755]" captionStart="FIGURE" captionStartId="10.[151,251,1604,1629]" captionTargetBox="[151,1436,417,1579]" captionTargetId="figure@10.[151,1436,417,1579]" captionText="FIGURE. 4. Lepthercus confusus sp. nov. (female paratype, NM 8787). A cephalothorax, dorsal view; B sternum, arrow indicates the anterior maxillary edge slightly concave, ventral view; C abdomen, dorsal view; D spermathecae." figureDoi="http://doi.org/10.5281/zenodo.3765475" httpUri="https://zenodo.org/record/3765475/files/figure.png" pageId="11" pageNumber="272">Fig. 4C</figureCitation>
): length 1.98, maximum width 1.64.
<emphasis box="[463,569,765,791]" italics="true" pageId="11" pageNumber="272">Abdomen</emphasis>
(
<figureCitation box="[583,670,765,791]" captionStart="FIGURE" captionStartId="10.[151,251,1604,1629]" captionTargetBox="[151,1436,417,1579]" captionTargetId="figure@10.[151,1436,417,1579]" captionText="FIGURE. 4. Lepthercus confusus sp. nov. (female paratype, NM 8787). A cephalothorax, dorsal view; B sternum, arrow indicates the anterior maxillary edge slightly concave, ventral view; C abdomen, dorsal view; D spermathecae." figureDoi="http://doi.org/10.5281/zenodo.3765475" httpUri="https://zenodo.org/record/3765475/files/figure.png" pageId="11" pageNumber="272">Fig. 4B</figureCitation>
): length 6.43, with small hairs in the dorsum. PMS: length 0.61; PLS: length of basal:medial:apical segments 1.23:0.79:1.16; total length 3.18.
<emphasis box="[986,1271,801,827]" italics="true" pageId="11" pageNumber="272">Lengths of legs and palp</emphasis>
: I: 2.63, 1.72, 1.85, 1.38, 0.98, 8.56. II: 2.41, 1.45, 1.51, 1.33, 0.98, 7.68. III: 2.21, 1.35, 1.36, 1.80, 1.02, 7.74. IV: 2.88, 1.66, 2.04, 1.74, 1.15, 9.47. Palp: 1.88, 1.21, 1.14, —, 1.24, 5.47.
</paragraph>
<paragraph blockId="11.[151,1437,153,1403]" pageId="11" pageNumber="272">
<emphasis box="[199,327,909,935]" italics="true" pageId="11" pageNumber="272">Chaetotaxy</emphasis>
: Leg I: femur, 1-1-1-1-1 d, 1 P A; patella, 0; tibia, 1 p, 1-1-2 v; metatarsus, 2-1-1 V; tarsus, 0. Leg II: femur, 1-1-1-1-1-1 d, 1 P A; patella, 0; tibia, 1 p, 2-2-3 v; metatarsus, 2-2-2 V; tarsus, 0. Leg III: femur, 1-1-1-1 d, 1 P A, 1-1 R A; patella, 1-1 P, 1 R; tibia, 1-1 P, 1-1-1 D, 1 R, 2-2-3 V (thins); metatarsus, 2-1-1-1 P, 1-1-1 DP, 2- 1-2 DR, 3-3-3 V; tarsus, 0. Leg IV: femur, 1-1-1-1-1 d, 1 R A; patella, 1 R; tibia, 1-1 P, 1-1 R, 2-2-3 V; metatarsus, 1-1-1-1 P, 1-1-1 DP, 1-1-0-1 DR, 1-1-1 R, 4-1-2-3 V; tarsus, 0. Palp: femur, 1-1-1 d, 1 pa; patella, 0; tibia, 1-1 P, 2-3 V (1:2A); tarsus, 2 V B.
</paragraph>
<paragraph blockId="11.[151,1437,153,1403]" pageId="11" pageNumber="272">
<emphasis box="[199,302,1125,1151]" italics="true" pageId="11" pageNumber="272">Scopulae</emphasis>
: Metatarsi: I, dense, uniformly distributed and divided by bristles; II, dense (1:2 A), divided by band of setae; IIIIV, no present. Tarsi: I, dense, uniformly distributed and divided by bristles; II, sparse, uniformly distributed and divided by setae; III, sparse (1:3 A) and divided by a narrow band of setae; IV, sparse (3:4 A) and divided by a wide band of setae.
<emphasis box="[432,588,1233,1259]" italics="true" pageId="11" pageNumber="272">Trichobothria</emphasis>
: Tibiae: palp 7-8; I 10-9; II 9-9; III 8-8; IV 10-9. Metatarsi: I (3)1(1)1(4)1; II-III (4)1(2)1(4)1; IV (4)1(2)1(3)1(5)1. Tarsi: palp 8; I 12; II-IV 11.
<emphasis box="[930,1111,1269,1295]" italics="true" pageId="11" pageNumber="272">Preening combs</emphasis>
: metatarsus: II 5 P V; III 3 P V3 R V; IV 2 P V3 R V.
<emphasis box="[445,604,1305,1331]" italics="true" pageId="11" pageNumber="272">Spermathecae</emphasis>
: as in
<figureCitation box="[675,760,1305,1331]" captionStart="FIGURE" captionStartId="10.[151,251,1604,1629]" captionTargetBox="[151,1436,417,1579]" captionTargetId="figure@10.[151,1436,417,1579]" captionText="FIGURE. 4. Lepthercus confusus sp. nov. (female paratype, NM 8787). A cephalothorax, dorsal view; B sternum, arrow indicates the anterior maxillary edge slightly concave, ventral view; C abdomen, dorsal view; D spermathecae." figureDoi="http://doi.org/10.5281/zenodo.3765475" httpUri="https://zenodo.org/record/3765475/files/figure.png" pageId="11" pageNumber="272">Fig. 4E</figureCitation>
.
<emphasis box="[770,835,1305,1331]" italics="true" pageId="11" pageNumber="272">Color</emphasis>
: Overall orange-yellowish, sternum lighter. Abdomen brown mottled with light spots on dorsal side, spinnerets light brown.
</paragraph>
</subSubSection>
<subSubSection box="[199,910,1376,1403]" pageId="11" pageNumber="272" type="distribution">
<paragraph blockId="11.[151,1437,153,1403]" box="[199,910,1376,1403]" pageId="11" pageNumber="272">
<emphasis bold="true" box="[199,353,1376,1403]" pageId="11" pageNumber="272">Distribution.</emphasis>
<collectingRegion box="[359,644,1377,1403]" country="South Africa" name="KwaZulu-Natal" pageId="11" pageNumber="272">KwaZulu-Natal Province</collectingRegion>
,
<collectingCountry box="[656,798,1377,1403]" name="South Africa" pageId="11" pageNumber="272">South Africa</collectingCountry>
(
<figureCitation box="[813,898,1377,1403]" captionStart="FIGURE" captionStartId="38.[151,251,1816,1841]" captionTargetBox="[185,1405,888,1784]" captionTargetId="figure@38.[176,1411,885,1790]" captionTargetPageId="38" captionText="FIGURE. 24. Distribution map of Lepthercus spp: ● Lepthercus confusus sp. nov., ○ Lepthercus dippenaarae sp. nov., Ɨ Lepthercus dregei Purcell, 1902, O Lepthercus engelbrechti sp. nov., A Lepthercus filmeri sp. nov., * Lepthercus haddadi sp. nov., D Lepthercus kwazuluensis sp. nov., ■ Lepthercus lawrencei sp. nov., v Lepthercus mandelai sp. nov., ▼ Lepthercus rattrayi Hewitt, 1917, ▲ Lepthercus sofiae sp. nov. Solid black line indicates the two species overlapped." figureDoi="http://doi.org/10.5281/zenodo.3765519" httpUri="https://zenodo.org/record/3765519/files/figure.png" pageId="11" pageNumber="272">Fig. 24</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>