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<document id="4C42D8B3D3043F8F649C4527CC78C9FD" ID-DOI="10.26107/RBZ-2022-0009" ID-ISSN="2345-7600" ID-Zenodo-Dep="7174234" ID-ZooBank="BE6164AE-1C24-4E01-8B7B-D80764F147B3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665290677448" checkinUser="felipe" docAuthor="Ng, Peter K. L. &amp; Ahyong, Shane T." docDate="2022" docId="03E887D8CE53FFF437FF18B2D09979FE" docLanguage="en" docName="RafflesBZool.70.134248.pdf" docOrigin="Raffles Bulletin of Zoology 70" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Arcotheres ocularius Komai, Kawai &amp; Ng 2020" docType="treatment" docVersion="4" lastPageNumber="171" masterDocId="FFD1FFA0CE71FFD137601D0FD0617E6B" masterDocTitle="The pea crab genus Arcotheres Manning, 1993 (Crustacea: Brachyura: Pinnotheridae) from Singapore and Peninsular Malaysia, with a reappraisal of diagnostic characters and descriptions of two new genera" masterLastPageNumber="248" masterPageNumber="134" pageNumber="168" updateTime="1699379806661" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="81F8F4428CA7B4D2DBC0829385C6A122">The pea crab genus Arcotheres Manning, 1993 (Crustacea: Brachyura: Pinnotheridae) from Singapore and Peninsular Malaysia, with a reappraisal of diagnostic characters and descriptions of two new genera</mods:title>
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<mods:namePart id="E360131CA3F96B68930E6CA5D1CF3D63">Ng, Peter K. L.</mods:namePart>
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<taxonomicName id="4C414D4DCE53FFF337FF18B2D2A27BBE" ID-CoL="84GPY" authority="Komai, Kawai &amp; Ng, 2020" authorityName="Komai, Kawai &amp; Ng" authorityYear="2020" box="[159,707,1469,1493]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="168" phylum="Arthropoda" rank="species" species="ocularius">
<emphasis id="B935EADCCE53FFF337FF18B2D2A27BBE" bold="true" box="[159,707,1469,1493]" pageId="34" pageNumber="168">
<emphasis id="B935EADCCE53FFF337FF18B2D1E67BBE" bold="true" box="[159,391,1469,1493]" italics="true" pageId="34" pageNumber="168">Arcotheres ocularius</emphasis>
<bibRefCitation id="EFD04B3FCE53FFF336F018B2D2A27BBE" author="Komai T &amp; Kawai K &amp; Ng PKL" box="[400,707,1469,1493]" pageId="34" pageNumber="168" pagination="221 - 247" refId="ref65974" refString="Komai T, Kawai K &amp; Ng PKL (2020) On the identity of the poorly known pea crab, Pinnothera obesa Dana, 1852, and description of a new species of Arcotheres Manning, 1993 from the southwest Pacific (Decapoda: Brachyura: Pinnotheridae). Zootaxa, 4822 (2): 221 - 247." type="journal article" year="2020">Komai, Kawai &amp; Ng, 2020</bibRefCitation>
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<subSubSection id="C35B6545CE53FFF3360718D2D19A7B9E" box="[359,507,1501,1525]" pageId="34" pageNumber="168" type="description">
<paragraph id="8BFE36CECE53FFF3360718D2D19A7B9E" blockId="34.[159,707,1469,1525]" box="[359,507,1501,1525]" pageId="34" pageNumber="168">
(
<figureCitation id="137A2A4BCE53FFF3360F18D2D1AB7B9E" box="[367,458,1501,1525]" captionStart="Fig" captionStartId="35.[136,171,1780,1802]" captionTargetBox="[136,1474,190,1765]" captionTargetId="figure-7@35.[136,1474,190,1765]" captionTargetPageId="35" captionText="Fig. 23. Arcotheres ocularius Komai, Kawai &amp; Ng, 2020, overall dorsal view. A, paratype ovigerous female (10.9 × 9.7 mm) (ZRC 2019.1877), Fiji; B, female (9.4 × 8.4 mm) (ZRC 2019.1025), Indonesia." figureDoi="http://doi.org/10.5281/zenodo.7174281" httpUri="https://zenodo.org/record/7174281/files/figure.png" pageId="34" pageNumber="168">Figs. 23</figureCitation>
,
<figureCitation id="137A2A4BCE53FFF336B718D2D1927B9E" box="[471,499,1501,1525]" captionStart="Fig" captionStartId="36.[113,148,1758,1780]" captionTargetBox="[116,1455,191,1741]" captionTargetId="figure-15@36.[103,1468,187,1753]" captionTargetPageId="36" captionText="Fig. 24. Arcotheres ocularius Komai, Kawai &amp; Ng, 2020. A, CK, paratype female (10.9 × 9.7 mm) (ZRC 2019.1877), Fiji; B, female (9.4 × 8.4 mm) (ZRC 2019.1025), Indonesia. A, B, left MXP3; C, outer view of right chela; DG, left P2P5, respectively; HK, right P2P5, respectively. Scales = A, B, 0.5 mm; CK, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174283" httpUri="https://zenodo.org/record/7174283/files/figure.png" pageId="34" pageNumber="168">24</figureCitation>
)
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<taxonomicName id="4C414D4DCE53FFF337111B13D2E5785A" ID-CoL="84GPY" authority="Komai, Kawai &amp; Ng, 2020: 232" authorityName="Komai, Kawai &amp; Ng" authorityPageNumber="232" authorityYear="2020" box="[113,644,1564,1586]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="168" phylum="Arthropoda" rank="species" species="ocularius">
<emphasis id="B935EADCCE53FFF337111B13D15C785A" box="[113,317,1564,1585]" italics="true" pageId="34" pageNumber="168">Arcotheres ocularius</emphasis>
<bibRefCitation id="EFD04B3FCE53FFF336251B13D2E5785A" author="Komai T &amp; Kawai K &amp; Ng PKL" box="[325,644,1564,1586]" pageId="34" pageNumber="168" pagination="221 - 247" refId="ref65974" refString="Komai T, Kawai K &amp; Ng PKL (2020) On the identity of the poorly known pea crab, Pinnothera obesa Dana, 1852, and description of a new species of Arcotheres Manning, 1993 from the southwest Pacific (Decapoda: Brachyura: Pinnotheridae). Zootaxa, 4822 (2): 221 - 247." type="journal article" year="2020">Komai, Kawai &amp; Ng, 2020: 232</bibRefCitation>
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, figs. 717 [
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locality:
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];
<bibRefCitation id="EFD04B3FCE53FFF3363F1B36D2347825" author="De Gier W &amp; Becker C" box="[351,597,1593,1615]" pageId="34" pageNumber="168" pagination="431" refId="ref64297" refString="De Gier W &amp; Becker C (2020) A review of the ecomorphology of pinnotherine pea crabs (Brachyura: Pinnotheridae), with an updated list of symbiont-host associations. Diversity, 12: 431." type="journal article" year="2020">De Gier &amp; Becker, 2020</bibRefCitation>
: tab. 1.
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: ovigerous female (11.8 ×
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15952), Kumi Village,
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Levu,
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, intertidal, with
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sp.
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(
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:
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),
<date id="FFFF100ECE53FFF337101BDBD1427887" box="[112,291,1748,1772]" pageId="34" pageNumber="168" value="2010-08-16">16 August 2010</date>
, coll. K. Kawai.
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:
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(8.9 ×
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), 3 ovigerous females (9.6 ×
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, 10.9 ×
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, 12.3 ×
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) (
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2019.1877), coll.
<date id="FFFF100ECE53FFF3351E1A1BD0CB7927" pageId="34" pageNumber="168" value="2010-08-16">16 August 2010</date>
;
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(5.5 × 5.0 mm, 6.9 ×
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), 17 ovigerous females (8.5 × 7.413.4 ×
<quantity id="4CB99B2BCE53FFF336EF1A5BD1827907" box="[399,483,1876,1900]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="34" pageNumber="168" unit="mm" value="9.6">9.6 mm</quantity>
) (
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15953), coll.
<date id="FFFF100ECE53FFF337121A7BD14379E7" box="[114,290,1908,1932]" pageId="34" pageNumber="168" value="2010-08-16">16 August 2010</date>
;
<specimenCount id="9D47FD47CE53FFF3364D1A7BD11779E7" box="[301,374,1908,1932]" pageId="34" pageNumber="168" type="male">1 male</specimenCount>
(6.2 ×
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) (
<collectionCode id="ED50AE0BCE53FFF3354E1A7BD2F779E7" box="[558,662,1908,1932]" pageId="34" pageNumber="168">CBM-ZC</collectionCode>
15954);
<specimenCount id="9D47FD47CE53FFF337121A9BD0AE79C7" box="[114,207,1940,1964]" pageId="34" pageNumber="168" type="female">1 female</specimenCount>
(5.2 ×
<quantity id="4CB99B2BCE53FFF3367C1A9BD10E79C7" box="[284,367,1940,1964]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="34" pageNumber="168" unit="mm" value="5.2">5.2 mm</quantity>
) (
<collectionCode id="ED50AE0BCE53FFF336E41A9BD18D79C7" box="[388,492,1940,1964]" pageId="34" pageNumber="168">CBM-ZC</collectionCode>
15956), coll.
<date id="FFFF100ECE53FFF335E21A9BD0CA79A7" pageId="34" pageNumber="168" value="2010-08-16">16 August 2010</date>
;
<specimenCount id="9D47FD47CE53FFF337D91ABBD16579A7" box="[185,260,1972,1996]" pageId="34" pageNumber="168" type="male">1 male</specimenCount>
(4.6 × 5.0 mm) (
<collectionCode id="ED50AE0BCE53FFF336AC1ABBD25479A7" box="[460,565,1972,1996]" pageId="34" pageNumber="168">CBM-ZC</collectionCode>
15957), coll.
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;
<specimenCount id="9D47FD47CE53FFF336681ADBD13B7987" box="[264,346,2004,2028]" pageId="34" pageNumber="168" type="male">2 males</specimenCount>
(3.2 ×
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, 3.3 ×
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) (
<collectionCode id="ED50AE0BCE53FFF335CA1ADBD0F27667" pageId="34" pageNumber="168">CBMZC</collectionCode>
15958), coll.
<date id="FFFF100ECE53FFF336511AFBD1847667" box="[305,485,2036,2060]" pageId="34" pageNumber="168" value="2010-08-16">16 August 2010</date>
;
<specimenCount id="9D47FD47CE53FFF336921AFBD25C7667" box="[498,573,2036,2060]" pageId="34" pageNumber="168" type="male">1 male</specimenCount>
(5.8 ×
<quantity id="4CB99B2BCE53FFF335F31AFBD2887667" box="[659,745,2036,2060]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="34" pageNumber="168" unit="mm" value="5.8">5.8 mm</quantity>
) (
<collectionCode id="ED50AE0BCE53FFF334541DB2D3FD7EBE" box="[820,924,189,213]" pageId="34" pageNumber="168">CBM-ZC</collectionCode>
15959);
<specimenCount id="9D47FD47CE53FFF333621DB2D42F7EBE" box="[1026,1102,189,213]" pageId="34" pageNumber="168" type="male">1 male</specimenCount>
(4.8 ×
<quantity id="4CB99B2BCE53FFF333C41DB2D49A7EBE" box="[1188,1275,189,213]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="34" pageNumber="168" unit="mm" value="5.4">5.4 mm</quantity>
) (
<collectionCode id="ED50AE0BCE53FFF332731DB2D5287EBE" box="[1299,1353,189,213]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="34" pageNumber="168">ZRC</collectionCode>
2020.7), coll.
<date id="FFFF100ECE53FFF334031DD2D4747E9E" box="[867,1045,221,246]" pageId="34" pageNumber="168" value="2010-08-16">16 August 2010</date>
; 1 ovigerous female (10.7 ×
<quantity id="4CB99B2BCE53FFF3322F1DD2D5C57E9E" box="[1359,1444,221,245]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="34" pageNumber="168" unit="mm" value="9.1">9.1 mm</quantity>
) (
<collectionCode id="ED50AE0BCE53FFF334551DF2D3C17F7E" box="[821,928,253,277]" pageId="34" pageNumber="168">CBM-ZC</collectionCode>
15960), coll.
<date id="FFFF100ECE53FFF333261DF2D5787F7E" box="[1094,1305,253,277]" pageId="34" pageNumber="168" value="2011-09-06">6 September 2011</date>
; 7 ovigerous females (8.6 × 7.210.7 ×
<quantity id="4CB99B2BCE53FFF3335A1C12D4EA7F5E" box="[1082,1163,285,309]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="34" pageNumber="168" unit="mm" value="9.5">9.5 mm</quantity>
) (
<collectionCode id="ED50AE0BCE53FFF333FE1C12D5647F5E" box="[1182,1285,285,309]" pageId="34" pageNumber="168">CBM-ZC</collectionCode>
15961),
<date id="FFFF100ECE53FFF3323F1C11D3057F3E" pageId="34" pageNumber="168" value="2018-08">August 2018</date>
; 5 ovigerous females (9.4 × 8.010.7 ×
<quantity id="4CB99B2BCE53FFF3326E1C32D5007F3E" box="[1294,1377,317,341]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="34" pageNumber="168" unit="mm" value="9.3">9.3 mm</quantity>
) (
<collectionCode id="ED50AE0BCE53FFF332161C32D5CD7F3E" box="[1398,1452,317,341]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="34" pageNumber="168">ZRC</collectionCode>
2020.8),
<date id="FFFF100ECE53FFF334EE1C51D4797F1E" box="[910,1048,349,374]" pageId="34" pageNumber="168" value="2018-08">August 2018</date>
(all from same locality as
<typeStatus id="54FA886CCE53FFF3325E1C52D5C17F1E" box="[1342,1440,349,373]" pageId="34" pageNumber="168" type="holotype">holotype</typeStatus>
); 1 ovigerous female (
<collectionCode id="ED50AE0BCE53FFF333691C72D4107FFE" box="[1033,1137,381,405]" pageId="34" pageNumber="168">CBM-ZC</collectionCode>
19562), Ucikumi, near Kumi Village, coll.
<date id="FFFF100ECE53FFF334A21C92D4E67FDE" box="[962,1159,413,437]" pageId="34" pageNumber="168" value="2019-02-22">22 February 2019</date>
.
</paragraph>
<paragraph id="8BFE36CECE53FFF3344D1CD2D51D7CFE" blockId="34.[811,1452,477,661]" pageId="34" pageNumber="168">
<emphasis id="B935EADCCE53FFF3344D1CD2D4E67F9E" bold="true" box="[813,1159,477,501]" pageId="34" pageNumber="168">
Other material examined.
<collectingCountry id="F356765ECE53FFF333361CD2D4E37F9E" box="[1110,1154,477,501]" name="Fiji" pageId="34" pageNumber="168">Fiji</collectingCountry>
:
</emphasis>
4 poorly preserved females (4.2 × 3.36.0 ×
<quantity id="4CB99B2BCE53FFF334961CF2D4317C7E" box="[1014,1104,509,533]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="34" pageNumber="168" unit="mm" value="4.6">4.6 mm</quantity>
) (
<collectionCode id="ED50AE0BCE53FFF3330A1CF2D4C07C7E" box="[1130,1185,509,533]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="34" pageNumber="168">ZRC</collectionCode>
2019.1878), intertidal, associated with
<taxonomicName id="4C414D4DCE53FFF334BB1F12D4017C5E" box="[987,1120,541,565]" class="Bivalvia" family="Arcidae" genus="Anadara" kingdom="Animalia" order="Arcida" pageId="34" pageNumber="168" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B935EADCCE53FFF334BB1F12D4597C5E" box="[987,1080,541,565]" italics="true" pageId="34" pageNumber="168">Anadara</emphasis>
sp.
</taxonomicName>
, coll. K. Kawai,
<date id="FFFF100ECE53FFF3327A1F11D5C97C5E" box="[1306,1448,541,566]" pageId="34" pageNumber="168" value="2018-08">August 2018</date>
.
<emphasis id="B935EADCCE53FFF3344E1F32D3C47C3E" bold="true" box="[814,933,573,597]" pageId="34" pageNumber="168">
<collectingCountry id="F356765ECE53FFF3344E1F32D3C17C3E" box="[814,928,573,597]" name="Indonesia" pageId="34" pageNumber="168">Indonesia</collectingCountry>
:
</emphasis>
<specimenCount id="9D47FD47CE53FFF334C81F32D4627C3E" box="[936,1027,573,597]" pageId="34" pageNumber="168" type="female">1 female</specimenCount>
(9.4 ×
<quantity id="4CB99B2BCE53FFF3332B1F32D4FD7C3E" box="[1099,1180,573,597]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="34" pageNumber="168" unit="mm" value="8.4">8.4 mm</quantity>
) (
<collectionCode id="ED50AE0BCE53FFF333CF1F32D4857C3E" box="[1199,1252,573,597]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="34" pageNumber="168">ZRC</collectionCode>
2019.1025), Teluk Kodek, Lombok, intertidal, in
<taxonomicName id="4C414D4DCE53FFF3330B1F52D3BA7CFE" authority="(Linnaeus, 1758)" authorityName="" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Bivalvia" family="Arcidae" genus="Anadara" kingdom="Animalia" order="Arcida" pageId="34" pageNumber="168" phylum="Mollusca" rank="species" species="antiquata">
<emphasis id="B935EADCCE53FFF3330B1F52D5527C1E" box="[1131,1331,605,629]" italics="true" pageId="34" pageNumber="168">Anadara antiquata</emphasis>
(Linnaeus, 1758) (Arcidae)
</taxonomicName>
, coll. S. Dwiono,
<date id="FFFF100ECE53FFF333C21F72D5197CFE" box="[1186,1400,637,661]" pageId="34" pageNumber="168" value="2013-11-12">12 November 2013</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C35B6545CE53FFF4344D1FB2D0A17C1E" lastPageId="37" lastPageNumber="171" pageId="34" pageNumber="168" type="description">
<paragraph id="8BFE36CECE53FFF3344D1FB2D3A07A7E" blockId="34.[811,1454,701,1045]" pageId="34" pageNumber="168">
<emphasis id="B935EADCCE53FFF3344D1FB2D3A07CBE" bold="true" box="[813,961,701,725]" pageId="34" pageNumber="168">Description.</emphasis>
Carapace and pereopods well chitinised. Female: Carapace subcircular, wider than long; dorsal and lateral surfaces smooth, glabrous; appearing domed in frontal view; front projecting anteriorly beyond orbits, margin gently convex to almost straight; anterolateral margin smooth, subparallel with frontal margin or gently sloping posteriorly, forming rounded angle with posterolateral margin (
<figureCitation id="137A2A4BCE53FFF3322E1E72D5FF7DFE" box="[1358,1438,893,917]" captionStart="Fig" captionStartId="35.[136,171,1780,1802]" captionTargetBox="[136,1474,190,1765]" captionTargetId="figure-7@35.[136,1474,190,1765]" captionTargetPageId="35" captionText="Fig. 23. Arcotheres ocularius Komai, Kawai &amp; Ng, 2020, overall dorsal view. A, paratype ovigerous female (10.9 × 9.7 mm) (ZRC 2019.1877), Fiji; B, female (9.4 × 8.4 mm) (ZRC 2019.1025), Indonesia." figureDoi="http://doi.org/10.5281/zenodo.7174281" httpUri="https://zenodo.org/record/7174281/files/figure.png" pageId="34" pageNumber="168">Fig. 23</figureCitation>
). Eyes clearly visible in dorsal view in adults; mobile, filling orbit (
<figureCitation id="137A2A4BCE53FFF334101EB2D3A37DBE" box="[880,962,957,981]" captionStart="Fig" captionStartId="35.[136,171,1780,1802]" captionTargetBox="[136,1474,190,1765]" captionTargetId="figure-7@35.[136,1474,190,1765]" captionTargetPageId="35" captionText="Fig. 23. Arcotheres ocularius Komai, Kawai &amp; Ng, 2020, overall dorsal view. A, paratype ovigerous female (10.9 × 9.7 mm) (ZRC 2019.1877), Fiji; B, female (9.4 × 8.4 mm) (ZRC 2019.1025), Indonesia." figureDoi="http://doi.org/10.5281/zenodo.7174281" httpUri="https://zenodo.org/record/7174281/files/figure.png" pageId="34" pageNumber="168">Fig. 23</figureCitation>
). Epistome with median part low, triangular, lateral margins wide, gently concave (
<bibRefCitation id="EFD04B3FCE53FFF333B51ED2D5CB7D9E" author="Komai T &amp; Kawai K &amp; Ng PKL" box="[1237,1450,989,1013]" pageId="34" pageNumber="168" pagination="221 - 247" refId="ref65974" refString="Komai T, Kawai K &amp; Ng PKL (2020) On the identity of the poorly known pea crab, Pinnothera obesa Dana, 1852, and description of a new species of Arcotheres Manning, 1993 from the southwest Pacific (Decapoda: Brachyura: Pinnotheridae). Zootaxa, 4822 (2): 221 - 247." type="journal article" year="2020">Komai et al., 2020</bibRefCitation>
: figs. 7D, 8B).
</paragraph>
<paragraph id="8BFE36CECE53FFF3344D1932D3A67B3E" blockId="34.[811,1453,1085,1365]" pageId="34" pageNumber="168">
MXP3 outer surface with scattered short setae; ischiomerus completely fused, subrhomboidal, inner margin rounded to gently angular at widest point; carpus short; propodus about 3 times as long as high, subspatulate, longer than carpus, tip rounded; dactylus digitiform, inserted submedially on flexor margin of propodus, tip reaching end of or slightly overreaching propodal apex; exopod relatively slender, about two-thirds length of ischiomerus, flagellum 2-segmented (
<figureCitation id="137A2A4BCE53FFF334551832D3DB7B3E" box="[821,954,1341,1365]" captionStart="Fig" captionStartId="36.[113,148,1758,1780]" captionTargetBox="[116,1455,191,1741]" captionTargetId="figure-15@36.[103,1468,187,1753]" captionTargetPageId="36" captionText="Fig. 24. Arcotheres ocularius Komai, Kawai &amp; Ng, 2020. A, CK, paratype female (10.9 × 9.7 mm) (ZRC 2019.1877), Fiji; B, female (9.4 × 8.4 mm) (ZRC 2019.1025), Indonesia. A, B, left MXP3; C, outer view of right chela; DG, left P2P5, respectively; HK, right P2P5, respectively. Scales = A, B, 0.5 mm; CK, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174283" httpUri="https://zenodo.org/record/7174283/files/figure.png" pageId="34" pageNumber="168">Fig. 24A, B</figureCitation>
).
</paragraph>
<paragraph id="8BFE36CECE53FFF3344D1872D424781E" blockId="34.[813,1452,1405,1653]" pageId="34" pageNumber="168">
Chela short, stout, dactylus about two-thirds palm length; palm relatively slender, medially widest; outer surfaces of palm, fingers (except for distal part) almost glabrous, with only scattered short setae; ventral margin of palm gently convex, with low bulge near midlength; dactylus occlusal margin with 1 subproximal tooth; pollex occlusal margin with 1 low sharp proximal tooth and denticles; tips of fingers sharp, hooked (
<figureCitation id="137A2A4BCE53FFF334B51B52D458781E" box="[981,1081,1629,1653]" captionStart="Fig" captionStartId="36.[113,148,1758,1780]" captionTargetBox="[116,1455,191,1741]" captionTargetId="figure-15@36.[103,1468,187,1753]" captionTargetPageId="36" captionText="Fig. 24. Arcotheres ocularius Komai, Kawai &amp; Ng, 2020. A, CK, paratype female (10.9 × 9.7 mm) (ZRC 2019.1877), Fiji; B, female (9.4 × 8.4 mm) (ZRC 2019.1025), Indonesia. A, B, left MXP3; C, outer view of right chela; DG, left P2P5, respectively; HK, right P2P5, respectively. Scales = A, B, 0.5 mm; CK, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174283" httpUri="https://zenodo.org/record/7174283/files/figure.png" pageId="34" pageNumber="168">Fig. 24C</figureCitation>
).
</paragraph>
<paragraph id="8BFE36CECE53FFF3344D1B92D4E2767E" blockId="34.[811,1452,1693,2069]" pageId="34" pageNumber="168">
P2P5 dorsally, ventrally unarmed; outer surface covered with scattered, very short setae or glabrous; ventral margins of propodus and dactylus slightly more setose; merus relatively short, stout, relative lengths of meri P4&gt;P3&gt;P2&gt;P5; left (sometimes right) P4 distinctly the longer; P2 and P3 dactyli short, subequal, tip gently hooked, half length of propodus; longer P4 dactylus elongate, almost straight, sparsely setose, longer than P5 dactylus; P5 merus 4.14.4 times longer than wide; P5 dactylus shorter than longer P5 dactylus, longer than shorter P4 dactylus, margins lined with short and long setae, denser on ventral margin, distoflexor margin without rows of spinules (
<figureCitation id="137A2A4BCE53FFF334901AF2D416767E" box="[1008,1143,2045,2069]" captionStart="Fig" captionStartId="36.[113,148,1758,1780]" captionTargetBox="[116,1455,191,1741]" captionTargetId="figure-15@36.[103,1468,187,1753]" captionTargetPageId="36" captionText="Fig. 24. Arcotheres ocularius Komai, Kawai &amp; Ng, 2020. A, CK, paratype female (10.9 × 9.7 mm) (ZRC 2019.1877), Fiji; B, female (9.4 × 8.4 mm) (ZRC 2019.1025), Indonesia. A, B, left MXP3; C, outer view of right chela; DG, left P2P5, respectively; HK, right P2P5, respectively. Scales = A, B, 0.5 mm; CK, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174283" httpUri="https://zenodo.org/record/7174283/files/figure.png" pageId="34" pageNumber="168">Fig. 24DK</figureCitation>
).
</paragraph>
<caption id="DF3E6646CE52FFF237E81BFBD336794C" ID-DOI="http://doi.org/10.5281/zenodo.7174281" ID-Zenodo-Dep="7174281" httpUri="https://zenodo.org/record/7174281/files/figure.png" pageId="35" pageNumber="169" startId="35.[136,171,1780,1802]" targetBox="[136,1474,190,1765]" targetPageId="35">
<paragraph id="8BFE36CECE52FFF237E81BFBD336794C" blockId="35.[136,1474,1780,1831]" pageId="35" pageNumber="169">
Fig. 23.
<taxonomicName id="4C414D4DCE52FFF237BF1BFBD2AC7962" authority="Komai, Kawai &amp; Ng, 2020" authorityName="Komai, Kawai &amp; Ng" authorityYear="2020" box="[223,717,1780,1802]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="169" phylum="Arthropoda" rank="species" species="ocularius">
<emphasis id="B935EADCCE52FFF237BF1BFBD1CF7962" box="[223,430,1780,1801]" italics="true" pageId="35" pageNumber="169">Arcotheres ocularius</emphasis>
<bibRefCitation id="EFD04B3FCE52FFF236D71BFBD2AC7962" author="Komai T &amp; Kawai K &amp; Ng PKL" box="[439,717,1780,1802]" pageId="35" pageNumber="169" pagination="221 - 247" refId="ref65974" refString="Komai T, Kawai K &amp; Ng PKL (2020) On the identity of the poorly known pea crab, Pinnothera obesa Dana, 1852, and description of a new species of Arcotheres Manning, 1993 from the southwest Pacific (Decapoda: Brachyura: Pinnotheridae). Zootaxa, 4822 (2): 221 - 247." type="journal article" year="2020">Komai, Kawai &amp; Ng, 2020</bibRefCitation>
</taxonomicName>
, overall dorsal view. A, paratype ovigerous female (10.9 × 9.7 mm) (ZRC 2019.1877), Fiji; B, female (9.4 × 8.4 mm) (ZRC 2019.1025), Indonesia.
</paragraph>
</caption>
<paragraph id="8BFE36CECE52FFF237E81A51D19E79DD" blockId="35.[136,775,1886,1974]" pageId="35" pageNumber="169">
Pleon extending to buccal region, covering bases of P2P5; telson gently recessed into concave distal margin of somite 6 (
<bibRefCitation id="EFD04B3FCE52FFF237C61A91D11979DD" author="Komai T &amp; Kawai K &amp; Ng PKL" box="[166,376,1950,1974]" pageId="35" pageNumber="169" pagination="221 - 247" refId="ref65974" refString="Komai T, Kawai K &amp; Ng PKL (2020) On the identity of the poorly known pea crab, Pinnothera obesa Dana, 1852, and description of a new species of Arcotheres Manning, 1993 from the southwest Pacific (Decapoda: Brachyura: Pinnotheridae). Zootaxa, 4822 (2): 221 - 247." type="journal article" year="2020">Komai et al., 2020</bibRefCitation>
: fig. 8D, E).
</paragraph>
<paragraph id="8BFE36CECE52FFF437E81AD1D29A7F7E" blockId="35.[136,775,2014,2102]" lastBlockId="37.[136,775,189,277]" lastPageId="37" lastPageNumber="171" pageId="35" pageNumber="169">
<collectingRegion id="4985F82CCE52FFF237E81AD1D0A2799D" box="[136,195,2014,2038]" country="Maldives" name="Maale" pageId="35" pageNumber="169">Male</collectingRegion>
: Carapace almost circular, slightly longer than wide; dorsal surface almost smooth, gently inflated, lateral surfaces with setae; front projecting anteriorly, margin gently convex to sinuous (
<bibRefCitation id="EFD04B3FCE52FFF234A41A51D4F9791D" author="Komai T &amp; Kawai K &amp; Ng PKL" box="[964,1176,1886,1910]" pageId="35" pageNumber="169" pagination="221 - 247" refId="ref65974" refString="Komai T, Kawai K &amp; Ng PKL (2020) On the identity of the poorly known pea crab, Pinnothera obesa Dana, 1852, and description of a new species of Arcotheres Manning, 1993 from the southwest Pacific (Decapoda: Brachyura: Pinnotheridae). Zootaxa, 4822 (2): 221 - 247." type="journal article" year="2020">Komai et al., 2020</bibRefCitation>
: figs. 12, 13A, 15A). Eyes distinctly visible in dorsal view (
<bibRefCitation id="EFD04B3FCE52FFF233D21A71D5E979FD" author="Komai T &amp; Kawai K &amp; Ng PKL" box="[1202,1416,1918,1942]" pageId="35" pageNumber="169" pagination="221 - 247" refId="ref65974" refString="Komai T, Kawai K &amp; Ng PKL (2020) On the identity of the poorly known pea crab, Pinnothera obesa Dana, 1852, and description of a new species of Arcotheres Manning, 1993 from the southwest Pacific (Decapoda: Brachyura: Pinnotheridae). Zootaxa, 4822 (2): 221 - 247." type="journal article" year="2020">Komai et al., 2020</bibRefCitation>
: figs. 12, 13A, 15A). MXP3 as in female except dactylus just reaching tip of propodus (
<bibRefCitation id="EFD04B3FCE52FFF2330C1AB1D52979BD" author="Komai T &amp; Kawai K &amp; Ng PKL" box="[1132,1352,1982,2006]" pageId="35" pageNumber="169" pagination="221 - 247" refId="ref65974" refString="Komai T, Kawai K &amp; Ng PKL (2020) On the identity of the poorly known pea crab, Pinnothera obesa Dana, 1852, and description of a new species of Arcotheres Manning, 1993 from the southwest Pacific (Decapoda: Brachyura: Pinnotheridae). Zootaxa, 4822 (2): 221 - 247." type="journal article" year="2020">Komai et al., 2020</bibRefCitation>
: figs. 13C, 16A). Anterior thoracic sternum wide, sternites 1, 2 fused, partially sunken into buccal cavity; suture between sternites 2 and 3 shallow; sternites 3, 4 completely fused, separated only by shallow grooves (
<bibRefCitation id="EFD04B3FCE55FFF5361C1A6AD2207916" author="Komai T &amp; Kawai K &amp; Ng PKL" box="[380,577,1893,1917]" pageId="36" pageNumber="170" pagination="221 - 247" refId="ref65974" refString="Komai T, Kawai K &amp; Ng PKL (2020) On the identity of the poorly known pea crab, Pinnothera obesa Dana, 1852, and description of a new species of Arcotheres Manning, 1993 from the southwest Pacific (Decapoda: Brachyura: Pinnotheridae). Zootaxa, 4822 (2): 221 - 247." type="journal article" year="2020">Komai et al., 2020</bibRefCitation>
: figs. 13D, 15B). Chela relatively stout, shorter than in female (
<bibRefCitation id="EFD04B3FCE55FFF535021A8AD0CD79D6" author="Komai T &amp; Kawai K &amp; Ng PKL" pageId="36" pageNumber="170" pagination="221 - 247" refId="ref65974" refString="Komai T, Kawai K &amp; Ng PKL (2020) On the identity of the poorly known pea crab, Pinnothera obesa Dana, 1852, and description of a new species of Arcotheres Manning, 1993 from the southwest Pacific (Decapoda: Brachyura: Pinnotheridae). Zootaxa, 4822 (2): 221 - 247." type="journal article" year="2020">Komai et al., 2020</bibRefCitation>
: figs. 14A, 16B). P2P5 dorsally, ventrally unarmed; outer surface covered with short setae; P3 and P4 carpus and propodus with long natatory setae; left and right meri equal, relative lengths of meri P4&gt;P3&gt;P2&gt;P5; dactyli of P2P4 progressively longer; left P4 dactylus slightly longer, more slender than right side; P4 and P5 dactylus subequal to that of P3, covered with short setae (
<bibRefCitation id="EFD04B3FCE55FFF533FB1A8AD51379F6" author="Komai T &amp; Kawai K &amp; Ng PKL" box="[1179,1394,1925,1949]" pageId="36" pageNumber="170" pagination="221 - 247" refId="ref65974" refString="Komai T, Kawai K &amp; Ng PKL (2020) On the identity of the poorly known pea crab, Pinnothera obesa Dana, 1852, and description of a new species of Arcotheres Manning, 1993 from the southwest Pacific (Decapoda: Brachyura: Pinnotheridae). Zootaxa, 4822 (2): 221 - 247." type="journal article" year="2020">Komai et al., 2020</bibRefCitation>
: figs. 14CG, 15A, 16DH). Pleon slender, triangular, widest at somite 3, lateral margins of somite 4 gently sinuous to almost straight; somite 6 subtrapezoidal; telson subhexagonal, much wider than long (
<bibRefCitation id="EFD04B3FCE55FFF53481150AD4CF7676" author="Komai T &amp; Kawai K &amp; Ng PKL" box="[993,1198,2053,2077]" pageId="36" pageNumber="170" pagination="221 - 247" refId="ref65974" refString="Komai T, Kawai K &amp; Ng PKL (2020) On the identity of the poorly known pea crab, Pinnothera obesa Dana, 1852, and description of a new species of Arcotheres Manning, 1993 from the southwest Pacific (Decapoda: Brachyura: Pinnotheridae). Zootaxa, 4822 (2): 221 - 247." type="journal article" year="2020">Komai et al., 2020</bibRefCitation>
: figs. 13D, H, 15F). G1 relatively stout, arcuate, curved outwards, tip subtruncate, without elongate process (
<bibRefCitation id="EFD04B3FCE54FFF436C91DB2D21C7EBE" author="Komai T &amp; Kawai K &amp; Ng PKL" box="[425,637,189,213]" pageId="37" pageNumber="171" pagination="221 - 247" refId="ref65974" refString="Komai T, Kawai K &amp; Ng PKL (2020) On the identity of the poorly known pea crab, Pinnothera obesa Dana, 1852, and description of a new species of Arcotheres Manning, 1993 from the southwest Pacific (Decapoda: Brachyura: Pinnotheridae). Zootaxa, 4822 (2): 221 - 247." type="journal article" year="2020">Komai et al., 2020</bibRefCitation>
: figs. 13I, J, 15G, H). G2 short, with spatuliform tip; exopod slightly longer than endopod (
<bibRefCitation id="EFD04B3FCE54FFF436171DF2D2297F7E" author="Komai T &amp; Kawai K &amp; Ng PKL" box="[375,584,253,277]" pageId="37" pageNumber="171" pagination="221 - 247" refId="ref65974" refString="Komai T, Kawai K &amp; Ng PKL (2020) On the identity of the poorly known pea crab, Pinnothera obesa Dana, 1852, and description of a new species of Arcotheres Manning, 1993 from the southwest Pacific (Decapoda: Brachyura: Pinnotheridae). Zootaxa, 4822 (2): 221 - 247." type="journal article" year="2020">Komai et al., 2020</bibRefCitation>
: figs. 13K, 15I).
</paragraph>
<caption id="DF3E6646CE55FFF537111BD1D2DC7945" ID-DOI="http://doi.org/10.5281/zenodo.7174283" ID-Zenodo-Dep="7174283" httpUri="https://zenodo.org/record/7174283/files/figure.png" pageId="36" pageNumber="170" startId="36.[113,148,1758,1780]" targetBox="[116,1455,191,1741]" targetPageId="36">
<paragraph id="8BFE36CECE55FFF537111BD1D2DC7945" blockId="36.[113,1451,1758,1838]" pageId="36" pageNumber="170">
Fig. 24.
<taxonomicName id="4C414D4DCE55FFF537A51BD1D2C8789F" authority="Komai, Kawai &amp; Ng, 2020" authorityName="Komai, Kawai &amp; Ng" authorityYear="2020" box="[197,681,1758,1780]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="170" phylum="Arthropoda" rank="species" species="ocularius">
<emphasis id="B935EADCCE55FFF537A51BD1D1F37898" box="[197,402,1758,1779]" italics="true" pageId="36" pageNumber="170">Arcotheres ocularius</emphasis>
<bibRefCitation id="EFD04B3FCE55FFF536F91BD1D2C8789F" author="Komai T &amp; Kawai K &amp; Ng PKL" box="[409,681,1758,1780]" pageId="36" pageNumber="170" pagination="221 - 247" refId="ref65974" refString="Komai T, Kawai K &amp; Ng PKL (2020) On the identity of the poorly known pea crab, Pinnothera obesa Dana, 1852, and description of a new species of Arcotheres Manning, 1993 from the southwest Pacific (Decapoda: Brachyura: Pinnotheridae). Zootaxa, 4822 (2): 221 - 247." type="journal article" year="2020">Komai, Kawai &amp; Ng, 2020</bibRefCitation>
</taxonomicName>
. A, CK, paratype female (10.9 × 9.7 mm) (ZRC 2019.1877), Fiji; B, female (9.4 × 8.4 mm) (ZRC 2019.1025), Indonesia. A, B, left MXP3; C, outer view of right chela; DG, left P2P5, respectively; HK, right P2P5, respectively. Scales = A, B, 0.5 mm; CK, 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BFE36CECE54FFF437E81C32D1107F9E" blockId="37.[135,776,317,501]" pageId="37" pageNumber="171">
<emphasis id="B935EADCCE54FFF437E81C32D09C7F3E" bold="true" box="[136,253,317,341]" pageId="37" pageNumber="171">Variation.</emphasis>
<bibRefCitation id="EFD04B3FCE54FFF436631C32D26A7F3E" author="Komai T &amp; Kawai K &amp; Ng PKL" box="[259,523,317,341]" pageId="37" pageNumber="171" pagination="221 - 247" refId="ref65974" refString="Komai T, Kawai K &amp; Ng PKL (2020) On the identity of the poorly known pea crab, Pinnothera obesa Dana, 1852, and description of a new species of Arcotheres Manning, 1993 from the southwest Pacific (Decapoda: Brachyura: Pinnotheridae). Zootaxa, 4822 (2): 221 - 247." type="journal article" year="2020">Komai et al. (2020: 241)</bibRefCitation>
noted that the carapace shape varied from broadly ovate (
<bibRefCitation id="EFD04B3FCE54FFF435611C52D2B67F1E" author="Komai T &amp; Kawai K &amp; Ng PKL" box="[513,727,349,373]" pageId="37" pageNumber="171" pagination="221 - 247" refId="ref65974" refString="Komai T, Kawai K &amp; Ng PKL (2020) On the identity of the poorly known pea crab, Pinnothera obesa Dana, 1852, and description of a new species of Arcotheres Manning, 1993 from the southwest Pacific (Decapoda: Brachyura: Pinnotheridae). Zootaxa, 4822 (2): 221 - 247." type="journal article" year="2020">Komai et al., 2020</bibRefCitation>
: fig. 7A) to almost circular (
<bibRefCitation id="EFD04B3FCE54FFF436E41C72D2337FFE" author="Komai T &amp; Kawai K &amp; Ng PKL" box="[388,594,381,405]" pageId="37" pageNumber="171" pagination="221 - 247" refId="ref65974" refString="Komai T, Kawai K &amp; Ng PKL (2020) On the identity of the poorly known pea crab, Pinnothera obesa Dana, 1852, and description of a new species of Arcotheres Manning, 1993 from the southwest Pacific (Decapoda: Brachyura: Pinnotheridae). Zootaxa, 4822 (2): 221 - 247." type="journal article" year="2020">Komai et al., 2020</bibRefCitation>
: fig. 7B, C). The condition of the dorsally exposed eyes and dactylus reaching end of or slightly overreaching MXP3 propodus, however, were always reliable.
</paragraph>
<paragraph id="8BFE36CECE54FFF437E81F12D0A17C1E" blockId="37.[136,776,541,629]" pageId="37" pageNumber="171">
<emphasis id="B935EADCCE54FFF437E81F12D0807C5E" bold="true" box="[136,225,541,565]" pageId="37" pageNumber="171">Colour.</emphasis>
“Entirely whitish; female ovary, visible through integument, orange; cornea of eye red” (
<bibRefCitation id="EFD04B3FCE54FFF435571F32D0D57C1E" author="Komai T &amp; Kawai K &amp; Ng PKL" pageId="37" pageNumber="171" pagination="221 - 247" refId="ref65974" refString="Komai T, Kawai K &amp; Ng PKL (2020) On the identity of the poorly known pea crab, Pinnothera obesa Dana, 1852, and description of a new species of Arcotheres Manning, 1993 from the southwest Pacific (Decapoda: Brachyura: Pinnotheridae). Zootaxa, 4822 (2): 221 - 247." type="journal article" year="2020">Komai et al., 2020: 241</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35B6545CE54FFF437E81F92D2827CBE" pageId="37" pageNumber="171" type="biology_ecology">
<paragraph id="8BFE36CECE54FFF437E81F92D2827CBE" blockId="37.[136,773,669,725]" pageId="37" pageNumber="171">
<emphasis id="B935EADCCE54FFF437E81F92D0A27CDE" bold="true" box="[136,195,669,693]" pageId="37" pageNumber="171">Host.</emphasis>
Known from
<taxonomicName id="4C414D4DCE54FFF436301F92D1B07CDE" box="[336,465,669,693]" class="Bivalvia" family="Arcidae" genus="Anadara" kingdom="Animalia" order="Arcida" pageId="37" pageNumber="171" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B935EADCCE54FFF436301F92D1CD7CDE" box="[336,428,669,693]" italics="true" pageId="37" pageNumber="171">Anadara</emphasis>
sp.
</taxonomicName>
in
<collectingCountry id="F356765ECE54FFF4368E1F92D2737CDE" box="[494,530,669,693]" name="Fiji" pageId="37" pageNumber="171">Fiji</collectingCountry>
and
<taxonomicName id="4C414D4DCE54FFF435211F92D1207CBE" authority="(Linnaeus, 1758)" authorityName="" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Bivalvia" family="Arcidae" genus="Anadara" kingdom="Animalia" order="Arcida" pageId="37" pageNumber="171" phylum="Mollusca" rank="species" species="antiquata">
<emphasis id="B935EADCCE54FFF435211F92D3647CDE" box="[577,773,669,693]" italics="true" pageId="37" pageNumber="171">Anadara antiquata</emphasis>
(Linnaeus, 1758)
</taxonomicName>
in Lombok (
<taxonomicName id="4C414D4DCE54FFF436B01FB2D2B77CBE" authority="Lamarck, 1809" authorityName="Lamarck" authorityYear="1809" box="[464,726,701,725]" class="Bivalvia" family="Arcidae" kingdom="Animalia" order="Arcida" pageId="37" pageNumber="171" phylum="Mollusca" rank="family">Arcidae Lamarck, 1809</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C35B6545CE54FFF437E81FF2D1F6795E" pageId="37" pageNumber="171" type="discussion">
<paragraph id="8BFE36CECE54FFF437E81FF2D1E47A9E" blockId="37.[134,777,765,1269]" pageId="37" pageNumber="171">
<emphasis id="B935EADCCE54FFF437E81FF2D0967D7E" bold="true" box="[136,247,765,789]" pageId="37" pageNumber="171">Remarks.</emphasis>
<bibRefCitation id="EFD04B3FCE54FFF4379C1FF2D1AB7D7E" author="Komai T &amp; Kawai K &amp; Ng PKL" box="[252,458,765,789]" pageId="37" pageNumber="171" pagination="221 - 247" refId="ref65974" refString="Komai T, Kawai K &amp; Ng PKL (2020) On the identity of the poorly known pea crab, Pinnothera obesa Dana, 1852, and description of a new species of Arcotheres Manning, 1993 from the southwest Pacific (Decapoda: Brachyura: Pinnotheridae). Zootaxa, 4822 (2): 221 - 247." type="journal article" year="2020">Komai et al. (2020)</bibRefCitation>
discussed the taxonomy of
<taxonomicName id="4C414D4DCE54FFF4358F1FF1D08B7D5E" authorityName="Komai, Kawai &amp; Ng" authorityYear="2020" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="171" phylum="Arthropoda" rank="species" species="ocularius">
<emphasis id="B935EADCCE54FFF4358F1FF1D08B7D5E" italics="true" pageId="37" pageNumber="171">A. ocularius</emphasis>
</taxonomicName>
at length, differentiating it from the morphologically similar
<taxonomicName id="4C414D4DCE54FFF437BC1E31D1037D3E" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[220,354,829,853]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="171" phylum="Arthropoda" rank="species" sensu="stricto" species="palaensis">
<emphasis id="B935EADCCE54FFF437BC1E31D1037D3E" box="[220,354,829,853]" italics="true" pageId="37" pageNumber="171">A. palaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20657A7CE54FFF4360C1E31D1C87D3E" box="[364,425,830,853]" pageId="37" pageNumber="171" sensu="stricto">s. str.</taxonomicNameLabel>
in the more strongly chitinised carapace and pereopods, eyes that are clearly visible in dorsal view in adult females, the prominently projecting front, the MXP3 dactylus being always long and reaching or overreaching the propodus apex, the male telson being subpentagonal or subhexagonal, and the G1 with a subtruncate tip (
<figureCitation id="137A2A4BCE54FFF437D31EF2D16E7A7E" box="[179,271,1021,1045]" captionStart="Fig" captionStartId="35.[136,171,1780,1802]" captionTargetBox="[136,1474,190,1765]" captionTargetId="figure-7@35.[136,1474,190,1765]" captionTargetPageId="35" captionText="Fig. 23. Arcotheres ocularius Komai, Kawai &amp; Ng, 2020, overall dorsal view. A, paratype ovigerous female (10.9 × 9.7 mm) (ZRC 2019.1877), Fiji; B, female (9.4 × 8.4 mm) (ZRC 2019.1025), Indonesia." figureDoi="http://doi.org/10.5281/zenodo.7174281" httpUri="https://zenodo.org/record/7174281/files/figure.png" pageId="37" pageNumber="171">Figs. 23</figureCitation>
,
<figureCitation id="137A2A4BCE54FFF4367C1EF2D10F7A7E" box="[284,366,1021,1045]" captionStart="Fig" captionStartId="36.[113,148,1758,1780]" captionTargetBox="[116,1455,191,1741]" captionTargetId="figure-15@36.[103,1468,187,1753]" captionTargetPageId="36" captionText="Fig. 24. Arcotheres ocularius Komai, Kawai &amp; Ng, 2020. A, CK, paratype female (10.9 × 9.7 mm) (ZRC 2019.1877), Fiji; B, female (9.4 × 8.4 mm) (ZRC 2019.1025), Indonesia. A, B, left MXP3; C, outer view of right chela; DG, left P2P5, respectively; HK, right P2P5, respectively. Scales = A, B, 0.5 mm; CK, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174283" httpUri="https://zenodo.org/record/7174283/files/figure.png" pageId="37" pageNumber="171">24A, B</figureCitation>
;
<bibRefCitation id="EFD04B3FCE54FFF4361C1EF2D22D7A7E" author="Komai T &amp; Kawai K &amp; Ng PKL" box="[380,588,1021,1045]" pageId="37" pageNumber="171" pagination="221 - 247" refId="ref65974" refString="Komai T, Kawai K &amp; Ng PKL (2020) On the identity of the poorly known pea crab, Pinnothera obesa Dana, 1852, and description of a new species of Arcotheres Manning, 1993 from the southwest Pacific (Decapoda: Brachyura: Pinnotheridae). Zootaxa, 4822 (2): 221 - 247." type="journal article" year="2020">Komai et al., 2020</bibRefCitation>
: figs. 7AD, 8C, F, G, 10B, E, 13D, HJ, 15FH, 16A). They also noted that unpublished genetic studies by Tsang Ling Ming from the Chinese University of
<collectingCountry id="F356765ECE54FFF436D21951D2557A1D" box="[434,564,1118,1142]" name="Hong Kong" pageId="37" pageNumber="171">Hong Kong</collectingCountry>
show both species to be distinct.
<taxonomicName id="4C414D4DCE54FFF436451972D2667AFE" authorityName="Komai, Kawai &amp; Ng" authorityYear="2020" box="[293,519,1149,1173]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="171" phylum="Arthropoda" rank="species" species="ocularius">
<emphasis id="B935EADCCE54FFF436451972D2667AFE" box="[293,519,1149,1173]" italics="true" pageId="37" pageNumber="171">Arcotheres ocularius</emphasis>
</taxonomicName>
is known from
<collectingCountry id="F356765ECE54FFF435D31972D2B97AFE" box="[691,728,1149,1173]" name="Fiji" pageId="37" pageNumber="171">Fiji</collectingCountry>
and Lombok,
<collectingCountry id="F356765ECE54FFF437931992D13C7ADE" box="[243,349,1181,1205]" name="Indonesia" pageId="37" pageNumber="171">Indonesia</collectingCountry>
, but not recorded from the main Sunda Shelf where
<taxonomicName id="4C414D4DCE54FFF4366B19B1D1ED7ABE" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[267,396,1213,1237]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="171" phylum="Arthropoda" rank="species" species="palaensis">
<emphasis id="B935EADCCE54FFF4366B19B1D1ED7ABE" box="[267,396,1213,1237]" italics="true" pageId="37" pageNumber="171">A. palaensis</emphasis>
</taxonomicName>
is common. Both species, however, are present in Lombok.
</paragraph>
<paragraph id="8BFE36CECE54FFF437E81812D1F6795E" blockId="37.[135,777,1309,1845]" pageId="37" pageNumber="171">
<bibRefCitation id="EFD04B3FCE54FFF437E81812D1C97B5E" author="Komai T &amp; Kawai K &amp; Ng PKL" box="[136,424,1309,1333]" pageId="37" pageNumber="171" pagination="221 - 247" refId="ref65974" refString="Komai T, Kawai K &amp; Ng PKL (2020) On the identity of the poorly known pea crab, Pinnothera obesa Dana, 1852, and description of a new species of Arcotheres Manning, 1993 from the southwest Pacific (Decapoda: Brachyura: Pinnotheridae). Zootaxa, 4822 (2): 221 - 247." type="journal article" year="2020">Komai et al. (2020: 244)</bibRefCitation>
observed that in the figure of the
<typeStatus id="54FA886CCE54FFF437D41832D1797B3E" box="[180,280,1341,1365]" pageId="37" pageNumber="171" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C414D4DCE54FFF436241831D1BA7B3E" authorityName="Burger" authorityYear="1895" box="[324,475,1341,1365]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="171" phylum="Arthropoda" rank="species" species="arcophilus">
<emphasis id="B935EADCCE54FFF436241831D1BA7B3E" box="[324,475,1341,1365]" italics="true" pageId="37" pageNumber="171">A. arcophilus</emphasis>
</taxonomicName>
in
<bibRefCitation id="EFD04B3FCE54FFF435651832D3627B3E" author="Ahyong ST &amp; Ng PKL" box="[517,771,1341,1366]" pageId="37" pageNumber="171" pagination="191 - 226" refId="ref62272" refString="Ahyong ST &amp; Ng PKL (2007 b) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16: 191 - 226." type="journal article" year="2007">Ahyong &amp; Ng (2007b</bibRefCitation>
: fig. 2A) (present
<figureCitation id="137A2A4BCE54FFF436271852D1C67B1E" box="[327,423,1373,1397]" captionStart="Fig" captionStartId="16.[113,148,1506,1528]" captionTargetBox="[113,1451,189,1491]" captionTargetId="figure-15@16.[113,1451,189,1491]" captionTargetPageId="16" captionText="Fig. 6. Arcotheres palaensis (Bürger, 1895). A, B, female (5.3 × 4.7 mm) (SMF-ZMG 172a) (lectotype of Pinnotheres arcophilus Bürger, 1895), Philippines; CF, ovigerous female (5.1 × 4.3 mm) (ZRC 1999.1202), Indonesia. A, C, overall dorsal view; B, D, dorsal view of carapace; E, frontal view of cephalothorax; F, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.7174246" httpUri="https://zenodo.org/record/7174246/files/figure.png" pageId="37" pageNumber="171">Figs. 6A</figureCitation>
,
<figureCitation id="137A2A4BCE54FFF436D61852D1867B1E" box="[438,487,1373,1397]" captionStart="Fig" captionStartId="29.[136,171,2006,2028]" captionTargetBox="[149,1465,190,1991]" captionTargetId="figure-8@29.[136,1482,181,1999]" captionTargetPageId="29" captionText="Fig. 19. Arcotheres palaensis (Bürger, 1895). AC, female (5.3 × 4.7 mm) (SMF-ZMG 172a) (lectotype of Pinnotheres arcophilus Bürger, 1895), Ubay, Philippines; DM ovigerous female (5.1 × 4.3 mm) (ZRC 1999.1202), Indonesia. A, overall dorsal view; B, right MXP3; C, right chela; D, left MXP3; EH, left P2P5, respectively; IL, right P2P5, respectively; M, frontal margin of carapace showing eyes in dorsal view. Scales = AL, 0.5 mm; M, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174273" httpUri="https://zenodo.org/record/7174273/files/figure.png" pageId="37" pageNumber="171">19A</figureCitation>
), the “female P4 merus is proportionately longer in
<taxonomicName id="4C414D4DCE54FFF436F31871D2727BFE" authorityName="Komai, Kawai &amp; Ng" authorityYear="2020" box="[403,531,1405,1429]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="171" phylum="Arthropoda" rank="species" species="ocularius">
<emphasis id="B935EADCCE54FFF436F31871D2727BFE" box="[403,531,1405,1429]" italics="true" pageId="37" pageNumber="171">A. ocularius</emphasis>
</taxonomicName>
” than in
<taxonomicName id="4C414D4DCE54FFF435111871D29E7BFE" authorityName="Burger" authorityYear="1895" box="[625,767,1405,1429]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="171" phylum="Arthropoda" rank="species" species="arcophilus">
<emphasis id="B935EADCCE54FFF435111871D29E7BFE" box="[625,767,1405,1429]" italics="true" pageId="37" pageNumber="171">A. arcophilus</emphasis>
</taxonomicName>
. We have re-examined the
<typeStatus id="54FA886CCE54FFF436D71892D27D7BDE" box="[439,540,1437,1461]" pageId="37" pageNumber="171" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C414D4DCE54FFF435271891D2817BDE" authorityName="Burger" authorityYear="1895" box="[583,736,1437,1461]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="171" phylum="Arthropoda" rank="species" species="arcophilus">
<emphasis id="B935EADCCE54FFF435271891D2817BDE" box="[583,736,1437,1461]" italics="true" pageId="37" pageNumber="171">A. arcophilus</emphasis>
</taxonomicName>
(at present synonymised under
<taxonomicName id="4C414D4DCE54FFF436D818B1D2587BBE" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[440,569,1469,1493]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="171" phylum="Arthropoda" rank="species" species="palaensis">
<emphasis id="B935EADCCE54FFF436D818B1D2587BBE" box="[440,569,1469,1493]" italics="true" pageId="37" pageNumber="171">A. palaensis</emphasis>
</taxonomicName>
) and the P4 merus is actually similar to
<taxonomicName id="4C414D4DCE54FFF4360018D1D1837B9E" authorityName="Komai, Kawai &amp; Ng" authorityYear="2020" box="[352,482,1501,1525]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="171" phylum="Arthropoda" rank="species" species="ocularius">
<emphasis id="B935EADCCE54FFF4360018D1D1837B9E" box="[352,482,1501,1525]" italics="true" pageId="37" pageNumber="171">A. ocularius</emphasis>
</taxonomicName>
and more like the condition in a more recent female of
<taxonomicName id="4C414D4DCE54FFF436FD18F1D27C787E" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[413,541,1533,1557]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="171" phylum="Arthropoda" rank="species" species="palaensis">
<emphasis id="B935EADCCE54FFF436FD18F1D27C787E" box="[413,541,1533,1557]" italics="true" pageId="37" pageNumber="171">A. palaensis</emphasis>
</taxonomicName>
from
<collectingCountry id="F356765ECE54FFF4353D18F2D2A5787E" box="[605,708,1533,1557]" name="Indonesia" pageId="37" pageNumber="171">Indonesia</collectingCountry>
(
<figureCitation id="137A2A4BCE54FFF435B118F2D0C9785E" captionStart="Fig" captionStartId="16.[113,148,1506,1528]" captionTargetBox="[113,1451,189,1491]" captionTargetId="figure-15@16.[113,1451,189,1491]" captionTargetPageId="16" captionText="Fig. 6. Arcotheres palaensis (Bürger, 1895). A, B, female (5.3 × 4.7 mm) (SMF-ZMG 172a) (lectotype of Pinnotheres arcophilus Bürger, 1895), Philippines; CF, ovigerous female (5.1 × 4.3 mm) (ZRC 1999.1202), Indonesia. A, C, overall dorsal view; B, D, dorsal view of carapace; E, frontal view of cephalothorax; F, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.7174246" httpUri="https://zenodo.org/record/7174246/files/figure.png" pageId="37" pageNumber="171">Figs. 6C</figureCitation>
,
<figureCitation id="137A2A4BCE54FFF437D21B12D09E785E" box="[178,255,1565,1589]" captionStart="Fig" captionStartId="29.[136,171,2006,2028]" captionTargetBox="[149,1465,190,1991]" captionTargetId="figure-8@29.[136,1482,181,1999]" captionTargetPageId="29" captionText="Fig. 19. Arcotheres palaensis (Bürger, 1895). AC, female (5.3 × 4.7 mm) (SMF-ZMG 172a) (lectotype of Pinnotheres arcophilus Bürger, 1895), Ubay, Philippines; DM ovigerous female (5.1 × 4.3 mm) (ZRC 1999.1202), Indonesia. A, overall dorsal view; B, right MXP3; C, right chela; D, left MXP3; EH, left P2P5, respectively; IL, right P2P5, respectively; M, frontal margin of carapace showing eyes in dorsal view. Scales = AL, 0.5 mm; M, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174273" httpUri="https://zenodo.org/record/7174273/files/figure.png" pageId="37" pageNumber="171">19EL</figureCitation>
). In the protruding front and dorsally visible eyes in adult females,
<taxonomicName id="4C414D4DCE54FFF4362C1B31D1B5783E" authorityName="Komai, Kawai &amp; Ng" authorityYear="2020" box="[332,468,1597,1621]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="171" phylum="Arthropoda" rank="species" species="ocularius">
<emphasis id="B935EADCCE54FFF4362C1B31D1B5783E" box="[332,468,1597,1621]" italics="true" pageId="37" pageNumber="171">A. ocularius</emphasis>
</taxonomicName>
resembles some specimens of
<taxonomicName id="4C414D4DCE54FFF437C71B51D14D781E" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[167,300,1629,1653]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="171" phylum="Arthropoda" rank="species" species="palaensis">
<emphasis id="B935EADCCE54FFF437C71B51D14D781E" box="[167,300,1629,1653]" italics="true" pageId="37" pageNumber="171">A. palaensis</emphasis>
</taxonomicName>
, but the carapace of
<taxonomicName id="4C414D4DCE54FFF435771B51D2FD781E" authorityName="Komai, Kawai &amp; Ng" authorityYear="2020" box="[535,668,1629,1653]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="171" phylum="Arthropoda" rank="species" species="ocularius">
<emphasis id="B935EADCCE54FFF435771B51D2FD781E" box="[535,668,1629,1653]" italics="true" pageId="37" pageNumber="171">A. ocularius</emphasis>
</taxonomicName>
is always more strongly chitinised, the MXP3 dactylus always long and overreaching the propodus apex, the longer female P4 merus relatively longer, the male telson subpentagonal or subhexagonal, and the G1 with a subtruncate tip (
<figureCitation id="137A2A4BCE54FFF435C71BD2D363789E" box="[679,770,1757,1781]" captionStart="Fig" captionStartId="35.[136,171,1780,1802]" captionTargetBox="[136,1474,190,1765]" captionTargetId="figure-7@35.[136,1474,190,1765]" captionTargetPageId="35" captionText="Fig. 23. Arcotheres ocularius Komai, Kawai &amp; Ng, 2020, overall dorsal view. A, paratype ovigerous female (10.9 × 9.7 mm) (ZRC 2019.1877), Fiji; B, female (9.4 × 8.4 mm) (ZRC 2019.1025), Indonesia." figureDoi="http://doi.org/10.5281/zenodo.7174281" httpUri="https://zenodo.org/record/7174281/files/figure.png" pageId="37" pageNumber="171">Figs. 23</figureCitation>
,
<figureCitation id="137A2A4BCE54FFF437EA1BF2D0BA797E" box="[138,219,1789,1813]" captionStart="Fig" captionStartId="36.[113,148,1758,1780]" captionTargetBox="[116,1455,191,1741]" captionTargetId="figure-15@36.[103,1468,187,1753]" captionTargetPageId="36" captionText="Fig. 24. Arcotheres ocularius Komai, Kawai &amp; Ng, 2020. A, CK, paratype female (10.9 × 9.7 mm) (ZRC 2019.1877), Fiji; B, female (9.4 × 8.4 mm) (ZRC 2019.1025), Indonesia. A, B, left MXP3; C, outer view of right chela; DG, left P2P5, respectively; HK, right P2P5, respectively. Scales = A, B, 0.5 mm; CK, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174283" httpUri="https://zenodo.org/record/7174283/files/figure.png" pageId="37" pageNumber="171">24A, B</figureCitation>
;
<bibRefCitation id="EFD04B3FCE54FFF437881BF2D1D6797E" author="Komai T &amp; Kawai K &amp; Ng PKL" box="[232,439,1789,1813]" pageId="37" pageNumber="171" pagination="221 - 247" refId="ref65974" refString="Komai T, Kawai K &amp; Ng PKL (2020) On the identity of the poorly known pea crab, Pinnothera obesa Dana, 1852, and description of a new species of Arcotheres Manning, 1993 from the southwest Pacific (Decapoda: Brachyura: Pinnotheridae). Zootaxa, 4822 (2): 221 - 247." type="journal article" year="2020">Komai et al., 2020</bibRefCitation>
: figs. 7AD, 8C, F, G, 10B, E, 13D, HJ, 15FH, 16A).
</paragraph>
</subSubSection>
<subSubSection id="C35B6545CE54FFF437E81A52D09979FE" pageId="37" pageNumber="171" type="distribution">
<paragraph id="8BFE36CECE54FFF437E81A52D09979FE" blockId="37.[136,774,1885,1941]" pageId="37" pageNumber="171">
<emphasis id="B935EADCCE54FFF437E81A52D17E791E" bold="true" box="[136,287,1885,1909]" pageId="37" pageNumber="171">Distribution.</emphasis>
Western Pacific, from
<collectingCountry id="F356765ECE54FFF4354C1A52D233791E" box="[556,594,1885,1909]" name="Fiji" pageId="37" pageNumber="171">Fiji</collectingCountry>
to southeastern
<collectingCountry id="F356765ECE54FFF437E81A72D09279FE" box="[136,243,1917,1941]" name="Indonesia" pageId="37" pageNumber="171">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>