treatments-xml/data/03/E8/87/03E887D2BA0AFE2BFDDBFF19611AF86B.xml

<document id="462D46A2A82E89FC033484F71C640FD9" ID-DOI="10.11646/zootaxa.5414.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10681263" ID-ZooBank="3B1DFADA-C20E-473B-A5E9-843548B54AD3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1708377054888" checkinUser="plazi" docAuthor="Paredes-Munguía, Williams, Brescovit, Antonio D. &amp; Teixeira, Renato A." docDate="2024" docId="03E887D2BA0AFE2BFDDBFF19611AF86B" docLanguage="en" docName="zootaxa.5414.1.1.pdf" docOrigin="Zootaxa 5414 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5414.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.18:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="18" docTitle="Arctosa ayaymama Paredes-Munguía &amp; Brescovit &amp; Teixeira 2024, sp. nov." docType="treatment" docVersion="1" lastPageNumber="8" masterDocId="FFD1FFAABA0DFE2CFD4CFF8E6559FFD2" masterDocTitle="Revision of Neotropical wolf spider genus Arctosa C. L. Koch, 1847 (Araneae: Lycosidae), with description of seven new species" masterLastPageNumber="83" masterPageNumber="1" pageNumber="8" updateTime="1708377875290" updateUser="ExternalLinkService">
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<mods:title id="DD0F5463D390EFC6E37EC1734D9C018C">Revision of Neotropical wolf spider genus Arctosa C. L. Koch, 1847 (Araneae: Lycosidae), with description of seven new species</mods:title>
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<mods:namePart id="CD15D04021B8B454AA44BFAABE225518">Paredes-Munguía, Williams</mods:namePart>
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<mods:affiliation id="01AEA4317FDA9BE63F4890295B7EB9FD">Pontifícia Universidade Católica do Rio Grande do Sul, Escola de Ciências da Saúde e da Vida, Museu de Ciências e Tecnologia, Laboratório de Aracnologia, Porto Alegre, Brazil. &amp; Museo de Historia Natural de la Universidad Nacional Mayor de San Marcos, Laboratorio de Entomología, Av. Arenales 1256, Lima 14, Peru. &amp; williams. paredes @ acad. pucrs. br; https: // orcid. org / 0000 - 0002 - 3320 - 4611</mods:affiliation>
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<mods:namePart id="A02C809FECF4A0B5F408D66059AFAFC6">Brescovit, Antonio D.</mods:namePart>
<mods:nameIdentifier id="E4A01AB5EBC2A8F09CC19A75C6DC15EF" type="ORCID">0000-0002-1511-5324</mods:nameIdentifier>
<mods:affiliation id="DE5DBBA06BD26FAE39C3C93D79113196">Laboratório de Coleções Zoológicas, Av. Vital Brasil, 1500, Butantã, 05503 - 900, São Paulo, SP, Brazil. &amp; antonio. brescovit @ butantan. gov. br; https: // orcid. org / 0000 - 0002 - 1511 - 5324</mods:affiliation>
<mods:nameIdentifier id="A97194509DD713E9B729E005A054150E" type="email">antonio.brescovit@butantan.gov.br</mods:nameIdentifier>
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<mods:namePart id="A4557702C122F6B236268C032CE2ED05">Teixeira, Renato A.</mods:namePart>
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<mods:affiliation id="17F8381347133B369C71436D75805D24">Pontifícia Universidade Católica do Rio Grande do Sul, Escola de Ciências da Saúde e da Vida, Museu de Ciências e Tecnologia, Laboratório de Aracnologia, Porto Alegre, Brazil. &amp; Pontifícia Universidade Católica do Rio Grande do Sul, Escola de Ciências da Saúde e da Vida, Museu de Ciências e Tecnologia, Laboratório de Aracnologia, Porto Alegre, Brazil. &amp; renatoaug. tx @ gmail. com; https: // orcid. org / 0000 - 0002 - 1756 - 9821</mods:affiliation>
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<taxonomicName id="4C414D47BA0AFE2BFDDBFF196424FF63" authority="Paredes-Munguía &amp; Brescovit &amp; Teixeira, 2024" authorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" authorityYear="2024" box="[151,381,151,177]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ayaymama" status="sp. nov.">
<emphasis id="B935EAD6BA0AFE2BFDDBFF196424FF63" bold="true" box="[151,381,151,177]" italics="true" pageId="7" pageNumber="8">Arctosa ayaymama</emphasis>
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<subSubSection id="C35B654FBA0AFE2BFDDBFF356415FF07" box="[151,332,187,214]" pageId="7" pageNumber="8" type="description">
<paragraph id="8BFE36C4BA0AFE2BFDDBFF356415FF07" blockId="7.[151,483,151,214]" box="[151,332,187,214]" pageId="7" pageNumber="8">
<figureCitation id="137A2A41BA0AFE2BFDDBFF3565B4FF07" box="[151,237,187,213]" captionStart="FIGURE 2" captionStartId="9.[151,250,1891,1915]" captionTargetBox="[185,1398,191,1857]" captionTargetId="figure-21@9.[180,1408,181,1867]" captionTargetPageId="9" captionText="FIGURE 2. Epigynum. a, c, f. Arctosa villa sp. nov (a, ventral; c, dorsal; f, posterolateral). b. Arctosa ayaymama sp. nov. d. Arctosa villa sp. nov. (anterior). e. Arctosa mineira sp. nov. (posterior). g. Arctosa jibarosa sp. nov. (posterolateral). Scale bars: a–g, 0.25 mm. AT, atrium, BS, base of spermatheca, CO, copulatory opening, FD, fertilization duct, H, hood, HS, head of spermatheca, MS, median septum, SS, stalk of spermatheca, TS, transversal septum, VC, vulval chamber." figureDoi="http://doi.org/10.5281/zenodo.10681267" httpUri="https://zenodo.org/record/10681267/files/figure.png" pageId="7" pageNumber="8">Figs 2b</figureCitation>
, 
<figureCitation id="137A2A41BA0AFE2BFDB4FF35647CFF04" box="[248,293,187,214]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStartId-0="10.[151,250,1933,1957]" captionStartId-1="11.[151,250,1614,1638]" captionStartId-2="12.[151,250,1317,1341]" captionTargetBox-0="[159,1428,1001,1903]" captionTargetBox-1="[151,1436,181,1591]" captionTargetBox-2="[151,1436,181,1293]" captionTargetId-0="figure-313@10.[151,1436,993,1910]" captionTargetId-1="figure-21@11.[151,1436,181,1591]" captionTargetId-2="figure-25@12.[151,1436,181,1293]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="FIGURE 3. Arctosa ayaymama sp. nov. Epigynum, holotype a–b (MUSM-ENT 500411) (a, ventral; b, dorsal); paratype c–d (MUSM-ENT 505143) (c, ventral; d, posterolateral) Scale bars: a–d, 0.25 mm. CO copulatory opening, FD fertilization duct, H hood, HS head of spermatheca, SS stalk of spermatheca, TS transversal septum." captionText-1="FIGURE 4. Arctosa ayaymama sp. nov. Female habitus (MUSM-ENT 500411) a, c, e–f (a, dorsal; c, ventral; e, lateral, f, frontal) Epigynum b, d (b, dorsal; d, ventral) Scale bars: a, c, e, 1 mm, b, d 100 µm, f, 500 µm." captionText-2="FIGURE 5. Arctosa ayaymama sp. nov. Epigynum (MUSM-ENT 506993) a–c (a, ventral; b, dorsal; c, dictynoid pore) d, Male leg I, prolateral. Scale bars: a, 150 µm, b, 400 µm, c, 15 µm, d, 200 µm. CO copulatory opening, FD fertilization duct, H hood, HS head of spermatheca, DP dictynoid pore, MS median septum." figureDoi-0="http://doi.org/10.5281/zenodo.10681269" figureDoi-1="http://doi.org/10.5281/zenodo.10681271" figureDoi-2="http://doi.org/10.5281/zenodo.10681273" httpUri-0="https://zenodo.org/record/10681269/files/figure.png" httpUri-1="https://zenodo.org/record/10681271/files/figure.png" httpUri-2="https://zenodo.org/record/10681273/files/figure.png" pageId="7" pageNumber="8">3‒5</figureCitation>
, 
<figureCitation id="137A2A41BA0AFE2BFC7CFF356415FF07" box="[304,332,187,213]" captionStart="FIGURE 62" captionStartId="72.[151,250,1618,1642]" captionTargetBox="[171,1418,564,1590]" captionTargetId="figure-197@72.[164,1423,561,1595]" captionTargetPageId="72" captionText="FIGURE 62. Geographic distribution records of Arctosa ayaymama sp. nov. (blue dot), Arctosa pacaya sp. nov. (red dot), Hogna andina comb. nov. (orange dot)." figureDoi="http://doi.org/10.5281/zenodo.10681391" httpUri="https://zenodo.org/record/10681391/files/figure.png" pageId="7" pageNumber="8">62</figureCitation>
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<emphasis id="B935EAD6BA0AFE2BFDDBFE8D6413FECF" bold="true" box="[151,330,259,285]" pageId="7" pageNumber="8">Type Material.</emphasis>
<materialsCitation id="3B293C99BA0AFE2BFC1FFE8D612FFEB4" collectingDate="2008-09-07" collectingDateMax="2008-09-11" collectingDateMin="2008-09-07" collectorName="W. Paredes &amp; C. Albujar" country="Peru" county="Moyobamba" elevation="805" latitude="-5.9017835" location="Female" longLatPrecision="1" longitude="-77.18859" municipality="Yuracyacu" pageId="7" pageNumber="8" specimenCode="MUSM-ENT 500411, MUSM-ENT 505143" specimenCount="2" stateProvince="San Martin" typeStatus="holotype">
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<typeStatus id="54FA8866BA0AFE2BFCE2FE8D6749FECF" box="[430,528,259,285]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
from 
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, 
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<geoCoordinate id="EE755003BA0AFE2BFE36FE8D6171FECC" box="[890,1064,259,286]" degrees="77" direction="west" minutes="08" orientation="longitude" pageId="7" pageNumber="8" precision="1" seconds="53.44" value="-77.14818">77°08'53.44&quot;W</geoCoordinate>
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, 
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<collectingDate id="EFBBE9ECBA0AFE2BF9C2FE8D61A3FECF" box="[1166,1274,259,285]" pageId="7" pageNumber="8" value="2008-09-07">7.ix.2008</collectingDate>
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, 
<collectingCounty id="629F4E48BA0AFE2BF844FE8D60C1FECF" box="[1288,1432,259,285]" pageId="7" pageNumber="8">Moyobamba</collectingCounty>
, 
<collectingRegion id="4985F826BA0AFE2BFDDBFEA96440FE93" box="[151,281,295,321]" country="Peru" name="San Martin" pageId="7" pageNumber="8">San Martín</collectingRegion>
, 
<collectingCountry id="F3567654BA0AFE2BFC6AFEA66405FE90" box="[294,348,296,322]" name="Peru" pageId="7" pageNumber="8">Peru</collectingCountry>
, 
<collectorName id="26B45312BA0AFE2BFC24FEA664BEFE93" box="[360,487,295,322]" pageId="7" pageNumber="8">W. Paredes</collectorName>
&amp; 
<collectorName id="26B45312BA0AFE2BFF42FEA967DCFE93" box="[526,645,295,321]" pageId="7" pageNumber="8">C. Albujar</collectorName>
<emphasis id="B935EAD6BA0AFE2BFFC1FEA967EFFE93" box="[653,694,295,321]" italics="true" pageId="7" pageNumber="8">leg.</emphasis>
(
<specimenCode id="DBE79EBFBA0AFE2BFF8AFEA966E7FE90" box="[710,958,295,322]" collectionCode="MUSM-ENT" pageId="7" pageNumber="8">MUSM-ENT 500411</specimenCode>
). Female 
<typeStatus id="54FA8866BA0AFE2BF961FEA661C9FE90" box="[1069,1168,296,322]" pageId="7" pageNumber="8" type="paratype">paratype</typeStatus>
, near the same locality, 
<geoCoordinate id="EE755003BA0AFE2BFDDBFEC56472FEB4" box="[151,299,331,358]" degrees="5" direction="south" minutes="54" orientation="latitude" pageId="7" pageNumber="8" precision="1" seconds="06.42" value="-5.9017835">5°54'06.42&quot;S</geoCoordinate>
<geoCoordinate id="EE755003BA0AFE2BFC7EFEC56486FEB7" box="[306,479,331,357]" degrees="77" direction="west" minutes="11" orientation="longitude" pageId="7" pageNumber="8" precision="1" seconds="18.92" value="-77.18859">77°11'18.92&quot;W</geoCoordinate>
, 
<quantity id="4CB99B21BA0AFE2BFCA6FEC5676AFEB4" box="[490,563,331,358]" metricMagnitude="2" metricUnit="m" metricValue="8.16" pageId="7" pageNumber="8" unit="m" value="816.0">
<elevation id="006CD1F7BA0AFE2BFCA6FEC5676AFEB4" box="[490,563,331,358]" metricMagnitude="2" metricUnit="m" metricValue="8.16" pageId="7" pageNumber="8" unit="m" value="816.0">816 m</elevation>
</quantity>
, 
<date id="FFFF1004BA0AFE2BFF73FEC567E0FEB7" box="[575,697,331,357]" pageId="7" pageNumber="8" value="2008-09-11">
<collectingDate id="EFBBE9ECBA0AFE2BFF73FEC567E0FEB7" box="[575,697,331,357]" pageId="7" pageNumber="8" value="2008-09-11">11.ix.2008</collectingDate>
</date>
. 
<collectorName id="26B45312BA0AFE2BFF8FFEC56660FEB7" box="[707,825,331,357]" pageId="7" pageNumber="8">C. Albujar</collectorName>
<emphasis id="B935EAD6BA0AFE2BFE73FEC56631FEB7" box="[831,872,331,357]" italics="true" pageId="7" pageNumber="8">leg.</emphasis>
(
<specimenCode id="DBE79EBFBA0AFE2BFE3BFEC56134FEB4" box="[887,1133,331,358]" collectionCode="MUSM-ENT" pageId="7" pageNumber="8">MUSM-ENT 505143</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BFE36C4BA0AFE2BFD8BFEE1678BFE24" blockId="7.[151,1437,259,790]" pageId="7" pageNumber="8">
<emphasis id="B935EAD6BA0AFE2BFD8BFEE164A4FE5B" bold="true" box="[199,509,367,393]" pageId="7" pageNumber="8">Other material examined.</emphasis>
<materialsCitation id="3B293C99BA0AFE2BFF49FEE16682FE7F" collectionCode="LEUA" collectorName="Vereda El Conquistador &amp; F. Arcos-Valencia" country="Colombia" county="Curillo" elevation="226" latitude="1.3377779" location="Laguna la Cocha" longLatPrecision="15" longitude="-75.91381" municipality="Vereda El Conquistador" pageId="7" pageNumber="8" specimenCode="LEUA 88163" specimenCount="1" specimenCount-female="1" stateProvince="Caqueta">
<emphasis id="B935EAD6BA0AFE2BFF49FEE1667AFE5B" bold="true" box="[517,803,366,393]" pageId="7" pageNumber="8">
<collectingCountry id="F3567654BA0AFE2BFF49FEE167F1FE5B" box="[517,680,367,393]" name="Colombia" pageId="7" pageNumber="8">COLOMBIA</collectingCountry>
: 
<collectingRegion id="4985F826BA0AFE2BFFF9FEE06647FE5B" box="[693,798,366,393]" country="Colombia" name="Caqueta" pageId="7" pageNumber="8">Caquetá</collectingRegion>
:
</emphasis>
<specimenCount id="9D47FD4DBA0AFE2BFE67FEE16617FE5B" box="[811,846,367,393]" count="1" pageId="7" pageNumber="8" type="female">1♀</specimenCount>
, 
<collectingCounty id="629F4E48BA0AFE2BFE11FEE166F6FE5B" box="[861,943,367,393]" pageId="7" pageNumber="8">Curillo</collectingCounty>
, 
<collectorName id="26B45312BA0AFE2BFEF7FEE16193FE5B" box="[955,1226,367,393]" pageId="7" pageNumber="8">
<collectingMunicipality id="6B9AACBEBA0AFE2BFEF7FEE16193FE5B" box="[955,1226,367,393]" pageId="7" pageNumber="8">Vereda El Conquistador</collectingMunicipality>
</collectorName>
, 
<location id="8E9E601FBA0AFE2BF99AFEFE60C1FE5B" LSID="urn:lsid:plazi:treatment:03E887D2BA0AFE2BFDDBFF19611AF86B:8E9E601FBA0AFE2BF99AFEFE60C1FE5B" box="[1238,1432,367,394]" country="Colombia" county="Curillo" latitude="1.3377779" longLatPrecision="15" longitude="-75.91381" municipality="Vereda El Conquistador" name="Laguna la Cocha" pageId="7" pageNumber="8" stateProvince="Caqueta">Laguna la Cocha</location>
, ca. 
<geoCoordinate id="EE755003BA0AFE2BFDF1FE1D646AFE7F" box="[189,307,403,429]" degrees="1" direction="north" minutes="20" orientation="latitude" pageId="7" pageNumber="8" precision="15" seconds="16" value="1.3377779">1°20'16&quot;N</geoCoordinate>
<geoCoordinate id="EE755003BA0AFE2BFC75FE1D6481FE7C" box="[313,472,403,430]" degrees="75" direction="west" minutes="54" orientation="longitude" pageId="7" pageNumber="8" precision="1" seconds="49.7" value="-75.91381">75°54'49.7&quot;W</geoCoordinate>
, 
<quantity id="4CB99B21BA0AFE2BFCAEFE1D6773FE7C" box="[482,554,403,430]" metricMagnitude="2" metricUnit="m" metricValue="2.26" pageId="7" pageNumber="8" unit="m" value="226.0">
<elevation id="006CD1F7BA0AFE2BFCAEFE1D6773FE7C" box="[482,554,403,430]" metricMagnitude="2" metricUnit="m" metricValue="2.26" pageId="7" pageNumber="8" unit="m" value="226.0">226 m</elevation>
</quantity>
, 
<collectorName id="26B45312BA0AFE2BFF79FE1A67A2FE7F" box="[565,763,403,430]" pageId="7" pageNumber="8">F. Arcos-Valencia</collectorName>
<emphasis id="B935EAD6BA0AFE2BFE4CFE1D6673FE7F" box="[768,810,403,429]" italics="true" pageId="7" pageNumber="8">leg.</emphasis>
(
<specimenCode id="DBE79EBFBA0AFE2BFE7BFE1D668AFE7F" box="[823,979,403,429]" collectionCode="LEUA" pageId="7" pageNumber="8">LEUA 88163</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="3B293C99BA0AFE2BFEA8FE1D607AFE00" collectingDate="2008-09-11" collectingDateMax="2008-09-12" collectingDateMin="2008-09-11" collectorName="C. Albujar" country="Peru" elevation="816" latitude="-5.9017835" location="Yuracyacu" longLatPrecision="1" longitude="-77.18859" municipality="Moyobamba" pageId="7" pageNumber="8" specimenCode="MUSM-ENT 500358" specimenCount="1" specimenCount-female="1" stateProvince="San Martin">
<emphasis id="B935EAD6BA0AFE2BFEA8FE1D618BFE7F" bold="true" box="[996,1234,402,429]" pageId="7" pageNumber="8">
<collectingCountry id="F3567654BA0AFE2BFEA8FE1D616CFE7F" box="[996,1077,403,429]" name="Peru" pageId="7" pageNumber="8">PERU</collectingCountry>
: 
<collectingRegion id="4985F826BA0AFE2BF90CFE1D6194FE7F" box="[1088,1229,402,429]" country="Peru" name="San Martin" pageId="7" pageNumber="8">San Martín</collectingRegion>
:
</emphasis>
<specimenCount id="9D47FD4DBA0AFE2BF994FE1D61A2FE7F" box="[1240,1275,403,429]" count="1" pageId="7" pageNumber="8" type="female">1♀</specimenCount>
, 
<collectingMunicipality id="6B9AACBEBA0AFE2BF844FE1D60C1FE7F" box="[1288,1432,403,429]" pageId="7" pageNumber="8">Moyobamba</collectingMunicipality>
, 
<location id="8E9E601FBA0AFE2BFDDBFE366449FE00" LSID="urn:lsid:plazi:treatment:03E887D2BA0AFE2BFDDBFF19611AF86B:8E9E601FBA0AFE2BFDDBFE366449FE00" box="[151,272,440,466]" country="Peru" latitude="-5.9017835" longLatPrecision="1" longitude="-77.18859" municipality="Moyobamba" name="Yuracyacu" pageId="7" pageNumber="8" stateProvince="San Martin">Yuracyacu</location>
, 
<geoCoordinate id="EE755003BA0AFE2BFC51FE3964FAFE00" box="[285,419,439,466]" degrees="5" direction="south" minutes="54" orientation="latitude" pageId="7" pageNumber="8" precision="1" seconds="6.42" value="-5.9017835">5°54'6.42&quot;S</geoCoordinate>
<geoCoordinate id="EE755003BA0AFE2BFCE7FE396701FE03" box="[427,600,439,465]" degrees="77" direction="west" minutes="11" orientation="longitude" pageId="7" pageNumber="8" precision="1" seconds="18.92" value="-77.18859">77°11'18.92&quot;W</geoCoordinate>
, 
<quantity id="4CB99B21BA0AFE2BFF28FE3967F7FE00" box="[612,686,439,466]" metricMagnitude="2" metricUnit="m" metricValue="8.16" pageId="7" pageNumber="8" unit="m" value="816.0">
<elevation id="006CD1F7BA0AFE2BFF28FE3967F7FE00" box="[612,686,439,466]" metricMagnitude="2" metricUnit="m" metricValue="8.16" pageId="7" pageNumber="8" unit="m" value="816.0">816 m</elevation>
</quantity>
, 
<date id="FFFF1004BA0AFE2BFFF0FE396607FE03" box="[700,862,439,465]" pageId="7" pageNumber="8" value="2008-09-11" valueMax="2008-09-12" valueMin="2008-09-11">
<collectingDate id="EFBBE9ECBA0AFE2BFFF0FE396607FE03" box="[700,862,439,465]" pageId="7" pageNumber="8" value="2008-09-11" valueMax="2008-09-12" valueMin="2008-09-11">11–12.ix.2008</collectingDate>
</date>
, 
<collectorName id="26B45312BA0AFE2BFE20FE3966BBFE03" box="[876,994,439,465]" pageId="7" pageNumber="8">C. Albujar</collectorName>
<emphasis id="B935EAD6BA0AFE2BFEA6FE39614AFE03" box="[1002,1043,439,465]" italics="true" pageId="7" pageNumber="8">leg.</emphasis>
(
<specimenCode id="DBE79EBFBA0AFE2BF96FFE396042FE00" box="[1059,1307,439,466]" collectionCode="MUSM-ENT" pageId="7" pageNumber="8">MUSM-ENT 500358</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="3B293C99BA0AFE2BF87CFE396797FE24" collectingDate="2008-09-11" collectingDateMax="2008-09-12" collectingDateMin="2008-09-11" collectorName="C. Albujar" country="Peru" elevation="816" latitude="-5.9017835" location="Yuracyacu" longLatPrecision="1" longitude="-77.18859" municipality="Moyobamba" pageId="7" pageNumber="8" specimenCode="MUSM-ENT 506993" specimenCount="4" specimenCount-female="4" stateProvince="San Martin">
<specimenCount id="9D47FD4DBA0AFE2BF87CFE39600DFE03" box="[1328,1364,439,465]" count="4" pageId="7" pageNumber="8" type="female">4♀</specimenCount>
, ditto, 
<date id="FFFF1004BA0AFE2BFDDBFE556449FE27" box="[151,272,475,501]" pageId="7" pageNumber="8" value="2008-09-11">
<collectingDate id="EFBBE9ECBA0AFE2BFDDBFE556449FE27" box="[151,272,475,501]" pageId="7" pageNumber="8" value="2008-09-11">11.ix.2008</collectingDate>
</date>
, 
<collectorName id="26B45312BA0AFE2BFC50FE5564C8FE27" box="[284,401,475,501]" pageId="7" pageNumber="8">C. Albujar</collectorName>
<emphasis id="B935EAD6BA0AFE2BFCD4FE556498FE27" box="[408,449,475,501]" italics="true" pageId="7" pageNumber="8">leg.</emphasis>
(
<specimenCode id="DBE79EBFBA0AFE2BFC83FE55679FFE24" box="[463,710,475,502]" collectionCode="MUSM-ENT" pageId="7" pageNumber="8">MUSM-ENT 506993</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35B654FBA0AFE2BFD8BFE71619EFDB3" pageId="7" pageNumber="8" type="etymology">
<paragraph id="8BFE36C4BA0AFE2BFD8BFE71619EFDB3" blockId="7.[151,1437,259,790]" pageId="7" pageNumber="8">
<emphasis id="B935EAD6BA0AFE2BFD8BFE716417FDCB" bold="true" box="[199,334,511,537]" pageId="7" pageNumber="8">Etymology.</emphasis>
The species name comes from a Peruvian Amazon folk tale. In this story, one stepmother in the middle of the jungle abandoned two stepsons, and their lament is heard on moonlit nights looking for his dead biological mother. The sound of these children's wailing is folklorically known as 
<taxonomicName id="4C414D47BA0AFE2BF963FDC961E2FDB3" authorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" authorityYear="2024" box="[1071,1211,583,609]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ayaymama">&quot;ayaymama</taxonomicName>
&quot;.
</paragraph>
</subSubSection>
<subSubSection id="C35B654FBA0AFE2BFD8BFDE564FBFCC4" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="8BFE36C4BA0AFE2BFD8BFDE564FBFCC4" blockId="7.[151,1437,259,790]" pageId="7" pageNumber="8">
<emphasis id="B935EAD6BA0AFE2BFD8BFDE5641BFD57" bold="true" box="[199,322,619,645]" pageId="7" pageNumber="8">Diagnosis.</emphasis>
Females of 
<taxonomicName id="4C414D47BA0AFE2BFC95FDE5672FFD57" authorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" authorityYear="2024" box="[473,630,619,645]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ayaymama">
<emphasis id="B935EAD6BA0AFE2BFC95FDE5672FFD57" box="[473,630,619,645]" italics="true" pageId="7" pageNumber="8">A. ayaymama</emphasis>
</taxonomicName>
can be recognized among all Neotropical species of 
<taxonomicName id="4C414D47BA0AFE2BF9BCFDE5601FFD57" authorityName="C. L. Koch" authorityYear="1847" box="[1264,1350,619,645]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B935EAD6BA0AFE2BF9BCFDE5601FFD57" box="[1264,1350,619,645]" italics="true" pageId="7" pageNumber="8">Arctosa</emphasis>
</taxonomicName>
by the clearly triangular median septum (
<figureCitation id="137A2A41BA0AFE2BFF55FD016736FD7B" box="[537,623,655,681]" captionStart="FIGURE 3" captionStartId="10.[151,250,1933,1957]" captionTargetBox="[159,1428,1001,1903]" captionTargetId="figure-313@10.[151,1436,993,1910]" captionTargetPageId="10" captionText="FIGURE 3. Arctosa ayaymama sp. nov. Epigynum, holotype a–b (MUSM-ENT 500411) (a, ventral; b, dorsal); paratype c–d (MUSM-ENT 505143) (c, ventral; d, posterolateral) Scale bars: a–d, 0.25 mm. CO copulatory opening, FD fertilization duct, H hood, HS head of spermatheca, SS stalk of spermatheca, TS transversal septum." figureDoi="http://doi.org/10.5281/zenodo.10681269" httpUri="https://zenodo.org/record/10681269/files/figure.png" pageId="7" pageNumber="8">Figs 3a</figureCitation>
, 
<figureCitation id="137A2A41BA0AFE2BFF36FD0167C3FD7B" box="[634,666,655,681]" captionStart="FIGURE 4" captionStartId="11.[151,250,1614,1638]" captionTargetBox="[151,1436,181,1591]" captionTargetId="figure-21@11.[151,1436,181,1591]" captionTargetPageId="11" captionText="FIGURE 4. Arctosa ayaymama sp. nov. Female habitus (MUSM-ENT 500411) a, c, e–f (a, dorsal; c, ventral; e, lateral, f, frontal) Epigynum b, d (b, dorsal; d, ventral) Scale bars: a, c, e, 1 mm, b, d 100 µm, f, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.10681271" httpUri="https://zenodo.org/record/10681271/files/figure.png" pageId="7" pageNumber="8">4d</figureCitation>
), like in 
<taxonomicName id="4C414D47BA0AFE2BFFB1FD01669EFD7B" authorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" authorityYear="2024" baseAuthorityName="Keyserling" baseAuthorityYear="1877" box="[765,967,655,681]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tenebrosa" status="comb. nov.">
<emphasis id="B935EAD6BA0AFE2BFFB1FD01669EFD7B" box="[765,967,655,681]" italics="true" pageId="7" pageNumber="8">Arctosa tenebrosa</emphasis>
</taxonomicName>
<emphasis id="B935EAD6BA0AFE2BFE83FD016116FD7B" bold="true" box="[975,1103,655,681]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="A20657ADBA0AFE2BFE83FD016116FD7B" box="[975,1103,655,681]" pageId="7" pageNumber="8" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, but with a deeper hood in 
<taxonomicName id="4C414D47BA0AFE2BF8C8FD016449FD1F" authorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" authorityYear="2024" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ayaymama">
<emphasis id="B935EAD6BA0AFE2BF8C8FD016449FD1F" italics="true" pageId="7" pageNumber="8">A. ayaymama</emphasis>
</taxonomicName>
(
<figureCitation id="137A2A41BA0AFE2BFC6DFD3D642CFD1F" box="[289,373,691,717]" captionStart="FIGURE 3" captionStartId="10.[151,250,1933,1957]" captionTargetBox="[159,1428,1001,1903]" captionTargetId="figure-313@10.[151,1436,993,1910]" captionTargetPageId="10" captionText="FIGURE 3. Arctosa ayaymama sp. nov. Epigynum, holotype a–b (MUSM-ENT 500411) (a, ventral; b, dorsal); paratype c–d (MUSM-ENT 505143) (c, ventral; d, posterolateral) Scale bars: a–d, 0.25 mm. CO copulatory opening, FD fertilization duct, H hood, HS head of spermatheca, SS stalk of spermatheca, TS transversal septum." figureDoi="http://doi.org/10.5281/zenodo.10681269" httpUri="https://zenodo.org/record/10681269/files/figure.png" pageId="7" pageNumber="8">Fig. 3b</figureCitation>
). Likewise, 
<taxonomicName id="4C414D47BA0AFE2BFCB3FD3D67C1FD1F" authorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" authorityYear="2024" box="[511,664,691,717]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ayaymama">
<emphasis id="B935EAD6BA0AFE2BFCB3FD3D67C1FD1F" box="[511,664,691,717]" italics="true" pageId="7" pageNumber="8">A. ayaymama</emphasis>
</taxonomicName>
can be distinguished by the longitudinal median band (LMB) of the carapace covered by white setae (
<figureCitation id="137A2A41BA0AFE2BFF47FD596703FD23" box="[523,602,727,753]" captionStart="FIGURE 4" captionStartId="11.[151,250,1614,1638]" captionTargetBox="[151,1436,181,1591]" captionTargetId="figure-21@11.[151,1436,181,1591]" captionTargetPageId="11" captionText="FIGURE 4. Arctosa ayaymama sp. nov. Female habitus (MUSM-ENT 500411) a, c, e–f (a, dorsal; c, ventral; e, lateral, f, frontal) Epigynum b, d (b, dorsal; d, ventral) Scale bars: a, c, e, 1 mm, b, d 100 µm, f, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.10681271" httpUri="https://zenodo.org/record/10681271/files/figure.png" pageId="7" pageNumber="8">Fig. 4a</figureCitation>
) and by the pyriform-shaped head of the spermatheca (HS), which narrows anteriorly (
<figureCitation id="137A2A41BA0AFE2BFC5FFD756430FCC7" box="[275,361,763,789]" captionStart="FIGURE 3" captionStartId="10.[151,250,1933,1957]" captionTargetBox="[159,1428,1001,1903]" captionTargetId="figure-313@10.[151,1436,993,1910]" captionTargetPageId="10" captionText="FIGURE 3. Arctosa ayaymama sp. nov. Epigynum, holotype a–b (MUSM-ENT 500411) (a, ventral; b, dorsal); paratype c–d (MUSM-ENT 505143) (c, ventral; d, posterolateral) Scale bars: a–d, 0.25 mm. CO copulatory opening, FD fertilization duct, H hood, HS head of spermatheca, SS stalk of spermatheca, TS transversal septum." figureDoi="http://doi.org/10.5281/zenodo.10681269" httpUri="https://zenodo.org/record/10681269/files/figure.png" pageId="7" pageNumber="8">Figs 3b</figureCitation>
, 
<figureCitation id="137A2A41BA0AFE2BFC3AFD7564CCFCC4" box="[374,405,763,790]" captionStart="FIGURE 5" captionStartId="12.[151,250,1317,1341]" captionTargetBox="[151,1436,181,1293]" captionTargetId="figure-25@12.[151,1436,181,1293]" captionTargetPageId="12" captionText="FIGURE 5. Arctosa ayaymama sp. nov. Epigynum (MUSM-ENT 506993) a–c (a, ventral; b, dorsal; c, dictynoid pore) d, Male leg I, prolateral. Scale bars: a, 150 µm, b, 400 µm, c, 15 µm, d, 200 µm. CO copulatory opening, FD fertilization duct, H hood, HS head of spermatheca, DP dictynoid pore, MS median septum." figureDoi="http://doi.org/10.5281/zenodo.10681273" httpUri="https://zenodo.org/record/10681273/files/figure.png" pageId="7" pageNumber="8">5b</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35B654FBA0AFE2BFDDBFCCD6416F847" pageId="7" pageNumber="8" type="description">
<paragraph id="8BFE36C4BA0AFE2BFDDBFCCD64B4FC8F" blockId="7.[151,1437,835,1978]" box="[151,493,835,861]" pageId="7" pageNumber="8">
<emphasis id="B935EAD6BA0AFE2BFDDBFCCD6470FC8F" bold="true" box="[151,297,835,861]" pageId="7" pageNumber="8">Description.</emphasis>
<emphasis id="B935EAD6BA0AFE2BFC7CFCCD6428FC8F" box="[304,369,835,861]" italics="true" pageId="7" pageNumber="8">Male.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BFE36C4BA0AFE2BFD8BFCE966CBFB3B" blockId="7.[151,1437,835,1978]" pageId="7" pageNumber="8">
<emphasis id="B935EAD6BA0AFE2BFD8BFCE967C4FC53" box="[199,669,871,897]" italics="true" pageId="7" pageNumber="8">
Female. 
<typeStatus id="54FA8866BA0AFE2BFC67FCE964CBFC53" box="[299,402,871,897]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
(MUSM-ENT 500411).
</emphasis>
Carapace brown with black radial pattern, covered with tiny black setae. Longitudinal median band dark orange, covered by white setae, and tapering towards the opisthosoma (
<figureCitation id="137A2A41BA0AFE2BF823FC056596FC1B" captionStart="FIGURE 4" captionStartId="11.[151,250,1614,1638]" captionTargetBox="[151,1436,181,1591]" captionTargetId="figure-21@11.[151,1436,181,1591]" captionTargetPageId="11" captionText="FIGURE 4. Arctosa ayaymama sp. nov. Female habitus (MUSM-ENT 500411) a, c, e–f (a, dorsal; c, ventral; e, lateral, f, frontal) Epigynum b, d (b, dorsal; d, ventral) Scale bars: a, c, e, 1 mm, b, d 100 µm, f, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.10681271" httpUri="https://zenodo.org/record/10681271/files/figure.png" pageId="7" pageNumber="8">Fig. 4a, e</figureCitation>
), submarginal bands pale. Ocular area light orange, covered by sparse white setae (
<figureCitation id="137A2A41BA0AFE2BF933FC216189FC1B" box="[1151,1232,943,969]" captionStart="FIGURE 4" captionStartId="11.[151,250,1614,1638]" captionTargetBox="[151,1436,181,1591]" captionTargetId="figure-21@11.[151,1436,181,1591]" captionTargetPageId="11" captionText="FIGURE 4. Arctosa ayaymama sp. nov. Female habitus (MUSM-ENT 500411) a, c, e–f (a, dorsal; c, ventral; e, lateral, f, frontal) Epigynum b, d (b, dorsal; d, ventral) Scale bars: a, c, e, 1 mm, b, d 100 µm, f, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.10681271" httpUri="https://zenodo.org/record/10681271/files/figure.png" pageId="7" pageNumber="8">Fig. 4f</figureCitation>
). Sternum yellow, sparsely covered by long brown setae, labium pale brown. Anterior row of eyes procurved, shorter than the middle row, anterior median eyes larger than anterior lateral eyes (
<figureCitation id="137A2A41BA0AFE2BFE7DFC7966DBFBC3" box="[817,898,1015,1041]" captionStart="FIGURE 4" captionStartId="11.[151,250,1614,1638]" captionTargetBox="[151,1436,181,1591]" captionTargetId="figure-21@11.[151,1436,181,1591]" captionTargetPageId="11" captionText="FIGURE 4. Arctosa ayaymama sp. nov. Female habitus (MUSM-ENT 500411) a, c, e–f (a, dorsal; c, ventral; e, lateral, f, frontal) Epigynum b, d (b, dorsal; d, ventral) Scale bars: a, c, e, 1 mm, b, d 100 µm, f, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.10681271" httpUri="https://zenodo.org/record/10681271/files/figure.png" pageId="7" pageNumber="8">Fig. 4f</figureCitation>
). Chelicerae brown, darker than the endites and labium (
<figureCitation id="137A2A41BA0AFE2BFDB4FB956414FBE7" box="[248,333,1051,1077]" captionStart="FIGURE 4" captionStartId="11.[151,250,1614,1638]" captionTargetBox="[151,1436,181,1591]" captionTargetId="figure-21@11.[151,1436,181,1591]" captionTargetPageId="11" captionText="FIGURE 4. Arctosa ayaymama sp. nov. Female habitus (MUSM-ENT 500411) a, c, e–f (a, dorsal; c, ventral; e, lateral, f, frontal) Epigynum b, d (b, dorsal; d, ventral) Scale bars: a, c, e, 1 mm, b, d 100 µm, f, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.10681271" httpUri="https://zenodo.org/record/10681271/files/figure.png" pageId="7" pageNumber="8">Fig. 4c</figureCitation>
), and condyle short. Legs, dorsum of femora yellowish with black annuli, patellae, tibiae, and metatarsi yellowish with black annuli; tarsi yellow, with no annuli (
<figureCitation id="137A2A41BA0AFE2BFEC1FBB166B9FB8B" box="[909,992,1087,1113]" captionStart="FIGURE 4" captionStartId="11.[151,250,1614,1638]" captionTargetBox="[151,1436,181,1591]" captionTargetId="figure-21@11.[151,1436,181,1591]" captionTargetPageId="11" captionText="FIGURE 4. Arctosa ayaymama sp. nov. Female habitus (MUSM-ENT 500411) a, c, e–f (a, dorsal; c, ventral; e, lateral, f, frontal) Epigynum b, d (b, dorsal; d, ventral) Scale bars: a, c, e, 1 mm, b, d 100 µm, f, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.10681271" httpUri="https://zenodo.org/record/10681271/files/figure.png" pageId="7" pageNumber="8">Fig. 4e</figureCitation>
). A sparse scopulae on legs I−II, absent in III− IV. Spinules present only on tarsi III−IV. All tibiae, metatarsi, and tarsi with trichobothria increasing in size distally. Opisthosoma dark dusky with arrow-shaped cardiac mark flanked by yellow bands, covered by tiny white setae, with an irregular yellow median band, distal portion with a chevron yellow pattern (
<figureCitation id="137A2A41BA0AFE2BF933FB256188FB17" box="[1151,1233,1195,1221]" captionStart="FIGURE 4" captionStartId="11.[151,250,1614,1638]" captionTargetBox="[151,1436,181,1591]" captionTargetId="figure-21@11.[151,1436,181,1591]" captionTargetPageId="11" captionText="FIGURE 4. Arctosa ayaymama sp. nov. Female habitus (MUSM-ENT 500411) a, c, e–f (a, dorsal; c, ventral; e, lateral, f, frontal) Epigynum b, d (b, dorsal; d, ventral) Scale bars: a, c, e, 1 mm, b, d 100 µm, f, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.10681271" httpUri="https://zenodo.org/record/10681271/files/figure.png" pageId="7" pageNumber="8">Fig. 4a</figureCitation>
). Venter yellow as the sternum with a dusky irregular band of spots. Spinnerets yellow.
</paragraph>
<paragraph id="8BFE36C4BA0AFE2BFD8BFB7D6459FA4F" blockId="7.[151,1437,835,1978]" pageId="7" pageNumber="8">
Genitalia (
<figureCitation id="137A2A41BA0AFE2BFC76FB7D64D7FADF" box="[314,398,1267,1293]" captionStart="FIGURE 2" captionStartId="9.[151,250,1891,1915]" captionTargetBox="[185,1398,191,1857]" captionTargetId="figure-21@9.[180,1408,181,1867]" captionTargetPageId="9" captionText="FIGURE 2. Epigynum. a, c, f. Arctosa villa sp. nov (a, ventral; c, dorsal; f, posterolateral). b. Arctosa ayaymama sp. nov. d. Arctosa villa sp. nov. (anterior). e. Arctosa mineira sp. nov. (posterior). g. Arctosa jibarosa sp. nov. (posterolateral). Scale bars: a–g, 0.25 mm. AT, atrium, BS, base of spermatheca, CO, copulatory opening, FD, fertilization duct, H, hood, HS, head of spermatheca, MS, median septum, SS, stalk of spermatheca, TS, transversal septum, VC, vulval chamber." figureDoi="http://doi.org/10.5281/zenodo.10681267" httpUri="https://zenodo.org/record/10681267/files/figure.png" pageId="7" pageNumber="8">Figs2b</figureCitation>
, 
<figureCitation id="137A2A41BA0AFE2BFCDAFB7D6496FADF" box="[406,463,1267,1293]" captionStart="FIGURE 3" captionStartId="10.[151,250,1933,1957]" captionTargetBox="[159,1428,1001,1903]" captionTargetId="figure-313@10.[151,1436,993,1910]" captionTargetPageId="10" captionText="FIGURE 3. Arctosa ayaymama sp. nov. Epigynum, holotype a–b (MUSM-ENT 500411) (a, ventral; b, dorsal); paratype c–d (MUSM-ENT 505143) (c, ventral; d, posterolateral) Scale bars: a–d, 0.25 mm. CO copulatory opening, FD fertilization duct, H hood, HS head of spermatheca, SS stalk of spermatheca, TS transversal septum." figureDoi="http://doi.org/10.5281/zenodo.10681269" httpUri="https://zenodo.org/record/10681269/files/figure.png" pageId="7" pageNumber="8">3a‒d</figureCitation>
, 
<figureCitation id="137A2A41BA0AFE2BFC9BFB7D6754FADF" box="[471,525,1267,1293]" captionStart="FIGURE 4" captionStartId="11.[151,250,1614,1638]" captionTargetBox="[151,1436,181,1591]" captionTargetId="figure-21@11.[151,1436,181,1591]" captionTargetPageId="11" captionText="FIGURE 4. Arctosa ayaymama sp. nov. Female habitus (MUSM-ENT 500411) a, c, e–f (a, dorsal; c, ventral; e, lateral, f, frontal) Epigynum b, d (b, dorsal; d, ventral) Scale bars: a, c, e, 1 mm, b, d 100 µm, f, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.10681271" httpUri="https://zenodo.org/record/10681271/files/figure.png" pageId="7" pageNumber="8">4b,d</figureCitation>
) with median and transversal septum of epigyne not distinguishable from each other in ventral view, TS with rounded lateral borders; atrium narrow, MS not depressed. Fertilization ducts membranous and fusiform (
<figureCitation id="137A2A41BA0AFE2BFC7BFAB564D2FA87" box="[311,395,1339,1365]" captionStart="FIGURE 3" captionStartId="10.[151,250,1933,1957]" captionTargetBox="[159,1428,1001,1903]" captionTargetId="figure-313@10.[151,1436,993,1910]" captionTargetPageId="10" captionText="FIGURE 3. Arctosa ayaymama sp. nov. Epigynum, holotype a–b (MUSM-ENT 500411) (a, ventral; b, dorsal); paratype c–d (MUSM-ENT 505143) (c, ventral; d, posterolateral) Scale bars: a–d, 0.25 mm. CO copulatory opening, FD fertilization duct, H hood, HS head of spermatheca, SS stalk of spermatheca, TS transversal septum." figureDoi="http://doi.org/10.5281/zenodo.10681269" httpUri="https://zenodo.org/record/10681269/files/figure.png" pageId="7" pageNumber="8">Fig. 3b</figureCitation>
). Copulatory ducts narrow. Spermatheca with stalk broad and curved outward, quite as thick as the base of spermatheca; head of spermatheca pear-shaped, narrowing anteriorly visible in posterolateral view (
<figureCitation id="137A2A41BA0AFE2BFDD3FA0D65AAFA4F" box="[159,243,1411,1437]" captionStart="FIGURE 3" captionStartId="10.[151,250,1933,1957]" captionTargetBox="[159,1428,1001,1903]" captionTargetId="figure-313@10.[151,1436,993,1910]" captionTargetPageId="10" captionText="FIGURE 3. Arctosa ayaymama sp. nov. Epigynum, holotype a–b (MUSM-ENT 500411) (a, ventral; b, dorsal); paratype c–d (MUSM-ENT 505143) (c, ventral; d, posterolateral) Scale bars: a–d, 0.25 mm. CO copulatory opening, FD fertilization duct, H hood, HS head of spermatheca, SS stalk of spermatheca, TS transversal septum." figureDoi="http://doi.org/10.5281/zenodo.10681269" httpUri="https://zenodo.org/record/10681269/files/figure.png" pageId="7" pageNumber="8">Fig. 3d</figureCitation>
).
</paragraph>
<paragraph id="8BFE36C4BA0AFE2BFD8BFA2661E8F983" blockId="7.[151,1437,835,1978]" pageId="7" pageNumber="8">
Leg formula IV&gt;I&gt;II&gt;III. Spination pattern: femur 
<emphasis id="B935EAD6BA0AFE2BFFB2FA296646FA13" bold="true" box="[766,799,1447,1473]" pageId="7" pageNumber="8">I p</emphasis>
0-0-2 
<emphasis id="B935EAD6BA0AFE2BFE2DFA296628FA13" bold="true" box="[865,881,1447,1473]" pageId="7" pageNumber="8">d</emphasis>
1-1-1 
<emphasis id="B935EAD6BA0AFE2BFEFFFA2966E6FA13" bold="true" box="[947,959,1447,1473]" pageId="7" pageNumber="8">r</emphasis>
0-d1-1ap, 
<emphasis id="B935EAD6BA0AFE2BF97DFA296100FA13" bold="true" box="[1073,1113,1447,1473]" pageId="7" pageNumber="8">II r</emphasis>
0-d1-d1 
<emphasis id="B935EAD6BA0AFE2BF9F4FA296191FA13" bold="true" box="[1208,1224,1447,1473]" pageId="7" pageNumber="8">d</emphasis>
1-1-1, 
<emphasis id="B935EAD6BA0AFE2BF85DFA296044FA13" bold="true" box="[1297,1309,1447,1473]" pageId="7" pageNumber="8">r</emphasis>
0-d1-d1 
<emphasis id="B935EAD6BA0AFE2BF830FA2965FEFA37" bold="true" pageId="7" pageNumber="8">III p</emphasis>
0-d1-d1 
<emphasis id="B935EAD6BA0AFE2BFC4BFA45644EFA37" bold="true" box="[263,279,1483,1509]" pageId="7" pageNumber="8">d</emphasis>
1-1-1 
<emphasis id="B935EAD6BA0AFE2BFC16FA45643FFA37" bold="true" box="[346,358,1483,1509]" pageId="7" pageNumber="8">r</emphasis>
1-1-1, 
<emphasis id="B935EAD6BA0AFE2BFCFDFA4564BEFA37" bold="true" box="[433,487,1483,1509]" pageId="7" pageNumber="8">IV p</emphasis>
d1-0-d1 
<emphasis id="B935EAD6BA0AFE2BFF0AFA45670FFA37" bold="true" box="[582,598,1483,1509]" pageId="7" pageNumber="8">d</emphasis>
1-1-1 
<emphasis id="B935EAD6BA0AFE2BFFD5FA4567FCFA37" bold="true" box="[665,677,1483,1509]" pageId="7" pageNumber="8">r</emphasis>
0-0-1; patella 
<emphasis id="B935EAD6BA0AFE2BFE0FFA45663CFA37" bold="true" box="[835,869,1483,1509]" pageId="7" pageNumber="8">I p</emphasis>
1 
<emphasis id="B935EAD6BA0AFE2BFE36FA4566D3FA37" bold="true" box="[890,906,1483,1509]" pageId="7" pageNumber="8">d</emphasis>
1bristle-1 bristle 
<emphasis id="B935EAD6BA0AFE2BF91CFA456105FA37" bold="true" box="[1104,1116,1483,1509]" pageId="7" pageNumber="8">r</emphasis>
1, 
<emphasis id="B935EAD6BA0AFE2BF935FA4561FFFA37" bold="true" box="[1145,1190,1483,1509]" pageId="7" pageNumber="8">II p</emphasis>
1 
<emphasis id="B935EAD6BA0AFE2BF9F6FA456193FA37" bold="true" box="[1210,1226,1483,1509]" pageId="7" pageNumber="8">d</emphasis>
1 bristle – 1 bristle 
<emphasis id="B935EAD6BA0AFE2BFDDBFA6165FAF9DB" bold="true" box="[151,163,1519,1545]" pageId="7" pageNumber="8">r</emphasis>
1, 
<emphasis id="B935EAD6BA0AFE2BFD8DFA6165A3F9DB" bold="true" box="[193,250,1519,1545]" pageId="7" pageNumber="8">III p</emphasis>
1 
<emphasis id="B935EAD6BA0AFE2BFC5CFA616445F9DB" bold="true" box="[272,284,1519,1545]" pageId="7" pageNumber="8">r</emphasis>
1 
<emphasis id="B935EAD6BA0AFE2BFC7FFA61641AF9DB" bold="true" box="[307,323,1519,1545]" pageId="7" pageNumber="8">d</emphasis>
1-1bristle ap, 
<emphasis id="B935EAD6BA0AFE2BFCACFA61674EF9DB" bold="true" box="[480,535,1519,1545]" pageId="7" pageNumber="8">IV p</emphasis>
1 
<emphasis id="B935EAD6BA0AFE2BFF60FA616761F9DB" bold="true" box="[556,568,1519,1545]" pageId="7" pageNumber="8">r</emphasis>
1 
<emphasis id="B935EAD6BA0AFE2BFF03FA616706F9DB" bold="true" box="[591,607,1519,1545]" pageId="7" pageNumber="8">d</emphasis>
1bristle - 1ap; tibia 
<emphasis id="B935EAD6BA0AFE2BFE73FA61663AF9DB" bold="true" box="[831,867,1519,1545]" pageId="7" pageNumber="8">I p</emphasis>
1-v1 
<emphasis id="B935EAD6BA0AFE2BFED2FA6166F3F9DB" bold="true" box="[926,938,1519,1545]" pageId="7" pageNumber="8">r</emphasis>
0-1-1 
<emphasis id="B935EAD6BA0AFE2BFEA3FA6166A4F9DB" bold="true" box="[1007,1021,1519,1545]" pageId="7" pageNumber="8">v</emphasis>
2-2-2ap, 
<emphasis id="B935EAD6BA0AFE2BF928FA6161CBF9DB" bold="true" box="[1124,1170,1519,1545]" pageId="7" pageNumber="8">II p</emphasis>
d1-1 
<emphasis id="B935EAD6BA0AFE2BF981FA616180F9DB" bold="true" box="[1229,1241,1519,1545]" pageId="7" pageNumber="8">r</emphasis>
1-1 
<emphasis id="B935EAD6BA0AFE2BF84BFA61604CF9DB" bold="true" box="[1287,1301,1519,1545]" pageId="7" pageNumber="8">v</emphasis>
2-2-2ap 
<emphasis id="B935EAD6BA0AFE2BF839FA6160CFF9DB" bold="true" box="[1397,1430,1519,1545]" pageId="7" pageNumber="8">III</emphasis>
, 
<emphasis id="B935EAD6BA0AFE2BFDDBF99D65FEF9FF" bold="true" box="[151,167,1555,1581]" pageId="7" pageNumber="8">p</emphasis>
1-1 
<emphasis id="B935EAD6BA0AFE2BFD9FF99D65BAF9FF" bold="true" box="[211,227,1555,1581]" pageId="7" pageNumber="8">d</emphasis>
1-1 
<emphasis id="B935EAD6BA0AFE2BFC43F99D6442F9FF" bold="true" box="[271,283,1555,1581]" pageId="7" pageNumber="8">r</emphasis>
1-1 
<emphasis id="B935EAD6BA0AFE2BFC04F99D640FF9FF" bold="true" box="[328,342,1555,1581]" pageId="7" pageNumber="8">v</emphasis>
1-2-2ap, 
<emphasis id="B935EAD6BA0AFE2BFCF7F99D64A9F9FF" bold="true" box="[443,496,1555,1581]" pageId="7" pageNumber="8">IV p</emphasis>
1-v1 
<emphasis id="B935EAD6BA0AFE2BFF66F99D6763F9FF" bold="true" box="[554,570,1555,1581]" pageId="7" pageNumber="8">d</emphasis>
1-1 
<emphasis id="B935EAD6BA0AFE2BFF2AF99D672BF9FF" bold="true" box="[614,626,1555,1581]" pageId="7" pageNumber="8">r</emphasis>
1-1 
<emphasis id="B935EAD6BA0AFE2BFFD3F99D67E2F9FF" bold="true" box="[671,699,1555,1581]" pageId="7" pageNumber="8">v1</emphasis>
-2-2ap; metatarsus 
<emphasis id="B935EAD6BA0AFE2BFEDDF99D66EAF9FF" bold="true" box="[913,947,1555,1581]" pageId="7" pageNumber="8">I p</emphasis>
0-1-1ap 
<emphasis id="B935EAD6BA0AFE2BF95DF99D6144F9FF" bold="true" box="[1041,1053,1555,1581]" pageId="7" pageNumber="8">r</emphasis>
0-1-2ap 
<emphasis id="B935EAD6BA0AFE2BF937F99D61D0F9FF" bold="true" box="[1147,1161,1555,1581]" pageId="7" pageNumber="8">v</emphasis>
2-2-1ap, 
<emphasis id="B935EAD6BA0AFE2BF9A2F99D6042F9FF" bold="true" box="[1262,1307,1555,1581]" pageId="7" pageNumber="8">II p</emphasis>
1-1-2ap 
<emphasis id="B935EAD6BA0AFE2BF834F99D60DDF9FF" bold="true" box="[1400,1412,1555,1581]" pageId="7" pageNumber="8">r</emphasis>
0- 1-2ap 
<emphasis id="B935EAD6BA0AFE2BFD92F9B965B5F983" bold="true" box="[222,236,1591,1617]" pageId="7" pageNumber="8">v</emphasis>
2-2-1ap, 
<emphasis id="B935EAD6BA0AFE2BFC1DF9B964D0F983" bold="true" box="[337,393,1591,1617]" pageId="7" pageNumber="8">III p</emphasis>
1-1-2ap 
<emphasis id="B935EAD6BA0AFE2BFCAAF9B964ABF983" bold="true" box="[486,498,1591,1617]" pageId="7" pageNumber="8">r</emphasis>
d1-1-2ap 
<emphasis id="B935EAD6BA0AFE2BFF13F9B96734F983" bold="true" box="[607,621,1591,1617]" pageId="7" pageNumber="8">v</emphasis>
2-2-1ap, 
<emphasis id="B935EAD6BA0AFE2BFF9EF9B96651F983" bold="true" box="[722,776,1591,1617]" pageId="7" pageNumber="8">IV p</emphasis>
d1-d1-2ap 
<emphasis id="B935EAD6BA0AFE2BFECDF9B966D4F983" bold="true" box="[897,909,1591,1617]" pageId="7" pageNumber="8">r</emphasis>
d1-v1-d1-0-2ap 
<emphasis id="B935EAD6BA0AFE2BF908F9B9610BF983" bold="true" box="[1092,1106,1591,1617]" pageId="7" pageNumber="8">v</emphasis>
2-2-1ap.
</paragraph>
<paragraph id="8BFE36C4BA0AFE2BFD8BF9D26731F89F" blockId="7.[151,1437,835,1978]" pageId="7" pageNumber="8">
Measurements: TL 5.90, CL 2.80, CCW 1.64, CTW 2.50, CCH 1.24, CTH 1.32. Eyes: AME 0.20, ALE 0.10, PME 0.38, PLE 0.30, interdistances: AME–AME 0.12, AME–ALE 0.08; PME–PME 0.30, PME–PLE 0.38, PLEPLE 0.88, POQ length 0.66, POQ posterior width 1.04, POQ anterior width 0.60. Anterior eye row procurved, as long as the PMEs. Chelicerae: 1.18 length. Opisthosoma: length 2.76, width 1.98. Legs: length of segments (femur + patella+tibia + metatarsus+tarsus = total length): 
<emphasis id="B935EAD6BA0AFE2BFF96F965661BF8D7" bold="true" box="[730,834,1771,1797]" pageId="7" pageNumber="8">pedipalp</emphasis>
1.10 + 0.56 + 0.80 + - + 0.90 = 3.36, 
<emphasis id="B935EAD6BA0AFE2BF9BDF96561A5F8D7" bold="true" box="[1265,1276,1771,1797]" pageId="7" pageNumber="8">I</emphasis>
2.30 + 1.10 + 1.94 + 1.78 + 1.22 = 8.34, 
<emphasis id="B935EAD6BA0AFE2BFCF3F881648CF8FB" bold="true" box="[447,469,1807,1833]" pageId="7" pageNumber="8">II</emphasis>
2.10 + 0.98 + 1.68 + 1.66 + 1.16 = 7.58, 
<emphasis id="B935EAD6BA0AFE2BFED3F8816699F8FB" bold="true" box="[927,960,1807,1833]" pageId="7" pageNumber="8">III</emphasis>
2. 00 
<emphasis id="B935EAD6BA0AFE2BF948F881614DF8FB" bold="true" box="[1028,1044,1807,1833]" pageId="7" pageNumber="8">+</emphasis>
0.82 +1.54 + 2.02+1.10 = 7.48, 
<emphasis id="B935EAD6BA0AFE2BF831F88160C5F8FB" bold="true" box="[1405,1436,1807,1833]" pageId="7" pageNumber="8">IV</emphasis>
2. 68 + 1.10 + 2. 38 + 3.20+1.48 = 10.84.
</paragraph>
<paragraph id="8BFE36C4BA0AFE2BFD8BF8D96416F847" blockId="7.[151,1437,835,1978]" pageId="7" pageNumber="8">
<emphasis id="B935EAD6BA0AFE2BFD8BF8D96466F8A3" bold="true" box="[199,319,1879,1905]" pageId="7" pageNumber="8">Variation.</emphasis>
Females (N = 6, range, mean±s.d.): TL 5.67–7.92, 6.51±0.33; CL 3.00–3.67, 3.31±0.09; CW 2.17– 2.58, 2.46±0.06.
</paragraph>
</subSubSection>
<subSubSection id="C35B654FBA0AFE2BFD8BF811611AF86B" box="[199,1091,1951,1978]" pageId="7" pageNumber="8" type="distribution">
<paragraph id="8BFE36C4BA0AFE2BFD8BF811611AF86B" blockId="7.[151,1437,835,1978]" box="[199,1091,1951,1978]" pageId="7" pageNumber="8">
<emphasis id="B935EAD6BA0AFE2BFD8BF8116438F86B" bold="true" box="[199,353,1951,1977]" pageId="7" pageNumber="8">Distribution.</emphasis>
Amazonian 
<collectingCountry id="F3567654BA0AFE2BFCA2F8116705F86B" box="[494,604,1951,1977]" name="Colombia" pageId="7" pageNumber="8">Colombia</collectingCountry>
(
<collectingRegion id="4985F826BA0AFE2BFF27F8116793F86B" box="[619,714,1951,1977]" country="Colombia" name="Caqueta" pageId="7" pageNumber="8">Caquetá</collectingRegion>
) and 
<collectingCountry id="F3567654BA0AFE2BFE44F82E6662F868" box="[776,827,1952,1978]" name="Peru" pageId="7" pageNumber="8">Peru</collectingCountry>
(
<collectingRegion id="4985F826BA0AFE2BFE06F8116692F86B" box="[842,971,1951,1977]" country="Peru" name="San Martin" pageId="7" pageNumber="8">San Martín</collectingRegion>
) (
<figureCitation id="137A2A41BA0AFE2BFEAEF811616FF86B" box="[994,1078,1951,1977]" captionStart="FIGURE 62" captionStartId="72.[151,250,1618,1642]" captionTargetBox="[171,1418,564,1590]" captionTargetId="figure-197@72.[164,1423,561,1595]" captionTargetPageId="72" captionText="FIGURE 62. Geographic distribution records of Arctosa ayaymama sp. nov. (blue dot), Arctosa pacaya sp. nov. (red dot), Hogna andina comb. nov. (orange dot)." figureDoi="http://doi.org/10.5281/zenodo.10681391" httpUri="https://zenodo.org/record/10681391/files/figure.png" pageId="7" pageNumber="8">Fig. 62</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>