247 lines
29 KiB
XML
247 lines
29 KiB
XML
<document id="E11D47D635136B0A95F56F0836573D98" ID-DOI="10.1080/00222933.2019.1640907" ID-GBIF-Dataset="f36dee28-a5db-4864-9b69-3df980679006" ID-ISSN="1464-5262" ID-Zenodo-Dep="3673561" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582027737237" checkinUser="valdenar" docAuthor="Pacheco, T. L. & Vaz-de-Mello, F. Z." docDate="2019" docId="03E84F54DE497A03FE90EC41FB186F5B" docLanguage="en" docName="JNATHIST.53.19–20.1247–1253.pdf.imf" docOrigin="Journal of Natural History 53 (19)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Isacanthon Pacheco & Vaz-de-Mello 2019" docType="treatment" docVersion="9" lastPageNumber="1249" masterDocId="FFD1372CDE4B7A00FFFFED2CFF9B6D32" masterDocTitle="New dung beetle genus and species from a cave in the Espinhaço mountain rangeı Brazil (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="1253" masterPageNumber="1247" pageNumber="1248" updateTime="1698718947287" updateUser="plazi">
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<mods:mods id="6E7E400D7634FF0654DA5FCD01DBF4B7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="3384F83811D3B8C82E125580113DED36">
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<mods:title id="E2EFDFBFDED050F47FAE171CD1E39693">New dung beetle genus and species from a cave in the Espinhaço mountain rangeı Brazil (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:name id="1FE24E3253A521940223958C58A7959C" type="personal">
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<mods:namePart id="F340E3785D073DC6E0A16684322C1D4B">Pacheco, T. L.</mods:namePart>
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<mods:name id="124ADC1FFB99CC46E9DEFD20A78C6DF0" type="personal">
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<mods:roleTerm id="91A536F8EB10239911165D5ED22A1E63">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0B9C54ACFDFC8B413B21D9F183D8FBBE">Vaz-de-Mello, F. Z.</mods:namePart>
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<mods:typeOfResource id="D293D52EC4E86440A93979F4607D73FE">text</mods:typeOfResource>
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<mods:title id="5FDF2E67C155EE25F0DCAF079D5E7209">Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part id="AB7BF7DDABC1C7FB921AEFAB08E16611">
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<mods:date id="B552DC4AB0C92CEB09499F36AE198583">2019</mods:date>
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<mods:detail id="FD7319825152A9232613E86158681CEE" type="pubDate">
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<mods:number id="B6250FAD8B09C6A2EF63571E27A87BFA">2019-07-17</mods:number>
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<mods:detail id="187F633AAD337DA2767A9AB7C6FBEA2E" type="volume">
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<mods:number id="68840688E89019584546C3EE17C5BC11">53</mods:number>
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</mods:detail>
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<mods:detail id="5021AF7468D41711DC10636F4CE83BA5" type="issue">
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<mods:number id="FBB2DF265B6591CB69B873031BA9A7A3">19</mods:number>
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<mods:classification id="96DAE7B90BFEFC071ED7D7BB14FA9CE7">journal article</mods:classification>
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<mods:identifier id="1E8D4F450CC0911BEB9A139ABFD3A143" type="DOI">10.1080/00222933.2019.1640907</mods:identifier>
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<mods:identifier id="2AA1FEB8F7E282104A4DD03AC4E7D3F6" type="ISSN">1464-5262</mods:identifier>
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<mods:identifier id="F5EEEE3311AF1482AD3B1C46F96ED614" type="Zenodo-Dep">3673561</mods:identifier>
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<treatment id="03E84F54DE497A03FE90EC41FB186F5B" ID-DOI="http://doi.org/10.5281/zenodo.3679855" ID-GBIF-Taxon="162161901" ID-Zenodo-Dep="3679855" LSID="urn:lsid:plazi:treatment:03E84F54DE497A03FE90EC41FB186F5B" httpUri="http://treatment.plazi.org/id/03E84F54DE497A03FE90EC41FB186F5B" lastPageId="3" lastPageNumber="1249" pageId="2" pageNumber="1248">
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<subSubSection id="C35BADC9DE497A02FE90EC41FC2E6CB4" box="[367,949,365,390]" pageId="2" pageNumber="1248" type="nomenclature">
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<paragraph id="8BFEFE42DE497A02FE90EC41FC2E6CB4" blockId="2.[367,949,365,390]" box="[367,949,365,390]" pageId="2" pageNumber="1248">
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<heading id="D0B6492EDE497A02FE90EC41FC2E6CB4" box="[367,949,365,390]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="1248" reason="2">
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<taxonomicName id="4C4185C1DE497A02FE90EC41FE6D6CB4" ID-CoL="554N" authority="Pacheco & Vaz-de-Mello, 2019" authorityName="Pacheco & Vaz-de-Mello" authorityYear="2019" box="[367,502,365,390]" class="Insecta" family="Scarabaeidae" genus="Isacanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1251" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B9352250DE497A02FE90EC41FE6D6CB4" bold="true" box="[367,502,365,390]" italics="true" pageId="2" pageNumber="1248">Isacanthon</emphasis>
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</taxonomicName>
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Pacheco and Vaz-de-Mello gen. Nov.
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35BADC9DE497A02FF5FEC91FDAE69C6" pageId="2" pageNumber="1248" type="diagnosis">
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<paragraph id="8BFEFE42DE497A02FF5FEC91FE8F6CE4" blockId="2.[160,1156,445,1268]" box="[160,276,445,470]" pageId="2" pageNumber="1248">
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<emphasis id="B9352250DE497A02FF5FEC91FE8F6CE4" bold="true" box="[160,276,445,470]" italics="true" pageId="2" pageNumber="1248">Diagnosis</emphasis>
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</paragraph>
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<paragraph id="8BFEFE42DE497A02FF5FECCDFDF26FFB" blockId="2.[160,1156,445,1268]" pageId="2" pageNumber="1248">Body very flattened, surface with microsculpture between ocellate punctuation, punctures with setae, clypeus with two teeth appearing to originate beneath clypeal margin, eyes slightly visible in dorsal view, elytra with eight striae, pseudoepipleuron large and with one stria, mesoventrite short, carina between mesoventrite and metaventrite almost straight, prolegs with anterior trochantofemoral pit, protibiae with three distinct teeth on external margin, metatibiae with one distinct tooth at its basal two-fifth, and aedeagus with asymmetrical parameres.</paragraph>
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<paragraph id="8BFEFE42DE497A02FF3FEFF8FC54699C" blockId="2.[160,1156,445,1268]" pageId="2" pageNumber="1248">
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To exclude the possibility that this genus can be confused with
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<taxonomicName id="4C4185C1DE497A02FC60EFF8FBBE6FDE" authorityName="Balthasar" authorityYear="1938" box="[927,1061,724,748]" class="Insecta" family="Scarabaeidae" genus="Paracanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1248" phylum="Arthropoda" rank="genus">
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<emphasis id="B9352250DE497A02FC60EFF8FBBE6FDE" box="[927,1061,724,748]" italics="true" pageId="2" pageNumber="1248">Paracanthon</emphasis>
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</taxonomicName>
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see the following characters: clypeus with two teeth appearing to originate beneath clypeal margin (
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<taxonomicName id="4C4185C1DE497A02FF58EE35FE816E03" authority="Pacheco & Vaz-de-Mello, 2019" authorityName="Pacheco & Vaz-de-Mello" authorityYear="2019" box="[167,282,793,817]" class="Insecta" family="Scarabaeidae" genus="Isacanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1251" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B9352250DE497A02FF58EE35FE816E03" box="[167,282,793,817]" italics="true" pageId="2" pageNumber="1248">Isacanthon</emphasis>
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</taxonomicName>
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) or originating from leading margin of clypeus (
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<taxonomicName id="4C4185C1DE497A02FCCEEE35FC2C6E03" authorityName="Balthasar" authorityYear="1938" box="[817,951,793,817]" class="Insecta" family="Scarabaeidae" genus="Paracanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1248" phylum="Arthropoda" rank="genus">
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<emphasis id="B9352250DE497A02FCCEEE35FC2C6E03" box="[817,951,793,817]" italics="true" pageId="2" pageNumber="1248">Paracanthon</emphasis>
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</taxonomicName>
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), hypomeron with (
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<taxonomicName id="4C4185C1DE497A02FF58EE10FEB66E66" authorityName="Balthasar" authorityYear="1938" box="[167,301,828,852]" class="Insecta" family="Scarabaeidae" genus="Paracanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1248" phylum="Arthropoda" rank="genus">
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<emphasis id="B9352250DE497A02FF58EE10FEB66E66" box="[167,301,828,852]" italics="true" pageId="2" pageNumber="1248">Paracanthon</emphasis>
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</taxonomicName>
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) or without (
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<taxonomicName id="4C4185C1DE497A02FE3FEE10FDA86E66" authority="Pacheco & Vaz-de-Mello, 2019" authorityName="Pacheco & Vaz-de-Mello" authorityYear="2019" box="[448,563,828,852]" class="Insecta" family="Scarabaeidae" genus="Isacanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1251" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B9352250DE497A02FE3FEE10FDA86E66" box="[448,563,828,852]" italics="true" pageId="2" pageNumber="1248">Isacanthon</emphasis>
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</taxonomicName>
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) carina parallel to the margin of the dorsal surface of pronotum, and by presence (
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<taxonomicName id="4C4185C1DE497A02FE27EE72FDD06E44" authority="Pacheco & Vaz-de-Mello, 2019" authorityName="Pacheco & Vaz-de-Mello" authorityYear="2019" box="[472,587,862,886]" class="Insecta" family="Scarabaeidae" genus="Isacanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1251" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B9352250DE497A02FE27EE72FDD06E44" box="[472,587,862,886]" italics="true" pageId="2" pageNumber="1248">Isacanthon</emphasis>
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</taxonomicName>
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) or absence (
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<taxonomicName id="4C4185C1DE497A02FD25EE72FCFB6E44" authorityName="Balthasar" authorityYear="1938" box="[730,864,862,886]" class="Insecta" family="Scarabaeidae" genus="Paracanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1248" phylum="Arthropoda" rank="genus">
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<emphasis id="B9352250DE497A02FD25EE72FCFB6E44" box="[730,864,862,886]" italics="true" pageId="2" pageNumber="1248">Paracanthon</emphasis>
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</taxonomicName>
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) of the unique armature of metatibiae. From
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<taxonomicName id="4C4185C1DE497A02FE9FEEADFDBC6EAB" box="[352,551,897,921]" class="Insecta" family="Scarabaeidae" genus="Paracryptocanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1248" phylum="Arthropoda" rank="genus">
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<emphasis id="B9352250DE497A02FE9FEEADFDBC6EAB" box="[352,551,897,921]" italics="true" pageId="2" pageNumber="1248">Paracryptocanthon</emphasis>
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</taxonomicName>
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is distinguished by clypeus with two teeth (
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<taxonomicName id="4C4185C1DE497A02FBF6EEADFBE76EAB" authority="Pacheco & Vaz-de-Mello, 2019" authorityName="Pacheco & Vaz-de-Mello" authorityYear="2019" box="[1033,1148,897,921]" class="Insecta" family="Scarabaeidae" genus="Isacanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1251" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B9352250DE497A02FBF6EEADFBE76EAB" box="[1033,1148,897,921]" italics="true" pageId="2" pageNumber="1248">Isacanthon</emphasis>
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</taxonomicName>
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) instead of six (
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<taxonomicName id="4C4185C1DE497A02FE99EE88FDB66E8E" box="[358,557,932,956]" class="Insecta" family="Scarabaeidae" genus="Paracryptocanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1248" phylum="Arthropoda" rank="genus">
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<emphasis id="B9352250DE497A02FE99EE88FDB66E8E" box="[358,557,932,956]" italics="true" pageId="2" pageNumber="1248">Paracryptocanthon</emphasis>
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</taxonomicName>
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), pygidium in vertical (
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<taxonomicName id="4C4185C1DE497A02FCA3EE88FC546E8E" authority="Pacheco & Vaz-de-Mello, 2019" authorityName="Pacheco & Vaz-de-Mello" authorityYear="2019" box="[860,975,932,956]" class="Insecta" family="Scarabaeidae" genus="Isacanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1251" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B9352250DE497A02FCA3EE88FC546E8E" box="[860,975,932,956]" italics="true" pageId="2" pageNumber="1248">Isacanthon</emphasis>
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</taxonomicName>
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) or horizontal (
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<taxonomicName id="4C4185C1DE497A02FF58EEEAFEF56EEC" box="[167,366,966,990]" class="Insecta" family="Scarabaeidae" genus="Paracryptocanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1248" phylum="Arthropoda" rank="genus">
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<emphasis id="B9352250DE497A02FF58EEEAFEF56EEC" box="[167,366,966,990]" italics="true" pageId="2" pageNumber="1248">Paracryptocanthon</emphasis>
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</taxonomicName>
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) position, eyes entirely visible (
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<taxonomicName id="4C4185C1DE497A02FD2AEEEAFCD36EEC" authority="Pacheco & Vaz-de-Mello, 2019" authorityName="Pacheco & Vaz-de-Mello" authorityYear="2019" box="[725,840,966,990]" class="Insecta" family="Scarabaeidae" genus="Isacanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1251" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B9352250DE497A02FD2AEEEAFCD36EEC" box="[725,840,966,990]" italics="true" pageId="2" pageNumber="1248">Isacanthon</emphasis>
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</taxonomicName>
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) or not (
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<taxonomicName id="4C4185C1DE497A02FC4AEEEAFBE76EEC" box="[949,1148,966,990]" class="Insecta" family="Scarabaeidae" genus="Paracryptocanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1248" phylum="Arthropoda" rank="genus">
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<emphasis id="B9352250DE497A02FC4AEEEAFBE76EEC" box="[949,1148,966,990]" italics="true" pageId="2" pageNumber="1248">Paracryptocanthon</emphasis>
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</taxonomicName>
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) dorsally, and the presence (
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<taxonomicName id="4C4185C1DE497A02FE1DEEC5FDCE6933" authority="Pacheco & Vaz-de-Mello, 2019" authorityName="Pacheco & Vaz-de-Mello" authorityYear="2019" box="[482,597,1001,1025]" class="Insecta" family="Scarabaeidae" genus="Isacanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1251" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B9352250DE497A02FE1DEEC5FDCE6933" box="[482,597,1001,1025]" italics="true" pageId="2" pageNumber="1248">Isacanthon</emphasis>
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</taxonomicName>
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) or absence (
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<taxonomicName id="4C4185C1DE497A02FD04EEC5FC596933" box="[763,962,1001,1025]" class="Insecta" family="Scarabaeidae" genus="Paracryptocanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1248" phylum="Arthropoda" rank="genus">
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<emphasis id="B9352250DE497A02FD04EEC5FC596933" box="[763,962,1001,1025]" italics="true" pageId="2" pageNumber="1248">Paracryptocanthon</emphasis>
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</taxonomicName>
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) of the spur on metatibiae. The entirely visible eyes (
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<taxonomicName id="4C4185C1DE497A02FDB4E920FD256916" authority="Pacheco & Vaz-de-Mello, 2019" authorityName="Pacheco & Vaz-de-Mello" authorityYear="2019" box="[587,702,1036,1060]" class="Insecta" family="Scarabaeidae" genus="Isacanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1251" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B9352250DE497A02FDB4E920FD256916" box="[587,702,1036,1060]" italics="true" pageId="2" pageNumber="1248">Isacanthon</emphasis>
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</taxonomicName>
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) or not (
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<taxonomicName id="4C4185C1DE497A02FCD1E920FC526916" authorityName="Balthasar" authorityYear="1942" box="[814,969,1036,1060]" class="Insecta" family="Scarabaeidae" genus="Cryptocanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1248" phylum="Arthropoda" rank="genus">
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<emphasis id="B9352250DE497A02FCD1E920FC526916" box="[814,969,1036,1060]" italics="true" pageId="2" pageNumber="1248">Cryptocanthon</emphasis>
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</taxonomicName>
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) in dorsal view, parameres asymmetrical (
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<taxonomicName id="4C4185C1DE497A02FE48E902FDB16974" authority="Pacheco & Vaz-de-Mello, 2019" authorityName="Pacheco & Vaz-de-Mello" authorityYear="2019" box="[439,554,1070,1094]" class="Insecta" family="Scarabaeidae" genus="Isacanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1251" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9352250DE497A02FE48E902FDB16974" box="[439,554,1070,1094]" italics="true" pageId="2" pageNumber="1248">Isacanthon</emphasis>
|
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</taxonomicName>
|
||
) or not (
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<taxonomicName id="4C4185C1DE497A02FD73E902FCBC6974" authorityName="Balthasar" authorityYear="1942" box="[652,807,1070,1094]" class="Insecta" family="Scarabaeidae" genus="Cryptocanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1248" phylum="Arthropoda" rank="genus">
|
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<emphasis id="B9352250DE497A02FD73E902FCBC6974" box="[652,807,1070,1094]" italics="true" pageId="2" pageNumber="1248">Cryptocanthon</emphasis>
|
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</taxonomicName>
|
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), and much larger size separate it from
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<taxonomicName id="4C4185C1DE497A02FF04E97DFE0D695B" authorityName="Balthasar" authorityYear="1942" box="[251,406,1105,1129]" class="Insecta" family="Scarabaeidae" genus="Cryptocanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1248" phylum="Arthropoda" rank="genus">
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<emphasis id="B9352250DE497A02FF04E97DFE0D695B" box="[251,406,1105,1129]" italics="true" pageId="2" pageNumber="1248">Cryptocanthon</emphasis>
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</taxonomicName>
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. Finally, the presence of just one stria (
|
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<taxonomicName id="4C4185C1DE497A02FC96E97DFC47695B" authority="Pacheco & Vaz-de-Mello, 2019" authorityName="Pacheco & Vaz-de-Mello" authorityYear="2019" box="[873,988,1105,1129]" class="Insecta" family="Scarabaeidae" genus="Isacanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1251" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
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<emphasis id="B9352250DE497A02FC96E97DFC47695B" box="[873,988,1105,1129]" italics="true" pageId="2" pageNumber="1248">Isacanthon</emphasis>
|
||
</taxonomicName>
|
||
) instead three (
|
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<emphasis id="B9352250DE497A02FF58E958FED769BE" box="[167,332,1140,1164]" italics="true" pageId="2" pageNumber="1248">
|
||
<taxonomicName id="4C4185C1DE497A02FF58E958FEDF69BE" authorityName="Vaz-De-Mello & Halffter" authorityYear="2006" box="[167,324,1140,1164]" class="Insecta" family="Scarabaeidae" genus="Tesserodoniella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1248" phylum="Arthropoda" rank="genus">Tesserodoniella</taxonomicName>
|
||
)
|
||
</emphasis>
|
||
on pseudoepipleuron, together with all previously mentioned characters under previous diagnoses, differentiate the new taxon from
|
||
<taxonomicName id="4C4185C1DE497A02FCD4E9BAFC52699C" authorityName="Vaz-De-Mello & Halffter" authorityYear="2006" box="[811,969,1174,1198]" class="Insecta" family="Scarabaeidae" genus="Tesserodoniella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1248" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9352250DE497A02FCD4E9BAFC52699C" box="[811,969,1174,1198]" italics="true" pageId="2" pageNumber="1248">Tesserodoniella</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BFEFE42DE497A02FF3FE995FDAE69C6" blockId="2.[160,1156,445,1268]" pageId="2" pageNumber="1248">As only one species is known so far, the description of the new genus and the new species shall be seen as equivalent.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35BADC9DE497A02FF5FE81CFB936883" pageId="2" pageNumber="1248" type="etymology">
|
||
<paragraph id="8BFEFE42DE497A02FF5FE81CFE86687B" blockId="2.[160,1155,1328,1457]" box="[160,285,1328,1353]" pageId="2" pageNumber="1248">
|
||
<emphasis id="B9352250DE497A02FF5FE81CFE86687B" bold="true" box="[160,285,1328,1353]" italics="true" pageId="2" pageNumber="1248">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFEFE42DE497A02FF5FE878FB936883" blockId="2.[160,1155,1328,1457]" pageId="2" pageNumber="1248">
|
||
The prefix
|
||
<taxonomicName id="4C4185C1DE497A02FEE4E878FEAC685E" authorityName="Packard" authorityYear="1864" box="[283,311,1364,1388]" class="Insecta" family="Limacodidae" genus="Isa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="1248" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9352250DE497A02FEE4E878FEAC685E" box="[283,311,1364,1388]" italics="true" pageId="2" pageNumber="1248">Isa</emphasis>
|
||
</taxonomicName>
|
||
is named after Isac da Silva
|
||
<collectingRegion id="498530A0DE497A02FD78E878FD2C685E" box="[647,695,1364,1388]" country="Venezuela" name="Lara" pageId="2" pageNumber="1248">Lara</collectingRegion>
|
||
, grandfather of the first author.
|
||
<taxonomicName id="4C4185C1DE497A02FBD6E878FEE468BC" authority="Hoffmannsegg, 1817" authorityName="Hoffmannsegg" authorityYear="1817" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1248" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9352250DE497A02FBD6E878FB18685E" box="[1065,1155,1364,1388]" italics="true" pageId="2" pageNumber="1248">Canthon</emphasis>
|
||
<bibRefCitation id="EFD083B3DE497A02FF5FE85AFEE468BC" author="Hoffmannsegg JCG" box="[160,383,1398,1422]" pageId="2" pageNumber="1248" pagination="8 - 56" refId="ref2604" refString="Hoffmannsegg JCG. 1817. Entomologische Bemerkungen bei Gelegenheit der Abhandlungen uber amerikanische Insecten, in der vierten bis sechsten Lieferung von den recueils d ' observations de zoologie et d ' anotomie comparee, oder dem 2 ten Theile der Reise, der Herren Al. v. Humboldt und A. Bonpland [Entomological remarks on occasion of treatises on American insects, in the fourth to sixth deliveries of the set of observations of zoology and comparative anatomy, from the second part of the voyage, Messrs. Al. v. Humboldt and A. Bonpland], nemlich: no. IX. in Livr. 4. p. 197 - 283. und No. XI. XII. in Livr. 5. 6. p. 294 - 397. Zoologisches Magazin. 1 (1): 8 - 56." type="journal article" year="1817">Hoffmannsegg, 1817</bibRefCitation>
|
||
</taxonomicName>
|
||
is a large New World genus of Scarabaeinae, with several (not necessarily phylogenetically related) derivatives, all lacking expanded meso- and metatibiae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35BADC9DE497A02FF5FE8C1FD146B86" pageId="2" pageNumber="1248" type="discussion">
|
||
<subSection id="E2CEE7A9DE497A02FF5FE8C1FEAF6B34" box="[160,308,1517,1542]" pageId="2" pageNumber="1248" type="multiple">
|
||
<paragraph id="8BFEFE42DE497A02FF5FE8C1FEAF6B34" blockId="2.[160,308,1517,1542]" box="[160,308,1517,1542]" pageId="2" pageNumber="1248">
|
||
<emphasis id="B9352250DE497A02FF5FE8C1FEAF6B34" bold="true" box="[160,308,1517,1542]" italics="true" pageId="2" pageNumber="1248">
|
||
<typeStatus id="54FA40E0DE497A02FF5FE8C1FF436B34" box="[160,216,1517,1542]" pageId="2" pageNumber="1248">Type</typeStatus>
|
||
species
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSection>
|
||
<paragraph id="8BFEFE42DE497A02FF45EB1CFBF16B7B" blockId="2.[186,1130,1584,1609]" box="[186,1130,1584,1609]" pageId="2" pageNumber="1248">
|
||
<heading id="D0B6492EDE497A02FF45EB1CFBF16B7B" box="[186,1130,1584,1609]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="1248" reason="2">
|
||
<taxonomicName id="4C4185C1DE497A02FF45EB1CFDD36B7B" authority="Pacheco" authorityName="Pacheco & Vaz-de-Mello" authorityYear="2019" box="[186,584,1584,1609]" class="Insecta" family="Scarabaeidae" genus="Isacanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1248" phylum="Arthropoda" rank="species" species="mariaelinae">
|
||
<emphasis id="B9352250DE497A02FF45EB1CFE476B7B" bold="true" box="[186,476,1584,1609]" italics="true" pageId="2" pageNumber="1248">Isacanthon mariaelinae</emphasis>
|
||
Pacheco
|
||
</taxonomicName>
|
||
and Vaz-de-Mello sp. nov.,
|
||
<emphasis id="B9352250DE497A02FC6AEB1CFBFF6B7B" bold="true" box="[917,1124,1584,1609]" pageId="2" pageNumber="1248">here designated</emphasis>
|
||
.
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BFEFE42DE497A02FF3FEB55FD146B86" blockId="2.[160,1155,1657,1716]" pageId="2" pageNumber="1248">
|
||
This genus can be separated from other Neotropical genera by modifying couplet 34 of
|
||
<bibRefCitation id="EFD083B3DE497A02FF40EBB0FE7A6B86" author="Vaz-de-Mello FZ & Edmonds WD & Ocampo FC & Schoolmeesters P" box="[191,481,1692,1716]" pageId="2" pageNumber="1248" pagination="1 - 73" refId="ref3144" refString="Vaz-de-Mello FZ, Edmonds WD, Ocampo FC, Schoolmeesters P. 2011. A multilingual key to the genera and subgenera of the subfamily Scarabaeinae of the New World (Coleoptera: Scarabaeidae). Zootaxa. 2854: 1 - 73." type="journal article" year="2011">Vaz-de-Mello et al. (2011)</bibRefCitation>
|
||
key as follows:
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35BADC9DE487A03FF40EDA3FB186F5B" pageId="3" pageNumber="1249" type="key">
|
||
<paragraph id="8BFEFE42DE487A03FF40EDA3FB1F6DF8" blockId="3.[191,1156,143,617]" pageId="3" pageNumber="1249">
|
||
34 (33)
|
||
<emphasis id="B9352250DE487A03FEE5EDA3FE9D6DF8" italics="true" pageId="3" pageNumber="1249">Length of metatarsomere 1 greater than combined length of metatarsomeres 2–3</emphasis>
|
||
.... ............................................................................................................................................... 34A
|
||
</paragraph>
|
||
<paragraph id="8BFEFE42DE487A03FF40EDF8FB186C3D" blockId="3.[191,1156,143,617]" pageId="3" pageNumber="1249">
|
||
<emphasis id="B9352250DE487A03FF40EDF8FE9D6C3D" italics="true" pageId="3" pageNumber="1249">– Length of metatarsomere 1 shorter than combined length of metatarsomeres 2–3</emphasis>
|
||
... ................................................................................................................................................... 35
|
||
</paragraph>
|
||
<paragraph id="8BFEFE42DE487A03FF40EC36FB186C8E" blockId="3.[191,1156,143,617]" pageId="3" pageNumber="1249">
|
||
34A (34) Clypeal teeth appearing to originate on clypeal margin. Hypomeron with posterior carina parallel to edge. If elytra with ocellate puncture, without microsculpture between punctuation. Metatibiae without distinct tooth at their two-fifth. Widely distributed across the Atlantic Forest in
|
||
<collectingCountry id="F356BED2DE487A03FD02ECAEFCA26CA8" box="[765,825,386,410]" name="Brazil" pageId="3" pageNumber="1249">Brazil</collectingCountry>
|
||
and
|
||
<collectingCountry id="F356BED2DE487A03FC89ECAEFC7E6CA8" box="[886,997,386,410]" name="Argentina" pageId="3" pageNumber="1249">Argentina</collectingCountry>
|
||
.. ....................... ...........................................................................................................................................
|
||
<taxonomicName id="4C4185C1DE487A03FC12EC88FB186C8E" ID-CoL="6DTT" authorityName="Balthasar" authorityYear="1938" box="[1005,1155,420,444]" class="Insecta" family="Scarabaeidae" genus="Paracanthon" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1249" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9352250DE487A03FC12EC88FB186C8E" bold="true" box="[1005,1155,420,444]" italics="true" pageId="3" pageNumber="1249">Paracanthon</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BFEFE42DE487A03FF40ECEBFB186F5B" blockId="3.[191,1156,143,617]" pageId="3" pageNumber="1249">
|
||
<emphasis id="B9352250DE487A03FF40ECEBFF576CED" box="[191,204,455,479]" italics="true" pageId="3" pageNumber="1249">–</emphasis>
|
||
Clypeal teeth appearing to originate beneath clypeal margin. Hypomeron without posterior carina parallel to edge (
|
||
<figureCitation id="137AE2C7DE487A03FD9CECC6FD256F30" box="[611,702,490,514]" captionStart="Figure 1" captionStartId="4.[160,225,1665,1688]" captionTargetBox="[171,1147,155,1624]" captionTargetId="figure@4.[163,1153,142,1634]" captionTargetPageId="4" captionText="Figure 1. Isacanthon mariaelinae sp. nov. (a) Lateral view. (b) Dorsal view. (c) Right view of the aedeagus. (d) Left view of the aedeagus. Scar bars: (a,b) 500 mm; (c,d) 500 μm." figureDoi="http://doi.org/10.5281/zenodo.3673564" httpUri="https://zenodo.org/record/3673564/files/figure.png" pageId="3" pageNumber="1249">Figure 1</figureCitation>
|
||
(a)). Elytra with ocellate puncture, with microsculpture between punctuation. Metatibiae with distinct tooth at their twofifth (
|
||
<figureCitation id="137AE2C7DE487A03FEE0EF03FEE06F75" box="[287,379,559,583]" captionStart="Figure 1" captionStartId="4.[160,225,1665,1688]" captionTargetBox="[171,1147,155,1624]" captionTargetId="figure@4.[163,1153,142,1634]" captionTargetPageId="4" captionText="Figure 1. Isacanthon mariaelinae sp. nov. (a) Lateral view. (b) Dorsal view. (c) Right view of the aedeagus. (d) Left view of the aedeagus. Scar bars: (a,b) 500 mm; (c,d) 500 μm." figureDoi="http://doi.org/10.5281/zenodo.3673564" httpUri="https://zenodo.org/record/3673564/files/figure.png" pageId="3" pageNumber="1249">Figure 1</figureCitation>
|
||
(b)). Know only from one place in the Espinhaço mountain range, southeastern
|
||
<collectingCountry id="F356BED2DE487A03FE86EF7DFE2E6F5B" box="[377,437,593,617]" name="Brazil" pageId="3" pageNumber="1249">Brazil</collectingCountry>
|
||
.. ..................................... ....................................
|
||
<emphasis id="B9352250DE487A03FC89EF7DFB186F5B" bold="true" box="[886,1155,593,617]" pageId="3" pageNumber="1249">
|
||
<taxonomicName id="4C4185C1DE487A03FC89EF7DFC6C6F5B" ID-CoL="554N" authority="Pacheco & Vaz-de-Mello, 2019" authorityName="Pacheco & Vaz-de-Mello" authorityYear="2019" box="[886,1015,593,617]" class="Insecta" family="Scarabaeidae" genus="Isacanthon" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1249" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9352250DE487A03FC89EF7DFC6C6F5B" bold="true" box="[886,1015,593,617]" italics="true" pageId="3" pageNumber="1249">Isacanthon</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2069F2BDE487A03FBFFEF7DFB186F5B" box="[1024,1155,593,617]" pageId="3" pageNumber="1249" rank="genus">new genus</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |