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<document id="4861A8F04CFFA14A344D3BB466AF9760" ID-CLB-Dataset="7679" ID-DOI="10.11646/zootaxa.4942.1.2" ID-GBIF-Dataset="25d73144-c90c-45c3-a9a2-2454f87a9aef" ID-ISSN="1175-5326" ID-Zenodo-Dep="4596097" ID-ZooBank="79353662-7653-4F41-8B39-40E6E4B2E005" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615448142785" checkinUser="plazi" docAuthor="Haddad, Charles R., Jin, Chi, Platnick, Norman I. &amp; Booysen, Ruan" docDate="2021" docId="03D67155FFBDFFB5FF25FCF8A11C2E2C" docLanguage="en" docName="zootaxa.4942.1.2.pdf" docOrigin="Zootaxa 4942 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Capobula ukhahlamba Haddad &amp; Jin &amp; Platnick &amp; Booysen 2021, spec. nov." docType="treatment" docVersion="6" lastPageNumber="60" masterDocId="FFEF092DFFAFFFA6FFB2FFBAA0202918" masterDocTitle="Capobula gen. nov., a new Afrotropical dark sac spider genus related to Orthobula Simon, 1897 (Araneae: Trachelidae)" masterLastPageNumber="71" masterPageNumber="41" pageNumber="59" updateTime="1698933034801" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C2074D1EF605304D401646890CD73898">Capobula gen. nov., a new Afrotropical dark sac spider genus related to Orthobula Simon, 1897 (Araneae: Trachelidae)</mods:title>
</mods:titleInfo>
<mods:name id="A6F7B0AF975EC921BA15CDBA641FF54A" type="personal">
<mods:role id="127C99F16011D1BBBE4DCFCEA0BA28FC">
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<mods:namePart id="7FB14378DD88F183C3CFB7BDC3B44C40">Haddad, Charles R.</mods:namePart>
<mods:nameIdentifier id="9A4A787A6756B7D5FBE1DC6DD604B0CB" type="ORCID">0000-0002-2317-7760</mods:nameIdentifier>
<mods:affiliation id="04409E3A138BB3FA0FE79A89602878D6">Department of Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein 9300, South Africa. &amp; haddadcr @ ufs. ac. za; https: // orcid. org / 0000 - 0002 - 2317 - 7760; &amp; booysenr @ ufs. ac. za; https: // orcid. org / 0000 - 0002 - 0841 - 9143</mods:affiliation>
<mods:nameIdentifier id="128595E5E30DC0307A12AECE6DFB292B" type="email">haddadcr@ufs.ac.za</mods:nameIdentifier>
</mods:name>
<mods:name id="46DE6B38CE42F41146390649D5276AE0" type="personal">
<mods:role id="F2C9D2477F3C29157744DD83CC3F4C7A">
<mods:roleTerm id="B465259F6984D78035AD3D5214681EFF">Author</mods:roleTerm>
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<mods:namePart id="60597C11F2A5E9C740C95ECB40391C28">Jin, Chi</mods:namePart>
<mods:nameIdentifier id="831D5393E7CD14C3E7C789F87FE2B71C" type="ORCID">0000-0002-4694-4870</mods:nameIdentifier>
<mods:affiliation id="E30EBD13EE1CFB44C27C15199FC1756B">College of Landscape and Ecological Engineering, Hebei University of Engineering, Handan, Hebei 056038, P. R. China. &amp; jinchi _ spider @ 163. com; https: // orcid. org / 0000 - 0002 - 4694 - 4870</mods:affiliation>
</mods:name>
<mods:name id="5439EB25F486E3D37D7C44B893EE7242" type="personal">
<mods:role id="BDCA83E1EF540FBA5024EECE006D39F5">
<mods:roleTerm id="68811C6B4EA7D2A11E70EDBAAB068072">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8CA2D2DFB292624DDBDBA1D3A8FC512A">Platnick, Norman I.</mods:namePart>
<mods:nameIdentifier id="9D3811BC58A9F65E7FADFAC66817BB81" type="ORCID">0000-0003-4868-7529</mods:nameIdentifier>
<mods:affiliation id="FDB625EDA01F3816BE57C41B67CBAAE2">Division of Invertebrate Zoology, American Museum of Natural History, Central Park West at 79 th Street, New York, NY 10024, U. S. A. &amp; https: // orcid. org / 0000 - 0003 - 4868 - 7529; deceased</mods:affiliation>
</mods:name>
<mods:name id="A6307711CACA327DC6D33A8C945F8DF5" type="personal">
<mods:role id="B9C9DFB7D2A4EB7376334201BB65ACA6">
<mods:roleTerm id="3A14D988C4AE8E82CEB5EE627DE990E9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D17CEFB293459A8B032973957A24B27A">Booysen, Ruan</mods:namePart>
<mods:nameIdentifier id="25E9300BBB46D38B126BFB552399A9F8" type="ORCID">0000-0002-2317-7760</mods:nameIdentifier>
<mods:affiliation id="444D21AF689FD225C23D51BFE5E98C24">Department of Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein 9300, South Africa. &amp; haddadcr @ ufs. ac. za; https: // orcid. org / 0000 - 0002 - 2317 - 7760; &amp; booysenr @ ufs. ac. za; https: // orcid. org / 0000 - 0002 - 0841 - 9143</mods:affiliation>
<mods:nameIdentifier id="7CBE1E1F001AEAA7C3D9D3AEBBCAB222" type="email">haddadcr@ufs.ac.za</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="7643C81D1593CDBAEFE18C5958E31FFA">text</mods:typeOfResource>
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<mods:title id="E1711DCB8E4F564FE8CF3D6DFC39E581">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="CF4A9D06DEC7058CE42D5B691AFFDFC1">
<mods:date id="8FBF708F71912C46A13E14569A3A5446">2021</mods:date>
<mods:detail id="49F19694C4EB40A8CB59C9274E3B03A1" type="pubDate">
<mods:number id="DEBC09CF66A4E9277E6EAF60ED8B563C">2021-03-11</mods:number>
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<mods:number id="E43B6EED1F588D516512ACEB9576FECA">4942</mods:number>
</mods:detail>
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<mods:number id="828CDFA4C1853718A05889AF56A1F39B">1</mods:number>
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<mods:start id="29988BCB4B9B026FB4A030F987B7CEA1">41</mods:start>
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<mods:classification id="5C360B417A85715E85A04D4440A75127">journal article</mods:classification>
<mods:identifier id="C38470B74436FFC3069778AE424587D5" type="CLB-Dataset">7679</mods:identifier>
<mods:identifier id="6EB2E9022DDE163895033A41197FB0DB" type="DOI">10.11646/zootaxa.4942.1.2</mods:identifier>
<mods:identifier id="F7F7571EAF71C0076B23BE71C806C859" type="GBIF-Dataset">25d73144-c90c-45c3-a9a2-2454f87a9aef</mods:identifier>
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<mods:identifier id="37A4BD5BD8FDD7E98A142F7A46162DB3" type="Zenodo-Dep">4596097</mods:identifier>
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<treatment id="03D67155FFBDFFB5FF25FCF8A11C2E2C" ID-DOI="http://doi.org/10.5281/zenodo.4596111" ID-GBIF-Taxon="180588635" ID-Zenodo-Dep="4596111" LSID="urn:lsid:plazi:treatment:03D67155FFBDFFB5FF25FCF8A11C2E2C" httpUri="http://treatment.plazi.org/id/03D67155FFBDFFB5FF25FCF8A11C2E2C" lastPageId="19" lastPageNumber="60" pageId="18" pageNumber="59">
<subSubSection id="C36593C8FFBDFFB4FF25FCF8A20C2A45" box="[151,556,834,861]" pageId="18" pageNumber="59" type="nomenclature">
<paragraph id="8BC0C043FFBDFFB4FF25FCF8A20C2A45" blockId="18.[151,556,834,897]" box="[151,556,834,861]" pageId="18" pageNumber="59">
<heading id="D088772FFFBDFFB4FF25FCF8A20C2A45" bold="true" box="[151,556,834,861]" fontSize="11" level="1" pageId="18" pageNumber="59" reason="1">
<emphasis id="B90B1C51FFBDFFB4FF25FCF8A20C2A45" bold="true" box="[151,556,834,861]" pageId="18" pageNumber="59">
<taxonomicName id="4C7FBBC0FFBDFFB4FF25FCF8A18D2A45" ID-CoL="98Z25" authority="Haddad &amp; Jin &amp; Platnick &amp; Booysen, 2021" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" box="[151,429,834,861]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="59" phylum="Arthropoda" rank="species" species="ukhahlamba" status="sp. nov.">
<emphasis id="B90B1C51FFBDFFB4FF25FCF8A18D2A45" bold="true" box="[151,429,834,861]" italics="true" pageId="18" pageNumber="59">Capobula ukhahlamba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238A12AFFBDFFB4FE06FCF9A20C2A45" box="[436,556,835,861]" pageId="18" pageNumber="59" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36593C8FFBDFFB4FF25FCDDA11E2A99" box="[151,318,871,897]" pageId="18" pageNumber="59" type="description">
<paragraph id="8BC0C043FFBDFFB4FF25FCDDA11E2A99" blockId="18.[151,556,834,897]" box="[151,318,871,897]" pageId="18" pageNumber="59">
<figureCitation id="1344DCC6FFBDFFB4FF25FCDDA0CC2A99" box="[151,236,871,897]" captionStart="FIGURES 3–10" captionStartId="5.[151,264,1869,1893]" captionTargetBox="[151,1436,946,1846]" captionTargetId="figure-265@5.[151,1436,946,1846]" captionTargetPageId="5" captionText="FIGURES 3–10. Capobula gen. nov. dorsal habitus photographs: 3 C. capensis spec. nov., female holotype from Pakhuis Pass; 4 Same, paratype male from Pakhuis Pass; 5 C. infima (Simon, 1896) comb. nov., female from Fisherhaven; 6 Same, male from De Hoop Nature Reserve; 7 C. montana spec. nov., female holotype from Mount Coke State Forest; 8 Same, paratype male from Shamwari Game Reserve; 9 C. neethlingi spec. nov., female holotype from Saasveld Pass; 10 C. ukhahlamba spec. nov., paratype female from Royal Natal National Park." figureDoi="http://doi.org/10.5281/zenodo.4596103" httpUri="https://zenodo.org/record/4596103/files/figure.png" pageId="18" pageNumber="59">Figs 10</figureCitation>
,
<figureCitation id="1344DCC6FFBDFFB4FF4AFCDDA11E2A99" box="[248,318,871,897]" captionStart="FIGURES 71–74" captionStartId="19.[151,264,1279,1303]" captionTargetBox="[196,1395,184,1245]" captionTargetId="figure-18@19.[189,1398,181,1255]" captionTargetPageId="19" captionText="FIGURES 71–74. Capobula neethlingi spec. nov. (71, 72) and C. ukhahlamba spec. nov. (73, 74), genitalic morphology of paratype females from Anysberg Nature Reserve and Cathedral Peak, respectively: 71, 73 Epigyne, ventral; 72, 74 Vulva, dorsal. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4596125" httpUri="https://zenodo.org/record/4596125/files/figure.png" pageId="18" pageNumber="59">73, 74</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36593C8FFBDFFB4FF25FC15A31E2C4D" pageId="18" pageNumber="59" type="materials_examined">
<paragraph id="8BC0C043FFBDFFB4FF25FC15A31E2C4D" blockId="18.[151,1437,942,1833]" pageId="18" pageNumber="59">
<emphasis id="B90B1C51FFBDFFB4FF25FC15A1722AD0" bold="true" box="[151,338,942,969]" pageId="18" pageNumber="59">Type material:</emphasis>
<materialsCitation id="3B17CA1EFFBDFFB4FED1FC15A1802D09" ID-GBIF-Occurrence="3053986347" collectingDate="2011-01-20" collectionCode="NCA" collectorName="C. Haddad" country="South Africa" elevation="1560" latitude="-29.039433" location="Hlathikulu Forest" longLatPrecision="1" longitude="29.39035" municipality="Champagne Castle" pageId="18" pageNumber="59" specimenCode="NCA 2018" specimenCount="1" specimenCount-female="1" stateProvince="KwaZulu-Natal" typeStatus="holotype">
<emphasis id="B90B1C51FFBDFFB4FED1FC15A1F02AD1" bold="true" box="[355,464,943,969]" pageId="18" pageNumber="59">
<typeStatus id="54C47EE1FFBDFFB4FED1FC15A1F02AD1" box="[355,464,943,969]" pageId="18" pageNumber="59" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D790BCAFFBDFFB4FE53FC15A1D82AD1" box="[481,504,943,969]" pageId="18" pageNumber="59" type="female">♀</specimenCount>
:
<emphasis id="B90B1C51FFBDFFB4FDBDFC14A2D82AD0" bold="true" box="[527,760,942,968]" pageId="18" pageNumber="59">
<collectingCountry id="F36880D3FFBDFFB4FDBDFC14A2D22AD0" box="[527,754,942,968]" name="South Africa" pageId="18" pageNumber="59">SOUTH AFRICA</collectingCountry>
:
</emphasis>
<emphasis id="B90B1C51FFBDFFB4FCBBFC15A3972AD1" box="[777,951,943,969]" italics="true" pageId="18" pageNumber="59">
<collectingRegion id="49BB0EA1FFBDFFB4FCBBFC15A3972AD1" box="[777,951,943,969]" country="South Africa" name="KwaZulu-Natal" pageId="18" pageNumber="59">KwaZulu-Natal</collectingRegion>
</emphasis>
:
<collectingMunicipality id="6BA45A39FFBDFFB4FC62FC15A4912AD1" box="[976,1201,943,969]" pageId="18" pageNumber="59">Champagne Castle</collectingMunicipality>
,
<location id="8EA09698FFBDFFB4FB74FC15A5B82AD1" LSID="urn:lsid:plazi:treatment:03D67155FFBDFFB5FF25FCF8A11C2E2C:8EA09698FFBDFFB4FB74FC15A5B82AD1" box="[1222,1432,943,969]" country="South Africa" latitude="-29.039433" longLatPrecision="1" longitude="29.39035" municipality="Champagne Castle" name="Hlathikulu Forest" pageId="18" pageNumber="59" stateProvince="KwaZulu-Natal">Hlathikulu Forest</location>
,
<geoCoordinate id="EE4BA684FFBDFFB4FF25FC69A1062AF5" box="[151,294,979,1005]" degrees="29" direction="south" minutes="02.366" orientation="latitude" pageId="18" pageNumber="59" precision="1" value="-29.039433">29°02.366’S</geoCoordinate>
,
<geoCoordinate id="EE4BA684FFBDFFB4FE86FC69A1E62AF5" box="[308,454,979,1005]" degrees="29" direction="east" minutes="23.421" orientation="longitude" pageId="18" pageNumber="59" precision="1" value="29.39035">29°23.421’E</geoCoordinate>
,
<elevation id="00522770FFBDFFB4FE61FC68A2492AF5" box="[467,617,978,1005]" metricMagnitude="3" metricUnit="m" metricValue="1.56" pageId="18" pageNumber="59" unit="m" value="1560.0">
<quantity id="4C876DA6FFBDFFB4FE61FC68A20A2AF5" box="[467,554,978,1005]" metricMagnitude="3" metricUnit="m" metricValue="1.56" pageId="18" pageNumber="59" unit="m" value="1560.0">1560 m</quantity>
a.s.l.
</elevation>
,
<date id="FFC1E683FFBDFFB4FDCAFC69A2C42AF5" box="[632,740,979,1005]" pageId="18" pageNumber="59" value="2011-01-20">
<collectingDate id="EF851F6BFFBDFFB4FDCAFC69A2C42AF5" box="[632,740,979,1005]" pageId="18" pageNumber="59" value="2011-01-20">20.I.2011</collectingDate>
</date>
, leg.
<collectorName id="268AA595FFBDFFB4FC96FC69A3BE2AF5" box="[804,926,979,1005]" pageId="18" pageNumber="59">C. Haddad</collectorName>
(base of grass tussocks, grassland-forest ecotone) (
<specimenCode id="DBD96838FFBDFFB4FF43FC4DA1532D09" box="[241,371,1014,1041]" collectionCode="NCA" pageId="18" pageNumber="59">NCA 2018</specimenCode>
/35)
</materialsCitation>
.
<materialsCitation id="3B17CA1EFFBDFFB4FE1CFC4DA1232D41" ID-GBIF-Occurrence="3053986326" collectingDate="1979-12-14" collectionCode="AMNH" collectorName="S. &amp; J. Peck" country="South Africa" elevation="1380" location="Cathedral Peak Forest Station" pageId="18" pageNumber="59" specimenCount="1" specimenCount-female="1" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<emphasis id="B90B1C51FFBDFFB4FE1CFC4DA2062D09" bold="true" box="[430,550,1015,1041]" pageId="18" pageNumber="59">
<typeStatus id="54C47EE1FFBDFFB4FE1CFC4DA2062D09" box="[430,550,1015,1041]" pageId="18" pageNumber="59" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<emphasis id="B90B1C51FFBDFFB4FD8BFC4CA33A2D08" bold="true" box="[569,794,1014,1040]" pageId="18" pageNumber="59">
<collectingCountry id="F36880D3FFBDFFB4FD8BFC4CA3352D08" box="[569,789,1014,1040]" name="South Africa" pageId="18" pageNumber="59">SOUTH AFRICA</collectingCountry>
:
</emphasis>
<emphasis id="B90B1C51FFBDFFB4FC97FC4DA3F32D09" box="[805,979,1015,1041]" italics="true" pageId="18" pageNumber="59">
<collectingRegion id="49BB0EA1FFBDFFB4FC97FC4DA3F32D09" box="[805,979,1015,1041]" country="South Africa" name="KwaZulu-Natal" pageId="18" pageNumber="59">KwaZulu-Natal</collectingRegion>
</emphasis>
:
<location id="8EA09698FFBDFFB4FC54FC4DA5622D09" LSID="urn:lsid:plazi:treatment:03D67155FFBDFFB5FF25FCF8A11C2E2C:8EA09698FFBDFFB4FC54FC4DA5622D09" box="[998,1346,1015,1041]" country="South Africa" name="Cathedral Peak Forest Station" pageId="18" pageNumber="59" stateProvince="KwaZulu-Natal">Cathedral Peak Forest Station</location>
,
<locationDeviation id="9A72D961FFBDFFB4FAE0FC4CA1622D2D" location="Estcourt" pageId="18" pageNumber="59">
<quantity id="4C876DA6FFBDFFB4FAE0FC4CA5BC2D09" box="[1362,1436,1014,1041]" metricMagnitude="4" metricUnit="m" metricValue="7.5" pageId="18" pageNumber="59" unit="km" value="75.0">75 km</quantity>
WSW Estcourt
</locationDeviation>
,
<elevation id="00522770FFBDFFB4FEFCFBA1A1FF2D2D" box="[334,479,1051,1077]" metricMagnitude="3" metricUnit="m" metricValue="1.38" pageId="18" pageNumber="59" unit="m" value="1380.0">
<quantity id="4C876DA6FFBDFFB4FEFCFBA1A1832D2D" box="[334,419,1051,1077]" metricMagnitude="3" metricUnit="m" metricValue="1.38" pageId="18" pageNumber="59" unit="m" value="1380.0">1380 m</quantity>
a.s.l.
</elevation>
,
<date id="FFC1E683FFBDFFB4FE5EFBA1A2572D2D" box="[492,631,1051,1077]" pageId="18" pageNumber="59" value="1979-12-14">
<collectingDate id="EF851F6BFFBDFFB4FE5EFBA1A2572D2D" box="[492,631,1051,1077]" pageId="18" pageNumber="59" value="1979-12-14">14.XII.1979</collectingDate>
</date>
, leg.
<collectorName id="268AA595FFBDFFB4FD01FBA1A31D2D2D" box="[691,829,1051,1077]" pageId="18" pageNumber="59">S. &amp; J. Peck</collectorName>
(
<taxonomicName id="4C7FBBC0FFBDFFB4FCFFFBA1A3ED2D2D" box="[845,973,1051,1077]" class="Magnoliopsida" family="Rosaceae" genus="Leucosidea" kingdom="Plantae" order="Rosales" pageId="18" pageNumber="59" phylum="Tracheophyta" rank="genus">
<emphasis id="B90B1C51FFBDFFB4FCFFFBA1A3ED2D2D" box="[845,973,1051,1077]" italics="true" pageId="18" pageNumber="59">Leucosidea</emphasis>
</taxonomicName>
boulder bush scrub, night sweeping),
<specimenCount id="9D790BCAFFBDFFB4FACBFBA1A5BC2D2D" box="[1401,1436,1051,1077]" pageId="18" pageNumber="59" type="female">1♀</specimenCount>
(
<collectionCode id="ED6E5886FFBDFFB4FF2DFB85A0DB2D41" box="[159,251,1087,1113]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="18" pageNumber="59">AMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B17CA1EFFBDFFB4FEBCFB85A2692DB9" ID-GBIF-Occurrence="3053986352" collectingDate="2011-01-19" collectionCode="CAS" collectorName="H. Wood &amp; C. Griswold" country="South Africa" county="Didima National Park" elevation="1518" latitude="-28.96" location="Rainbow Gorge" longLatPrecision="1" longitude="29.22725" municipality="Cathedral Peak area" pageId="18" pageNumber="59" specimenCode="CASENT 9043022, SA11-041" specimenCount="1" specimenCount-female="1" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<collectingCounty id="62A1B8CFFFBDFFB4FEBCFB85A2262D41" box="[270,518,1087,1113]" pageId="18" pageNumber="59">Didima National Park</collectingCounty>
,
<collectingMunicipality id="6BA45A39FFBDFFB4FDA0FB85A2D42D41" box="[530,756,1087,1113]" pageId="18" pageNumber="59">Cathedral Peak area</collectingMunicipality>
,
<location id="8EA09698FFBDFFB4FD4DFB85A3942D41" LSID="urn:lsid:plazi:treatment:03D67155FFBDFFB5FF25FCF8A11C2E2C:8EA09698FFBDFFB4FD4DFB85A3942D41" box="[767,948,1087,1113]" country="South Africa" county="Didima National Park" latitude="-28.96" longLatPrecision="1" longitude="29.22725" municipality="Cathedral Peak area" name="Rainbow Gorge" pageId="18" pageNumber="59" stateProvince="KwaZulu-Natal">Rainbow Gorge</location>
,
<locationDeviation id="9A72D961FFBDFFB4FC0DFB84A4DD2D41" box="[959,1277,1086,1113]" location="Winterton" pageId="18" pageNumber="59">
<quantity id="4C876DA6FFBDFFB4FC0DFB84A43B2D41" box="[959,1051,1086,1113]" metricMagnitude="4" metricUnit="m" metricValue="3.35" pageId="18" pageNumber="59" unit="km" value="33.5">33.5 km</quantity>
WSW of Winterton
</locationDeviation>
,
<geoCoordinate id="EE4BA684FFBDFFB4FABAFB84A5B72D41" box="[1288,1431,1086,1113]" degrees="28" direction="south" minutes="57.600" orientation="latitude" pageId="18" pageNumber="59" precision="1" value="-28.96">28°57.600’S</geoCoordinate>
,
<geoCoordinate id="EE4BA684FFBDFFB4FF25FBD8A1092D65" box="[151,297,1122,1149]" degrees="29" direction="east" minutes="13.635" orientation="longitude" pageId="18" pageNumber="59" precision="1" value="29.22725">29°13.635’E</geoCoordinate>
,
<elevation id="00522770FFBDFFB4FE85FBD8A1ED2D65" box="[311,461,1122,1149]" metricMagnitude="3" metricUnit="m" metricValue="1.518" pageId="18" pageNumber="59" unit="m" value="1518.0">
<quantity id="4C876DA6FFBDFFB4FE85FBD8A1AE2D65" box="[311,398,1122,1149]" metricMagnitude="3" metricUnit="m" metricValue="1.518" pageId="18" pageNumber="59" unit="m" value="1518.0">1518 m</quantity>
a.s.l.
</elevation>
,
<date id="FFC1E683FFBDFFB4FE6EFBD9A2692D65" box="[476,585,1123,1149]" pageId="18" pageNumber="59" value="2011-01-19">
<collectingDate id="EF851F6BFFBDFFB4FE6EFBD9A2692D65" box="[476,585,1123,1149]" pageId="18" pageNumber="59" value="2011-01-19">19.I.2011</collectingDate>
</date>
, leg.
<collectorName id="268AA595FFBDFFB4FD38FBD9A2D12D65" box="[650,753,1123,1149]" pageId="18" pageNumber="59">H. Wood</collectorName>
&amp;
<collectorName id="268AA595FFBDFFB4FCA8FBD9A3852D65" box="[794,933,1123,1149]" pageId="18" pageNumber="59">C. Griswold</collectorName>
(sifting leaf litter,
<location id="8EA09698FFBDFFB4FB32FBD9A5432D66" LSID="urn:lsid:plazi:treatment:03D67155FFBDFFB5FF25FCF8A11C2E2C:8EA09698FFBDFFB4FB32FBD9A5432D66" box="[1152,1379,1123,1150]" country="South Africa" county="Didima National Park" latitude="-28.96" longLatPrecision="1" longitude="29.22725" municipality="Cathedral Peak area" name="Afromontane forest" pageId="18" pageNumber="59" stateProvince="KwaZulu-Natal">Afromontane forest</location>
),
<specimenCount id="9D790BCAFFBDFFB4FACBFBD9A5BC2D65" box="[1401,1436,1123,1149]" pageId="18" pageNumber="59" type="female">1♀</specimenCount>
(
<collectionCode id="ED6E5886FFBDFFB4FF2DFB3DA0F92DB9" box="[159,217,1159,1185]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="18" pageNumber="59" type="Museum">CAS</collectionCode>
,
<specimenCode id="DBD96838FFBDFFB4FF54FB3DA19F2DB9" box="[230,447,1159,1185]" collectionCode="CASENT" pageId="18" pageNumber="59">CASENT 9043022</specimenCode>
,
<specimenCode id="DBD96838FFBDFFB4FE79FB3DA2622DB9" box="[459,578,1159,1185]" collectionCode="SA" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="18" pageNumber="59">SA11-041</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B17CA1EFFBDFFB4FDE4FB3DA2992DDD" ID-GBIF-Occurrence="3053986338" collectingDate="2011-01-19" collectionCode="MHBU" collectorName="C. Haddad" country="South Africa" elevation="1400" latitude="-28.9497" location="Same" longLatPrecision="1" longitude="29.231234" pageId="18" pageNumber="59" specimenCount="1" specimenCount-female="1" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<location id="8EA09698FFBDFFB4FDE4FB3DA2B42DB9" LSID="urn:lsid:plazi:treatment:03D67155FFBDFFB5FF25FCF8A11C2E2C:8EA09698FFBDFFB4FDE4FB3DA2B42DB9" box="[598,660,1159,1185]" country="South Africa" latitude="-28.9497" longLatPrecision="1" longitude="29.231234" name="Same" pageId="18" pageNumber="59" stateProvince="KwaZulu-Natal">Same</location>
locality,
<geoCoordinate id="EE4BA684FFBDFFB4FD4CFB3CA3AD2DB9" box="[766,909,1158,1185]" degrees="28" direction="south" minutes="56.982" orientation="latitude" pageId="18" pageNumber="59" precision="1" value="-28.9497">28°56.982’S</geoCoordinate>
,
<geoCoordinate id="EE4BA684FFBDFFB4FC28FB3DA40B2DB9" box="[922,1067,1159,1185]" degrees="29" direction="east" minutes="13.874" orientation="longitude" pageId="18" pageNumber="59" precision="1" value="29.231234">29°13.874’E</geoCoordinate>
,
<elevation id="00522770FFBDFFB4FB85FB3DA4EB2DB9" box="[1079,1227,1159,1185]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="18" pageNumber="59" unit="m" value="1400.0">
<quantity id="4C876DA6FFBDFFB4FB85FB3DA4AD2DB9" box="[1079,1165,1159,1185]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="18" pageNumber="59" unit="m" value="1400.0">1400 m</quantity>
a.s.l.
</elevation>
,
<date id="FFC1E683FFBDFFB4FB6AFB3DA5652DB9" box="[1240,1349,1159,1185]" pageId="18" pageNumber="59" value="2011-01-19">
<collectingDate id="EF851F6BFFBDFFB4FB6AFB3DA5652DB9" box="[1240,1349,1159,1185]" pageId="18" pageNumber="59" value="2011-01-19">19.I.2011</collectingDate>
</date>
, leg.
<collectorName id="268AA595FFBDFFB4FA30FB3DA0CE2DDD" pageId="18" pageNumber="59">C. Haddad</collectorName>
(base of grasses and ferns),
<specimenCount id="9D790BCAFFBDFFB4FD94FB11A2692DDD" box="[550,585,1195,1221]" pageId="18" pageNumber="59" type="female">1♀</specimenCount>
(
<collectionCode id="ED6E5886FFBDFFB4FDE5FB11A2912DDD" box="[599,689,1195,1221]" pageId="18" pageNumber="59">MHBU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B17CA1EFFBDFFB4FD76FB11A2C82C16" ID-GBIF-Occurrence="3053986323" collectingDate="2011-01-21" collectionCode="CAS" collectorName="H. Wood &amp; C. Haddad &amp; C. Griswold" country="South Africa" elevation="1403" latitude="-28.685617" location="Royal Natal National Park" longLatPrecision="1" longitude="28.957083" pageId="18" pageNumber="59" specimenCode="CASENT 9043423, SA11-046" specimenCount="1" specimenCount-female="1" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<location id="8EA09698FFBDFFB4FD76FB11A3CC2DDD" LSID="urn:lsid:plazi:treatment:03D67155FFBDFFB5FF25FCF8A11C2E2C:8EA09698FFBDFFB4FD76FB11A3CC2DDD" box="[708,1004,1195,1221]" country="South Africa" latitude="-28.685617" longLatPrecision="1" longitude="28.957083" name="Royal Natal National Park" pageId="18" pageNumber="59" stateProvince="KwaZulu-Natal">Royal Natal National Park</location>
,
<locationDeviation id="9A72D961FFBDFFB4FC44FB11A4DE2DDD" box="[1014,1278,1195,1221]" location="Bergville" pageId="18" pageNumber="59">
<quantity id="4C876DA6FFBDFFB4FC44FB11A4712DDD" box="[1014,1105,1195,1221]" metricMagnitude="4" metricUnit="m" metricValue="3.86" pageId="18" pageNumber="59" unit="km" value="38.6">38.6 km</quantity>
W of Bergville
</locationDeviation>
,
<geoCoordinate id="EE4BA684FFBDFFB4FABAFB11A5B72DDD" box="[1288,1431,1195,1221]" degrees="28" direction="south" minutes="41.137" orientation="latitude" pageId="18" pageNumber="59" precision="1" value="-28.685617">28°41.137’S</geoCoordinate>
,
<geoCoordinate id="EE4BA684FFBDFFB4FF25FB74A1092DF1" box="[151,297,1230,1257]" degrees="28" direction="east" minutes="57.425" orientation="longitude" pageId="18" pageNumber="59" precision="1" value="28.957083">28°57.425’E</geoCoordinate>
,
<elevation id="00522770FFBDFFB4FE80FB75A1E12DF1" box="[306,449,1231,1257]" metricMagnitude="3" metricUnit="m" metricValue="1.403" pageId="18" pageNumber="59" unit="m" value="1403.0">
<quantity id="4C876DA6FFBDFFB4FE80FB75A1A62DF1" box="[306,390,1231,1257]" metricMagnitude="3" metricUnit="m" metricValue="1.403" pageId="18" pageNumber="59" unit="m" value="1403.0">1403 m</quantity>
a.s.l.
</elevation>
,
<date id="FFC1E683FFBDFFB4FE7EFB75A2192DF1" box="[460,569,1231,1257]" pageId="18" pageNumber="59" value="2011-01-21">
<collectingDate id="EF851F6BFFBDFFB4FE7EFB75A2192DF1" box="[460,569,1231,1257]" pageId="18" pageNumber="59" value="2011-01-21">21.I.2011</collectingDate>
</date>
, leg.
<collectorName id="268AA595FFBDFFB4FDC0FB75A2F82DF1" box="[626,728,1231,1257]" pageId="18" pageNumber="59">H. Wood</collectorName>
,
<collectorName id="268AA595FFBDFFB4FD53FB75A3782DF1" box="[737,856,1231,1257]" pageId="18" pageNumber="59">C. Haddad</collectorName>
&amp;
<collectorName id="268AA595FFBDFFB4FCCBFB75A4202DF1" box="[889,1024,1231,1257]" pageId="18" pageNumber="59">C. Griswold</collectorName>
(general collecting in ferns and grass tussocks),
<specimenCount id="9D790BCAFFBDFFB4FEBCFB49A1102C15" box="[270,304,1267,1293]" pageId="18" pageNumber="59" type="female">1♀</specimenCount>
(
<collectionCode id="ED6E5886FFBDFFB4FEF2FB49A15A2C15" box="[320,378,1267,1293]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="18" pageNumber="59" type="Museum">CAS</collectionCode>
,
<specimenCode id="DBD96838FFBDFFB4FE34FB49A27E2C15" box="[390,606,1267,1293]" collectionCode="CASENT" pageId="18" pageNumber="59">CASENT 9043423</specimenCode>
,
<specimenCode id="DBD96838FFBDFFB4FDD8FB49A2C02C16" box="[618,736,1267,1294]" collectionCode="SA" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="18" pageNumber="59">SA11-046</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B17CA1EFFBDFFB4FD46FB49A13C2C29" ID-GBIF-Occurrence="3053986343" collectingDate="2006-09" collectionCode="NCA" collectorName="D. Prentice" country="South Africa" location="Sani Pass" pageId="18" pageNumber="59" specimenCode="NCA 2008" specimenCount="1" specimenCount-female="1" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<location id="8EA09698FFBDFFB4FD46FB49A3412C15" LSID="urn:lsid:plazi:treatment:03D67155FFBDFFB5FF25FCF8A11C2E2C:8EA09698FFBDFFB4FD46FB49A3412C15" box="[756,865,1267,1293]" country="South Africa" name="Sani Pass" pageId="18" pageNumber="59" stateProvince="KwaZulu-Natal">Sani Pass</location>
,
<date id="FFC1E683FFBDFFB4FCDFFB49A3EC2C15" box="[877,972,1267,1293]" pageId="18" pageNumber="59" value="2006-09">
<collectingDate id="EF851F6BFFBDFFB4FCDFFB49A3EC2C15" box="[877,972,1267,1293]" pageId="18" pageNumber="59" value="2006-09">IX.2006</collectingDate>
</date>
, leg.
<collectorName id="268AA595FFBDFFB4FBBAFB49A4A72C15" box="[1032,1159,1267,1293]" pageId="18" pageNumber="59">D. Prentice</collectorName>
(
<collectingMethod id="523EB854FFBDFFB4FB27FB49A53B2C15" box="[1173,1307,1267,1293]" pageId="18" pageNumber="59">pitfall traps</collectingMethod>
),
<specimenCount id="9D790BCAFFBDFFB4FA9DFB49A5722C15" box="[1327,1362,1267,1293]" pageId="18" pageNumber="59" type="female">1♀</specimenCount>
(
<specimenCode id="DBD96838FFBDFFB4FAD3FB49A0F22C29" collectionCode="NCA" pageId="18" pageNumber="59">NCA 2008</specimenCode>
/1982)
</materialsCitation>
;
<materialsCitation id="3B17CA1EFFBDFFB4FE9BFAADA31B2C4D" ID-GBIF-Occurrence="3053986327" collectingDate="2009-01-01" collectionCode="NCA" country="South Africa" elevation="900" latitude="-30.1835" location="Ixopo" longLatPrecision="1" longitude="30.152166" municipality="Sani Pass" pageId="18" pageNumber="59" specimenCode="NCA 2011" specimenCount="1" specimenCount-female="1" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<collectingMunicipality id="6BA45A39FFBDFFB4FE9BFAADA1B52C29" box="[297,405,1303,1329]" pageId="18" pageNumber="59">Sani Pass</collectingMunicipality>
elevational project,
<location id="8EA09698FFBDFFB4FDCDFAADA2E32C29" LSID="urn:lsid:plazi:treatment:03D67155FFBDFFB5FF25FCF8A11C2E2C:8EA09698FFBDFFB4FDCDFAADA2E32C29" box="[639,707,1303,1329]" country="South Africa" latitude="-30.1835" longLatPrecision="1" longitude="30.152166" municipality="Sani Pass" name="Ixopo" pageId="18" pageNumber="59" stateProvince="KwaZulu-Natal">Ixopo</location>
,
<geoCoordinate id="EE4BA684FFBDFFB4FD62FAADA37E2C29" box="[720,862,1303,1329]" degrees="30" direction="south" minutes="11.010" orientation="latitude" pageId="18" pageNumber="59" precision="1" value="-30.1835">30°11.010’S</geoCoordinate>
,
<geoCoordinate id="EE4BA684FFBDFFB4FCDEFAADA3DD2C29" box="[876,1021,1303,1329]" degrees="30" direction="east" minutes="09.130" orientation="longitude" pageId="18" pageNumber="59" precision="1" value="30.152166">30°09.130’E</geoCoordinate>
,
<elevation id="00522770FFBDFFB4FBB9FAADA4B22C29" box="[1035,1170,1303,1329]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="18" pageNumber="59" unit="m" value="900.0">
<quantity id="4C876DA6FFBDFFB4FBB9FAADA4732C29" box="[1035,1107,1303,1329]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="18" pageNumber="59" unit="m" value="900.0">900 m</quantity>
a.s.l.
</elevation>
,
<date id="FFC1E683FFBDFFB4FB12FAADA4DF2C29" box="[1184,1279,1303,1329]" pageId="18" pageNumber="59" value="2009-01-01">
<collectingDate id="EF851F6BFFBDFFB4FB12FAADA4DF2C29" box="[1184,1279,1303,1329]" pageId="18" pageNumber="59" value="2009-01-01">1.I.2009</collectingDate>
</date>
, leg. Univ. of Pretoria students (
<collectingMethod id="523EB854FFBDFFB4FED3FA81A1C62C4D" box="[353,486,1339,1365]" pageId="18" pageNumber="59">pitfall traps</collectingMethod>
, site 8a),
<specimenCount id="9D790BCAFFBDFFB4FDE3FA81A2542C4D" box="[593,628,1339,1365]" pageId="18" pageNumber="59" type="female">1♀</specimenCount>
(
<specimenCode id="DBD96838FFBDFFB4FD30FA81A2DE2C4D" box="[642,766,1339,1365]" collectionCode="NCA" pageId="18" pageNumber="59">NCA 2011</specimenCode>
/774)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36593C8FFBDFFB4FF75FAE4A5742CD9" pageId="18" pageNumber="59" type="diagnosis">
<paragraph id="8BC0C043FFBDFFB4FF75FAE4A5742CD9" blockId="18.[151,1437,942,1833]" pageId="18" pageNumber="59">
<emphasis id="B90B1C51FFBDFFB4FF75FAE4A1622C60" bold="true" box="[199,322,1374,1400]" pageId="18" pageNumber="59">Diagnosis.</emphasis>
Females can be easily distinguished by the large circular ridges surrounding the copulatory openings (
<figureCitation id="1344DCC6FFBDFFB4FF12FA39A0D12C85" box="[160,241,1411,1437]" captionStart="FIGURES 71–74" captionStartId="19.[151,264,1279,1303]" captionTargetBox="[196,1395,184,1245]" captionTargetId="figure-18@19.[189,1398,181,1255]" captionTargetPageId="19" captionText="FIGURES 71–74. Capobula neethlingi spec. nov. (71, 72) and C. ukhahlamba spec. nov. (73, 74), genitalic morphology of paratype females from Anysberg Nature Reserve and Cathedral Peak, respectively: 71, 73 Epigyne, ventral; 72, 74 Vulva, dorsal. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4596125" httpUri="https://zenodo.org/record/4596125/files/figure.png" pageId="18" pageNumber="59">Fig. 73</figureCitation>
), the oval rather than teardrop-shaped primary spermathecae (
<figureCitation id="1344DCC6FFBDFFB4FC29FA39A3CD2C85" box="[923,1005,1411,1437]" captionStart="FIGURES 71–74" captionStartId="19.[151,264,1279,1303]" captionTargetBox="[196,1395,184,1245]" captionTargetId="figure-18@19.[189,1398,181,1255]" captionTargetPageId="19" captionText="FIGURES 71–74. Capobula neethlingi spec. nov. (71, 72) and C. ukhahlamba spec. nov. (73, 74), genitalic morphology of paratype females from Anysberg Nature Reserve and Cathedral Peak, respectively: 71, 73 Epigyne, ventral; 72, 74 Vulva, dorsal. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4596125" httpUri="https://zenodo.org/record/4596125/files/figure.png" pageId="18" pageNumber="59">Fig. 74</figureCitation>
), and the presence of a cream Y-shaped marking on the abdominal dorsum (
<figureCitation id="1344DCC6FFBDFFB4FD9AFA1DA25B2CD9" box="[552,635,1447,1473]" captionStart="FIGURES 3–10" captionStartId="5.[151,264,1869,1893]" captionTargetBox="[151,1436,946,1846]" captionTargetId="figure-265@5.[151,1436,946,1846]" captionTargetPageId="5" captionText="FIGURES 3–10. Capobula gen. nov. dorsal habitus photographs: 3 C. capensis spec. nov., female holotype from Pakhuis Pass; 4 Same, paratype male from Pakhuis Pass; 5 C. infima (Simon, 1896) comb. nov., female from Fisherhaven; 6 Same, male from De Hoop Nature Reserve; 7 C. montana spec. nov., female holotype from Mount Coke State Forest; 8 Same, paratype male from Shamwari Game Reserve; 9 C. neethlingi spec. nov., female holotype from Saasveld Pass; 10 C. ukhahlamba spec. nov., paratype female from Royal Natal National Park." figureDoi="http://doi.org/10.5281/zenodo.4596103" httpUri="https://zenodo.org/record/4596103/files/figure.png" pageId="18" pageNumber="59">Fig. 10</figureCitation>
), which is absent in the other species (
<figureCitation id="1344DCC6FFBDFFB4FB98FA1DA4B02CD9" box="[1066,1168,1447,1473]" captionStart="FIGURES 3–10" captionStartId="5.[151,264,1869,1893]" captionTargetBox="[151,1436,946,1846]" captionTargetId="figure-265@5.[151,1436,946,1846]" captionTargetPageId="5" captionText="FIGURES 3–10. Capobula gen. nov. dorsal habitus photographs: 3 C. capensis spec. nov., female holotype from Pakhuis Pass; 4 Same, paratype male from Pakhuis Pass; 5 C. infima (Simon, 1896) comb. nov., female from Fisherhaven; 6 Same, male from De Hoop Nature Reserve; 7 C. montana spec. nov., female holotype from Mount Coke State Forest; 8 Same, paratype male from Shamwari Game Reserve; 9 C. neethlingi spec. nov., female holotype from Saasveld Pass; 10 C. ukhahlamba spec. nov., paratype female from Royal Natal National Park." figureDoi="http://doi.org/10.5281/zenodo.4596103" httpUri="https://zenodo.org/record/4596103/files/figure.png" pageId="18" pageNumber="59">Figs 3–9</figureCitation>
). Male unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36593C8FFBDFFB4FF75FA71A5412F35" pageId="18" pageNumber="59" type="discussion">
<paragraph id="8BC0C043FFBDFFB4FF75FA71A5412F35" blockId="18.[151,1437,942,1833]" pageId="18" pageNumber="59">
<emphasis id="B90B1C51FFBDFFB4FF75FA71A11B2CFD" bold="true" box="[199,315,1483,1509]" pageId="18" pageNumber="59">Remarks.</emphasis>
Although the male of this species is unknown and the female has an inverted Y-shaped cream marking that it absent in congeners, we place this species in
<taxonomicName id="4C7FBBC0FFBDFFB4FCB1FA55A3512F11" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" box="[771,881,1519,1545]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B90B1C51FFBDFFB4FCB1FA55A3512F11" box="[771,881,1519,1545]" italics="true" pageId="18" pageNumber="59">Capobula</emphasis>
</taxonomicName>
and not
<taxonomicName id="4C7FBBC0FFBDFFB4FC61FA55A4682F11" authorityName="Simon" authorityYear="1897" box="[979,1096,1519,1545]" class="Arachnida" family="Corinnidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B90B1C51FFBDFFB4FC61FA55A4682F11" box="[979,1096,1519,1545]" italics="true" pageId="18" pageNumber="59">Orthobula</emphasis>
</taxonomicName>
because of the absence of pits along the dorsal midline of the carapace and the anterolaterally-positioned spermathecae; name in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C36593C8FFBDFFB5FF75F98DA5272FF3" lastPageId="19" lastPageNumber="60" pageId="18" pageNumber="59" type="etymology">
<paragraph id="8BC0C043FFBDFFB4FF75F98DA3902F81" blockId="18.[151,1437,942,1833]" pageId="18" pageNumber="59">
<emphasis id="B90B1C51FFBDFFB4FF75F98DA16E2F49" bold="true" box="[199,334,1591,1617]" pageId="18" pageNumber="59">Etymology.</emphasis>
Taken from the name of the Ukhahlamba-Drakensberg Transfrontier Conservation Park, a conservation initiative between
<collectingCountry id="F36880D3FFBDFFB4FE1DF9E1A21C2F6D" box="[431,572,1627,1653]" name="South Africa" pageId="18" pageNumber="59">South Africa</collectingCountry>
and
<collectingCountry id="F36880D3FFBDFFB4FDC2F9E1A2ED2F6D" box="[624,717,1627,1653]" name="Lesotho" pageId="18" pageNumber="59">Lesotho</collectingCountry>
, which includes the distribution range of this species. Ukhahlamba is the isiZulu word meaning “barrier of spears”; noun in apposition.
</paragraph>
<paragraph id="8BC0C043FFBDFFB4FF75F919A1F32E31" blockId="18.[151,1437,942,1833]" pageId="18" pageNumber="59">
<emphasis id="B90B1C51FFBDFFB4FF75F919A26F2FA4" bold="true" box="[199,591,1698,1725]" pageId="18" pageNumber="59">
Female (
<typeStatus id="54C47EE1FFBDFFB4FE9EF918A1B62FA4" box="[300,406,1698,1724]" pageId="18" pageNumber="59" type="holotype">holotype</typeStatus>
, NCA 2018/35).
</emphasis>
Measurements: CL 0.89, CW 0.67, AL 1.06, AW 0.89, TL 2.15 (2.03–2.15), PERW 0.39, MOQAW 0.12, MOQPW 0.16, MOQL 0.15. Length of leg segments: I 0.54 + 0.22 + 0.48 + 0.41 + 0.27 = 1.92; II 0.48 + 0.22 + 0.37 + 0.37 + 0.25 = 1.69; III 0.41 + 0.19 + 0.33 + 0.33 + 0.24 = 1.50; IV 0.57 + 0.24 + 0.48 + 0.54 + 0.30 = 2.13.
</paragraph>
<caption id="DF0090CBFFBCFFB5FF25FB45A1B72C47" ID-DOI="http://doi.org/10.5281/zenodo.4596125" ID-Zenodo-Dep="4596125" httpUri="https://zenodo.org/record/4596125/files/figure.png" pageId="19" pageNumber="60" startId="19.[151,264,1279,1303]" targetBox="[196,1395,184,1245]" targetPageId="19">
<paragraph id="8BC0C043FFBCFFB5FF25FB45A1B72C47" blockId="19.[151,1436,1279,1375]" pageId="19" pageNumber="60">
<emphasis id="B90B1C51FFBCFFB5FF25FB45A1762C0F" bold="true" box="[151,342,1279,1303]" pageId="19" pageNumber="60">FIGURES 71–74.</emphasis>
<taxonomicName id="4C7FBBC0FFBCFFB5FEEFFB45A20E2C0F" authority="Haddad &amp; Jin &amp; Platnick &amp; Booysen, 2021" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" box="[349,558,1279,1303]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="60" phylum="Arthropoda" rank="species" species="neethlingi" status="sp. nov.">
<emphasis id="B90B1C51FFBCFFB5FEEFFB45A20E2C0F" box="[349,558,1279,1303]" italics="true" pageId="19" pageNumber="60">Capobula neethlingi</emphasis>
</taxonomicName>
<emphasis id="B90B1C51FFBCFFB5FD85FB45A2BF2C0F" bold="true" box="[567,671,1279,1303]" pageId="19" pageNumber="60">
<taxonomicNameLabel id="A238A12AFFBCFFB5FD85FB45A2BF2C0F" box="[567,671,1279,1303]" pageId="19" pageNumber="60" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(71, 72) and
<taxonomicName id="4C7FBBC0FFBCFFB5FC9CFB45A3EA2C0F" authority="Haddad &amp; Jin &amp; Platnick &amp; Booysen, 2021" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" box="[814,970,1279,1303]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="60" phylum="Arthropoda" rank="species" species="ukhahlamba" status="sp. nov.">
<emphasis id="B90B1C51FFBCFFB5FC9CFB45A3EA2C0F" box="[814,970,1279,1303]" italics="true" pageId="19" pageNumber="60">C. ukhahlamba</emphasis>
</taxonomicName>
<emphasis id="B90B1C51FFBCFFB5FC61FB45A41B2C0F" bold="true" box="[979,1083,1279,1303]" pageId="19" pageNumber="60">
<taxonomicNameLabel id="A238A12AFFBCFFB5FC61FB45A41B2C0F" box="[979,1083,1279,1303]" pageId="19" pageNumber="60" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(73, 74), genitalic morphology of paratype females from Anysberg Nature Reserve and Cathedral Peak, respectively: 71, 73 Epigyne, ventral; 72, 74 Vulva, dorsal. Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BC0C043FFBCFFB5FF75FA37A5B92F0C" blockId="19.[151,1437,1421,1844]" pageId="19" pageNumber="60">Colour: carapace orange-brown, pits and lateral margins brown; chelicerae yellow-brown; endites and labium yellow-brown proximally, distal third cream; sternum yellow, pit margins and lateral margins yellow-brown; palps and legs creamy-yellow; abdomen dark grey dorsally and laterally, dorsally with diverging cream Y-shaped marking in anterior half, narrow cream chevrons in posterior half; venter cream, with faint grey mottling; spinnerets cream.</paragraph>
<paragraph id="8BC0C043FFBCFFB5FF75F9A4A2602F43" blockId="19.[151,1437,1421,1844]" pageId="19" pageNumber="60">Leg spination: femora and patellae: spineless; tibiae: I plv 6 rlv 5, II plv 5 rlv 4; metatarsi: I plv 4 rlv 4, II plv 4 rlv 4; tarsi: I plv 3 rlv 3, II plv 3 rlv 2.</paragraph>
<paragraph id="8BC0C043FFBCFFB5FF75F9DFA2422FDF" blockId="19.[151,1437,1421,1844]" pageId="19" pageNumber="60">
Epigyne with tiny copulatory openings in round epigynal ridges (
<figureCitation id="1344DCC6FFBCFFB5FC07F9DFA42A2F67" box="[949,1034,1637,1663]" captionStart="FIGURES 71–74" captionStartId="19.[151,264,1279,1303]" captionTargetBox="[196,1395,184,1245]" captionTargetId="figure-18@19.[189,1398,181,1255]" captionTargetPageId="19" captionText="FIGURES 71–74. Capobula neethlingi spec. nov. (71, 72) and C. ukhahlamba spec. nov. (73, 74), genitalic morphology of paratype females from Anysberg Nature Reserve and Cathedral Peak, respectively: 71, 73 Epigyne, ventral; 72, 74 Vulva, dorsal. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4596125" httpUri="https://zenodo.org/record/4596125/files/figure.png" pageId="19" pageNumber="60">Fig. 73</figureCitation>
); copulatory ducts initially curving medially, looping posteriorly, then laterally, before entering anterolateral primary spermathecae along mesal margin; bursae small and spherical (
<figureCitation id="1344DCC6FFBCFFB5FDB3F917A2762FDF" box="[513,598,1709,1735]" captionStart="FIGURES 71–74" captionStartId="19.[151,264,1279,1303]" captionTargetBox="[196,1395,184,1245]" captionTargetId="figure-18@19.[189,1398,181,1255]" captionTargetPageId="19" captionText="FIGURES 71–74. Capobula neethlingi spec. nov. (71, 72) and C. ukhahlamba spec. nov. (73, 74), genitalic morphology of paratype females from Anysberg Nature Reserve and Cathedral Peak, respectively: 71, 73 Epigyne, ventral; 72, 74 Vulva, dorsal. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4596125" httpUri="https://zenodo.org/record/4596125/files/figure.png" pageId="19" pageNumber="60">Fig. 74</figureCitation>
).
</paragraph>
<paragraph id="8BC0C043FFBCFFB5FF75F96BA5272FF3" blockId="19.[151,1437,1421,1844]" box="[199,1287,1745,1772]" pageId="19" pageNumber="60">
<emphasis id="B90B1C51FFBCFFB5FF75F96BA19B2FF3" bold="true" box="[199,443,1745,1771]" pageId="19" pageNumber="60">Habitat and biology.</emphasis>
Distributed in montane grasslands at elevations between
<quantity id="4C876DA6FFBCFFB5FB8DF96BA4EC2FF4" box="[1087,1228,1745,1772]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.6" metricValueMin="0.9" pageId="19" pageNumber="60" unit="m" value="1250.0" valueMax="1600.0" valueMin="900.0">900–1600 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="C36593C8FFBCFFB5FF75F94FA11C2E2C" pageId="19" pageNumber="60" type="distribution">
<paragraph id="8BC0C043FFBCFFB5FF75F94FA11C2E2C" blockId="19.[151,1437,1421,1844]" pageId="19" pageNumber="60">
<emphasis id="B90B1C51FFBCFFB5FF75F94FA1412E17" bold="true" box="[199,353,1781,1807]" pageId="19" pageNumber="60">Distribution.</emphasis>
Only known from
<collectingRegion id="49BB0EA1FFBCFFB5FD81F94FA2C62E17" box="[563,742,1781,1807]" country="South Africa" name="KwaZulu-Natal" pageId="19" pageNumber="60">KwaZulu-Natal</collectingRegion>
,
<collectingCountry id="F36880D3FFBCFFB5FD42F94FA3A02E17" box="[752,896,1781,1807]" name="South Africa" pageId="19" pageNumber="60">South Africa</collectingCountry>
, with most records from the Drakensberg Mountains (
<figureCitation id="1344DCC6FFBCFFB5FF69F8A3A1102E2C" box="[219,304,1817,1844]" captionStart="FIGURE 75" captionStartId="20.[151,250,1541,1565]" captionTargetBox="[189,1398,561,1517]" captionTargetId="figure-129@20.[189,1398,561,1517]" captionTargetPageId="20" captionText="FIGURE 75. Distribution of Capobula capensis spec. nov. (circles), C. infima (Simon, 1896) comb. nov. (triangles), C. montana spec. nov. (squares), C. neethlingi spec. nov. (stars) and C. ukhahlamba spec. nov. (inverted triangles) in southern Africa." figureDoi="http://doi.org/10.5281/zenodo.4596127" httpUri="https://zenodo.org/record/4596127/files/figure.png" pageId="19" pageNumber="60">Fig. 75</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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