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<mods:title id="81F7DA2073CA05E1F34665B3E576DBB6">A systematic revision of Draculoides (Schizomida: Hubbardiidae) of the Pilbara, Western Australia, Part I: the Western Pilbara</mods:title>
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<emphasis id="B913EADDFFB3FFE6CC8B023AFE6BFDED" bold="true" box="[151,395,582,609]" italics="true" pageId="53" pageNumber="54">Draculoides kryptus</emphasis>
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(
<figureCitation id="135C2A4AFFB3FFE6CC830217FEE9FD09" box="[159,265,619,645]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStartId-0="16.[151,250,1868,1892]" captionStartId-1="17.[151,250,1910,1934]" captionStartId-2="18.[151,250,1547,1571]" captionStartId-3="19.[151,250,1882,1906]" captionStartId-4="20.[151,250,1216,1240]" captionStartId-5="21.[151,250,1709,1733]" captionStartId-6="22.[151,250,1755,1779]" captionStartId-7="23.[151,250,1919,1943]" captionTargetBox-0="[346,1243,188,1844]" captionTargetBox-1="[332,1254,181,1881]" captionTargetBox-2="[158,1420,181,1521]" captionTargetBox-3="[151,1425,182,1858]" captionTargetBox-4="[158,1424,183,1189]" captionTargetBox-5="[155,1423,181,1683]" captionTargetBox-6="[159,1425,181,1729]" captionTargetBox-7="[151,1430,181,1893]" captionTargetId-0="figure-20@16.[339,1249,181,1844]" captionTargetId-1="figure-17@17.[332,1254,181,1886]" captionTargetId-2="figure-20@18.[151,1436,181,1524]" captionTargetId-3="figure-17@19.[151,1436,181,1859]" captionTargetId-4="figure-20@20.[151,1436,181,1192]" captionTargetId-5="figure-17@21.[155,1432,181,1685]" captionTargetId-6="figure-20@22.[151,1436,181,1732]" captionTargetId-7="figure-17@23.[151,1436,181,1894]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionTargetPageId-3="19" captionTargetPageId-4="20" captionTargetPageId-5="21" captionTargetPageId-6="22" captionTargetPageId-7="23" captionText-0="FIGURE 1. Map of an area of the Pilbara described as “West Pilbara” for the purposes of this study, showing the distribution of the named Draculoides species. Inset maps show species distributions in finer detail encompassing areas A: Warramboo, Mesas A, B and C and “BudgieBore”; B: Bungaroo, Mesas G, H, J, K and L; C: Cochrane and Jewell, Kens Bore, Cane and Upper Cane River, Trinity Bore and Catho Well, Mt. Stuart Station; D: Middle Robe, Robe Valley; E: Bungaroo South, West Pit, Buckland Hills and Dragon deposit." captionText-1="FIGURE 2. Maximum Likelihood tree, based on the reduced dataset. See Methods for details. Each terminal represents a single species or OTU, with named species in bold. Clades are identified using dashed boxes and shaded boxes encompass the species discussed in this study. Nodes with bootstrap support &lt;80 are not shown, except for the common ancestor of the Draculoides radiation." captionText-2="FIGURE 3. COI mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913085). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-3="FIGURE 4. Maximum Likelihood tree based on COI mini-barcodes (Fig. 3) showing similarity of mini-barcode sequences, relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." captionText-4="FIGURE 5. 12S mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913012). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-5="FIGURE 6. Maximum Likelihood tree based on COI mini-barcodes (Fig. 5) showing similarity of mini-barcode sequences relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." captionText-6="FIGURE 7. ITS2 mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913105). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-7="FIGURE 8. Maximum Likelihood tree based on ITS2 mini-barcodes (Fig. 7) showing similarity of mini-barcode sequences, relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." figureDoi-0="http://doi.org/10.5281/zenodo.4416851" figureDoi-1="http://doi.org/10.5281/zenodo.4416855" figureDoi-2="http://doi.org/10.5281/zenodo.4416857" figureDoi-3="http://doi.org/10.5281/zenodo.4416859" figureDoi-4="http://doi.org/10.5281/zenodo.4416863" figureDoi-5="http://doi.org/10.5281/zenodo.4416865" figureDoi-6="http://doi.org/10.5281/zenodo.4416867" figureDoi-7="http://doi.org/10.5281/zenodo.4416869" httpUri-0="https://zenodo.org/record/4416851/files/figure.png" httpUri-1="https://zenodo.org/record/4416855/files/figure.png" httpUri-2="https://zenodo.org/record/4416857/files/figure.png" httpUri-3="https://zenodo.org/record/4416859/files/figure.png" httpUri-4="https://zenodo.org/record/4416863/files/figure.png" httpUri-5="https://zenodo.org/record/4416865/files/figure.png" httpUri-6="https://zenodo.org/record/4416867/files/figure.png" httpUri-7="https://zenodo.org/record/4416869/files/figure.png" pageId="53" pageNumber="54">Figs. 18</figureCitation>
)
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http://zoobank.org/NomenclaturalActs/
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<bibRefCitation id="EFF64B3EFFB3FFE6CD8502A8FC2BFD67" author="Harvey, M. S. &amp; Berry, O. &amp; Edward, K. L. &amp; Humphreys, G." box="[409,971,723,748]" pageId="53" pageNumber="54" pagination="167 - 194" refId="ref46967" refString="Harvey, M. S., Berry, O., Edward, K. L. &amp; Humphreys, G. (2008) Molecular and morphological systematics of hypogean schizomids (Schizomida: Hubbardiidae) in semi-arid Australia. Invertebrate Systematics, 22 (2), 167 - 194. https: // doi. org / 10.1071 / IS 07026" type="journal article" year="2008">Harvey, Berry, Edward and Humphreys 2008: 190191</bibRefCitation>
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, figs. 10, 12, 4753.
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<taxonomicName id="4C674D4CFFB3FFE6CC8B028EFCF1FC86" authority="(Harvey, Berry, Edward and Humphreys)" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[151,785,754,778]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="kryptus">
<emphasis id="B913EADDFFB3FFE6CC8B028EFE86FC86" box="[151,358,754,778]" italics="true" pageId="53" pageNumber="54">Draculoides kryptus</emphasis>
(Harvey, Berry, Edward and Humphreys)
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:
<bibRefCitation id="EFF64B3EFFB3FFE6CF05028EFC3CFC86" author="Abrams, K. M. &amp; Huey, J. A. &amp; Hillyer, M. J. &amp; Humphreys, W. F. &amp; Didham, R. K. &amp; Harvey, M. S." box="[793,988,754,778]" pageId="53" pageNumber="54" pagination="106532" refId="ref46122" refString="Abrams, K. M., Huey, J. A., Hillyer, M. J., Humphreys, W. F., Didham, R. K. &amp; Harvey, M. S. (2019) Too hot to handle: Tertiary aridification drives multiple independent incursions of Schizomida (Hubbardiidae) into hypogean environments. Molecular Phylogenetics and Evolution, 139, 106532. https: // doi. org / 10.1016 / j. ympev. 2019.106532" type="journal article" year="2019">
Abrams
<emphasis id="B913EADDFFB3FFE6CF6C028FFC7CFC86" box="[880,924,754,778]" italics="true" pageId="53" pageNumber="54">et al</emphasis>
. 2019
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MPE 106532: 8, fig. 2.
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<emphasis id="B913EADDFFB3FFE6CC8B0344FE61FCDE" bold="true" box="[151,385,824,850]" pageId="53" pageNumber="54">Material examined.</emphasis>
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male.
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<collectorName id="26925319FFB3FFE6CFDD0345FBFAFCDF" box="[961,1050,825,851]" pageId="53" pageNumber="54">Mesa K</collectorName>
(Borehole 1685 trap 1), S of
<location id="8EB86014FFB3FFE6C9750345FEE5FCFA" LSID="urn:lsid:plazi:treatment:03CE87D9FFB3FFE6CC8B023AFE33F902:8EB86014FFB3FFE6C9750345FEE5FCFA" country="Australia" latitude="-21.718334" longLatPrecision="21" longitude="116.26" name="Pannawonica" pageId="53" pageNumber="54" stateProvince="Western Australia">Pannawonica</location>
,
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,
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,
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<collectingDate id="EF9DE9E7FFB3FFE6CE200320FD5EFCFB" box="[572,702,860,887]" pageId="53" pageNumber="54" value="2005-07-12">12.vii.2005</collectingDate>
</date>
,
<collectorName id="26925319FFB3FFE6CED60321FC89FCFA" box="[714,873,860,887]" pageId="53" pageNumber="54">M. Greenham</collectorName>
,
<collectorName id="26925319FFB3FFE6CF6F0321FC0EFCFA" box="[883,1006,860,887]" pageId="53" pageNumber="54">D. Kamien</collectorName>
and
<collectorName id="26925319FFB3FFE6C8380321FB6FFCFA" box="[1060,1167,860,887]" pageId="53" pageNumber="54">L. Mould</collectorName>
(
<collectionCode id="ED76AE0AFFB3FFE6C8800320FB03FCFA" box="[1180,1251,860,886]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="53" pageNumber="54" type="Museum">WAM</collectionCode>
<collectionCode id="ED76AE0AFFB3FFE6C8F50320FB1BFCFB" box="[1257,1275,860,887]" name="Tavera, Department of Geology and Geophysics" pageId="53" pageNumber="54">T</collectionCode>
65801).
</materialsCitation>
</paragraph>
<paragraph id="8BD836CFFFB3FFE6CCDB03FDFDB8FBC3" blockId="53.[151,1437,824,1679]" pageId="53" pageNumber="54">
<materialsCitation id="3B0F3C92FFB3FFE6CCDB03FDFB18FC33" ID-GBIF-Occurrence="3013449434" accessionNumber="T65795" collectingDate="2005-07-12" collectionCode="WAM, T" collectorName="M. Greenham &amp; D. Kamien &amp; L. Mould" country="Australia" latitude="-21.715557" location="Borehole" longLatPrecision="21" longitude="116.26583" municipality="Mesa" pageId="53" pageNumber="54" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia" typeStatus="paratype">
<emphasis id="B913EADDFFB3FFE6CCDB03FDFED9FC16" box="[199,313,897,922]" italics="true" pageId="53" pageNumber="54">
<typeStatus id="54DC886DFFB3FFE6CCDB03FDFED9FC16" box="[199,313,897,922]" pageId="53" pageNumber="54" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<collectingCountry id="F370765FFFB3FFE6CD5703FCFE0EFC16" box="[331,494,896,922]" name="Australia" pageId="53" pageNumber="54">AUSTRALIA</collectingCountry>
:
<emphasis id="B913EADDFFB3FFE6CDE303FDFD2DFC16" box="[511,717,896,922]" italics="true" pageId="53" pageNumber="54">
<collectingRegion id="49A3F82DFFB3FFE6CDE303FDFD2DFC16" box="[511,717,896,922]" country="Australia" name="Western Australia" pageId="53" pageNumber="54">Western Australia</collectingRegion>
</emphasis>
:
<specimenCount id="9D61FD46FFB3FFE6CEFE03FCFCF4FC17" box="[738,788,896,923]" pageId="53" pageNumber="54" type="male">1 ♂</specimenCount>
,
<collectingMunicipality id="6BBCACB5FFB3FFE6CF3A03FDFC83FC17" box="[806,867,897,923]" pageId="53" pageNumber="54">Mesa</collectingMunicipality>
K,
<location id="8EB86014FFB3FFE6CF8403FCFC1FFC16" LSID="urn:lsid:plazi:treatment:03CE87D9FFB3FFE6CC8B023AFE33F902:8EB86014FFB3FFE6CF8403FCFC1FFC16" box="[920,1023,896,922]" country="Australia" latitude="-21.715557" longLatPrecision="21" longitude="116.26583" municipality="Mesa" name="Borehole" pageId="53" pageNumber="54" stateProvince="Western Australia">Borehole</location>
1721 (trap 3),
<locationDeviation id="9A6A2FEDFFB3FFE6C8A303FCFA78FC16" box="[1215,1432,896,922]" location="Pannawonica" pageId="53" pageNumber="54">S of Pannawonica</locationDeviation>
,
<geoCoordinate id="EE535008FFB3FFE6CC8B03D8FEFEFC33" box="[151,286,932,959]" degrees="21" direction="south" minutes="42" orientation="latitude" pageId="53" pageNumber="54" precision="15" seconds="56" value="-21.715557">21°4256”S</geoCoordinate>
,
<geoCoordinate id="EE535008FFB3FFE6CD3703D8FE20FC33" box="[299,448,932,959]" degrees="116" direction="east" minutes="15" orientation="longitude" pageId="53" pageNumber="54" precision="15" seconds="57" value="116.26583">116°1557”E</geoCoordinate>
,
<date id="FFD9100FFFB3FFE6CDD003D8FD82FC33" box="[460,610,932,959]" pageId="53" pageNumber="54" value="2005-07-12">
<collectingDate id="EF9DE9E7FFB3FFE6CDD003D8FD82FC33" box="[460,610,932,959]" pageId="53" pageNumber="54" value="2005-07-12">12 July 2005</collectingDate>
</date>
,
<collectorName id="26925319FFB3FFE6CE7203D9FCEDFC32" box="[622,781,932,959]" pageId="53" pageNumber="54">M. Greenham</collectorName>
,
<collectorName id="26925319FFB3FFE6CF0503D9FC75FC32" box="[793,917,932,959]" pageId="53" pageNumber="54">D. Kamien</collectorName>
and
<collectorName id="26925319FFB3FFE6CFD103D9FBD8FC32" box="[973,1080,932,959]" pageId="53" pageNumber="54">L. Mould</collectorName>
(
<collectionCode id="ED76AE0AFFB3FFE6C85A03D8FB6DFC32" box="[1094,1165,932,958]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="53" pageNumber="54" type="Museum">WAM</collectionCode>
<collectionCode id="ED76AE0AFFB3FFE6C88803D8FB46FC33" box="[1172,1190,932,959]" name="Tavera, Department of Geology and Geophysics" pageId="53" pageNumber="54">T</collectionCode>
65795)
</materialsCitation>
;
<materialsCitation id="3B0F3C92FFB3FFE6C91803D8FE15FB8B" ID-GBIF-Occurrence="3013449444" accessionNumber="T65796" collectingDate="2005-07-12" collectionCode="WAM, T" collectorName="M. Greenham &amp; D. Kamien &amp; L. Mould" country="Australia" latitude="-21.716946" location="Borehole" longLatPrecision="21" longitude="116.264725" municipality="Mesa" pageId="53" pageNumber="54" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia" typeStatus="paratype">
<specimenCount id="9D61FD46FFB3FFE6C91803D8FAB3FC32" box="[1284,1363,932,958]" pageId="53" pageNumber="54" type="male">1 male</specimenCount>
,
<collectingMunicipality id="6BBCACB5FFB3FFE6C97C03D9FA7DFC33" box="[1376,1437,933,959]" pageId="53" pageNumber="54">Mesa</collectingMunicipality>
K,
<location id="8EB86014FFB3FFE6CCA603B4FEC1FC6E" LSID="urn:lsid:plazi:treatment:03CE87D9FFB3FFE6CC8B023AFE33F902:8EB86014FFB3FFE6CCA603B4FEC1FC6E" box="[186,289,968,994]" country="Australia" latitude="-21.716946" longLatPrecision="21" longitude="116.264725" municipality="Mesa" name="Borehole" pageId="53" pageNumber="54" stateProvince="Western Australia">Borehole</location>
1728 (trap 1),
<locationDeviation id="9A6A2FEDFFB3FFE6CDD503B4FD77FC6E" box="[457,663,968,994]" location="Pannawonica" pageId="53" pageNumber="54">S of Pannawonica</locationDeviation>
,
<geoCoordinate id="EE535008FFB3FFE6CEBF03B4FCC9FC6E" box="[675,809,968,994]" degrees="21" direction="south" minutes="43" orientation="latitude" pageId="53" pageNumber="54" precision="15" seconds="01" value="-21.716946">21°4301”S</geoCoordinate>
,
<geoCoordinate id="EE535008FFB3FFE6CF2903B4FC2BFC6F" box="[821,971,968,995]" degrees="116" direction="east" minutes="15" orientation="longitude" pageId="53" pageNumber="54" precision="15" seconds="53" value="116.264725">116°1553”E</geoCoordinate>
,
<date id="FFD9100FFFB3FFE6CFCB03B4FB8BFC6F" box="[983,1131,968,995]" pageId="53" pageNumber="54" value="2005-07-12">
<collectingDate id="EF9DE9E7FFB3FFE6CFCB03B4FB8BFC6F" box="[983,1131,968,995]" pageId="53" pageNumber="54" value="2005-07-12">12 July 2005</collectingDate>
</date>
,
<collectorName id="26925319FFB3FFE6C86B03B5FAF6FC6E" box="[1143,1302,968,995]" pageId="53" pageNumber="54">M. Greenham</collectorName>
,
<collectorName id="26925319FFB3FFE6C93D03B5FA7DFC6E" box="[1313,1437,968,995]" pageId="53" pageNumber="54">D. Kamien</collectorName>
and
<collectorName id="26925319FFB3FFE6CCD40391FED3FB8A" box="[200,307,1004,1031]" pageId="53" pageNumber="54">L. Mould</collectorName>
(
<collectionCode id="ED76AE0AFFB3FFE6CD5E0390FE69FB8A" box="[322,393,1004,1030]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="53" pageNumber="54" type="Museum">WAM</collectionCode>
<collectionCode id="ED76AE0AFFB3FFE6CD8D0390FE43FB8B" box="[401,419,1004,1031]" name="Tavera, Department of Geology and Geophysics" pageId="53" pageNumber="54">T</collectionCode>
65796)
</materialsCitation>
;
<materialsCitation id="3B0F3C92FFB3FFE6CE1D0390FB15FB8B" ID-GBIF-Occurrence="3013449375" accessionNumber="T65802, T65998" box="[513,1269,1004,1031]" collectingDate="2005-07-12" collectionCode="WAM" collectorName="M. Greenham &amp; D. Kamien &amp; L. Mould" country="Australia" latitude="-21.718334" location="Pannawonica" longLatPrecision="21" longitude="116.26" pageId="53" pageNumber="54" specimenCount="2" specimenCount-female="2" stateProvince="Western Australia" typeStatus="paratype">
<specimenCount id="9D61FD46FFB3FFE6CE1D0390FD8EFB8A" box="[513,622,1004,1030]" pageId="53" pageNumber="54" type="female">2 females</specimenCount>
(trap 2) same data as holotype (
<collectionCode id="ED76AE0AFFB3FFE6CFC10390FBC4FB8A" box="[989,1060,1004,1030]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="53" pageNumber="54" type="Museum">WAM</collectionCode>
T65802, T65998)
</materialsCitation>
;
<materialsCitation id="3B0F3C92FFB3FFE6C91E0390FDB4FBC3" ID-GBIF-Occurrence="3013449388" accessionNumber="T65798, T65997" collectingDate="2005-07-12" collectionCode="WAM" collectorName="M. Greenham &amp; D. Kamien &amp; L. Mould" country="Australia" latitude="-21.719168" location="Borehole" longLatPrecision="21" longitude="116.26444" municipality="Mesa" pageId="53" pageNumber="54" specimenCount="2" specimenCount-female="2" stateProvince="Western Australia" typeStatus="paratype">
<specimenCount id="9D61FD46FFB3FFE6C91E0390FACDFB8B" box="[1282,1325,1004,1031]" pageId="53" pageNumber="54" type="female">2 ♀</specimenCount>
,
<collectingMunicipality id="6BBCACB5FFB3FFE6C9200391FA99FB8B" box="[1340,1401,1005,1031]" pageId="53" pageNumber="54">Mesa</collectingMunicipality>
K,
<location id="8EB86014FFB3FFE6CC8B046CFF1EFBA6" LSID="urn:lsid:plazi:treatment:03CE87D9FFB3FFE6CC8B023AFE33F902:8EB86014FFB3FFE6CC8B046CFF1EFBA6" box="[151,254,1040,1066]" country="Australia" latitude="-21.719168" longLatPrecision="21" longitude="116.26444" municipality="Mesa" name="Borehole" pageId="53" pageNumber="54" stateProvince="Western Australia">Borehole</location>
1694 (traps 1, 2),
<locationDeviation id="9A6A2FEDFFB3FFE6CDD0046CFD79FBA6" box="[460,665,1040,1066]" location="Pannawonica" pageId="53" pageNumber="54">S of Pannawonica</locationDeviation>
,
<geoCoordinate id="EE535008FFB3FFE6CEB9046CFCCBFBA6" box="[677,811,1040,1066]" degrees="21" direction="south" minutes="43" orientation="latitude" pageId="53" pageNumber="54" precision="15" seconds="09" value="-21.719168">21°4309”S</geoCoordinate>
,
<geoCoordinate id="EE535008FFB3FFE6CF2B046CFC2DFBA7" box="[823,973,1040,1067]" degrees="116" direction="east" minutes="15" orientation="longitude" pageId="53" pageNumber="54" precision="15" seconds="52" value="116.26444">116°1552”E</geoCoordinate>
,
<date id="FFD9100FFFB3FFE6CFC5046CFB8CFBA7" box="[985,1132,1040,1067]" pageId="53" pageNumber="54" value="2005-07-12">
<collectingDate id="EF9DE9E7FFB3FFE6CFC5046CFB8CFBA7" box="[985,1132,1040,1067]" pageId="53" pageNumber="54" value="2005-07-12">12 July 2005</collectingDate>
</date>
,
<collectorName id="26925319FFB3FFE6C864046DFAF7FBA6" box="[1144,1303,1040,1067]" pageId="53" pageNumber="54">M. Greenham</collectorName>
,
<collectorName id="26925319FFB3FFE6C93D046DFA7DFBA6" box="[1313,1437,1040,1067]" pageId="53" pageNumber="54">D. Kamien</collectorName>
and
<collectorName id="26925319FFB3FFE6CCDB0449FED1FBC2" box="[199,305,1076,1103]" pageId="53" pageNumber="54">L. Mould</collectorName>
(
<collectionCode id="ED76AE0AFFB3FFE6CD230448FE66FBC2" box="[319,390,1076,1102]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="53" pageNumber="54" type="Museum">WAM</collectionCode>
T65798, T65997)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD836CFFFB3FFE6CCD20424FADEFBFE" blockId="53.[151,1437,824,1679]" box="[206,1342,1112,1139]" pageId="53" pageNumber="54">
<emphasis id="B913EADDFFB3FFE6CCD20424FE9DFBFE" box="[206,381,1112,1138]" italics="true" pageId="53" pageNumber="54">Other material.</emphasis>
<collectingCountry id="F370765FFFB3FFE6CD9E0424FDC5FBFE" box="[386,549,1112,1138]" name="Australia" pageId="53" pageNumber="54">AUSTRALIA</collectingCountry>
:
<emphasis id="B913EADDFFB3FFE6CE330425FD17FBFE" box="[559,759,1112,1138]" italics="true" pageId="53" pageNumber="54">
<collectingRegion id="49A3F82DFFB3FFE6CE330425FD17FBFE" box="[559,759,1112,1138]" country="Australia" name="Western Australia" pageId="53" pageNumber="54">Western Australia</collectingRegion>
</emphasis>
: See
<bibRefCitation id="EFF64B3EFFB3FFE6CF290425FBF9FBFE" author="Harvey, M. S. &amp; Berry, O. &amp; Edward, K. L. &amp; Humphreys, G." box="[821,1049,1112,1139]" pageId="53" pageNumber="54" pagination="167 - 194" refId="ref46967" refString="Harvey, M. S., Berry, O., Edward, K. L. &amp; Humphreys, G. (2008) Molecular and morphological systematics of hypogean schizomids (Schizomida: Hubbardiidae) in semi-arid Australia. Invertebrate Systematics, 22 (2), 167 - 194. https: // doi. org / 10.1071 / IS 07026" type="journal article" year="2008">
Harvey
<emphasis id="B913EADDFFB3FFE6CF930425FC20FBFE" box="[911,960,1112,1138]" italics="true" pageId="53" pageNumber="54">et al</emphasis>
. (2008)
</bibRefCitation>
and
<bibRefCitation id="EFF64B3EFFB3FFE6C8510424FADAFBFE" author="Abrams, K. M. &amp; Huey, J. A. &amp; Hillyer, M. J. &amp; Humphreys, W. F. &amp; Didham, R. K. &amp; Harvey, M. S." box="[1101,1338,1112,1139]" pageId="53" pageNumber="54" pagination="106532" refId="ref46122" refString="Abrams, K. M., Huey, J. A., Hillyer, M. J., Humphreys, W. F., Didham, R. K. &amp; Harvey, M. S. (2019) Too hot to handle: Tertiary aridification drives multiple independent incursions of Schizomida (Hubbardiidae) into hypogean environments. Molecular Phylogenetics and Evolution, 139, 106532. https: // doi. org / 10.1016 / j. ympev. 2019.106532" type="journal article" year="2019">
Abrams
<emphasis id="B913EADDFFB3FFE6C8B10425FB3EFBFE" box="[1197,1246,1112,1138]" italics="true" pageId="53" pageNumber="54">et al</emphasis>
. (2019)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37D6544FFB3FFE6CCDB0400FE7CF9AE" pageId="53" pageNumber="54" type="diagnosis">
<paragraph id="8BD836CFFFB3FFE6CCDB0400FE7CF9AE" blockId="53.[151,1437,824,1679]" pageId="53" pageNumber="54">
<emphasis id="B913EADDFFB3FFE6CCDB0400FEA2FB1A" bold="true" box="[199,322,1148,1174]" pageId="53" pageNumber="54">Diagnosis.</emphasis>
The shape of the male flagellum of
<taxonomicName id="4C674D4CFFB3FFE6CEC50400FC5DFB1A" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[729,957,1148,1174]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="kryptus">
<emphasis id="B913EADDFFB3FFE6CEC50400FC5DFB1A" box="[729,957,1148,1174]" italics="true" pageId="53" pageNumber="54">Draculoides kryptus</emphasis>
</taxonomicName>
most closely resembles
<emphasis id="B913EADDFFB3FFE6C8CC0401FF16FB36" italics="true" pageId="53" pageNumber="54">
<taxonomicName id="4C674D4CFFB3FFE6C8CC0401FADBFB1A" baseAuthorityName="Framenau, Hamilton, Finston, Humphreys, Abrams, Huey and Harvey" baseAuthorityYear="2018" box="[1232,1339,1148,1174]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="affinis">D. affinis</taxonomicName>
,
<taxonomicName id="4C674D4CFFB3FFE6C95A0401FF16FB36" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="anachoretus">D. anachoretus</taxonomicName>
</emphasis>
,
<taxonomicName id="4C674D4CFFB3FFE6CD1904DDFE99FB36" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[261,377,1184,1210]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="bythius">
<emphasis id="B913EADDFFB3FFE6CD1904DDFE99FB36" box="[261,377,1184,1210]" italics="true" pageId="53" pageNumber="54">D. bythius</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFB3FFE6CD9B04DDFDFFFB36" baseAuthorityName="Framenau, Hamilton, Finston, Humphreys, Abrams, Huey and Harvey" baseAuthorityYear="2018" box="[391,543,1184,1210]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="cochranus">
<emphasis id="B913EADDFFB3FFE6CD9B04DDFDFFFB36" box="[391,543,1184,1210]" italics="true" pageId="53" pageNumber="54">D. cochranus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFB3FFE6CE3104DDFD48FB36" baseAuthorityName="Abrams and Harvey" baseAuthorityYear="2015" box="[557,680,1185,1210]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="eremius">
<emphasis id="B913EADDFFB3FFE6CE3104DDFD48FB36" box="[557,680,1185,1210]" italics="true" pageId="53" pageNumber="54">D. eremius</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFB3FFE6CEAA04DDFCB7FB36" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[694,855,1184,1210]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="gnophicola">
<emphasis id="B913EADDFFB3FFE6CEAA04DDFCB7FB36" box="[694,855,1184,1210]" italics="true" pageId="53" pageNumber="54">D. gnophicola</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFB3FFE6CF7904DDFBD9FB36" authorityName="Abrams and Harvey" authorityYear="2020" box="[869,1081,1184,1210]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="mckechnieorum">
<emphasis id="B913EADDFFB3FFE6CF7904DDFBD9FB36" box="[869,1081,1184,1210]" italics="true" pageId="53" pageNumber="54">D. mckechnieorum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFB3FFE6C85404DDFAEDFB36" box="[1096,1293,1185,1210]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="noctigrassator">
<emphasis id="B913EADDFFB3FFE6C85404DDFAEDFB36" box="[1096,1293,1185,1210]" italics="true" pageId="53" pageNumber="54">D. noctigrassator</emphasis>
</taxonomicName>
and
<emphasis id="B913EADDFFB3FFE6C95904DDFF15FB52" italics="true" pageId="53" pageNumber="54">
<taxonomicName id="4C674D4CFFB3FFE6C95904DDFF11FB52" authorityName="Abrams and Harvey" authorityYear="2020" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="warramboo">D. warramboo</taxonomicName>
,
</emphasis>
especially in the presence of a broad base. It differs from
<taxonomicName id="4C674D4CFFB3FFE6CF9404B9FC11FB52" baseAuthorityName="Framenau, Hamilton, Finston, Humphreys, Abrams, Huey and Harvey" baseAuthorityYear="2018" box="[904,1009,1220,1246]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="B913EADDFFB3FFE6CF9404B9FC11FB52" box="[904,1009,1220,1246]" italics="true" pageId="53" pageNumber="54">D. affinis</emphasis>
</taxonomicName>
,
<emphasis id="B913EADDFFB3FFE6C81C04B9FAB5FB52" box="[1024,1365,1220,1246]" italics="true" pageId="53" pageNumber="54">
<taxonomicName id="4C674D4CFFB3FFE6C81C04B9FB9BFB52" baseAuthorityName="Abrams and Harvey" baseAuthorityYear="2015" box="[1024,1147,1221,1246]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="eremius">D. eremius</taxonomicName>
and
<taxonomicName id="4C674D4CFFB3FFE6C8A804B9FAB5FB52" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[1204,1365,1220,1246]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="gnophicola">D. gnophicola</taxonomicName>
</emphasis>
in the presence of a distally tapered tip (blunt tip in the aforementioned species). It differs from
<taxonomicName id="4C674D4CFFB3FFE6C86F0495FAFDFA8E" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[1139,1309,1256,1282]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="anachoretus">
<emphasis id="B913EADDFFB3FFE6C86F0495FAFDFA8E" box="[1139,1309,1256,1282]" italics="true" pageId="53" pageNumber="54">D. anachoretus</emphasis>
</taxonomicName>
,
<emphasis id="B913EADDFFB3FFE6C9360495FECEFAAA" italics="true" pageId="53" pageNumber="54">
<taxonomicName id="4C674D4CFFB3FFE6C9360495FA7CFA8E" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[1322,1436,1256,1282]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="bythius">D. bythius</taxonomicName>
<taxonomicName id="4C674D4CFFB3FFE6CC8B0571FECEFAAA" baseAuthorityName="Framenau, Hamilton, Finston, Humphreys, Abrams, Huey and Harvey" baseAuthorityYear="2018" box="[151,302,1292,1318]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="cochranus">D. cochranus</taxonomicName>
</emphasis>
,
<taxonomicName id="4C674D4CFFB3FFE6CD200571FDEEFAAA" authorityName="Abrams and Harvey" authorityYear="2020" box="[316,526,1292,1318]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="mckechnieorum">
<emphasis id="B913EADDFFB3FFE6CD200571FDEEFAAA" box="[316,526,1292,1318]" italics="true" pageId="53" pageNumber="54">D. mckechnieorum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFB3FFE6CE000571FD01FAAA" box="[540,737,1293,1318]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="noctigrassator">
<emphasis id="B913EADDFFB3FFE6CE000571FD01FAAA" box="[540,737,1293,1318]" italics="true" pageId="53" pageNumber="54">D. noctigrassator</emphasis>
</taxonomicName>
and
<taxonomicName id="4C674D4CFFB3FFE6CF040571FC5DFAAA" authorityName="Abrams and Harvey" authorityYear="2020" box="[792,957,1292,1318]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="warramboo">
<emphasis id="B913EADDFFB3FFE6CF040571FC5DFAAA" box="[792,957,1292,1318]" italics="true" pageId="53" pageNumber="54">D. warramboo</emphasis>
</taxonomicName>
in the posterior position of dm
<quantity id="4C9F9B2AFFB3FFE6C90A0570FAA0FAAA" box="[1302,1344,1292,1318]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="53" pageNumber="54" unit="in" value="4.0">4 in</quantity>
relation to dl3 (dm4 anterior to dl
<quantity id="4C9F9B2AFFB3FFE6CDAD054CFE3BFAC6" box="[433,475,1328,1354]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="53" pageNumber="54" unit="in" value="3.0">3 in</quantity>
all of the aforementioned species except
<taxonomicName id="4C674D4CFFB3FFE6CFB4054DFBFBFAC6" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[936,1051,1328,1354]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="bythius">
<emphasis id="B913EADDFFB3FFE6CFB4054DFBFBFAC6" box="[936,1051,1328,1354]" italics="true" pageId="53" pageNumber="54">D. bythius</emphasis>
</taxonomicName>
, in which dm4 is in line with dl3). It is currently not possible to differentiate females of this species on morphology alone.
<taxonomicName id="4C674D4CFFB3FFE6C8760528FAADFAE2" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[1130,1357,1364,1390]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="kryptus">
<emphasis id="B913EADDFFB3FFE6C8760528FAADFAE2" box="[1130,1357,1364,1390]" italics="true" pageId="53" pageNumber="54">Draculoides kryptus</emphasis>
</taxonomicName>
can be diagnosed from all other
<taxonomicName id="4C674D4CFFB3FFE6CDB70504FDD5FA1E" box="[427,565,1400,1426]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B913EADDFFB3FFE6CDB70504FDD5FA1E" box="[427,565,1400,1426]" italics="true" pageId="53" pageNumber="54">Draculoides</emphasis>
</taxonomicName>
species that were sequenced at COI and 12S by the 50bp mini-barcodes shown in
<figureCitation id="135C2A4AFFB3FFE6CCAF05E0FEFBFA3A" box="[179,283,1436,1462]" captionStart="FIGURE 3" captionStartId="18.[151,250,1547,1571]" captionTargetBox="[158,1420,181,1521]" captionTargetId="figure-20@18.[151,1436,181,1524]" captionTargetPageId="18" captionText="FIGURE 3. COI mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913085). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." figureDoi="http://doi.org/10.5281/zenodo.4416857" httpUri="https://zenodo.org/record/4416857/files/figure.png" pageId="53" pageNumber="54">Figures 3</figureCitation>
and
<figureCitation id="135C2A4AFFB3FFE6CD4C05E0FE80FA3B" box="[336,352,1436,1463]" captionStart="FIGURE 5" captionStartId="20.[151,250,1216,1240]" captionTargetBox="[158,1424,183,1189]" captionTargetId="figure-20@20.[151,1436,181,1192]" captionTargetPageId="20" captionText="FIGURE 5. 12S mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913012). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." figureDoi="http://doi.org/10.5281/zenodo.4416863" httpUri="https://zenodo.org/record/4416863/files/figure.png" pageId="53" pageNumber="54">5</figureCitation>
.
<taxonomicName id="4C674D4CFFB3FFE6CD7705E0FDAEFA3A" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[363,590,1436,1462]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="kryptus">
<emphasis id="B913EADDFFB3FFE6CD7705E0FDAEFA3A" box="[363,590,1436,1462]" italics="true" pageId="53" pageNumber="54">Draculoides kryptus</emphasis>
</taxonomicName>
can be diagnosed from all other
<taxonomicName id="4C674D4CFFB3FFE6CFA505E0FBA3FA3A" box="[953,1091,1436,1462]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B913EADDFFB3FFE6CFA505E0FBA3FA3A" box="[953,1091,1436,1462]" italics="true" pageId="53" pageNumber="54">Draculoides</emphasis>
</taxonomicName>
species that were sequenced at ITS2 (
<figureCitation id="135C2A4AFFB3FFE6CCC205BCFEC1FA57" box="[222,289,1472,1499]" captionStart="FIGURE 7" captionStartId="22.[151,250,1755,1779]" captionTargetBox="[159,1425,181,1729]" captionTargetId="figure-20@22.[151,1436,181,1732]" captionTargetPageId="22" captionText="FIGURE 7. ITS2 mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913105). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." figureDoi="http://doi.org/10.5281/zenodo.4416867" httpUri="https://zenodo.org/record/4416867/files/figure.png" pageId="53" pageNumber="54">Fig. 7</figureCitation>
) except for
<taxonomicName id="4C674D4CFFB3FFE6CDBA05BDFDB0FA56" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[422,592,1472,1498]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="anachoretus">
<emphasis id="B913EADDFFB3FFE6CDBA05BDFDB0FA56" box="[422,592,1472,1498]" italics="true" pageId="53" pageNumber="54">D. anachoretus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFB3FFE6CE4205BDFD30FA56" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[606,720,1472,1498]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="bythius">
<emphasis id="B913EADDFFB3FFE6CE4205BDFD30FA56" box="[606,720,1472,1498]" italics="true" pageId="53" pageNumber="54">D. bythius</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFB3FFE6CEC205BDFCB7FA56" baseAuthorityName="Abrams and Harvey" baseAuthorityYear="2015" box="[734,855,1473,1498]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="eremius">
<emphasis id="B913EADDFFB3FFE6CEC205BDFCB7FA56" box="[734,855,1473,1498]" italics="true" pageId="53" pageNumber="54">D. eremius</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFB3FFE6CF7805BDFBE4FA56" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[868,1028,1472,1498]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="gnophicola">
<emphasis id="B913EADDFFB3FFE6CF7805BDFBE4FA56" box="[868,1028,1472,1498]" italics="true" pageId="53" pageNumber="54">D. gnophicola</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFB3FFE6C80D05BDFB03FA56" authorityName="Abrams and Harvey" authorityYear="2020" box="[1041,1251,1472,1498]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="mckechnieorum">
<emphasis id="B913EADDFFB3FFE6C80D05BDFB03FA56" box="[1041,1251,1472,1498]" italics="true" pageId="53" pageNumber="54">D. mckechnieorum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFB3FFE6C8ED05BDFA76FA56" authorityName="Abrams and Harvey" authorityYear="2020" box="[1265,1430,1472,1498]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="warramboo">
<emphasis id="B913EADDFFB3FFE6C8ED05BDFA76FA56" box="[1265,1430,1472,1498]" italics="true" pageId="53" pageNumber="54">D. warramboo</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFB3FFE6CC8B0599FED1FA72" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" box="[151,305,1508,1534]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="immortalis">
<emphasis id="B913EADDFFB3FFE6CC8B0599FED1FA72" box="[151,305,1508,1534]" italics="true" pageId="53" pageNumber="54">D. immortalis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFB3FFE6CD220599FE3BFA72" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" box="[318,475,1508,1534]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="belalugosii">
<emphasis id="B913EADDFFB3FFE6CD220599FE3BFA72" box="[318,475,1508,1534]" italics="true" pageId="53" pageNumber="54">D. belalugosii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFB3FFE6CDF40599FD53FA72" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" box="[488,691,1508,1534]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="christopherleei">
<emphasis id="B913EADDFFB3FFE6CDF40599FD53FA72" box="[488,691,1508,1534]" italics="true" pageId="53" pageNumber="54">D. christopherleei</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFB3FFE6CEA30599FCB8FA72" authorityName="Abrams and Harvey" authorityYear="2020" box="[703,856,1508,1534]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="piscivultus">
<emphasis id="B913EADDFFB3FFE6CEA30599FCB8FA72" box="[703,856,1508,1534]" italics="true" pageId="53" pageNumber="54">D. piscivultus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C674D4CFFB3FFE6CF900599FBE9FA72" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" box="[908,1033,1508,1534]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="akashae">
<emphasis id="B913EADDFFB3FFE6CF900599FBE9FA72" box="[908,1033,1508,1534]" italics="true" pageId="53" pageNumber="54">D. akashae</emphasis>
</taxonomicName>
, which are not distinguishable using the ITS2 mini-barcode.
</paragraph>
</subSubSection>
<subSubSection id="C37D6544FFB3FFE6CCDB0650FD9AF9CA" box="[199,634,1580,1607]" pageId="53" pageNumber="54" type="description">
<paragraph id="8BD836CFFFB3FFE6CCDB0650FD9AF9CA" blockId="53.[151,1437,824,1679]" box="[199,634,1580,1607]" pageId="53" pageNumber="54">
<emphasis id="B913EADDFFB3FFE6CCDB0650FEB9F9CA" bold="true" box="[199,345,1580,1606]" pageId="53" pageNumber="54">Description.</emphasis>
See
<bibRefCitation id="EFF64B3EFFB3FFE6CD930651FD95F9CA" author="Harvey, M. S. &amp; Berry, O. &amp; Edward, K. L. &amp; Humphreys, G." box="[399,629,1580,1607]" pageId="53" pageNumber="54" pagination="167 - 194" refId="ref46967" refString="Harvey, M. S., Berry, O., Edward, K. L. &amp; Humphreys, G. (2008) Molecular and morphological systematics of hypogean schizomids (Schizomida: Hubbardiidae) in semi-arid Australia. Invertebrate Systematics, 22 (2), 167 - 194. https: // doi. org / 10.1071 / IS 07026" type="journal article" year="2008">
Harvey
<emphasis id="B913EADDFFB3FFE6CDF40651FDFAF9CA" box="[488,538,1580,1606]" italics="true" pageId="53" pageNumber="54">et al</emphasis>
. (2008)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37D6544FFB3FFE6CCDB062CFE33F902" pageId="53" pageNumber="54" type="discussion">
<paragraph id="8BD836CFFFB3FFE6CCDB062CFE33F902" blockId="53.[151,1437,824,1679]" pageId="53" pageNumber="54">
<emphasis id="B913EADDFFB3FFE6CCDB062CFEDBF9E6" bold="true" box="[199,315,1616,1642]" pageId="53" pageNumber="54">Remarks.</emphasis>
<taxonomicName id="4C674D4CFFB3FFE6CD59062CFDCCF9E6" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[325,556,1616,1642]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="kryptus">
<emphasis id="B913EADDFFB3FFE6CD59062CFDCCF9E6" box="[325,556,1616,1642]" italics="true" pageId="53" pageNumber="54">Draculoides kryptus</emphasis>
</taxonomicName>
is found in Mesa K situated south of Pannawonica in the Pilbara region of
<collectingRegion id="49A3F82DFFB3FFE6CC8B0609FEBFF902" box="[151,351,1652,1679]" country="Australia" name="Western Australia" pageId="53" pageNumber="54">Western Australia</collectingRegion>
(
<figureCitation id="135C2A4AFFB3FFE6CD720608FE24F902" box="[366,452,1652,1678]" captionStart="FIGURE 1" captionStartId="16.[151,250,1868,1892]" captionTargetBox="[346,1243,188,1844]" captionTargetId="figure-20@16.[339,1249,181,1844]" captionTargetPageId="16" captionText="FIGURE 1. Map of an area of the Pilbara described as “West Pilbara” for the purposes of this study, showing the distribution of the named Draculoides species. Inset maps show species distributions in finer detail encompassing areas A: Warramboo, Mesas A, B and C and “BudgieBore”; B: Bungaroo, Mesas G, H, J, K and L; C: Cochrane and Jewell, Kens Bore, Cane and Upper Cane River, Trinity Bore and Catho Well, Mt. Stuart Station; D: Middle Robe, Robe Valley; E: Bungaroo South, West Pit, Buckland Hills and Dragon deposit." figureDoi="http://doi.org/10.5281/zenodo.4416851" httpUri="https://zenodo.org/record/4416851/files/figure.png" pageId="53" pageNumber="54">Fig. 1B</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>