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<document id="B358D97201B9198D770F3C3C869DFC9E" ID-DOI="10.11646/zootaxa.3652.3.2" ID-GBIF-Dataset="9cb4b047-ae44-419e-898a-eb3bc27c3d15" ID-ISSN="1175-5326" ID-Zenodo-Dep="224020" ID-ZooBank="B334624E-A7F5-4077-BF64-D2AD6DB215D5" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460609979718" checkinUser="plazi" docAuthor="Mattos, Ingrid &amp; Mermudes, José Ricardo M." docDate="2013" docId="03CE87A7FFBF694ECFEFFAD8FCB7F8CF" docLanguage="en" docName="zt03652p342.pdf" docOrigin="Zootaxa 3652 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paxillus manausensis Mattos &amp; Mermudes, new species" docType="treatment" docVersion="8" lastPageNumber="336" masterDocId="FFF7FFDFFFB76947CF78FFA2FFF8FFF8" masterDocTitle="Synopsis of Paxillus MacLeay, 1819 (Coleoptera: Passalidae): distributional records and descriptions of four new species from Brazil" masterLastPageNumber="342" masterPageNumber="327" pageNumber="335" updateTime="1698328243075" updateUser="plazi">
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<mods:title id="E1CC6064826586DCF1D1CF5F91250D3D">Synopsis of Paxillus MacLeay, 1819 (Coleoptera: Passalidae): distributional records and descriptions of four new species from Brazil</mods:title>
</mods:titleInfo>
<mods:name id="AF55E182D14150BC427F5FC1474E2523" type="personal">
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<mods:namePart id="DB74742345A82090A9A8C9F6F9544378">Mattos, Ingrid</mods:namePart>
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<mods:namePart id="2D2B3CDB890E45B87C7F81B038C34ABD">Mermudes, José Ricardo M.</mods:namePart>
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<mods:typeOfResource id="2CA84A7AE3DE47953D0C370CCF9FB6F0">text</mods:typeOfResource>
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<mods:title id="092A02027A7601B5A2E7DD4CE769FD48">Zootaxa</mods:title>
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<mods:part id="652E85061F85033CAE12661D88694D76">
<mods:date id="794EF307A483ACA4AA9110891E999D61">2013</mods:date>
<mods:detail id="FA84114207746BACDC158A12CF386F56" type="volume">
<mods:number id="D0D47787BA61F33F25964355FC3C00BD">3652</mods:number>
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<mods:identifier id="C2052A0FD15296E46C7AC3DC39A92109" type="DOI">10.11646/zootaxa.3652.3.2</mods:identifier>
<mods:identifier id="7E1C0116CDE46DFDDABBF1EB0C0A0E16" type="GBIF-Dataset">9cb4b047-ae44-419e-898a-eb3bc27c3d15</mods:identifier>
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<treatment id="03CE87A7FFBF694ECFEFFAD8FCB7F8CF" ID-DOI="http://doi.org/10.5281/zenodo.6154349" ID-GBIF-Taxon="119568116" ID-Zenodo-Dep="6154349" LSID="urn:lsid:plazi:treatment:03CE87A7FFBF694ECFEFFAD8FCB7F8CF" httpUri="http://treatment.plazi.org/id/03CE87A7FFBF694ECFEFFAD8FCB7F8CF" lastPageId="9" lastPageNumber="336" pageId="8" pageNumber="335">
<subSubSection id="C37D653AFFBF694FCFEFFAD8FED6FA4D" pageId="8" pageNumber="335" type="nomenclature">
<paragraph id="8BD836B1FFBF694FCFEFFAD8FCB2FA6C" blockId="8.[151,842,1402,1461]" box="[151,842,1402,1428]" pageId="8" pageNumber="335">
<heading id="D09081DDFFBF694FCFEFFAD8FCB2FA6C" bold="true" box="[151,842,1402,1428]" fontSize="11" level="1" pageId="8" pageNumber="335" reason="1">
<emphasis id="B913EAA3FFBF694FCFEFFAD8FCB2FA6C" bold="true" box="[151,842,1402,1428]" pageId="8" pageNumber="335">
<taxonomicName id="4C674D32FFBF694FCFEFFAD8FD55FA6C" authority="Mattos &amp; Mermudes" authorityName="Mattos &amp; Mermudes" box="[151,685,1402,1428]" class="Insecta" family="Passallidae" genus="Paxillus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="335" phylum="Arthropoda" rank="species" species="manausensis" status="sp. nov.">
<emphasis id="B913EAA3FFBF694FCFEFFAD8FE65FA6C" bold="true" box="[151,413,1402,1428]" italics="true" pageId="8" pageNumber="335">Paxillus manausensis</emphasis>
Mattos &amp; Mermudes
</taxonomicName>
,
<taxonomicNameLabel id="A22057D8FFBF694FCDC2FAD8FCB2FA6C" box="[698,842,1402,1428]" pageId="8" pageNumber="335" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD836B1FFBF694FCFEFFA3FFED6FA4D" blockId="8.[151,842,1402,1461]" box="[151,302,1437,1461]" pageId="8" pageNumber="335">
(
<figureCitation id="135C2A34FFBF694FCFE7FA3FFEDEFA4D" box="[159,294,1437,1461]" captionStart="FIGURES 17 21" captionStartId="9.[151,270,1366,1390]" captionTargetBox="[199,1400,495,1345]" captionTargetId="figure@9.[151,1436,482,1345]" captionTargetPageId="9" captionText="FIGURES 17 21. Paxillus manausensis, male: 17, head and anterior margin of pronotum, dorsal; 18, antennae, dorsal. 19 21. Aedeagus: 19, dorsal; 20, lateral; 21, ventral. Scale = 0.5 mm (Figs. 19 21 to the same scale)." httpUri="https://zenodo.org/record/224025/files/figure.png" pageId="8" pageNumber="335">Figs. 1721</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37D653AFFBF694ECFEFFA47FBA4FA0B" lastPageId="9" lastPageNumber="336" pageId="8" pageNumber="335" type="description">
<paragraph id="8BD836B1FFBF694ECFEFFA47FBA4FE54" blockId="8.[151,1436,1509,2038]" lastBlockId="9.[151,1436,151,428]" lastPageId="9" lastPageNumber="336" pageId="8" pageNumber="335">
<emphasis id="B913EAA3FFBF694FCFEFFA47FED9FA06" bold="true" box="[151,289,1509,1534]" pageId="8" pageNumber="335">Description</emphasis>
. Length
<quantity id="4C9F9B54FFBF694FCEE9FA47FDEAFA05" box="[401,530,1509,1533]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="8" pageNumber="335" unit="mm" value="15.5" valueMax="16.0" valueMin="15.0">1516 mm</quantity>
. Head (
<figureCitation id="135C2A34FFBF694FCD0AFA47FD33FA05" box="[626,715,1509,1533]" captionStart="FIGURES 17 21" captionStartId="9.[151,270,1366,1390]" captionTargetBox="[199,1400,495,1345]" captionTargetId="figure@9.[151,1436,482,1345]" captionTargetPageId="9" captionText="FIGURES 17 21. Paxillus manausensis, male: 17, head and anterior margin of pronotum, dorsal; 18, antennae, dorsal. 19 21. Aedeagus: 19, dorsal; 20, lateral; 21, ventral. Scale = 0.5 mm (Figs. 19 21 to the same scale)." httpUri="https://zenodo.org/record/224025/files/figure.png" pageId="8" pageNumber="335">Fig. 17</figureCitation>
): labrum with anterior margin almost straight, with fine and moderately dense setiferous punctures; anterior frontal area irregularly punctate, with coarse and shallow punctures, somewhat dense; opaque inside the punctures; median frontal area irregularly elevated, opaque; anterior frontal ridges elevated, almost straight, reaching median frontal tubercles; posterior ridges forming an obtuse angle. Inner tubercles evident and prominent, smaller than medial frontal tubercles, medial frontal tubercles developed, elevated, and acuminate. Lateral frontal area deep with irregular elevations; elevations with coarse, shallow punctures, microsculpturing inside punctures. Bridge strong and smooth. Central tubercle short, not free, somewhat less dilated at base and merged with posterior lateral tubercles; posterior lateral tubercles small and distinct at the apex. Lateral post-frontal area smooth or punctate. Antennal club (
<figureCitation id="135C2A34FFBF694FCCF7F8A7FC1BF8E5" box="[911,995,1797,1821]" captionStart="FIGURES 17 21" captionStartId="9.[151,270,1366,1390]" captionTargetBox="[199,1400,495,1345]" captionTargetId="figure@9.[151,1436,482,1345]" captionTargetPageId="9" captionText="FIGURES 17 21. Paxillus manausensis, male: 17, head and anterior margin of pronotum, dorsal; 18, antennae, dorsal. 19 21. Aedeagus: 19, dorsal; 20, lateral; 21, ventral. Scale = 0.5 mm (Figs. 19 21 to the same scale)." httpUri="https://zenodo.org/record/224025/files/figure.png" pageId="8" pageNumber="335">Fig. 18</figureCitation>
): first antennomere reduced, ½ shorter than second antennomere, narrower and acuminate at apex, other antennomeres subequal. Pronotum: anterior and lateral margins straight, anterior marginal groove short, usually reaching ¼ of anterior margin; rarely long, narrow and with fine punctures, weakly separated from anterior margin. Lateral marginal groove of pronotum narrow, with coarse, contiguous and dense punctures; lateral area with irregular punctation, sometimes confluent; indistinct scar; posterior margin with or without short pubescence or tuft of setae. Prosternal process, not flattened, posteriorly wide and truncate. Scutellum flattened, with wide median groove. Elytra: humeri with tuft of setae (females) or few short setae (males). Mesosternum smooth, short scar, reaching mid length of side of mesosternum, usually suboval; lateral anterior area opaque. Metasternum: disc evident, anterior lateral area with coarse punctures, confluent and microsculptured, reaching lateral median area near the disc; posterolateral area of metasternum with fine and denser punctures. Mesotibia dorsally pubescent, sometimes with one small spine. Aedeagus (
<figureCitation id="135C2A34FFBE694ECBC3FEA6FAC0FEE4" box="[1211,1336,260,284]" captionStart="FIGURES 17 21" captionStartId="9.[151,270,1366,1390]" captionTargetBox="[199,1400,495,1345]" captionTargetId="figure@9.[151,1436,482,1345]" captionTargetPageId="9" captionText="FIGURES 17 21. Paxillus manausensis, male: 17, head and anterior margin of pronotum, dorsal; 18, antennae, dorsal. 19 21. Aedeagus: 19, dorsal; 20, lateral; 21, ventral. Scale = 0.5 mm (Figs. 19 21 to the same scale)." httpUri="https://zenodo.org/record/224025/files/figure.png" pageId="9" pageNumber="336">Fig. 1921</figureCitation>
) slightly longer than wide and less sclerotized (dorsal view). Phallus globular, longer than parameres and phallobase together (lateral view); phallobase and parameres fused (lateral view), lacking division between them; parameres shortened, abruptly acuminate at apex, with inner right angle (lateral view). Phallobase strongly transverse and narrowed in the middle; ventral distal margin wide and deeply notched (ventral view).
</paragraph>
<caption id="DF186639FFBE694ECFEFFAF4FB36FA76" httpUri="https://zenodo.org/record/224025/files/figure.png" pageId="9" pageNumber="336" targetBox="[199,1400,495,1345]" targetPageId="9">
<paragraph id="8BD836B1FFBE694ECFEFFAF4FB36FA76" blockId="9.[151,1437,1366,1422]" pageId="9" pageNumber="336">
<emphasis id="B913EAA3FFBE694ECFEFFAF4FEA6FA96" bold="true" box="[151,350,1366,1390]" pageId="9" pageNumber="336">FIGURES 1721.</emphasis>
<taxonomicName id="4C674D32FFBE694ECE1DFAF5FDB4FA96" box="[357,588,1367,1390]" class="Insecta" family="Passallidae" genus="Paxillus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="336" phylum="Arthropoda" rank="species" species="manausensis">
<emphasis id="B913EAA3FFBE694ECE1DFAF5FDB4FA96" box="[357,588,1367,1390]" italics="true" pageId="9" pageNumber="336">Paxillus manausensis</emphasis>
</taxonomicName>
, male: 17, head and anterior margin of pronotum, dorsal; 18, antennae, dorsal. 1921. Aedeagus: 19, dorsal; 20, lateral; 21, ventral. Scale = 0.5 mm (Figs. 1921 to the same scale).
</paragraph>
</caption>
<paragraph id="8BD836B1FFBE694ECFBFFA14FBA4FA0B" blockId="9.[151,1436,1462,1848]" pageId="9" pageNumber="336">
<emphasis id="B913EAA3FFBE694ECFBFFA14FEAAFA37" bold="true" box="[199,338,1462,1487]" pageId="9" pageNumber="336">Dimensions</emphasis>
(mm). Total length (from anterior margin of labrum to apex of elytra) 1516; elytral length (on median line) 9; pronotal length (on median line) 3; pronotal width 4; humeral width 4.
</paragraph>
</subSubSection>
<subSubSection id="C37D653AFFBE694ECFBFFA5DFB3EF9C3" pageId="9" pageNumber="336" type="materials_examined">
<paragraph id="8BD836B1FFBE694ECFBFFA5DFB3EF9C3" blockId="9.[151,1436,1462,1848]" pageId="9" pageNumber="336">
<emphasis id="B913EAA3FFBE694ECFBFFA5DFE8DF9EF" bold="true" box="[199,373,1534,1559]" pageId="9" pageNumber="336">
<typeStatus id="54DC8813FFBE694ECFBFFA5DFEFAF9EF" box="[199,258,1535,1559]" pageId="9" pageNumber="336">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54DC8813FFBE694ECE04FA5DFE1FF9EF" box="[380,487,1535,1559]" pageId="9" pageNumber="336" type="holotype">Holotype</typeStatus>
, male from
<collectingCountry id="F3707621FFBE694ECD17FA5DFD40F9EF" box="[623,696,1535,1559]" name="Brazil" pageId="9" pageNumber="336">Brazil</collectingCountry>
:
<emphasis id="B913EAA3FFBE694ECDBBFA5DFCCEF9EF" box="[707,822,1535,1559]" italics="true" pageId="9" pageNumber="336">Amazonas</emphasis>
:
<taxonomicName id="4C674D32FFBE694ECC3CF9A2FC67F9EF" box="[836,927,1536,1559]" class="Insecta" family="Passallidae" genus="Paxillus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="327" phylum="Arthropoda" rank="species" species="manaus" status="sp. nov.">Manaus</taxonomicName>
,
<date id="FFD91071FFBE694ECCD4FA5DFBEDF9EF" box="[940,1045,1535,1559]" pageId="9" pageNumber="336" value="1959-07">VII.1959</date>
, C. Elias leg. Mattos &amp; Mermudes det. 2012 (DZUP, nº 187918).
<typeStatus id="54DC8813FFBE694ECE94F986FDAAF9C4" box="[492,594,1572,1596]" pageId="9" pageNumber="336" type="paratype">Paratype</typeStatus>
: female with same data as
<typeStatus id="54DC8813FFBE694ECC05F981FC27F9C3" box="[893,991,1571,1595]" pageId="9" pageNumber="336" type="holotype">holotype</typeStatus>
(DZUP, nº 187916).
</paragraph>
</subSubSection>
<subSubSection id="C37D653AFFBE694ECFBFF9E5FECCF8EB" pageId="9" pageNumber="336" type="discussion">
<paragraph id="8BD836B1FFBE694ECFBFF9E5FECCF8EB" blockId="9.[151,1436,1462,1848]" pageId="9" pageNumber="336">
<emphasis id="B913EAA3FFBE694ECFBFF9E5FEC3F9A7" bold="true" box="[199,315,1607,1631]" pageId="9" pageNumber="336">Remarks.</emphasis>
<taxonomicName id="4C674D32FFBE694ECE3BF9E5FDCFF9A7" box="[323,567,1607,1631]" class="Insecta" family="Passallidae" genus="Paxillus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="336" phylum="Arthropoda" rank="species" species="manausensis">
<emphasis id="B913EAA3FFBE694ECE3BF9E5FDCFF9A7" box="[323,567,1607,1631]" italics="true" pageId="9" pageNumber="336">Paxillus manausensis</emphasis>
</taxonomicName>
is easily distinguished from
<emphasis id="B913EAA3FFBE694ECCFAF9EAFB54F9A7" box="[898,1196,1607,1631]" italics="true" pageId="9" pageNumber="336">
<taxonomicName id="4C674D32FFBE694ECCFAF9EAFC0BF9A7" box="[898,1011,1607,1631]" class="Insecta" family="Passallidae" genus="Paxillus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="336" phylum="Arthropoda" rank="species" species="borellii">P. borellii</taxonomicName>
,
<taxonomicName id="4C674D32FFBE694ECB78F9EAFB54F9A7" box="[1024,1196,1607,1631]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="336" phylum="Arthropoda" rank="species" species="pentaphyllus">P. pentaphyllus</taxonomicName>
</emphasis>
, and
<taxonomicName id="4C674D32FFBE694ECB95F9EAFAB4F9A7" box="[1261,1356,1608,1631]" class="Insecta" family="Passallidae" genus="Paxillus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="336" phylum="Arthropoda" rank="species" species="reyesi">
<emphasis id="B913EAA3FFBE694ECB95F9EAFAB4F9A7" box="[1261,1356,1608,1631]" italics="true" pageId="9" pageNumber="336">P. reyesi</emphasis>
</taxonomicName>
by the inner tubercles evident and prominent, smaller than the median frontal tubercles, the latter developed, elevated and acuminate; central tubercle short and merged with the posterior lateral tubercles, these small and distinct at the apex (
<figureCitation id="135C2A34FFBE694ECFA3F911FED5F933" box="[219,301,1715,1739]" captionStart="FIGURES 17 21" captionStartId="9.[151,270,1366,1390]" captionTargetBox="[199,1400,495,1345]" captionTargetId="figure@9.[151,1436,482,1345]" captionTargetPageId="9" captionText="FIGURES 17 21. Paxillus manausensis, male: 17, head and anterior margin of pronotum, dorsal; 18, antennae, dorsal. 19 21. Aedeagus: 19, dorsal; 20, lateral; 21, ventral. Scale = 0.5 mm (Figs. 19 21 to the same scale)." httpUri="https://zenodo.org/record/224025/files/figure.png" pageId="9" pageNumber="336">Fig. 17</figureCitation>
). In addition, the phallobase is strongly transverse and narrowed in the middle (
<figureCitation id="135C2A34FFBE694ECBDDF911FAD5F933" box="[1189,1325,1715,1739]" captionStart="FIGURES 17 21" captionStartId="9.[151,270,1366,1390]" captionTargetBox="[199,1400,495,1345]" captionTargetId="figure@9.[151,1436,482,1345]" captionTargetPageId="9" captionText="FIGURES 17 21. Paxillus manausensis, male: 17, head and anterior margin of pronotum, dorsal; 18, antennae, dorsal. 19 21. Aedeagus: 19, dorsal; 20, lateral; 21, ventral. Scale = 0.5 mm (Figs. 19 21 to the same scale)." httpUri="https://zenodo.org/record/224025/files/figure.png" pageId="9" pageNumber="336">Figs. 1921</figureCitation>
); with the distal margin wide and deeply notched (ventral view), and the parameres short, abruptly acuminate at the apex (lateral view).
</paragraph>
</subSubSection>
<subSubSection id="C37D653AFFBE694ECFBFF8BDFCB7F8CF" box="[199,847,1823,1848]" pageId="9" pageNumber="336" type="etymology">
<paragraph id="8BD836B1FFBE694ECFBFF8BDFCB7F8CF" blockId="9.[151,1436,1462,1848]" box="[199,847,1823,1848]" pageId="9" pageNumber="336">
<emphasis id="B913EAA3FFBE694ECFBFF8BDFEB6F8CF" bold="true" box="[199,334,1823,1847]" pageId="9" pageNumber="336">Etymology.</emphasis>
The name refers to the
<typeStatus id="54DC8813FFBE694ECD2FF882FD7FF8C0" box="[599,647,1824,1848]" pageId="9" pageNumber="336">type</typeStatus>
locality,
<taxonomicName id="4C674D32FFBE694ECD97F882FCB3F8CF" box="[751,843,1824,1847]" class="Insecta" family="Passallidae" genus="Paxillus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="327" phylum="Arthropoda" rank="species" species="manaus" status="sp. nov.">Manaus</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>