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<document id="754EF0112A895AEE4339EAF584A05B39" ID-CLB-Dataset="51159" ID-DOI="10.11646/zootaxa.4067.2.3" ID-GBIF-Dataset="65b27f71-5ad2-4183-893c-65bfc6189df7" ID-ISSN="1175-5326" ID-Zenodo-Dep="262371" ID-ZooBank="1331311A-4DF6-4414-A5FF-6E5152F593EA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459693015290" checkinUser="plazi" docAuthor="Cho, Hee-Wook &amp; Borowiec, Lech" docDate="2016" docId="03AE7B0CF54C9D3BF1C9FB41FE39FEDA" docLanguage="en" docName="zootaxa.4067.2.3.pdf" docOrigin="Zootaxa 4067 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gonioctena (Brachyphytodecta) arunensis Cho &amp; Borowiec, sp. nov." docType="treatment" docVersion="9" lastPageNumber="176" masterDocId="FF970374F5499D33F15EFF84FF8BFFBF" masterDocTitle="On the genus Gonioctena Chevrolat (Coleoptera: Chrysomelidae: Chrysomelinae), with descriptions of seven new species from the Oriental region and Palaearctic China" masterLastPageNumber="184" masterPageNumber="168" pageNumber="173" updateTime="1698209477154" updateUser="ExternalLinkService">
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<mods:title id="54C05BCF01F9A9517E800BB9016BDE65">On the genus Gonioctena Chevrolat (Coleoptera: Chrysomelidae: Chrysomelinae), with descriptions of seven new species from the Oriental region and Palaearctic China</mods:title>
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<mods:namePart id="991B9543F59ACA23B2502345CF965362">Cho, Hee-Wook</mods:namePart>
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<mods:namePart id="C90B69139E268CF2A03EA8792A1A2E86">Borowiec, Lech</mods:namePart>
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<mods:title id="38F50D43A59CEBC572849A112D0DBD61">Zootaxa</mods:title>
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<mods:date id="9B1F1D759C0FE208ABB26DA991D3EC06">2016</mods:date>
<mods:detail id="C25106B36BC20FA9826303EC18C8FD79" type="volume">
<mods:number id="CC91EBBC336FD4F7BD3B20579A616CB5">4067</mods:number>
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<mods:identifier id="96F4193A872C2FF2A81BED105124BEF3" type="DOI">10.11646/zootaxa.4067.2.3</mods:identifier>
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<treatment id="03AE7B0CF54C9D3BF1C9FB41FE39FEDA" ID-DOI="http://doi.org/10.5281/zenodo.6080899" ID-GBIF-Taxon="119333513" ID-Zenodo-Dep="6080899" LSID="urn:lsid:plazi:treatment:03AE7B0CF54C9D3BF1C9FB41FE39FEDA" httpUri="http://treatment.plazi.org/id/03AE7B0CF54C9D3BF1C9FB41FE39FEDA" lastPageId="8" lastPageNumber="176" pageId="5" pageNumber="173">
<subSubSection id="C31D9991F54C9D36F1C9FB41FEF1FABF" pageId="5" pageNumber="173" type="nomenclature">
<paragraph id="8BB8CA1AF54C9D36F1C9FB41FC59FB60" blockId="5.[151,978,1221,1281]" box="[151,978,1221,1247]" pageId="5" pageNumber="173">
<heading id="D0F07D76F54C9D36F1C9FB41FC59FB60" bold="true" box="[151,978,1221,1247]" fontSize="11" level="1" pageId="5" pageNumber="173" reason="1">
<emphasis id="B9731608F54C9D36F1C9FB41FC59FB60" bold="true" box="[151,978,1221,1247]" pageId="5" pageNumber="173">
<taxonomicName id="4C07B199F54C9D36F1C9FB41FCE6FB60" authority="Cho &amp; Borowiec" authorityName="Cho &amp; Borowiec" box="[151,877,1221,1247]" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="173" phylum="Arthropoda" rank="species" species="arunensis" status="sp. nov." subGenus="Brachyphytodecta">
<emphasis id="B9731608F54C9D36F1C9FB41FEA8FB60" bold="true" box="[151,291,1221,1247]" italics="true" pageId="5" pageNumber="173">Gonioctena</emphasis>
(
<emphasis id="B9731608F54C9D36F06AFB41FD85FB60" bold="true" box="[308,526,1221,1247]" italics="true" pageId="5" pageNumber="173">Brachyphytodecta</emphasis>
)
<emphasis id="B9731608F54C9D36F341FB41FD1DFB60" bold="true" box="[543,662,1221,1247]" italics="true" pageId="5" pageNumber="173">arunensis</emphasis>
Cho &amp; Borowiec
</taxonomicName>
<taxonomicNameLabel id="A240AB73F54C9D36F22AFB41FC59FB60" box="[884,978,1221,1247]" pageId="5" pageNumber="173" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB8CA1AF54C9D36F1C9FB6CFEF1FABF" blockId="5.[151,978,1221,1281]" box="[151,378,1256,1281]" pageId="5" pageNumber="173">
(
<figureCitation id="133CD69FF54C9D36F1C1FB6CFE94FABF" box="[159,287,1256,1281]" captionStart="FIGURES 7 – 12" captionStartId="3.[151,263,1983,2006]" captionTargetBox="[151,1436,1057,1962]" captionTargetId="figure@3.[151,1436,1057,1963]" captionTargetPageId="3" captionText="FIGURES 7 – 12. Dorsal habitus (7, 9, 11) and head and pronotum (8, 10, 12). 7 – 8. Gonioctena (A.) tonkinensis stat. nov.; 9 – 10. G. (A.) sapaensis nom. nov.; 11 – 12. G. (B.) arunensis sp. nov. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/262373/files/figure.png" pageId="5" pageNumber="173">Figs 11–12</figureCitation>
,
<figureCitation id="133CD69FF54C9D36F074FB6CFEFAFABF" box="[298,369,1256,1280]" captionStart="FIGURES 33 – 39" captionStartId="7.[151,263,1951,1974]" captionTargetBox="[202,1384,602,1910]" captionTargetId="figure@7.[185,1401,589,1930]" captionTargetPageId="7" captionText="FIGURES 33 – 39. Antenna (33, 36, 38) and aedeagus (34 – 35, 37, 39, dorsal, apical and lateral views). 33 – 34. Gonioctena (A.) oudai sp. nov.; 35. G. (A.) coccinella; 36 – 37. G. (A.) sapaensis nom. nov.; 38 – 39. G. (B.) arunensis sp. nov. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/262376/files/figure.png" pageId="5" pageNumber="173">38–39</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31D9991F54C9D36F1C9FAB5FB51FAF9" box="[151,1242,1328,1350]" pageId="5" pageNumber="173" type="reference_group">
<paragraph id="8BB8CA1AF54C9D36F1C9FAB5FB51FAF9" blockId="5.[151,1242,1328,1350]" box="[151,1242,1328,1350]" pageId="5" pageNumber="173">
<treatmentCitationGroup id="AB17ED34F54C9D36F1C9FAB5FB51FAF9" box="[151,1242,1328,1350]" pageId="5" pageNumber="173">
<taxonomicName id="4C07B199F54C9D36F1C9FAB5FCC6FAF9" authority="Medvedev &amp; Sprecher-Uebersachs 1998: 26" authorityName="Medvedev &amp; Sprecher-Uebersachs" authorityPageNumber="26" authorityYear="1998" box="[151,845,1328,1350]" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="173" phylum="Arthropoda" rank="species" species="inconditus">
<emphasis id="B9731608F54C9D36F1C9FAB5FEF7FAF9" box="[151,380,1328,1350]" italics="true" pageId="5" pageNumber="173">Gonioctena inconditus</emphasis>
:
<treatmentCitation id="0AA6EC0BF54C9D36F0D4FAB4FCC6FAF9" author="Medvedev" box="[394,845,1328,1350]" page="26" pageId="5" pageNumber="173" year="1998">
<bibRefCitation id="EF96B7EBF54C9D36F0D4FAB4FCA3FAF9" author="Medvedev" box="[394,808,1328,1350]" pageId="5" pageNumber="184" refString="Medvedev, L. N. &amp; Sprecher-Uebersachs, E. (1998) New data on Chrysomelidae of Nepal (Insecta, Coleoptera). Spixiana, 21 (1), 25 - 42." type="journal article" year="1998">Medvedev &amp; Sprecher-Uebersachs 1998</bibRefCitation>
: 26
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0AA6EC0BF54C9D36F206FAB4FB8EFAF9" author="Kimoto" box="[856,1029,1328,1350]" page="41" pageId="5" pageNumber="173" year="2005">
<bibRefCitation id="EF96B7EBF54C9D36F206FAB4FC69FAF9" author="Kimoto" box="[856,994,1328,1350]" pageId="5" pageNumber="183" refString="Kimoto, S. (2005) Systematic catalog of the Chrysomelidae (Coleoptera) from Nepal and Bhutan. Bulletin of the Kitakyushu Museum of Natural History and Human History, Series A, 3, 13 - 114." type="journal article" year="2005">Kimoto 2005</bibRefCitation>
: 41
</treatmentCitation>
(misidentifications).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31D9991F54C9D36F1C9FAF7FC71FA10" pageId="5" pageNumber="173" type="materials_examined">
<paragraph id="8BB8CA1AF54C9D36F1C9FAF7FC71FA10" blockId="5.[151,1437,1395,2031]" pageId="5" pageNumber="173">
<emphasis id="B9731608F54C9D36F1C9FAF7FF53FA33" bold="true" box="[151,216,1395,1420]" pageId="5" pageNumber="173">
<typeStatus id="54BC74B8F54C9D36F1C9FAF7FF5FFA33" box="[151,212,1395,1420]" pageId="5" pageNumber="173">Type</typeStatus>
.
</emphasis>
<typeStatus id="54BC74B8F54C9D36F1B8FAF7FED8FA33" box="[230,339,1395,1420]" pageId="5" pageNumber="173" type="holotype">Holotype</typeStatus>
: male (
<collectionCode id="ED1652DFF54C9D36F0F1FAF7FD80FA34" box="[431,523,1395,1419]" pageId="5" pageNumber="173">LMCM</collectionCode>
), Num-Chichila, 15/
<quantity id="4CFF67FFF54C9D36F3A5FAF7FCD3FA34" box="[763,856,1395,1420]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="173" unit="m" value="1900.0">1900 m</quantity>
17.6. // O-Nepal Arun
<date id="FFB9ECDAF54C9D36F522FAF7FB44FA34" box="[1148,1231,1395,1419]" pageId="5" pageNumber="173" value="1980-05">V.1980</date>
, C. Holzschuh //
<typeStatus id="54BC74B8F54C9D36F1C9FA13FEB6FA10" box="[151,317,1431,1455]" pageId="5" pageNumber="173" type="holotype">HOLOTYPUS</typeStatus>
<taxonomicName id="4C07B199F54C9D36F01AFA13FDBDFA10" authority="Brachy." authorityName="Brachy." box="[324,566,1431,1455]" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="173" phylum="Arthropoda" rank="genus">
<emphasis id="B9731608F54C9D36F01AFA13FE4CFA11" box="[324,455,1431,1454]" italics="true" pageId="5" pageNumber="173">Gonioctena</emphasis>
(
<emphasis id="B9731608F54C9D36F089FA13FDADFA10" box="[471,550,1431,1455]" italics="true" pageId="5" pageNumber="173">Brachy</emphasis>
.)
</taxonomicName>
<taxonomicName id="4C07B199F54C9D36F363FA1CFD21FA10" box="[573,682,1432,1455]" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="173" phylum="Arthropoda" rank="species" species="arunensis" status="sp. nov.">
<emphasis id="B9731608F54C9D36F363FA1CFD21FA10" box="[573,682,1432,1455]" italics="true" pageId="5" pageNumber="173">arunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A240AB73F54C9D36F3EFFA1CFD66FA10" box="[689,749,1432,1455]" pageId="5" pageNumber="173" rank="species">sp. n.</taxonomicNameLabel>
<bibRefCitation id="EF96B7EBF54C9D36F3AAFA12FC7CFA10" author="Cho" box="[756,1015,1430,1455]" pageId="5" pageNumber="183" refString="Cho, H. W. &amp; Borowiec, L. (2014) Two new species of the genus Gonioctena Chevrolat (Coleoptera, Chrysomelidae, Chrysomelinae) from Sichuan, China. Zootaxa, 3765 (3), 295 - 300. http: // dx. doi. org / 10.11646 / zootaxa. 3765.3.6" type="journal article" year="2014">Cho &amp; Borowiec 2014</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31D9991F54C9D36F199FA3FFE6FF94C" pageId="5" pageNumber="173" type="diagnosis">
<paragraph id="8BB8CA1AF54C9D36F199FA3FFE6FF94C" blockId="5.[151,1437,1395,2031]" pageId="5" pageNumber="173">
<emphasis id="B9731608F54C9D36F199FA3FFECAFA6B" bold="true" box="[199,321,1467,1492]" pageId="5" pageNumber="173">Diagnosis.</emphasis>
<taxonomicName id="4C07B199F54C9D36F015FA3FFCA6FA6C" box="[331,813,1467,1491]" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="173" phylum="Arthropoda" rank="species" species="arunensis" status="sp. nov." subGenus="Brachyphytodecta">
<emphasis id="B9731608F54C9D36F015FA3FFE45FA6D" box="[331,462,1467,1490]" italics="true" pageId="5" pageNumber="173">Gonioctena</emphasis>
(
<emphasis id="B9731608F54C9D36F0BFFA3FFD26FA6C" box="[481,685,1467,1491]" italics="true" pageId="5" pageNumber="173">Brachyphytodecta</emphasis>
)
<emphasis id="B9731608F54C9D36F39EFA38FCA6FA6C" box="[704,813,1468,1491]" italics="true" pageId="5" pageNumber="173">arunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A240AB73F54C9D36F269FA3FFC1FFA6B" box="[823,916,1467,1492]" pageId="5" pageNumber="173" rank="species">
<emphasis id="B9731608F54C9D36F269FA3FFC1FFA6B" bold="true" box="[823,916,1467,1492]" pageId="5" pageNumber="173">sp. nov.</emphasis>
</taxonomicNameLabel>
is similar to
<taxonomicName id="4C07B199F54C9D36F56CFA3FFA17FA6C" authority="Weise, 1898" authorityName="Weise" authorityYear="1898" box="[1074,1436,1467,1492]" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="173" phylum="Arthropoda" rank="species" species="incondita" subGenus="Brachyphytodecta">
<emphasis id="B9731608F54C9D36F56CFA3FFBC8FA6D" box="[1074,1091,1467,1490]" italics="true" pageId="5" pageNumber="173">G</emphasis>
. (
<emphasis id="B9731608F54C9D36F503FA38FBE5FA6C" box="[1117,1134,1468,1491]" italics="true" pageId="5" pageNumber="173">B</emphasis>
.)
<emphasis id="B9731608F54C9D36F5D7FA3FFB78FA6C" box="[1161,1267,1467,1491]" italics="true" pageId="5" pageNumber="173">incondita</emphasis>
(Weise, 1898)
</taxonomicName>
occurring in Sikkim,
<collectingCountry id="F3108A8AF54C9D36F0D0FA5AFE4CFA48" box="[398,455,1502,1527]" name="India" pageId="5" pageNumber="173">India</collectingCountry>
in having large body size (more than
<quantity id="4CFF67FFF54C9D36F2D7FA5BFC5AFA48" box="[905,977,1503,1527]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="173" unit="mm" value="9.0">9 mm</quantity>
), black head and pronotum and reddish brown elytra. However,
<taxonomicName id="4C07B199F54C9D36F0F6F987FD14F9A4" box="[424,671,1539,1563]" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="173" phylum="Arthropoda" rank="species" species="arunensis" status="sp. nov.">
<emphasis id="B9731608F54C9D36F0F6F987FD14F9A4" box="[424,671,1539,1563]" italics="true" pageId="5" pageNumber="173">Gonioctena arunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A240AB73F54C9D36F3F8F987FC8BF9A3" box="[678,768,1539,1564]" pageId="5" pageNumber="173" rank="species">
<emphasis id="B9731608F54C9D36F3F8F987FC8BF9A3" bold="true" box="[678,768,1539,1564]" pageId="5" pageNumber="173">sp. nov.</emphasis>
</taxonomicNameLabel>
can be distinguished by black antennae (antennomeres 1–5 generally yellowish brown in
<taxonomicName id="4C07B199F54C9D36F0A2F9A3FD00F980" box="[508,651,1575,1599]" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="173" phylum="Arthropoda" rank="species" species="incondita">
<emphasis id="B9731608F54C9D36F0A2F9A3FD86F981" box="[508,525,1575,1598]" italics="true" pageId="5" pageNumber="173">G</emphasis>
.
<emphasis id="B9731608F54C9D36F37FF9A3FD00F980" box="[545,651,1575,1599]" italics="true" pageId="5" pageNumber="173">incondita</emphasis>
</taxonomicName>
, Fig. 13), black scutellum (reddish brown in
<taxonomicName id="4C07B199F54C9D36F5EAF9A3FAC9F980" box="[1204,1346,1575,1599]" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="173" phylum="Arthropoda" rank="species" species="incondita">
<emphasis id="B9731608F54C9D36F5EAF9A3FB4EF981" box="[1204,1221,1575,1598]" italics="true" pageId="5" pageNumber="173">G</emphasis>
.
<emphasis id="B9731608F54C9D36F586F9A3FAC9F980" box="[1240,1346,1575,1599]" italics="true" pageId="5" pageNumber="173">incondita</emphasis>
</taxonomicName>
), black abdomen (abdominal ventrites 2–5 reddish brown in
<taxonomicName id="4C07B199F54C9D36F3A6F9CFFC08F9DC" box="[760,899,1611,1635]" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="173" phylum="Arthropoda" rank="species" species="incondita">
<emphasis id="B9731608F54C9D36F3A6F9CFFC82F9DD" box="[760,777,1611,1634]" italics="true" pageId="5" pageNumber="173">G</emphasis>
.
<emphasis id="B9731608F54C9D36F247F9CFFC08F9DC" box="[793,899,1611,1635]" italics="true" pageId="5" pageNumber="173">incondita</emphasis>
</taxonomicName>
), pronotum with very large punctures laterally (much smaller in
<taxonomicName id="4C07B199F54C9D36F005F9EBFE6FF938" box="[347,484,1647,1671]" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="173" phylum="Arthropoda" rank="species" species="incondita">
<emphasis id="B9731608F54C9D36F005F9EBFEE7F939" box="[347,364,1647,1670]" italics="true" pageId="5" pageNumber="173">G</emphasis>
.
<emphasis id="B9731608F54C9D36F024F9EBFE6FF938" box="[378,484,1647,1671]" italics="true" pageId="5" pageNumber="173">incondita</emphasis>
</taxonomicName>
, Fig. 14) and aedeagus with apical process thin, roundly widened before apex (thick, apex widely truncate in
<taxonomicName id="4C07B199F54C9D36F0F6F917FDB9F914" box="[424,562,1683,1707]" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="173" phylum="Arthropoda" rank="species" species="incondita">
<emphasis id="B9731608F54C9D36F0F6F917FE32F915" box="[424,441,1683,1706]" italics="true" pageId="5" pageNumber="173">G</emphasis>
.
<emphasis id="B9731608F54C9D36F096F917FDB9F914" box="[456,562,1683,1707]" italics="true" pageId="5" pageNumber="173">incondita</emphasis>
</taxonomicName>
, Fig. 41).
<taxonomicName id="4C07B199F54C9D36F3F4F917FB72F914" authority="Chen, 1931" authorityName="Chen" authorityYear="1931" box="[682,1273,1683,1708]" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="173" phylum="Arthropoda" rank="species" species="lesnei" subGenus="Brachyphytodecta">
<emphasis id="B9731608F54C9D36F3F4F917FCA6F915" box="[682,813,1683,1706]" italics="true" pageId="5" pageNumber="173">Gonioctena</emphasis>
(
<emphasis id="B9731608F54C9D36F260F917FB81F914" box="[830,1034,1683,1707]" italics="true" pageId="5" pageNumber="173">Brachyphytodecta</emphasis>
)
<emphasis id="B9731608F54C9D36F545F917FBD6F914" box="[1051,1117,1683,1707]" italics="true" pageId="5" pageNumber="173">lesnei</emphasis>
(Chen, 1931)
</taxonomicName>
also has large body size and similar shape of aedeagus, but differs in having reddish brown head and pronotum and aedeagus with apical process thick (Fig. 42).
</paragraph>
</subSubSection>
<subSubSection id="C31D9991F54C9D3BF199F97AFABCFEA3" lastPageId="8" lastPageNumber="176" pageId="5" pageNumber="173" type="description">
<paragraph id="8BB8CA1AF54C9D36F199F97AFC68F8E0" blockId="5.[151,1437,1395,2031]" pageId="5" pageNumber="173">
<emphasis id="B9731608F54C9D36F199F97AFED2F8A8" bold="true" box="[199,345,1790,1815]" pageId="5" pageNumber="173">Description.</emphasis>
Measurements in mm (n = 1): length of body: 9.60; width of body: 6.40; height of body: 4.30; width of head: 2.55; interocular distance: 1.75; width of apex of pronotum: 3.05; width of base of pronotum: 5.20; length of pronotum along midline: 2.40; length of elytra along suture: 7.20.
</paragraph>
<paragraph id="8BB8CA1AF54C9D36F199F8EFFEB3F850" blockId="5.[151,1437,1395,2031]" pageId="5" pageNumber="173">
Body oval and strongly convex (
<figureCitation id="133CD69FF54C9D36F367F8EFFD07F83C" box="[569,652,1899,1924]" captionStart="FIGURES 7 – 12" captionStartId="3.[151,263,1983,2006]" captionTargetBox="[151,1436,1057,1962]" captionTargetId="figure@3.[151,1436,1057,1963]" captionTargetPageId="3" captionText="FIGURES 7 – 12. Dorsal habitus (7, 9, 11) and head and pronotum (8, 10, 12). 7 – 8. Gonioctena (A.) tonkinensis stat. nov.; 9 – 10. G. (A.) sapaensis nom. nov.; 11 – 12. G. (B.) arunensis sp. nov. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/262373/files/figure.png" pageId="5" pageNumber="173">Fig. 11</figureCitation>
). Head black, with dark reddish brown band near apex of mandibles. Antennomeres 1–4 black, partially blackish brown, 5–11 black. Pronotum and scutellum black. Elytra reddish brown, basal margin black. Venter black, with apical margin of last abdominal ventrite reddish brown. Legs entirely black.
</paragraph>
<caption id="DF789A92F54F9D35F1C9FA78FC92F9EF" httpUri="https://zenodo.org/record/262375/files/figure.png" pageId="6" pageNumber="174" targetBox="[170,1417,204,1500]" targetPageId="6">
<paragraph id="8BB8CA1AF54F9D35F1C9FA78FC92F9EF" blockId="6.[151,1436,1532,1616]" pageId="6" pageNumber="174">
<emphasis id="B9731608F54F9D35F1C9FA78FEDFF9AD" bold="true" box="[151,340,1532,1555]" pageId="6" pageNumber="174">FIGURES 25–32.</emphasis>
Antenna (25, 28), aedeagus (26–27, 29, 31, dorsal, apical and lateral views) and spermatheca (30, 32). 25– 26.
<taxonomicName id="4C07B199F54F9D35F19EF998FE79F98D" box="[192,498,1564,1586]" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="174" phylum="Arthropoda" rank="species" species="mausonensis" status="sp. nov." subGenus="Asiphytodecta">
<emphasis id="B9731608F54F9D35F19EF998FEBDF98E" box="[192,310,1564,1585]" italics="true" pageId="6" pageNumber="174">Gonioctena</emphasis>
(
<emphasis id="B9731608F54F9D35F017F998FED4F98E" box="[329,351,1564,1585]" italics="true" pageId="6" pageNumber="174">A.</emphasis>
)
<emphasis id="B9731608F54F9D35F02FF999FE79F98D" box="[369,498,1565,1586]" italics="true" pageId="6" pageNumber="174">mausonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A240AB73F54F9D35F0A3F998FDDAF98E" box="[509,593,1564,1585]" pageId="6" pageNumber="174" rank="species">
<emphasis id="B9731608F54F9D35F0A3F998FDDAF98E" bold="true" box="[509,593,1564,1585]" pageId="6" pageNumber="174">sp. nov.</emphasis>
</taxonomicNameLabel>
; 27.
<taxonomicName id="4C07B199F54F9D35F3D2F998FCEFF98D" box="[652,868,1564,1586]" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="174" phylum="Arthropoda" rank="species" species="flavoplagiata" subGenus="Asiphytodecta">
<emphasis id="B9731608F54F9D35F3D2F998FD2BF98E" box="[652,672,1564,1585]" italics="true" pageId="6" pageNumber="174">G.</emphasis>
(
<emphasis id="B9731608F54F9D35F3EAF998FD48F98E" box="[692,707,1564,1585]" italics="true" pageId="6" pageNumber="174">A</emphasis>
.)
<emphasis id="B9731608F54F9D35F383F998FCEFF98D" box="[733,868,1564,1586]" italics="true" pageId="6" pageNumber="174">flavoplagiata</emphasis>
</taxonomicName>
; 28–30.
<taxonomicName id="4C07B199F54F9D35F29AF998FB0CF98D" box="[964,1159,1564,1586]" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="174" phylum="Arthropoda" rank="species" species="tonkinensis" status="stat. nov." subGenus="Asiphytodecta">
<emphasis id="B9731608F54F9D35F29AF998FC53F98E" box="[964,984,1564,1585]" italics="true" pageId="6" pageNumber="174">G.</emphasis>
(
<emphasis id="B9731608F54F9D35F2B2F998FB88F98E" box="[1004,1027,1564,1585]" italics="true" pageId="6" pageNumber="174">A.</emphasis>
)
<emphasis id="B9731608F54F9D35F54BF998FB0CF98D" box="[1045,1159,1564,1586]" italics="true" pageId="6" pageNumber="174">tonkinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A240AB73F54F9D35F5CFF998FB7FF98E" box="[1169,1268,1564,1585]" pageId="6" pageNumber="174">
<emphasis id="B9731608F54F9D35F5CFF998FB7FF98E" bold="true" box="[1169,1268,1564,1585]" pageId="6" pageNumber="174">stat. nov.</emphasis>
</taxonomicNameLabel>
; 31–32.
<taxonomicName id="4C07B199F54F9D35F408F998FE93F9EF" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="174" phylum="Arthropoda" rank="species" species="subgeminata" subGenus="Asiphytodecta">
<emphasis id="B9731608F54F9D35F408F998FAE2F98E" box="[1366,1385,1564,1585]" italics="true" pageId="6" pageNumber="174">G.</emphasis>
(
<emphasis id="B9731608F54F9D35F420F998FA1FF98E" box="[1406,1428,1564,1585]" italics="true" pageId="6" pageNumber="174">A.</emphasis>
)
<emphasis id="B9731608F54F9D35F1C9F9BEFE93F9EF" box="[151,280,1594,1616]" italics="true" pageId="6" pageNumber="174">subgeminata</emphasis>
</taxonomicName>
. Scale bars: 1.0 mm (except Figs 30, 32: 0.5 mm).
</paragraph>
</caption>
<paragraph id="8BB8CA1AF54F9D35F199F9FCFF48F833" blockId="6.[151,1437,1655,2004]" pageId="6" pageNumber="174">
<emphasis id="B9731608F54F9D35F199F9FCFE8FF92F" box="[199,260,1656,1680]" italics="true" pageId="6" pageNumber="174">Head</emphasis>
. Vertex distinctly depressed near frontal suture, covered with sparse punctures, becoming coarser and denser toward sides. Frontal suture depressed, V-shaped, coronal suture indistinct. Frons flat, strongly depressed at anterior margin, covered with dense punctures. Clypeus very narrow and trapezoidal. Anterior margin of labrum weakly concave. Mandibles with 2 sharp apical teeth, a large excavation for maxillary palp at outer side. Maxillary palp 4-segmented, with apical palpomere slightly widened, truncate apically. Antennae reaching base of pronotum; antennomere 1 robust; antennomere 2 shorter than 3; antennomere 3 longer than 4; antennomeres 7–10 widened, 7– 8 each as long as wide, 9–10 each wider than long; antennomere 11 longest, about 1.20 times as long as wide (
<figureCitation id="133CD69FF54F9D35F431F8CBFF3CF833" captionStart="FIGURES 33 – 39" captionStartId="7.[151,263,1951,1974]" captionTargetBox="[202,1384,602,1910]" captionTargetId="figure@7.[185,1401,589,1930]" captionTargetPageId="7" captionText="FIGURES 33 – 39. Antenna (33, 36, 38) and aedeagus (34 – 35, 37, 39, dorsal, apical and lateral views). 33 – 34. Gonioctena (A.) oudai sp. nov.; 35. G. (A.) coccinella; 36 – 37. G. (A.) sapaensis nom. nov.; 38 – 39. G. (B.) arunensis sp. nov. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/262376/files/figure.png" pageId="6" pageNumber="174">Fig. 38</figureCitation>
).
</paragraph>
<paragraph id="8BB8CA1AF54F9D34F199F81DFE4CFF47" blockId="6.[151,1437,1655,2004]" lastBlockId="7.[151,1437,151,536]" lastPageId="7" lastPageNumber="175" pageId="6" pageNumber="174">
<emphasis id="B9731608F54F9D35F199F81DFEBDF80F" box="[199,310,1945,1968]" italics="true" pageId="6" pageNumber="174">Pronotum</emphasis>
. Lateral sides widest at base, roundly narrowed anteriorly, anterior angles strongly produced (
<figureCitation id="133CD69FF54F9D35F431F813FF3DF86B" captionStart="FIGURES 7 – 12" captionStartId="3.[151,263,1983,2006]" captionTargetBox="[151,1436,1057,1962]" captionTargetId="figure@3.[151,1436,1057,1963]" captionTargetPageId="3" captionText="FIGURES 7 – 12. Dorsal habitus (7, 9, 11) and head and pronotum (8, 10, 12). 7 – 8. Gonioctena (A.) tonkinensis stat. nov.; 9 – 10. G. (A.) sapaensis nom. nov.; 11 – 12. G. (B.) arunensis sp. nov. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/262373/files/figure.png" pageId="6" pageNumber="174">Fig. 12</figureCitation>
). Anterior and lateral margins bordered, lateral margins barely visible in dorsal view. Trichobothria absent on both anterior and posterior angles. Disc covered with very sparse punctures; lateral sides covered with very large and moderately dense punctures; interspaces covered with fine and sparse punctures. Scutellum slightly wider than long, narrowed posteriorly.
</paragraph>
<paragraph id="8BB8CA1AF54E9D34F199FE80FC72FE37" blockId="7.[151,1437,151,536]" pageId="7" pageNumber="175">
<emphasis id="B9731608F54E9D34F199FE80FE86FEA3" box="[199,269,260,284]" italics="true" pageId="7" pageNumber="175">Elytra</emphasis>
. Lateral sides subparallel, widest near middle, thence roundly narrowed posteriorly. Humeral calli well developed. Disc covered with 11 regular rows of large punctures, including a short scutellar row; punctures partially irregular, with additional punctures between striae; interspaces covered with fine and sparse punctures. Epipleura visible except near base in lateral view.
<collectingCountry id="F3108A8AF54E9D34F397FEEBFC8AFE37" box="[713,769,367,392]" name="India" pageId="7" pageNumber="175">Hind</collectingCountry>
wing well developed.
</paragraph>
<paragraph id="8BB8CA1AF54E9D34F199FE11FAC5FDA8" blockId="7.[151,1437,151,536]" pageId="7" pageNumber="175">
<emphasis id="B9731608F54E9D34F199FE11FE87FE13" box="[199,268,405,428]" italics="true" pageId="7" pageNumber="175">Venter</emphasis>
. Hypomera rugose, with a few punctures near anterolateral corners of prosternum. Prosternum covered with coarse and dense punctures bearing long setae; prosternal process enlarged apically, bordered laterally, with sparse punctures. Metasternum covered with small and sparse punctures in median region, large and dense punctures in lateral region. Abdominal ventrites covered with sparse or dense punctures bearing short setae.
</paragraph>
<caption id="DF789A92F54E9D34F1C9F81BFA16F868" httpUri="https://zenodo.org/record/262376/files/figure.png" pageId="7" pageNumber="175" targetBox="[202,1384,602,1910]" targetPageId="7">
<paragraph id="8BB8CA1AF54E9D34F1C9F81BFA16F868" blockId="7.[151,1437,1951,2007]" pageId="7" pageNumber="175">
<emphasis id="B9731608F54E9D34F1C9F81BFEDFF80A" bold="true" box="[151,340,1951,1974]" pageId="7" pageNumber="175">FIGURES 33–39.</emphasis>
Antenna (33, 36, 38) and aedeagus (34–35, 37, 39, dorsal, apical and lateral views). 33–34.
<taxonomicName id="4C07B199F54E9D34F5A7F825FF5AF868" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="175" phylum="Arthropoda" rank="species" species="oudai" status="sp. nov." subGenus="Asiphytodecta">
<emphasis id="B9731608F54E9D34F5A7F825FAE4F809" box="[1273,1391,1953,1974]" italics="true" pageId="7" pageNumber="175">Gonioctena</emphasis>
(
<emphasis id="B9731608F54E9D34F420F825FA1FF809" box="[1406,1428,1953,1974]" italics="true" pageId="7" pageNumber="175">A.</emphasis>
)
<emphasis id="B9731608F54E9D34F1C9F845FF5AF868" box="[151,209,1985,2007]" italics="true" pageId="7" pageNumber="175">oudai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A240AB73F54E9D34F188F845FEADF869" box="[214,294,1985,2006]" pageId="7" pageNumber="175" rank="species">
<emphasis id="B9731608F54E9D34F188F845FEADF869" bold="true" box="[214,294,1985,2006]" pageId="7" pageNumber="175">sp. nov.</emphasis>
</taxonomicNameLabel>
; 35.
<taxonomicName id="4C07B199F54E9D34F006F846FD83F868" box="[344,520,1985,2007]" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="175" phylum="Arthropoda" rank="species" species="coccinella" subGenus="Asiphytodecta">
<emphasis id="B9731608F54E9D34F006F846FEE7F868" box="[344,364,1986,2007]" italics="true" pageId="7" pageNumber="175">G.</emphasis>
(
<emphasis id="B9731608F54E9D34F022F846FE00F868" box="[380,395,1986,2007]" italics="true" pageId="7" pageNumber="175">A</emphasis>
.)
<emphasis id="B9731608F54E9D34F0FEF845FD83F868" box="[416,520,1985,2007]" italics="true" pageId="7" pageNumber="175">coccinella</emphasis>
</taxonomicName>
; 36–37.
<taxonomicName id="4C07B199F54E9D34F33EF846FC81F868" box="[608,778,1985,2007]" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="175" phylum="Arthropoda" rank="species" species="sapaensis" subGenus="Asiphytodecta">
<emphasis id="B9731608F54E9D34F33EF846FDFFF868" box="[608,628,1986,2007]" italics="true" pageId="7" pageNumber="175">G.</emphasis>
(
<emphasis id="B9731608F54E9D34F3DAF846FD11F868" box="[644,666,1986,2007]" italics="true" pageId="7" pageNumber="175">A.</emphasis>
)
<emphasis id="B9731608F54E9D34F3F9F846FC81F868" box="[679,778,1986,2007]" italics="true" pageId="7" pageNumber="175">sapaensis</emphasis>
</taxonomicName>
<emphasis id="B9731608F54E9D34F24EF845FCFCF869" bold="true" box="[784,887,1985,2006]" pageId="7" pageNumber="175">nom. nov.</emphasis>
; 38–39.
<taxonomicName id="4C07B199F54E9D34F28EF846FBF2F868" box="[976,1145,1985,2007]" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="175" phylum="Arthropoda" rank="species" species="arunensis" status="sp. nov." subGenus="Brachyphytodecta">
<emphasis id="B9731608F54E9D34F28EF846FC68F868" box="[976,995,1986,2007]" italics="true" pageId="7" pageNumber="175">G.</emphasis>
(
<emphasis id="B9731608F54E9D34F2ADF846FB81F868" box="[1011,1034,1986,2007]" italics="true" pageId="7" pageNumber="175">B.</emphasis>
)
<emphasis id="B9731608F54E9D34F549F846FBF2F868" box="[1047,1145,1986,2007]" italics="true" pageId="7" pageNumber="175">arunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A240AB73F54E9D34F521F845FB44F869" box="[1151,1231,1985,2006]" pageId="7" pageNumber="175" rank="species">
<emphasis id="B9731608F54E9D34F521F845FB44F869" bold="true" box="[1151,1231,1985,2006]" pageId="7" pageNumber="175">sp. nov.</emphasis>
</taxonomicNameLabel>
Scale bars: 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BB8CA1AF5419D3BF199FF1DFD71FF6A" blockId="8.[151,1436,151,357]" pageId="8" pageNumber="176">
<emphasis id="B9731608F5419D3BF199FF1DFF77FF0F" box="[199,252,153,176]" italics="true" pageId="8" pageNumber="176">Legs</emphasis>
. Moderately robust. Tibiae widened apically, with a tooth-like projection. Fore legs with tarsomere 1 distinctly narrower than 3. Tarsal claws appendiculate.
</paragraph>
<paragraph id="8BB8CA1AF5419D3BF199FF64FABCFEA3" blockId="8.[151,1436,151,357]" pageId="8" pageNumber="176">
<emphasis id="B9731608F5419D3BF199FF64FEBAFF47" box="[199,305,224,248]" italics="true" pageId="8" pageNumber="176">Genitalia</emphasis>
. Aedeagus moderately swollen laterally, with apical process roundly widened before apex in dorsal view; almost straight except base, tapering apically, with apical process recurved in lateral view (
<figureCitation id="133CD69FF5419D3BF588FE80FAA0FEA3" box="[1238,1323,260,285]" captionStart="FIGURES 33 – 39" captionStartId="7.[151,263,1951,1974]" captionTargetBox="[202,1384,602,1910]" captionTargetId="figure@7.[185,1401,589,1930]" captionTargetPageId="7" captionText="FIGURES 33 – 39. Antenna (33, 36, 38) and aedeagus (34 – 35, 37, 39, dorsal, apical and lateral views). 33 – 34. Gonioctena (A.) oudai sp. nov.; 35. G. (A.) coccinella; 36 – 37. G. (A.) sapaensis nom. nov.; 38 – 39. G. (B.) arunensis sp. nov. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/262376/files/figure.png" pageId="8" pageNumber="176">Fig. 39</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31D9991F5419D3BF199FEA3FCBEFEFF" box="[199,821,295,320]" pageId="8" pageNumber="176" type="etymology">
<paragraph id="8BB8CA1AF5419D3BF199FEA3FCBEFEFF" blockId="8.[151,1436,151,357]" box="[199,821,295,320]" pageId="8" pageNumber="176">
<emphasis id="B9731608F5419D3BF199FEA3FEC5FEFF" bold="true" box="[199,334,295,320]" pageId="8" pageNumber="176">Etymology.</emphasis>
Named after the
<typeStatus id="54BC74B8F5419D3BF34EFEACFDCBFEFF" box="[528,576,296,320]" pageId="8" pageNumber="176">type</typeStatus>
locality, Arun Valley.
</paragraph>
</subSubSection>
<subSubSection id="C31D9991F5419D3BF199FEC8FE39FEDA" box="[199,434,332,357]" pageId="8" pageNumber="176" type="distribution">
<paragraph id="8BB8CA1AF5419D3BF199FEC8FE39FEDA" blockId="8.[151,1436,151,357]" box="[199,434,332,357]" pageId="8" pageNumber="176">
<emphasis id="B9731608F5419D3BF199FEC8FEEBFEDA" bold="true" box="[199,352,332,357]" pageId="8" pageNumber="176">Distribution.</emphasis>
<collectingCountry id="F3108A8AF5419D3BF039FEC8FE25FEDA" box="[359,430,332,357]" name="Nepal" pageId="8" pageNumber="176">Nepal</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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