191 lines
28 KiB
XML
191 lines
28 KiB
XML
<document id="12C0E413F0C0A1DC7BF723850143491E" ID-DOI="10.5281/zenodo.183126" ID-GBIF-Dataset="5ba6cec4-345d-4f95-8064-dd9dc33dac4b" ID-ISSN="1175-5326" ID-Zenodo-Dep="183126" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459892214864" checkinUser="plazi" docAuthor="Miller, Kelly B. & Spangler, Paul J." docDate="2008" docId="03AD2179AE5DFFC0FF78F8BCFE46F96B" docLanguage="en" docName="zt01827p052.pdf" docOrigin="Zootaxa 1827" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Fontidessus wheeleri Miller, new species" docType="treatment" docVersion="9" lastPageNumber="51" masterDocId="FF945901AE58FFC6FFEFFFDBFFA9FFC9" masterDocTitle="Fontidessus Miller and Spangler, a new genus of Bidessini from Venezuela (Coleoptera: Dytiscidae: Hydroporinae) with three new species" masterLastPageNumber="52" masterPageNumber="45" pageNumber="50" updateTime="1698228095007" updateUser="plazi">
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<mods:mods id="BA5120955CF1CE34676E42591D0F575E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="22F01D3B10B3B520AEFD201D23A153DF">
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<mods:title id="6B2BC91338A64C0256E92522B4143B4B">Fontidessus Miller and Spangler, a new genus of Bidessini from Venezuela (Coleoptera: Dytiscidae: Hydroporinae) with three new species</mods:title>
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</mods:titleInfo>
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<mods:name id="B6E36F57C7E814B6F6FB599DFFAC592B" type="personal">
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<mods:role id="390565FFD8C751790191112626B6454D">
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<mods:roleTerm id="70A3FF1EED553DDD11CCBDC2850C7C06">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="46726533C0666277AC198A6657E08C15">Miller, Kelly B.</mods:namePart>
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</mods:name>
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<mods:name id="47E5C28CBAB21EC425C5CC26213B09C2" type="personal">
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<mods:role id="062543FC2AE1CED085A54D83AC9A24EF">
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<mods:roleTerm id="50927F8A92D8448FB503B385BF8BF61F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9B9487DEB785A7064543C09E504854B7">Spangler, Paul J.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="65F07D7BA3FA2BA26410057D939F4EF8">text</mods:typeOfResource>
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<mods:relatedItem id="D7E0BF287BA2886454F1A44452A82A76" type="host">
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<mods:titleInfo id="2899F309CAD26FC8EC9626DEC0232A0D">
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<mods:title id="901802440010E17817440D557479349B">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="AC45C93FDBC29D45692CA5704EAFDDF4">
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<mods:date id="49D3AECFEF2D8FF31497F0F5B7DE41E2">2008</mods:date>
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<mods:detail id="72708CDFB6EBA3C76730DC4D2DE39729" type="volume">
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<mods:number id="7B1C5571784AF8F31CD5CB8109725ED6">1827</mods:number>
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<mods:extent id="8ECCA6FCCC332C416BBB8FB02AB49A4F" unit="page">
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<mods:start id="DAAA137FAD90FCA7AAD22B85971180FF">45</mods:start>
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<mods:end id="128A35E0F448529ED8A7695119EEC482">52</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="7D7AD15C61566660163F29BAA95490AD">journal article</mods:classification>
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<mods:identifier id="1B771F2BC389846B5EA77E41A6EC1ADE" type="DOI">10.5281/zenodo.183126</mods:identifier>
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<mods:identifier id="BEB11E9D47EDB9FE612C16FF14BEC26C" type="GBIF-Dataset">5ba6cec4-345d-4f95-8064-dd9dc33dac4b</mods:identifier>
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<mods:identifier id="23EC3441BBF58551697947517C837A20" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="2EDBA4F862E07FCAAE1EE96797A57117" type="Zenodo-Dep">183126</mods:identifier>
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<treatment id="03AD2179AE5DFFC0FF78F8BCFE46F96B" ID-DOI="http://doi.org/10.5281/zenodo.6233628" ID-GBIF-Taxon="119372127" ID-Zenodo-Dep="6233628" LSID="urn:lsid:plazi:treatment:03AD2179AE5DFFC0FF78F8BCFE46F96B" httpUri="http://treatment.plazi.org/id/03AD2179AE5DFFC0FF78F8BCFE46F96B" lastPageId="6" lastPageNumber="51" pageId="5" pageNumber="50">
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<subSubSection id="C31EC3E4AE5DFFC3FF78F8BCFEC1F86D" pageId="5" pageNumber="50" type="nomenclature">
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<paragraph id="8BBB906FAE5DFFC3FF78F8BCFD2EF848" blockId="5.[151,647,1895,1956]" box="[151,647,1895,1921]" pageId="5" pageNumber="50">
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<heading id="D0F32703AE5DFFC3FF78F8BCFD2EF848" bold="true" box="[151,647,1895,1921]" fontSize="11" level="1" pageId="5" pageNumber="50" reason="1">
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<emphasis id="B9704C7DAE5DFFC3FF78F8BCFD2EF848" bold="true" box="[151,647,1895,1921]" pageId="5" pageNumber="50">
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<taxonomicName id="4C04EBECAE5DFFC3FF78F8BCFE43F848" ID-CoL="6JDL5" authority="Miller" authorityName="Miller" box="[151,490,1895,1921]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="wheeleri" status="sp. nov.">
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<emphasis id="B9704C7DAE5DFFC3FF78F8BCFE3FF848" bold="true" box="[151,406,1895,1921]" italics="true" pageId="5" pageNumber="50">Fontidessus wheeleri</emphasis>
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Miller
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</taxonomicName>
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,
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<taxonomicNameLabel id="A243F106AE5DFFC3FE18F8BCFD2EF848" box="[503,647,1895,1921]" pageId="5" pageNumber="50" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBB906FAE5DFFC3FF78F851FEC1F86D" blockId="5.[151,647,1895,1956]" box="[151,360,1930,1956]" pageId="5" pageNumber="50">
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(
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<figureCitation id="133F8CEAAE5DFFC3FF70F851FEA0F86D" box="[159,265,1930,1956]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Fontidessus species. 1 – 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 – 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 – 4. Scale bar b = 0.1 mm for Figs 5 – 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="5" pageNumber="50">Figs 3, 7</figureCitation>
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,
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<figureCitation id="133F8CEAAE5DFFC3FEFAF851FEC9F86D" box="[277,352,1930,1956]" captionStart="FIGURES 8 – 17" captionStartId="4.[151,269,1053,1077]" captionTargetBox="[469,1097,360,1021]" captionTargetId="figure@4.[463,1103,354,1031]" captionTargetPageId="4" captionText="FIGURES 8 – 17. Fontidessus species. 8 – 10) F. toboganensis, male genitalia, 8) median lobe, dorsal aspect, 9) median lobe, right lateral aspect, 10) right lateral lobe, right lateral aspect. 11 – 13) F. ornatus, male genitalia, 11) median lobe and left lateral lobe, dorsal aspect, 12) median lobe, right lateral aspect, 13) right lateral lobe, right lateral aspect. 14 – 16) F. wheeleri, male genitalia, 14) median lobe and left lateral lobe, dorsal aspect, 15) median lobe, left lateral aspect, 16) right lateral lobe, right lateral aspect. 17) F. toboganensis, female genitalia, ventral aspect. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/183128/files/figure.png" pageId="5" pageNumber="50">14–16</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31EC3E4AE5DFFC3FF78F801FA3CF83D" box="[151,1429,2010,2036]" pageId="5" pageNumber="50" type="materials_examined">
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<paragraph id="8BBB906FAE5DFFC3FF78F801FA3CF83D" blockId="5.[151,1429,2010,2036]" box="[151,1429,2010,2036]" pageId="5" pageNumber="50">
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<emphasis id="B9704C7DAE5DFFC3FF78F801FE94F83D" bold="true" box="[151,317,2010,2036]" pageId="5" pageNumber="50">
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<typeStatus id="54BF2ECDAE5DFFC3FF78F801FF7DF83D" box="[151,212,2010,2036]" pageId="5" pageNumber="50">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F313D0FFAE5DFFC3FEABF801FE16F83D" box="[324,447,2010,2036]" name="Venezuela" pageId="5" pageNumber="50">Venezuela</collectingCountry>
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, Territorio Federal Amazonas,
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<quantity id="4CFC3D8AAE5DFFC3FCC4F801FCC7F83D" box="[811,878,2010,2036]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="50" unit="km" value="40.0">40km</quantity>
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S Puerto Ayacucho, at El Tobogán, Coromoto.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31EC3E4AE5EFFC0FF2AFF4CFDC9FEB3" pageId="6" pageNumber="51" type="diagnosis">
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<paragraph id="8BBB906FAE5EFFC0FF2AFF4CFDC9FEB3" blockId="6.[151,1436,151,1698]" pageId="6" pageNumber="51">
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<emphasis id="B9704C7DAE5EFFC0FF2AFF4CFEECFF78" bold="true" box="[197,325,151,177]" pageId="6" pageNumber="51">Diagnosis.</emphasis>
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This species is smaller than
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<taxonomicName id="4C04EBECAE5EFFC0FD40FF43FCC3FF78" box="[687,874,152,177]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="toboganensis">
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<emphasis id="B9704C7DAE5EFFC0FD40FF43FCC3FF78" box="[687,874,152,177]" italics="true" pageId="6" pageNumber="51">F. toboganensis</emphasis>
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</taxonomicName>
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(TL <
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<quantity id="4CFC3D8AAE5EFFC0FC25FF43FB88FF7B" box="[970,1057,152,178]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="51" unit="mm" value="1.4">1.4mm</quantity>
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), but is similar in size (though slightly larger) and external appearance to
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<emphasis id="B9704C7DAE5EFFC0FD78FF1BFCA5FF10" box="[663,780,192,217]" italics="true" pageId="6" pageNumber="51">F. o r n a t u s</emphasis>
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.
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<taxonomicName id="4C04EBECAE5EFFC0FCF0FF1BFBB0FF10" box="[799,1049,192,217]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="wheeleri">
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<emphasis id="B9704C7DAE5EFFC0FCF0FF1BFBB0FF10" box="[799,1049,192,217]" italics="true" pageId="6" pageNumber="51">Fontidessus wheeleri</emphasis>
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</taxonomicName>
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has the prosternal process elongate with the lateral margins slightly concave and tapered to sharply pointed apex (
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<figureCitation id="133F8CEAAE5EFFC0FBB5FF33FB08FECB" box="[1114,1185,232,258]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Fontidessus species. 1 – 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 – 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 – 4. Scale bar b = 0.1 mm for Figs 5 – 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="6" pageNumber="51">Fig. 7</figureCitation>
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). The pale regions on the elytron are brighter yellow and more clearly demarcated in
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<emphasis id="B9704C7DAE5EFFC0FC6BFECBFBAAFEE0" box="[900,1027,272,297]" italics="true" pageId="6" pageNumber="51">F. w h e e l e r i</emphasis>
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than in
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<taxonomicName id="4C04EBECAE5EFFC0FB86FECBFB49FEE0" box="[1129,1248,272,297]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="ornatus">
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<emphasis id="B9704C7DAE5EFFC0FB86FECBFB49FEE0" box="[1129,1248,272,297]" italics="true" pageId="6" pageNumber="51">F. ornatus</emphasis>
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</taxonomicName>
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which have the elytral maculae more pale and diffuse. The male genitalia are diagnostic with the median lobe broad and deeply bifid in ventral aspect (
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<figureCitation id="133F8CEAAE5EFFC0FE14FEBBFDFBFEB3" box="[507,594,352,378]" captionStart="FIGURES 8 – 17" captionStartId="4.[151,269,1053,1077]" captionTargetBox="[469,1097,360,1021]" captionTargetId="figure@4.[463,1103,354,1031]" captionTargetPageId="4" captionText="FIGURES 8 – 17. Fontidessus species. 8 – 10) F. toboganensis, male genitalia, 8) median lobe, dorsal aspect, 9) median lobe, right lateral aspect, 10) right lateral lobe, right lateral aspect. 11 – 13) F. ornatus, male genitalia, 11) median lobe and left lateral lobe, dorsal aspect, 12) median lobe, right lateral aspect, 13) right lateral lobe, right lateral aspect. 14 – 16) F. wheeleri, male genitalia, 14) median lobe and left lateral lobe, dorsal aspect, 15) median lobe, left lateral aspect, 16) right lateral lobe, right lateral aspect. 17) F. toboganensis, female genitalia, ventral aspect. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/183128/files/figure.png" pageId="6" pageNumber="51">Fig. 14</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C31EC3E4AE5EFFC0FF2AFE5CFEFEFAF3" pageId="6" pageNumber="51" type="description">
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<paragraph id="8BBB906FAE5EFFC0FF2AFE5CFDE9FDD3" blockId="6.[151,1436,151,1698]" pageId="6" pageNumber="51">
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<emphasis id="B9704C7DAE5EFFC0FF2AFE5CFD8CFE68" bold="true" box="[197,549,391,417]" pageId="6" pageNumber="51">Description. Measurements.</emphasis>
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TL =
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<quantity id="4CFC3D8AAE5EFFC0FD9BFE53FCACFE6B" box="[628,773,392,418]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="6" pageNumber="51" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.2–1.3 mm</quantity>
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, GW =
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<quantity id="4CFC3D8AAE5EFFC0FC8AFE53FC5EFE6B" box="[869,1015,392,418]" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="6" pageNumber="51" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.6–0.7 mm</quantity>
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, PW =
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<quantity id="4CFC3D8AAE5EFFC0FBBDFE53FB4DFE6B" box="[1106,1252,392,418]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="6" pageNumber="51" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.5–0.6 mm</quantity>
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, HW =
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<quantity id="4CFC3D8AAE5EFFC0FAABFE53FF61FE03" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="6" pageNumber="51" unit="mm" value="0.45" valueMax="0.5" valueMin="0.4">0.4–0.5 mm</quantity>
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, EW =
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<quantity id="4CFC3D8AAE5EFFC0FECBFE6BFE1DFE03" box="[292,436,432,458]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="6" pageNumber="51" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 mm</quantity>
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, TL/GW = 1.7–1.8, HW/EW = 1.6–1.7. Body (
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<figureCitation id="133F8CEAAE5EFFC0FC05FE6BFB98FE03" box="[1002,1073,432,458]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Fontidessus species. 1 – 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 – 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 – 4. Scale bar b = 0.1 mm for Figs 5 – 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="6" pageNumber="51">Fig. 3</figureCitation>
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) oval, elongate; lateral outline slightly continuous between pronotum and elytron; lateral margins of pronotum gently curved; lateral margins of elytron evenly and gently curved.
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</paragraph>
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<paragraph id="8BBB906FAE5EFFC0FF29FDF3FD0AFCFB" blockId="6.[151,1436,151,1698]" pageId="6" pageNumber="51">
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<emphasis id="B9704C7DAE5EFFC0FF29FDF3FEEEFD88" box="[198,327,552,577]" italics="true" pageId="6" pageNumber="51">Coloration</emphasis>
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. Head yellow brown, brown along posterior margins and medial margins of eyes; pronotum broadly brown along posterior margin, narrowly dark brown along anterior margin; elytron brown with the following yellow maculae: 1) a basal macula in an irregular band extending posteriorly along suture at medial margin, 2) a lateral longitudinal macula posterior to humeral angle, broader anteriorly and extending posteriorly in narrow band to near apex, 3) a subtriangular subapical macula (
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<figureCitation id="133F8CEAAE5EFFC0FC34FD13FB8AFD2B" box="[987,1059,712,738]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Fontidessus species. 1 – 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 – 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 – 4. Scale bar b = 0.1 mm for Figs 5 – 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="6" pageNumber="51">Fig. 3</figureCitation>
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). Ventral surfaces of thorax and abdomen dark brown except prosternum, prosternal process, propleuron and pronotal epipleuron yellowbrown; appendages yellow to yellow-brown.
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</paragraph>
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<paragraph id="8BBB906FAE5EFFC0FF29FC9BFBBFFBBB" blockId="6.[151,1436,151,1698]" pageId="6" pageNumber="51">
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<emphasis id="B9704C7DAE5EFFC0FF29FC9BFE73FC90" box="[198,474,832,857]" italics="true" pageId="6" pageNumber="51">Sculpture and structure</emphasis>
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. Head with very fine, inconspicuous, irregular punctation, surface between punctures shiny, without distinct microsculpture; eyes medium in size (
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<figureCitation id="133F8CEAAE5EFFC0FC40FCB3FC5EFC4B" box="[943,1015,872,898]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Fontidessus species. 1 – 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 – 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 – 4. Scale bar b = 0.1 mm for Figs 5 – 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="6" pageNumber="51">Fig. 3</figureCitation>
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, HW/EW = 1.6–1.7). Pronotal surface similar to that of head, with posterior surface with fine microsculpture; with posterior angles obtuse; lateral bead slender, of even width throughout; pronotal striae finely incised, extending nearly 1/2 distance across pronotum (
|
||
<figureCitation id="133F8CEAAE5EFFC0FE80FC3BFE10FC33" box="[367,441,992,1018]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Fontidessus species. 1 – 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 – 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 – 4. Scale bar b = 0.1 mm for Figs 5 – 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="6" pageNumber="51">Fig. 3</figureCitation>
|
||
). Elytron with anterolateral angle obtuse, not extended anteriorly (
|
||
<figureCitation id="133F8CEAAE5EFFC0FB0DFC3BFA85FC33" box="[1250,1324,992,1018]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Fontidessus species. 1 – 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 – 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 – 4. Scale bar b = 0.1 mm for Figs 5 – 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="6" pageNumber="51">Fig. 3</figureCitation>
|
||
); surface similar to pronotum, with microsculpture indistinct. Prosternal process elongate and slender, lateral margins slightly concave, apex of process rounded (
|
||
<figureCitation id="133F8CEAAE5EFFC0FD7BFBEBFD75FB83" box="[660,732,1072,1098]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Fontidessus species. 1 – 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 – 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 – 4. Scale bar b = 0.1 mm for Figs 5 – 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="6" pageNumber="51">Fig. 7</figureCitation>
|
||
); metacoxal process with lateral lobe small. Pro- and mesotarsi relatively narrow in both male and female, but slightly broader in male.
|
||
</paragraph>
|
||
<paragraph id="8BBB906FAE5EFFC0FF29FB5BFEFEFAF3" blockId="6.[151,1436,151,1698]" pageId="6" pageNumber="51">
|
||
<emphasis id="B9704C7DAE5EFFC0FF29FB5BFEDAFB50" box="[198,371,1152,1177]" italics="true" pageId="6" pageNumber="51">Male genitalia</emphasis>
|
||
. Median lobe in ventral aspect very broad, deeply bifid, with branches broad, subapically strongly constricted on lateral margins, apices sharply pointed (
|
||
<figureCitation id="133F8CEAAE5EFFC0FC62FB73FC4FFB0B" box="[909,998,1192,1218]" captionStart="FIGURES 8 – 17" captionStartId="4.[151,269,1053,1077]" captionTargetBox="[469,1097,360,1021]" captionTargetId="figure@4.[463,1103,354,1031]" captionTargetPageId="4" captionText="FIGURES 8 – 17. Fontidessus species. 8 – 10) F. toboganensis, male genitalia, 8) median lobe, dorsal aspect, 9) median lobe, right lateral aspect, 10) right lateral lobe, right lateral aspect. 11 – 13) F. ornatus, male genitalia, 11) median lobe and left lateral lobe, dorsal aspect, 12) median lobe, right lateral aspect, 13) right lateral lobe, right lateral aspect. 14 – 16) F. wheeleri, male genitalia, 14) median lobe and left lateral lobe, dorsal aspect, 15) median lobe, left lateral aspect, 16) right lateral lobe, right lateral aspect. 17) F. toboganensis, female genitalia, ventral aspect. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/183128/files/figure.png" pageId="6" pageNumber="51">Fig. 14</figureCitation>
|
||
); in lateral broad at base, irregularly narrowed to sharply pointed apex (
|
||
<figureCitation id="133F8CEAAE5EFFC0FDD4FB0BFD3DFB23" box="[571,660,1232,1258]" captionStart="FIGURES 8 – 17" captionStartId="4.[151,269,1053,1077]" captionTargetBox="[469,1097,360,1021]" captionTargetId="figure@4.[463,1103,354,1031]" captionTargetPageId="4" captionText="FIGURES 8 – 17. Fontidessus species. 8 – 10) F. toboganensis, male genitalia, 8) median lobe, dorsal aspect, 9) median lobe, right lateral aspect, 10) right lateral lobe, right lateral aspect. 11 – 13) F. ornatus, male genitalia, 11) median lobe and left lateral lobe, dorsal aspect, 12) median lobe, right lateral aspect, 13) right lateral lobe, right lateral aspect. 14 – 16) F. wheeleri, male genitalia, 14) median lobe and left lateral lobe, dorsal aspect, 15) median lobe, left lateral aspect, 16) right lateral lobe, right lateral aspect. 17) F. toboganensis, female genitalia, ventral aspect. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/183128/files/figure.png" pageId="6" pageNumber="51">Fig. 15</figureCitation>
|
||
). Lateral lobe narrow at base, medially expanded, apical portion parallel-sided to slightly expanded, broadly rounded apex, with small series of elongate setae along dorsal margin (
|
||
<figureCitation id="133F8CEAAE5EFFC0FF16FAFBFEE0FAF3" box="[249,329,1312,1338]" captionStart="FIGURES 8 – 17" captionStartId="4.[151,269,1053,1077]" captionTargetBox="[469,1097,360,1021]" captionTargetId="figure@4.[463,1103,354,1031]" captionTargetPageId="4" captionText="FIGURES 8 – 17. Fontidessus species. 8 – 10) F. toboganensis, male genitalia, 8) median lobe, dorsal aspect, 9) median lobe, right lateral aspect, 10) right lateral lobe, right lateral aspect. 11 – 13) F. ornatus, male genitalia, 11) median lobe and left lateral lobe, dorsal aspect, 12) median lobe, right lateral aspect, 13) right lateral lobe, right lateral aspect. 14 – 16) F. wheeleri, male genitalia, 14) median lobe and left lateral lobe, dorsal aspect, 15) median lobe, left lateral aspect, 16) right lateral lobe, right lateral aspect. 17) F. toboganensis, female genitalia, ventral aspect. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/183128/files/figure.png" pageId="6" pageNumber="51">Fig 16</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31EC3E4AE5EFFC0FF2AFA9CFD99FA43" pageId="6" pageNumber="51" type="etymology">
|
||
<paragraph id="8BBB906FAE5EFFC0FF2AFA9CFD99FA43" blockId="6.[151,1436,151,1698]" pageId="6" pageNumber="51">
|
||
<emphasis id="B9704C7DAE5EFFC0FF2AFA9CFEFBFAA8" bold="true" box="[197,338,1351,1377]" pageId="6" pageNumber="51">Etymology.</emphasis>
|
||
This species is named in honor of my good friend and fellow coleopterist, Dr. Quentin D. Wheeler, Arizona State University.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31EC3E4AE5EFFC0FF2AFA4CFD5EFA13" pageId="6" pageNumber="51" type="distribution">
|
||
<paragraph id="8BBB906FAE5EFFC0FF2AFA4CFD5EFA13" blockId="6.[151,1436,151,1698]" pageId="6" pageNumber="51">
|
||
<emphasis id="B9704C7DAE5EFFC0FF2AFA4CFDA9FA78" bold="true" box="[197,512,1431,1457]" pageId="6" pageNumber="51">Distribution and habitat.</emphasis>
|
||
The species is known only from the
|
||
<typeStatus id="54BF2ECDAE5EFFC0FC54FA43FC44FA7B" box="[955,1005,1432,1458]" pageId="6" pageNumber="51">type</typeStatus>
|
||
locality. Label data indicate the following habitats; “seep,” and “bare rock streambed.”
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31EC3E4AE5EFFC0FF2AFA3CFE46F96B" pageId="6" pageNumber="51" type="materials_examined">
|
||
<paragraph id="8BBB906FAE5EFFC0FF2AFA3CFE46F96B" blockId="6.[151,1436,151,1698]" pageId="6" pageNumber="51">
|
||
<emphasis id="B9704C7DAE5EFFC0FF2AFA3CFE69F9C8" bold="true" box="[197,448,1511,1537]" pageId="6" pageNumber="51">Material examined.</emphasis>
|
||
<typeStatus id="54BF2ECDAE5EFFC0FE20FA33FDC3F9CB" box="[463,618,1512,1538]" pageId="6" pageNumber="51" type="holotype">HOLOTYPE</typeStatus>
|
||
in
|
||
<collectionCode id="ED1508AAAE5EFFC0FD72FA33FD5FF9CB" LSID="urn:lsid:biocol.org:col:34871" box="[669,758,1512,1538]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="51">USNM</collectionCode>
|
||
: male labeled, “
|
||
<collectingCountry id="F313D0FFAE5EFFC0FC2CFA33FBD0F9CB" box="[963,1145,1512,1538]" name="Venezuela" pageId="6" pageNumber="51">VENEZUELA</collectingCountry>
|
||
: T.F. Amazonas Puerto Ayacucho (
|
||
<quantity id="4CFC3D8AAE5EFFC0FEF5F9CBFECCF9E3" box="[282,357,1552,1578]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="51" unit="km" value="40.0">40km</quantity>
|
||
.S) at Tobogán,
|
||
<date id="FFBAB6AFAE5EFFC0FDF7F9CBFD35F9E3" box="[536,668,1552,1578]" pageId="6" pageNumber="51" value="1986-02-22">22Feb1986</date>
|
||
P.J. Spangler, Colln.#6/ Bare rock streambed in sunlight; beneath wet leaves at margin/
|
||
<typeStatus id="54BF2ECDAE5EFFC0FE74F9E3FD93F99B" box="[411,570,1592,1618]" pageId="6" pageNumber="51" type="holotype">HOLOTYPE</typeStatus>
|
||
:
|
||
<taxonomicName id="4C04EBECAE5EFFC0FDA9F9E3FC73F99B" authority="Miller, 2008" authorityName="Miller" authorityYear="2008" box="[582,986,1592,1618]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="wheeleri">
|
||
<emphasis id="B9704C7DAE5EFFC0FDA9F9E3FCE9F998" box="[582,832,1592,1617]" italics="true" pageId="6" pageNumber="51">Fontidessus wheeleri</emphasis>
|
||
Miller, 2008
|
||
</taxonomicName>
|
||
[red label with double black line bor- der].”
|
||
<typeStatus id="54BF2ECDAE5EFFC0FF0CF9BBFE27F9B3" box="[227,398,1632,1658]" pageId="6" pageNumber="51" type="paratype">PARATYPES</typeStatus>
|
||
, 1, same as
|
||
<typeStatus id="54BF2ECDAE5EFFC0FDCEF9BBFD2EF9B3" box="[545,647,1632,1658]" pageId="6" pageNumber="51" type="holotype">holotype</typeStatus>
|
||
except
|
||
<date id="FFBAB6AFAE5EFFC0FD0AF9BBFC05F9B3" box="[741,940,1632,1658]" pageId="6" pageNumber="51" value="1989-01-23">23 January 1989</date>
|
||
, seep at upper shelter, P.J. Spangler, R.A. Faitoute and C.B. Barr, colrs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |