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<mods:title id="9A8BAD9A2CDFADA5748BE8AB95504030">A review of the gekkotan lizards of Bénin, with the description of a new species of Hemidactylus (Squamata: Gekkonidae)</mods:title>
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<mods:namePart id="AFEFD98BBD5A6F86A12017872BDB3BBE">Tchibozo, Sévérin</mods:namePart>
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<mods:namePart id="82D75E0BA2B9DD255DFE5A6F745F4DC4">Lenglet, Georges</mods:namePart>
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<treatment id="03A9E573FFA8165C057FF8E27788FBAD" ID-DOI="http://doi.org/10.5281/zenodo.6255693" ID-GBIF-Taxon="119345730" ID-Zenodo-Dep="6255693" LSID="urn:lsid:plazi:treatment:03A9E573FFA8165C057FF8E27788FBAD" httpUri="http://treatment.plazi.org/id/03A9E573FFA8165C057FF8E27788FBAD" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<subSubSection id="C31A07EEFFA8165D057FF8E27617F89B" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8BBF5465FFA8165D057FF8E271F5F8E0" blockId="4.[264,714,1793,1819]" box="[264,714,1793,1819]" pageId="4" pageNumber="5">
<heading id="D0F7E309FFA8165D057FF8E271F5F8E0" bold="true" box="[264,714,1793,1819]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<taxonomicName id="4C002FE6FFA8165D057FF8E271F5F8E0" authority="Anderson, 1898" authorityName="Anderson" authorityYear="1898" box="[264,714,1793,1819]" class="Reptilia" family="Phyllodactylidae" genus="Ptyodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="ragazzi">
<emphasis id="B9748877FFA8165D057FF8E271F5F8E0" bold="true" box="[264,714,1793,1819]" pageId="4" pageNumber="5">
<emphasis id="B9748877FFA8165D057FF8E272C1F8E0" bold="true" box="[264,510,1793,1819]" italics="true" pageId="4" pageNumber="5">Ptyodactylus ragazzi</emphasis>
Anderson, 1898
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BBF5465FFA8165D057FF8A47617F89B" blockId="4.[264,1320,1863,1889]" box="[264,1320,1863,1889]" pageId="4" pageNumber="5">
<bibRefCitation id="EF912994FFA8165D057FF8A472DAF89A" author="Loveridge" box="[264,485,1863,1889]" pageId="4" pageNumber="20" refString="Loveridge, A. (1947) Revision of the African lizards of the family Gekkonidae. Bulletin of the Museum of Comparative Zoology, 98, 1 - 469, pls. 1 - 7." type="journal article" year="1947">Loveridge (1947)</bibRefCitation>
regarded
<taxonomicName id="4C002FE6FFA8165D0601F8A47032F89B" box="[630,781,1863,1888]" class="Reptilia" family="Phyllodactylidae" genus="Ptyodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
<emphasis id="B9748877FFA8165D0601F8A47032F89B" box="[630,781,1863,1888]" italics="true" pageId="4" pageNumber="5">Ptyodactylus</emphasis>
</taxonomicName>
from northern
<collectingCountry id="F31714F5FFA8165D079CF8A47719F89A" box="[1003,1062,1863,1889]" name="Togo" pageId="4" pageNumber="5">Togo</collectingCountry>
as referable to
<emphasis id="B9748877FFA8165D0162F8A47617F89B" box="[1301,1320,1863,1888]" italics="true" pageId="4" pageNumber="5">P.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C31A07EEFFA9165C057FFEFF7244FC26" pageId="5" pageNumber="6" type="reference_group">
<paragraph id="8BBF5465FFA9165C057FFEFF7037FC75" blockId="5.[264,1324,284,910]" pageId="5" pageNumber="6">
<emphasis id="B9748877FFA9165C057FFEFF712EFECE" box="[264,529,284,309]" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="4C002FE6FFA9165C057FFEFF72A5FECE" box="[264,410,284,309]" class="Reptilia" family="Phyllodactylidae" genus="Ptyodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="hasselquistii">hasselquistii</taxonomicName>
togoensis
</emphasis>
<bibRefCitation id="EF912994FFA9165C066EFEFF7187FECD" author="Tornier" box="[537,696,284,310]" pageId="5" pageNumber="20" refString="Tornier, G. (1901) Die Crocodile, Schildkroten und Eidechsen in Togo. Archiv fur Naturgeschichte, 1901 Beiheft, 65 - 88." type="journal article" year="1901">Tornier, 1901</bibRefCitation>
and referred
<emphasis id="B9748877FFA9165C0720FEFF70D6FECE" box="[855,1001,284,309]" italics="true" pageId="5" pageNumber="6">
P. h.
<taxonomicName id="4C002FE6FFA9165C07E4FEFF70D6FECE" box="[915,1001,284,309]" class="Reptilia" family="Phyllodactylidae" genus="Ptyodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="ragazzi">ragazzi</taxonomicName>
</emphasis>
, with its
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from
<collectingCountry id="F31714F5FFA9165C00A4FEFF7619FECD" box="[1235,1318,284,310]" name="Eritrea" pageId="5" pageNumber="6">Eritrea</collectingCountry>
, to the synonymy of
<emphasis id="B9748877FFA9165C0580FEA771F8FEA6" box="[503,711,324,349]" italics="true" pageId="5" pageNumber="6">
P. h.
<taxonomicName id="4C002FE6FFA9165C0643FEA771F8FEA6" authority="Donndorff. However, Heimes (1987)" authorityName="Donndorff. However, Heimes" authorityYear="1987" box="[564,711,324,349]" class="Reptilia" family="Phyllodactylidae" genus="Ptyodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="hasselquistii">hasselquistii</taxonomicName>
</emphasis>
Donndorff. However,
<bibRefCitation id="EF912994FFA9165C07A2FEA777B9FEA5" author="Heimes" box="[981,1158,324,350]" pageId="5" pageNumber="19" refString="Heimes, P. (1987) Beitrag zur Systematik der Facherfinger (Sauria: Gekkonidae: Ptyodactylus). Salamandra, 23, 212 - 235." type="journal article" year="1987">Heimes (1987)</bibRefCitation>
demonstrated that
<emphasis id="B9748877FFA9165C054BFE8F72FAFE7E" box="[316,453,364,389]" italics="true" pageId="5" pageNumber="6">
P.h.
<taxonomicName id="4C002FE6FFA9165C0518FE8F72FAFE7E" box="[367,453,364,389]" class="Reptilia" family="Phyllodactylidae" genus="Ptyodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="ragazzi">ragazzi</taxonomicName>
</emphasis>
and
<emphasis id="B9748877FFA9165C0677FE8F719DFE7E" box="[512,674,364,389]" italics="true" pageId="5" pageNumber="6">P.h. togoensis</emphasis>
were synonyms and resurrected the older name for the subspecies ranging across the central Sahara and the Sahel. He considered the range in West Africa to span from
<collectingCountry id="F31714F5FFA9165C0633FE5F71DBFE2D" box="[580,740,444,470]" name="Ivory Coast" pageId="5" pageNumber="6">Côte dIvoire</collectingCountry>
to
<collectingCountry id="F31714F5FFA9165C0778FE5F70B3FE2D" box="[783,908,444,470]" name="Cameroon" pageId="5" pageNumber="6">Cameroon</collectingCountry>
, but did not examine any material from
<collectingCountry id="F31714F5FFA9165C053EFE0772ACFE05" box="[329,403,484,510]" name="Benin" pageId="5" pageNumber="6">Bénin</collectingCountry>
.
<bibRefCitation id="EF912994FFA9165C05E9FE0771A2FE05" box="[414,669,484,510]" pageId="5" pageNumber="20" refString="Schleich, H. - H., Kastle, W. &amp; Kabisch, K. (1996) Amphibians and Reptiles of North Africa. Koeltz Scientific, Koenigstein, 627 pp." type="book">
Schleich
<emphasis id="B9748877FFA9165C067BFE077178FE06" box="[524,583,484,509]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
(1996)
</bibRefCitation>
subsequently accorded full species status to
<taxonomicName id="4C002FE6FFA9165C00D8FE07761CFE06" box="[1199,1315,484,509]" class="Reptilia" family="Phyllodactylidae" genus="Ptyodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="ragazzi">
<emphasis id="B9748877FFA9165C00D8FE07761CFE06" box="[1199,1315,484,509]" italics="true" pageId="5" pageNumber="6">P. ragazzi</emphasis>
</taxonomicName>
, but without explicit justification.
<bibRefCitation id="EF912994FFA9165C06D0FDEF70A0FDDD" box="[679,927,524,550]" pageId="5" pageNumber="18" refString="Baha El Din, S. (1999) On the specific status of Ptyodactylus ragazzii Anderson, 1898 and its occurrence in Egypt (Squamata: Sauria: Gekkonidae). Herpetozoa, 12, 39 - 43." type="journal article">Baha El Din (1999)</bibRefCitation>
provided evidence of sympatry between
<taxonomicName id="4C002FE6FFA9165C0501FDD77114FDB6" box="[374,555,564,589]" class="Reptilia" family="Phyllodactylidae" genus="Ptyodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="hasselquistii">
<emphasis id="B9748877FFA9165C0501FDD77114FDB6" box="[374,555,564,589]" italics="true" pageId="5" pageNumber="6">P. hasselquistii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C002FE6FFA9165C061AFDD771DBFDB6" box="[621,740,564,589]" class="Reptilia" family="Phyllodactylidae" genus="Ptyodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="ragazzi">
<emphasis id="B9748877FFA9165C061AFDD771DBFDB6" box="[621,740,564,589]" italics="true" pageId="5" pageNumber="6">P. ragazzi</emphasis>
</taxonomicName>
in southeastern
<collectingCountry id="F31714F5FFA9165C07C4FDD770C3FDB5" box="[947,1020,564,590]" name="Egypt" pageId="5" pageNumber="6">Egypt</collectingCountry>
, demonstrating that they should indeed be considered distinct species. The species is widespread in northern areas of
<collectingCountry id="F31714F5FFA9165C055CFD6772BCFD65" box="[299,387,644,670]" name="Nigeria" pageId="5" pageNumber="6">Nigeria</collectingCountry>
(
<bibRefCitation id="EF912994FFA9165C05E2FD677108FD65" author="Dunger" box="[405,567,644,670]" pageId="5" pageNumber="19" refString="Dunger, G. T. (1968) The lizards and snakes of Nigeria. Part 4. Geckos of Nigeria. The Nigerian Field, 33, 18 - 47." type="journal article" year="1968">Dunger 1968</bibRefCitation>
;
<bibRefCitation id="EF912994FFA9165C0631FD6771E9FD65" author="Butler" box="[582,726,644,670]" pageId="5" pageNumber="19" refString="Butler, J. A. (1986) Notes on collections and observations of lizards in Nigeria. The Nigerian Field, 51, 85 - 94." type="journal article" year="1986">Butler 1986</bibRefCitation>
) and
<collectingCountry id="F31714F5FFA9165C076AFD6770A8FD65" box="[797,919,644,670]" name="Cameroon" pageId="5" pageNumber="6">Cameroon</collectingCountry>
(
<bibRefCitation id="EF912994FFA9165C07DEFD67775CFD65" author="LeBreton" box="[937,1123,644,670]" pageId="5" pageNumber="20" refString="LeBreton, M. (1999) A Working Checklist of the Herpetofauna of Cameroon, with Localities for Species Occurring in Southern Cameroon and a List of Herpetofauna for the Dja Faunal Reserve. Netherlands Committee for IUCN, Amsterdam, 139 pp." type="book" year="1999">LeBreton 1999</bibRefCitation>
).
<bibRefCitation id="EF912994FFA9165C000CFD677614FD65" author="Dunger" box="[1147,1323,644,670]" pageId="5" pageNumber="19" refString="Dunger, G. T. (1968) The lizards and snakes of Nigeria. Part 4. Geckos of Nigeria. The Nigerian Field, 33, 18 - 47." type="journal article" year="1968">Dunger (1968)</bibRefCitation>
considered
<taxonomicName id="4C002FE6FFA9165C05E4FD4F7135FD3E" box="[403,522,684,709]" class="Reptilia" family="Phyllodactylidae" genus="Ptyodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="ragazzi">
<emphasis id="B9748877FFA9165C05E4FD4F7135FD3E" box="[403,522,684,709]" italics="true" pageId="5" pageNumber="6">P. ragazzi</emphasis>
</taxonomicName>
to be a rupicolous or crevice­dwelling savanna species, although
<bibRefCitation id="EF912994FFA9165C057FFD377298FD15" author="Butler" box="[264,423,724,750]" pageId="5" pageNumber="19" refString="Butler, J. A. (1986) Notes on collections and observations of lizards in Nigeria. The Nigerian Field, 51, 85 - 94." type="journal article" year="1986">Butler (1986)</bibRefCitation>
collected it at the Jebba crossing of the
<collectingCountry id="F31714F5FFA9165C07F0FD3770F5FD15" box="[903,970,724,750]" name="Niger" pageId="5" pageNumber="6">Niger</collectingCountry>
River in western
<collectingCountry id="F31714F5FFA9165C00E9FD3777C9FD15" box="[1182,1270,724,750]" name="Nigeria" pageId="5" pageNumber="6">Nigeria</collectingCountry>
in a small shed. There appear to be no prior published records of this species from
<collectingCountry id="F31714F5FFA9165C00DFFD1F77CDFCED" box="[1192,1266,764,790]" name="Benin" pageId="5" pageNumber="6">Bénin</collectingCountry>
. Our material was collected on boulders (
<figureCitation id="133B48E0FFA9165C06CFFCC7703FFCC5" box="[696,768,804,830]" captionStart="FIGURE 3" captionStartId="6.[264,368,1469,1493]" captionTargetBox="[333,1252,332,1444]" captionTargetId="figure@6.[327,1252,326,1445]" captionTargetPageId="6" captionText="FIGURE 3. Boulder habitat of Ptyodactylus ragazzi at Chutes de Koudou, Parc National du W du Bénin, Département de lAlibori." httpUri="https://zenodo.org/record/273467/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
) or abandoned buildings adjacent to boulders, confirming earlier ecological observations from
<collectingCountry id="F31714F5FFA9165C073DFCAF7099FC9D" box="[842,934,844,870]" name="Nigeria" pageId="5" pageNumber="6">Nigeria</collectingCountry>
. The largest individual collected (IRSNB 17151­3) measured
<quantity id="4CF8F980FFA9165C062EFC9771FDFC75" box="[601,706,884,910]" metricMagnitude="-2" metricUnit="m" metricValue="7.77" pageId="5" pageNumber="6" unit="mm" value="77.7">77.7 mm</quantity>
SVL.
</paragraph>
<paragraph id="8BBF5465FFA9165C057FFC277244FC26" blockId="5.[264,1322,964,1110]" box="[264,379,964,989]" pageId="5" pageNumber="6">
<emphasis id="B9748877FFA9165C057FFC277244FC26" box="[264,379,964,989]" italics="true" pageId="5" pageNumber="6">Localities</emphasis>
</paragraph>
</subSubSection>
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<paragraph id="8BBF5465FFA9165C054FFC0F7788FBAD" blockId="5.[264,1322,964,1110]" pageId="5" pageNumber="6">
Chutes de Koudou, Parc National du W du
<collectingCountry id="F31714F5FFA9165C0745FC0F7044FBFD" box="[818,891,1004,1030]" name="Benin" pageId="5" pageNumber="6">Bénin</collectingCountry>
, Département de lAlibori (
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,
<geoCoordinate id="EE3432A2FFA9165C057FFBF77251FBD5" box="[264,366,1044,1070]" direction="east" orientation="longitude" pageId="5" pageNumber="6" precision="925" value="2.3">02°18E</geoCoordinate>
): IRSNB 17151­1, 17151­2, 17151­3; Parc National du W du
<collectingCountry id="F31714F5FFA9165C0048FBF777B7FBD5" box="[1087,1160,1044,1070]" name="Benin" pageId="5" pageNumber="6">Bénin</collectingCountry>
, Département de lAlibori: IRSNB 17154; without specific locality: IRSNB 17171­1, 17171­4.
</paragraph>
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