405 lines
49 KiB
XML
405 lines
49 KiB
XML
<document id="807E264939386BC4A0D23EB3419247B6" ID-CLB-Dataset="50634" ID-DOI="10.5281/zenodo.273464" ID-GBIF-Dataset="3301c485-76c8-496d-93fd-a360f691a89b" ID-ISSN="11755326" ID-Zenodo-Dep="273464" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459764780349" checkinUser="plazi" docAuthor="Bauer, Aaron M., Tchibozo, Sévérin, Pauwels, Olivier S. G. & Lenglet, Georges" docDate="2006" docId="03A9E573FFA61656057FFBBD7109FC9D" docLanguage="en" docName="zt01242p020.pdf" docOrigin="Zootaxa 1242" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Hemidactylus beninensis Bauer, Tchibozo, Pauwels & Lenglet, 2006, sp. nov." docType="treatment" docVersion="8" lastPageNumber="16" masterDocId="FF909D0BFFAC16590477FFE3733FFFFB" masterDocTitle="A review of the gekkotan lizards of Bénin, with the description of a new species of Hemidactylus (Squamata: Gekkonidae)" masterLastPageNumber="20" masterPageNumber="1" pageNumber="11" updateTime="1698222181484" updateUser="ExternalLinkService">
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<mods:titleInfo id="EBC2C6696200EE4643D8D745E7C65E09">
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<mods:title id="24F1506419EC8FA5302633E63727114C">A review of the gekkotan lizards of Bénin, with the description of a new species of Hemidactylus (Squamata: Gekkonidae)</mods:title>
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<mods:namePart id="919CAECC97F38E331CFB646551147DDE">Bauer, Aaron M.</mods:namePart>
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<mods:namePart id="62F3BDDDC2003AD499EB9136C5A6B20A">Tchibozo, Sévérin</mods:namePart>
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<mods:namePart id="A2625FD75658B2CA62D9AD9BC0A18BAE">Pauwels, Olivier S. G.</mods:namePart>
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<mods:namePart id="F8796CED516E26392717BBC6F0E5157E">Lenglet, Georges</mods:namePart>
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<mods:typeOfResource id="62EAD5E95793C35E681C05BD08FD6F9D">text</mods:typeOfResource>
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<mods:title id="3347C3A18CD4BDC7DB575053EBF2E4C8">Zootaxa</mods:title>
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<mods:date id="CCCE3A7F8F4664C133498A11784C4A94">2006</mods:date>
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<mods:detail id="051D5C3C0E712C703B9DF4321560C5FD" type="volume">
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<treatment id="03A9E573FFA61656057FFBBD7109FC9D" ID-DOI="http://doi.org/10.5281/zenodo.6255717" ID-GBIF-Taxon="119345746" ID-Zenodo-Dep="6255717" LSID="urn:lsid:plazi:treatment:03A9E573FFA61656057FFBBD7109FC9D" httpUri="http://treatment.plazi.org/id/03A9E573FFA61656057FFBBD7109FC9D" lastPageId="15" lastPageNumber="16" pageId="10" pageNumber="11">
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<subSubSection id="C31A07EEFFA61653057FFBBD72ACFB60" pageId="10" pageNumber="11" type="nomenclature">
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<paragraph id="8BBF5465FFA61653057FFBBD71A4FB83" blockId="10.[264,667,1118,1179]" box="[264,667,1118,1144]" pageId="10" pageNumber="11">
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<heading id="D0F7E309FFA61653057FFBBD71A4FB83" bold="true" box="[264,667,1118,1144]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
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<emphasis id="B9748877FFA61653057FFBBD71A4FB83" bold="true" box="[264,667,1118,1144]" pageId="10" pageNumber="11">
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<taxonomicName id="4C002FE6FFA61653057FFBBD7109FB83" box="[264,566,1118,1144]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="beninensis" status="sp. nov.">
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<emphasis id="B9748877FFA61653057FFBBD7109FB83" bold="true" box="[264,566,1118,1144]" italics="true" pageId="10" pageNumber="11">Hemidactylus beninensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247350CFFA61653064AFBBD71A4FB83" box="[573,667,1118,1144]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="8BBF5465FFA61653057FFB6272ACFB60" blockId="10.[264,667,1118,1179]" box="[264,403,1153,1179]" pageId="10" pageNumber="11">
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<figureCitation id="133B48E0FFA61653057FFB6272ACFB60" box="[264,403,1153,1179]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="12.[264,367,1446,1470]" captionStartId-1="13.[264,368,1601,1625]" captionTargetBox-0="[267,1319,646,1403]" captionTargetBox-1="[294,1278,797,1593]" captionTargetId-0="figure@12.[267,1319,646,1422]" captionTargetId-1="figure@13.[292,1311,797,1597]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="FIGURE 4. Dorsal view of the holotype of Hemidactylus beninensis sp. nov. (IRSNB 2617). Scale bar = 20 mm." captionText-1="FIGURE 5. Dorsal view of the paratype of Hemidactylus beninensis sp. nov. (IRSNB 2618). Scale bar = 20 mm." httpUri-0="https://zenodo.org/record/273469/files/figure.png" httpUri-1="https://zenodo.org/record/273470/files/figure.png" pageId="10" pageNumber="11">Figures 4–5</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C31A07EEFFA61653057FFB327249FA70" pageId="10" pageNumber="11" type="materials_examined">
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<paragraph id="8BBF5465FFA61653057FFB32724BFB11" blockId="10.[264,1324,1233,1419]" box="[264,372,1233,1258]" pageId="10" pageNumber="11">
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<emphasis id="B9748877FFA61653057FFB32724BFB11" box="[264,372,1233,1258]" italics="true" pageId="10" pageNumber="11">
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<typeStatus id="54BBEAC7FFA61653057FFB32724BFB11" box="[264,372,1233,1258]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
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</emphasis>
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</paragraph>
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<paragraph id="8BBF5465FFA61653054FFB1A7249FA70" blockId="10.[264,1324,1233,1419]" pageId="10" pageNumber="11">
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Institut Royal des Sciences Naturelles de
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<collectingCountry id="F31714F5FFA616530754FB1A70B2FAE8" box="[803,909,1273,1299]" name="Belgium" pageId="10" pageNumber="11">Belgique</collectingCountry>
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(
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<collectionCode id="ED11CCA0FFA6165307EAFB1A70C8FAE8" box="[925,1015,1273,1299]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="10" pageNumber="11">IRSNB</collectionCode>
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) 2617, adult male;
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<collectingCountry id="F31714F5FFA6165300A9FB1A7618FAE8" box="[1246,1319,1273,1299]" name="Benin" pageId="10" pageNumber="11">Bénin</collectingCountry>
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, Département des Collines, Collines de DassaZoumè, (
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<geoCoordinate id="EE3432A2FFA6165307D4FAC27733FAC0" box="[931,1036,1313,1339]" direction="north" orientation="latitude" pageId="10" pageNumber="11" precision="925" value="7.75">07°45’N</geoCoordinate>
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,
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<geoCoordinate id="EE3432A2FFA61653006CFAC277BEFAC0" box="[1051,1153,1313,1339]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="925" value="2.1666667">02°10’E</geoCoordinate>
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); collected by Sévérin Tchibozo,
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<date id="FFBE72A5FFA61653059DFAAA71AFFA98" box="[490,656,1353,1379]" pageId="10" pageNumber="11" value="2005-06-13">13 June 2005</date>
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.
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<emphasis id="B9748877FFA6165306DFFAAA7025FA98" bold="true" box="[680,794,1353,1379]" pageId="10" pageNumber="11">
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<typeStatus id="54BBEAC7FFA6165306DFFAAA7025FA98" box="[680,794,1353,1379]" pageId="10" pageNumber="11" type="paratype">Paratype</typeStatus>
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</emphasis>
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.
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<collectionCode id="ED11CCA0FFA61653075CFAAA70BDFA98" box="[811,898,1353,1379]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="10" pageNumber="11">IRSNB</collectionCode>
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2618, subadult male; same data as
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<typeStatus id="54BBEAC7FFA61653057FFA92724DFA70" box="[264,370,1393,1419]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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||
</subSubSection>
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||
<paragraph id="8BBF5465FFA61653057FFA227269F9D0" pageId="10" pageNumber="11">
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<table id="F900A6C5FFA6E9A6057FFA227615F9D0" box="[264,1322,1473,1579]" gridcols="14" gridrows="3" pageId="10" pageNumber="11">
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<tr id="35305627FFA6E9A6057FFA227615FA21" box="[264,1322,1473,1498]" gridrow="0" pageId="10" pageNumber="11" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
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<th id="76E13F5BFFA6E9A6057FFA2272F6FA21" box="[264,457,1473,1498]" gridcol="0" gridrow="0" pageId="10" pageNumber="11">
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<emphasis id="B9748877FFA61653057FFA2272BAFA21" box="[264,389,1473,1498]" italics="true" pageId="10" pageNumber="11">Etymology</emphasis>
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||
</th>
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||
</tr>
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<tr id="35305627FFA6E9A6057FFA0A7615F9F8" box="[264,1322,1513,1539]" gridrow="1" pageId="10" pageNumber="11">
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<th id="76E13F5BFFA6E9A6057FFA0A72F6F9F8" box="[264,457,1513,1539]" gridcol="0" gridrow="1" pageId="10" pageNumber="11">The specific</th>
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<td id="76E13F5BFFA6E9A605A6FA0A711EF9F8" box="[465,545,1513,1539]" gridcol="1" gridrow="1" pageId="10" pageNumber="11">epithet</td>
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<td id="76E13F5BFFA6E9A6065EFA0A7153F9F8" box="[553,620,1513,1539]" gridcol="2" gridrow="1" pageId="10" pageNumber="11">refers</td>
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||
<td id="76E13F5BFFA6E9A60603FA0A71B4F9F8" box="[628,651,1513,1539]" gridcol="3" gridrow="1" pageId="10" pageNumber="11">to</td>
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<td id="76E13F5BFFA6E9A606E4FA0A7188F9F8" box="[659,695,1513,1539]" gridcol="4" gridrow="1" pageId="10" pageNumber="11">the</td>
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<td id="76E13F5BFFA6E9A606C8FA0A7000F9F8" box="[703,831,1513,1539]" gridcol="5" gridrow="1" pageId="10" pageNumber="11">occurrence</td>
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<td id="76E13F5BFFA6E9A6073FFA0A705FF9F8" box="[840,864,1513,1539]" gridcol="6" gridrow="1" pageId="10" pageNumber="11">of</td>
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<td id="76E13F5BFFA6E9A6071FFA0A70B3F9F8" box="[872,908,1513,1539]" gridcol="7" gridrow="1" pageId="10" pageNumber="11">the</td>
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<td id="76E13F5BFFA6E9A607E2FA0A70F9F9F8" box="[917,966,1513,1539]" gridcol="8" gridrow="1" pageId="10" pageNumber="11">new</td>
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<td id="76E13F5BFFA6E9A607B9FA0A771CF9F8" box="[974,1059,1513,1539]" gridcol="9" gridrow="1" pageId="10" pageNumber="11">species</td>
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<td id="76E13F5BFFA6E9A6005CFA0A777DF9F8" box="[1067,1090,1513,1539]" gridcol="10" gridrow="1" pageId="10" pageNumber="11">in</td>
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||
<td id="76E13F5BFFA6E9A6003DFA0A7751F9F8" box="[1098,1134,1513,1539]" gridcol="11" gridrow="1" pageId="10" pageNumber="11">the</td>
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<td id="76E13F5BFFA6E9A60009FA0A763AF9F8" box="[1150,1285,1513,1539]" gridcol="12" gridrow="1" pageId="10" pageNumber="11">République</td>
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<td id="76E13F5BFFA6E9A6017AFA0A7615F9F8" box="[1293,1322,1513,1539]" gridcol="13" gridrow="1" pageId="10" pageNumber="11">du</td>
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</tr>
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||
<tr id="35305627FFA6E9A6057FF9F27615F9D0" box="[264,1322,1553,1579]" gridrow="2" pageId="10" pageNumber="11" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
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<th id="76E13F5BFFA6E9A6057FF9F272F6F9D0" box="[264,457,1553,1579]" gridcol="0" gridrow="2" pageId="10" pageNumber="11">Bénin.</th>
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||
</tr>
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||
</table>
|
||
</paragraph>
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||
<subSubSection id="C31A07EEFFA61652057FF96A7140FB85" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="diagnosis">
|
||
<paragraph id="8BBF5465FFA61653057FF96A7240F959" blockId="10.[264,1323,1673,1899]" box="[264,383,1673,1698]" pageId="10" pageNumber="11">
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||
<emphasis id="B9748877FFA61653057FF96A7240F959" box="[264,383,1673,1698]" italics="true" pageId="10" pageNumber="11">Diagnosis</emphasis>
|
||
</paragraph>
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||
<paragraph id="8BBF5465FFA61652054FF95270FAFE7D" blockId="10.[264,1323,1673,1899]" lastBlockId="11.[264,1324,284,1150]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
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A moderatesized
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<taxonomicName id="4C002FE6FFA616530679F9527190F931" box="[526,687,1713,1738]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
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||
<emphasis id="B9748877FFA616530679F9527190F931" box="[526,687,1713,1738]" italics="true" pageId="10" pageNumber="11">Hemidactylus</emphasis>
|
||
</taxonomicName>
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||
, snoutvent length at least
|
||
<quantity id="4CF8F980FFA61653079CF952777DF930" box="[1003,1090,1713,1739]" metricMagnitude="-2" metricUnit="m" metricValue="6.8" pageId="10" pageNumber="11" unit="mm" value="68.0">68 mm</quantity>
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||
. One pair of greatly enlarged postmentals, in contact behind mental. 14–16 rows of relatively small tubercles. 49 scale rows across venter between lowest rows of tubercles. Six to seven enlarged paired scansors beneath fourth toes of both manus and pes. A single row of 38 precloacalfemoral pores in adult males. Original tail atuberculate, with median subcaudal scales forming broad transverse plates. Dorsal pattern with a series of four complete, dark crossmarkings between nape and sacrum, alternating with incomplete transverse bands extending dorsally form the flanks but failing to meet along the dorsal midline.
|
||
</paragraph>
|
||
<paragraph id="8BBF5465FFA71652054FFE777140FB85" blockId="11.[264,1324,284,1150]" pageId="11" pageNumber="12">
|
||
<taxonomicName id="4C002FE6FFA71652054FFE777161FE56" box="[312,606,404,429]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="beninensis">
|
||
<emphasis id="B9748877FFA71652054FFE777161FE56" box="[312,606,404,429]" italics="true" pageId="11" pageNumber="12">Hemidactylus beninensis</emphasis>
|
||
</taxonomicName>
|
||
may be distinguished from all other mainland West African congeners on the basis of (sympatric taxa with differing or nonoverlapping character states indicated parenthetically): 14–16 rows of dorsal tubercles (0 rows in
|
||
<taxonomicName id="4C002FE6FFA7165200FAFE07761CFE06" box="[1165,1315,484,509]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="matschiei">
|
||
<emphasis id="B9748877FFA7165200FAFE07761CFE06" box="[1165,1315,484,509]" italics="true" pageId="11" pageNumber="12">H. matschiei</emphasis>
|
||
</taxonomicName>
|
||
, 0–4 rows in
|
||
<taxonomicName id="4C002FE6FFA7165205EBFDEF716FFDDE" box="[412,592,524,549]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="richardsonii">
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||
<emphasis id="B9748877FFA7165205EBFDEF716FFDDE" box="[412,592,524,549]" italics="true" pageId="11" pageNumber="12">H. richardsonii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="4CF8F980FFA716520628FDEF7187FDDD" box="[607,696,524,550]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" metricValueMax="2.54" metricValueMin="2.032" pageId="11" pageNumber="12" unit="in" value="9.0" valueMax="10.0" valueMin="8.0">8–10 in</quantity>
|
||
<taxonomicName id="4C002FE6FFA7165206B7FDEF707AFDDE" box="[704,837,524,549]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="ansorgii">
|
||
<emphasis id="B9748877FFA7165206B7FDEF707AFDDE" box="[704,837,524,549]" italics="true" pageId="11" pageNumber="12">H. ansorgii</emphasis>
|
||
</taxonomicName>
|
||
, 18–24 rows in
|
||
<taxonomicName id="4C002FE6FFA716520073FDEF77AEFDDE" box="[1028,1169,524,549]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="fasciatus">
|
||
<emphasis id="B9748877FFA716520073FDEF77AEFDDE" box="[1028,1169,524,549]" italics="true" pageId="11" pageNumber="12">H. fasciatus</emphasis>
|
||
</taxonomicName>
|
||
, 18–20 rows in
|
||
<emphasis id="B9748877FFA716520550FDD7727BFDB6" box="[295,324,564,589]" italics="true" pageId="11" pageNumber="12">H.</emphasis>
|
||
n. sp. from
|
||
<collectingCountry id="F31714F5FFA7165205A2FDD7716BFDB5" box="[469,596,564,590]" name="Cameroon" pageId="11" pageNumber="12">Cameroon</collectingCountry>
|
||
), precloacalfemoral pores in male in single row of 38 (fewer than
|
||
<quantity id="4CF8F980FFA716520536FDBF7243FD8D" box="[321,380,604,630]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="11" pageNumber="12" unit="in" value="12.0">12 in</quantity>
|
||
<taxonomicName id="4C002FE6FFA7165205F4FDBF713FFD8E" box="[387,512,604,629]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="echinus">
|
||
<emphasis id="B9748877FFA7165205F4FDBF713FFD8E" box="[387,512,604,629]" italics="true" pageId="11" pageNumber="12">H. echinus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C002FE6FFA716520679FDBF71A2FD8E" box="[526,669,604,629]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="muriceus">
|
||
<emphasis id="B9748877FFA716520679FDBF71A2FD8E" box="[526,669,604,629]" italics="true" pageId="11" pageNumber="12">H. muriceus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C002FE6FFA7165206ABFDBF705EFD8E" box="[732,865,604,629]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="ansorgii">
|
||
<emphasis id="B9748877FFA7165206ABFDBF705EFD8E" box="[732,865,604,629]" italics="true" pageId="11" pageNumber="12">H. ansorgii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="4CF8F980FFA716520719FDBF70E9FD8D" box="[878,982,604,630]" metricMagnitude="-1" metricUnit="m" metricValue="3.937" metricValueMax="4.318" metricValueMin="3.556" pageId="11" pageNumber="12" unit="in" value="15.5" valueMax="17.0" valueMin="14.0">14–17 in</quantity>
|
||
<taxonomicName id="4C002FE6FFA7165207ABFDBF7781FD8E" box="[988,1214,604,629]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pseudomuriceus">
|
||
<emphasis id="B9748877FFA7165207ABFDBF7781FD8E" box="[988,1214,604,629]" italics="true" pageId="11" pageNumber="12">H. pseudomuriceus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="4CF8F980FFA7165200BBFDBF7638FD8D" box="[1228,1287,604,630]" metricMagnitude="0" metricUnit="m" metricValue="1.143" pageId="11" pageNumber="12" unit="in" value="45.0">45 in</quantity>
|
||
<emphasis id="B9748877FFA716520179FDBF7614FD8E" box="[1294,1323,604,629]" italics="true" pageId="11" pageNumber="12">H.</emphasis>
|
||
n. sp. from
|
||
<collectingCountry id="F31714F5FFA7165205ECFD677125FD65" box="[411,538,644,670]" name="Cameroon" pageId="11" pageNumber="12">Cameroon</collectingCountry>
|
||
), median subcaudal scale row greatly enlarged transversely (not enlarged or hexagonally enlarged in
|
||
<taxonomicName id="4C002FE6FFA7165206B5FD4F707DFD3E" box="[706,834,684,709]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="echinus">
|
||
<emphasis id="B9748877FFA7165206B5FD4F707DFD3E" box="[706,834,684,709]" italics="true" pageId="11" pageNumber="12">H. echinus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C002FE6FFA716520724FD4F7712FD3E" box="[851,1069,684,709]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="kamdemtohami">
|
||
<emphasis id="B9748877FFA716520724FD4F7712FD3E" box="[851,1069,684,709]" italics="true" pageId="11" pageNumber="12">H. kamdemtohami</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C002FE6FFA716520049FD4F761CFD3E" box="[1086,1315,684,709]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pseudomuriceus">
|
||
<emphasis id="B9748877FFA716520049FD4F761CFD3E" box="[1086,1315,684,709]" italics="true" pageId="11" pageNumber="12">H. pseudomuriceus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C002FE6FFA71652057FFD3772A4FD16" box="[264,411,724,749]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="muriceus">
|
||
<emphasis id="B9748877FFA71652057FFD3772A4FD16" box="[264,411,724,749]" italics="true" pageId="11" pageNumber="12">H. muriceus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C002FE6FFA716520591FD377150FD16" box="[486,623,724,749]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="ansorgii">
|
||
<emphasis id="B9748877FFA716520591FD377150FD16" box="[486,623,724,749]" italics="true" pageId="11" pageNumber="12">H. ansorgii</emphasis>
|
||
</taxonomicName>
|
||
). Both
|
||
<taxonomicName id="4C002FE6FFA7165206A9FD377048FD16" box="[734,887,724,749]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="angularis">
|
||
<emphasis id="B9748877FFA7165206A9FD377048FD16" box="[734,887,724,749]" italics="true" pageId="11" pageNumber="12">H. angularis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C002FE6FFA7165207CEFD377777FD16" box="[953,1096,724,749]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="mabouia">
|
||
<emphasis id="B9748877FFA7165207CEFD377777FD16" box="[953,1096,724,749]" italics="true" pageId="11" pageNumber="12">H. mabouia</emphasis>
|
||
</taxonomicName>
|
||
share overlapping precloacalfemoral pore counts and transversely enlarged subcaudal scales with
|
||
<taxonomicName id="4C002FE6FFA716520179FD1F72BBFCC6" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="beninensis">
|
||
<emphasis id="B9748877FFA716520179FD1F72BBFCC6" italics="true" pageId="11" pageNumber="12">H. beninensis</emphasis>
|
||
</taxonomicName>
|
||
; however, both species have conspicuous tubercles on the dorsum of the tail, whereas that in
|
||
<taxonomicName id="4C002FE6FFA7165205B3FCAF715BFC9E" box="[452,612,844,869]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="beninensis">
|
||
<emphasis id="B9748877FFA7165205B3FCAF715BFC9E" box="[452,612,844,869]" italics="true" pageId="11" pageNumber="12">H. beninensis</emphasis>
|
||
</taxonomicName>
|
||
. The new species is most similar in general appearance to a newly described species of
|
||
<taxonomicName id="4C002FE6FFA71652062CFC9771C3FC76" box="[603,764,884,909]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9748877FFA71652062CFC9771C3FC76" box="[603,764,884,909]" italics="true" pageId="11" pageNumber="12">Hemidactylus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F31714F5FFA71652073CFC9770FAFC75" box="[843,965,884,910]" name="Cameroon" pageId="11" pageNumber="12">Cameroon</collectingCountry>
|
||
(Bauer et al., in press), with which it shares a mostly smooth dorsal tail surface (atuberculate in
|
||
<taxonomicName id="4C002FE6FFA716520035FC7F77DBFC4E" box="[1090,1252,924,949]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="beninensis">
|
||
<emphasis id="B9748877FFA716520035FC7F77DBFC4E" box="[1090,1252,924,949]" italics="true" pageId="11" pageNumber="12">H. beninensis</emphasis>
|
||
</taxonomicName>
|
||
, very weakly tuberculate in
|
||
<emphasis id="B9748877FFA71652066DFC277110FC26" box="[538,559,964,989]" italics="true" pageId="11" pageNumber="12">H</emphasis>
|
||
. n. sp.), transversely enlarged subcaudal scales, and a similar dorsal color pattern. However, these taxa differ with respect to (
|
||
<taxonomicName id="4C002FE6FFA716520780FC0F77A8FBFE" box="[1015,1175,1004,1029]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="beninensis">
|
||
<emphasis id="B9748877FFA716520780FC0F77A8FBFE" box="[1015,1175,1004,1029]" italics="true" pageId="11" pageNumber="12">H. beninensis</emphasis>
|
||
</taxonomicName>
|
||
versus
|
||
<emphasis id="B9748877FFA716520087FC0F7632FBFE" box="[1264,1293,1004,1029]" italics="true" pageId="11" pageNumber="12">H.</emphasis>
|
||
n. sp.): size (68 versus
|
||
<quantity id="4CF8F980FFA716520677FBF7715BFBD5" box="[512,612,1044,1070]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="12" unit="mm" value="100.0">100 mm</quantity>
|
||
SVL), scansors beneath the fourth toe (6–7 versus 10–11), precloacalfemoral pores (38 versus
|
||
<quantity id="4CF8F980FFA7165206B7FBDF71C0FBAD" box="[704,767,1084,1110]" metricMagnitude="0" metricUnit="m" metricValue="1.143" pageId="11" pageNumber="12" unit="in" value="45.0">45 in</quantity>
|
||
the respective
|
||
<typeStatus id="54BBEAC7FFA7165207CFFBDF7710FBAD" box="[952,1071,1084,1110]" pageId="11" pageNumber="12" type="holotype">holotypes</typeStatus>
|
||
), and rows of dorsal tubercles (14–16 versus 18–20).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A07EEFFA71656057FFB5777A1FE2D" lastPageId="15" lastPageNumber="16" pageId="11" pageNumber="12" type="description">
|
||
<paragraph id="8BBF5465FFA71652057FFB577072FB36" blockId="11.[264,1324,1204,1870]" box="[264,845,1204,1229]" pageId="11" pageNumber="12">
|
||
<emphasis id="B9748877FFA71652057FFB577072FB36" box="[264,845,1204,1229]" italics="true" pageId="11" pageNumber="12">
|
||
Description (based on the
|
||
<typeStatus id="54BBEAC7FFA716520649FB577199FB36" box="[574,678,1204,1229]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
|
||
, IRSNB 2617)
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BBF5465FFA71655054FFB3F77CAFDB5" blockId="11.[264,1324,1204,1870]" lastBlockId="12.[264,1323,284,590]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
|
||
Adult male, SVL
|
||
<quantity id="4CF8F980FFA71652066FFB3F71B6FB0D" box="[536,649,1244,1270]" metricMagnitude="-2" metricUnit="m" metricValue="6.83" pageId="11" pageNumber="12" unit="mm" value="68.3">68.3 mm</quantity>
|
||
. Head long (HeadL/SVL ratio 0.29), relatively narrow (HeadW/HeadL ratio 0.67), somewhat depressed (HeadH/HL ratio 0.44), strongly distinct from neck. Lores and interorbital region slightly inflated. Snout moderate (SnEye/HeadL ratio 0.41), less than twice eye diameter (OrbD/SnEye ratio 0.57); scales on snout and forehead small, granular; scales on snout much larger than those on occipital region. Eye relatively large (OrbD/HeadL ratio 0.24); pupil vertical with crenelated margins; supraciliaries short, without spines. Ear opening oval, relatively small (EarL/HeadL ratio 0.06); eye to ear distance greater than diameter of eyes (EyeEar/OrbD ratio 1.22). Rostral approximately 1.5 times wider (3.0 mm) than deep (
|
||
<quantity id="4CF8F980FFA7165207DFF9FF7730F9CD" box="[936,1039,1564,1590]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="11" pageNumber="12" unit="mm" value="1.9">1.9 mm</quantity>
|
||
), incompletely divided dorsally by a deeplyincised rostral groove; two enlarged supranasals separated by a single internasal; rostral in contact with supralabial I, supranasals, and internasal; nostrils oval, each surrounded by supranasal, rostral, first supralabial (narrow contact), and three postnasals, the middle one of which smallest, with very narrow contact with nostril rim; each nostril with a recessed valvular scale at posteromedial border; 3–4 rows of scales separate orbit from supralabials. Mental triangular, minimally deeper (
|
||
<quantity id="4CF8F980FFA71652003FF8EF7798F8DD" box="[1096,1191,1804,1830]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="11" pageNumber="12" unit="mm" value="3.2">3.2 mm</quantity>
|
||
) than wide (
|
||
<quantity id="4CF8F980FFA716520567F8D7724CF8B5" box="[272,371,1844,1870]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="11" pageNumber="12" unit="mm" value="2.9">2.9 mm</quantity>
|
||
) and much deeper than infralabials; one pair of greatly enlarged postmentals meeting at a point behind the mental, each postmental bordered anteriorly by first infralabial, medially by mental, and laterally and posteriorly by a series of four chin shields, the lateralmost much enlarged and also bordering first and second infralabials; remaining three chinshields smaller, approximately four times the size of a granular gular scale. Infralabials bordered by a row of enlarged and axially elongated scales, decreasing in size posteriorly. Enlarged supralabials to midpoint of orbit 8 (left)–9 (right); supralabials to angle of jaws 10 (right)–13 (left); infralabials 10 (left)–9 (right); interorbital scale rows across narrowest point of frontal 14, between supraciliaries 36.
|
||
</paragraph>
|
||
<caption id="DF7F04EDFFA01655057FFA4572A7FA1A" httpUri="https://zenodo.org/record/273469/files/figure.png" pageId="12" pageNumber="13" targetBox="[267,1319,646,1403]" targetPageId="12">
|
||
<paragraph id="8BBF5465FFA01655057FFA4572A7FA1A" blockId="12.[264,1323,1446,1505]" pageId="12" pageNumber="13">
|
||
<emphasis id="B9748877FFA01655057FFA4572B5FA45" bold="true" box="[264,394,1446,1470]" pageId="12" pageNumber="13">FIGURE 4.</emphasis>
|
||
Dorsal view of the holotype of
|
||
<taxonomicName id="4C002FE6FFA0165506ABFA4470DAFA45" box="[732,997,1447,1470]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="beninensis" status="sp. nov.">
|
||
<emphasis id="B9748877FFA0165506ABFA4470DAFA45" box="[732,997,1447,1470]" italics="true" pageId="12" pageNumber="13">Hemidactylus beninensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9748877FFA01655079BFA45777DFA45" bold="true" box="[1004,1090,1446,1470]" pageId="12" pageNumber="13">
|
||
<taxonomicNameLabel id="A247350CFFA01655079BFA45777DFA45" box="[1004,1090,1446,1470]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(IRSNB 2617). Scale bar = 20 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBF5465FFA01654054FF9F7704EFE55" blockId="12.[264,1323,1556,1902]" lastBlockId="13.[264,1325,284,750]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">Body moderately slender, trunk relatively long (TrunkL/SVL ratio 0.40), oval in crosssection, with very feebly developed ventrolateral folds without denticulate scales. Dorsal scales heterogeneous, granular; regularly arranged, small (4 times size of granules), weakly conical to slightly keeled, erect to posteriorlydirected tubercles extending from behind occiput to tail base; no tubercles on temporal region; tubercles more or less uniform across dorsum, somewhat more prominent on flanks; tubercles in 14 rows at midbody. Ventral scales larger than dorsal tubercles, subimbricate; somewhat larger on abdomen than on chest, granular and much smaller in gular region; midbody scale rows across belly to lowest row of tubercles 49. Precloacalfemoral pores in a single row of 38, with a single poreless scales in midline separating continuous series of 19 pored scales on each side, extending nearly to knees. Scales on palm and sole smooth, flat, rounded; scales on dorsal aspects of limbs heterogeneous – granular, intermixed with larger domed to conical tubercules, particularly on knees and shanks.</paragraph>
|
||
<paragraph id="8BBF5465FFA11654054FFE5F77BFFD15" blockId="13.[264,1325,284,750]" pageId="13" pageNumber="14">Fore and hindlimbs moderately long, stout; forearm and tibia moderately long (ForeaL/SVL ratio 0.15; CrusL/SVL ratio 0.17); digits relatively short, strongly clawed; all digits of manus and pes webbed proximally only; distal portions of digits curved, arising from distal portion of expanded subdigital pad; scansors beneath each toe divided, except for distalmost and a few, basal scansors (usually 0–1); scansors from proximalmost at least twice diameter of palmar scales to distalmost divided scansor: 56777 (left manus), 57677 (right manus), 57876 (left pes), 47766 (right pes). Relative length of digits of manus: IV ~ III> V> II> I; of pes: IV> III> V> II> I.</paragraph>
|
||
<caption id="DF7F04EDFFA11654057FF9A272A7F980" httpUri="https://zenodo.org/record/273470/files/figure.png" pageId="13" pageNumber="14" targetBox="[294,1278,797,1593]" targetPageId="13">
|
||
<paragraph id="8BBF5465FFA11654057FF9A272A7F980" blockId="13.[264,1323,1601,1659]" pageId="13" pageNumber="14">
|
||
<emphasis id="B9748877FFA11654057FF9A272B5F9A2" bold="true" box="[264,394,1601,1625]" pageId="13" pageNumber="14">FIGURE 5.</emphasis>
|
||
Dorsal view of the paratype of
|
||
<taxonomicName id="4C002FE6FFA1165406ABF9A170DAF9A2" box="[732,997,1602,1625]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="beninensis" status="sp. nov.">
|
||
<emphasis id="B9748877FFA1165406ABF9A170DAF9A2" box="[732,997,1602,1625]" italics="true" pageId="13" pageNumber="14">Hemidactylus beninensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9748877FFA11654079BF9A2777EF9A2" bold="true" box="[1004,1089,1601,1625]" pageId="13" pageNumber="14">
|
||
<taxonomicNameLabel id="A247350CFFA11654079BF9A2777EF9A2" box="[1004,1089,1601,1625]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(IRSNB 2618). Scale bar = 20 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBF5465FFA11657054FF94C72F8FE55" blockId="13.[264,1323,1711,1897]" lastBlockId="14.[264,1325,284,630]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">Original portion of tail depressed; regenerated portion more slender; length of partly regenerated tail approximately equal to snoutvent length (TailL/SVL ratio 0.97); basal part of original portion of tail covered dorsally with small, posteriorlypointed, subimbricate to imbricate scales, atuberculate; scales forming regular transverse rows, about 8 such rows per distinct caudal segment (corresponding to underlying muscle segments); ventral scales much larger, smooth, imbricate, with a median row of greatly enlarged subcaudal plates extending nearly across the width of tail. Regenerated portion of tail like original but without segmentation. One small, rounded postcloacal spur on each side of tail base.</paragraph>
|
||
<paragraph id="8BBF5465FFA21657054FFE5F70F3FD8D" blockId="14.[264,1325,284,630]" pageId="14" pageNumber="15">
|
||
Mensural features (
|
||
<typeStatus id="54BBEAC7FFA21657065FFE5F71AEFE2D" box="[552,657,444,470]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
|
||
/
|
||
<typeStatus id="54BBEAC7FFA2165706ECFE5F703BFE2D" box="[667,772,444,470]" pageId="14" pageNumber="15" type="paratype">paratype</typeStatus>
|
||
). SVL 68.3/
|
||
<quantity id="4CF8F980FFA2165707D7FE5F772CFE2D" box="[928,1043,444,470]" metricMagnitude="-2" metricUnit="m" metricValue="5.43" pageId="14" pageNumber="15" unit="mm" value="54.3">54.3 mm</quantity>
|
||
, ForeaL 10.5/
|
||
<quantity id="4CF8F980FFA2165700B2FE5F7617FE2D" box="[1221,1320,444,470]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="14" pageNumber="15" unit="mm" value="8.3">8.3 mm</quantity>
|
||
, CrusL 11.5/
|
||
<quantity id="4CF8F980FFA2165705E3FE0772CDFE05" box="[404,498,484,510]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="14" pageNumber="15" unit="mm" value="9.9">9.9 mm</quantity>
|
||
, TailL 66.5 (31.3 regenerated)/51.3 (5.9 regenerated) mm, TailW 7.3/
|
||
<quantity id="4CF8F980FFA21657057FFDEF725BFDDD" box="[264,356,524,550]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="14" pageNumber="15" unit="mm" value="5.1">5.1 mm</quantity>
|
||
, TrunkL 27.2/21.0 mm, HeadL 19.7/
|
||
<quantity id="4CF8F980FFA216570760FDEF70BCFDDD" box="[791,899,524,550]" metricMagnitude="-2" metricUnit="m" metricValue="1.64" pageId="14" pageNumber="15" unit="mm" value="16.4">16.4 mm</quantity>
|
||
, HeadW 13.2/
|
||
<quantity id="4CF8F980FFA21657005BFDEF77A8FDDD" box="[1068,1175,524,550]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="14" pageNumber="15" unit="mm" value="10.3">10.3 mm</quantity>
|
||
, HeadH 8.7/
|
||
<quantity id="4CF8F980FFA21657057FFDD7725AFDB5" box="[264,357,564,590]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="14" pageNumber="15" unit="mm" value="6.1">6.1 mm</quantity>
|
||
, OrbD 4.6/
|
||
<quantity id="4CF8F980FFA21657059BFDD77176FDB5" box="[492,585,564,590]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="14" pageNumber="15" unit="mm" value="4.1">4.1 mm</quantity>
|
||
, EyeEar 5.6/
|
||
<quantity id="4CF8F980FFA216570695FDD7707FFDB5" box="[738,832,564,590]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="14" pageNumber="15" unit="mm" value="4.3">4.3 mm</quantity>
|
||
, SnEye 8.1/
|
||
<quantity id="4CF8F980FFA2165707BAFDD77713FDB5" box="[973,1068,564,590]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="14" pageNumber="15" unit="mm" value="6.4">6.4 mm</quantity>
|
||
, NarEye 5.5/
|
||
<quantity id="4CF8F980FFA2165700B0FDD77619FDB5" box="[1223,1318,564,590]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="14" pageNumber="15" unit="mm" value="4.2">4.2 mm</quantity>
|
||
, Interorb 2.4/
|
||
<quantity id="4CF8F980FFA2165705EAFDBF72C8FD8D" box="[413,503,604,630]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="14" pageNumber="15" unit="mm" value="1.7">1.7 mm</quantity>
|
||
, EarL 1.3/
|
||
<quantity id="4CF8F980FFA216570605FDBF71F1FD8D" box="[626,718,604,630]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="15" unit="mm" value="1.5">1.5 mm</quantity>
|
||
, Internar 2.1/
|
||
<quantity id="4CF8F980FFA21657071BFDBF70F7FD8D" box="[876,968,604,630]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="15" unit="mm" value="1.5">1.5 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBF5465FFA21657057FFD4F7241FD3E" blockId="14.[264,1323,684,1033]" box="[264,382,684,709]" pageId="14" pageNumber="15">
|
||
<emphasis id="B9748877FFA21657057FFD4F7241FD3E" box="[264,382,684,709]" italics="true" pageId="14" pageNumber="15">Osteology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BBF5465FFA21657054FFD377637FBF2" blockId="14.[264,1323,684,1033]" pageId="14" pageNumber="15">
|
||
Parietal bones paired. Stapes imperforate. Phalangeal formulae 23453 for manus and 23454 for pes. Presacral vertebrae 26, including 3 anterior cervical (without ribs), 1 lumbar, and 2 sacral vertebrae; 5 pygal and 8.5 post pygal caudal vertebrae to point of regeneration (
|
||
<quantity id="4CF8F980FFA2165705DAFCAF713CFC9D" box="[429,515,844,870]" metricMagnitude="-1" metricUnit="m" metricValue="4.699" pageId="14" pageNumber="15" unit="in" value="18.5">18.5 in</quantity>
|
||
<typeStatus id="54BBEAC7FFA21657067AFCAF7148FC9D" box="[525,631,844,870]" pageId="14" pageNumber="15" type="paratype">paratype</typeStatus>
|
||
). One pair of slender, crescentic cloacal bones present in both
|
||
<typeStatus id="54BBEAC7FFA21657053EFC97728FFC75" box="[329,432,884,910]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="54BBEAC7FFA216570582FC977161FC75" box="[501,606,884,910]" pageId="14" pageNumber="15" type="paratype">paratype</typeStatus>
|
||
. Endolymphatic sacs enlarged extracranially, extending to level of 4th–6th vertebrae in both specimens. Radiographs reveal that the
|
||
<typeStatus id="54BBEAC7FFA2165700D5FC7C7636FC42" box="[1186,1289,927,953]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
|
||
is skeletally mature, as is confirmed by its welldeveloped precloacalfemoral pores, whereas the subadult
|
||
<typeStatus id="54BBEAC7FFA2165705EAFC0C713DFBF2" box="[413,514,1007,1033]" pageId="14" pageNumber="15" type="paratype">paratype</typeStatus>
|
||
exhibits incomplete ossification of the carpal and tarsal elements.
|
||
</paragraph>
|
||
<paragraph id="8BBF5465FFA21657057FFBDC716EFBA3" blockId="14.[264,1325,1087,1756]" box="[264,593,1087,1112]" pageId="14" pageNumber="15">
|
||
<emphasis id="B9748877FFA21657057FFBDC716EFBA3" box="[264,593,1087,1112]" italics="true" pageId="14" pageNumber="15">Coloration (in preservative)</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BBF5465FFA21657054FFB8471B7F927" blockId="14.[264,1325,1087,1756]" pageId="14" pageNumber="15">Body grayish with a series of dark grayishbrown crossbands with darker brown borders; one across nape, one behind axilla, one at midtrunk, and one in lumbar position. These complete dorsal bands alternating with bands extending dorsally from flanks but not meeting in the dorsal midline, first over scapula. Tips of tubercles whitish, particularly on the flanks. Parietal and temporal regions of head with grayishbrown markings without darker margins. A thick white to cream stripe from lateral surface of rostral and anterior of first supralabial to anterodorsal corner of orbit continuing behind orbit and on to occiput as a series of irregular pale blotches. This stripe bordered above by a grayishbrown stripe extending from nostril to top of orbit and below by a similarly colored triangular marking extending posteriorly from the 2nd to 3rd supralabial scales to ventral half of the anterior margin of the orbit. Limbs mottled with alternating brown and pale gray blotches, particularly distal to the knees and elbows, basal limb segments mostly grayish with faint darker irregular markings. Tail with alternating markings as on dorsum; two complete, darkbordered brown bands on original portion of postpygal tail; regenerated portion grayishbrown with darker speckles, without distinct pattern. Venter grayishcream becoming pale gray subcaudally.</paragraph>
|
||
<paragraph id="8BBF5465FFA21657057FF8F1724EF8D0" blockId="14.[264,1323,1810,1876]" box="[264,369,1810,1835]" pageId="14" pageNumber="15">
|
||
<emphasis id="B9748877FFA21657057FF8F1724EF8D0" box="[264,369,1810,1835]" italics="true" pageId="14" pageNumber="15">Va r i a t i o n</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BBF5465FFA21656054FF8D977A1FE2D" blockId="14.[264,1323,1810,1876]" lastBlockId="15.[264,1323,284,470]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
|
||
The subadult
|
||
<typeStatus id="54BBEAC7FFA2165705A0F8D97103F8AF" box="[471,572,1850,1876]" pageId="14" pageNumber="15" type="paratype">paratype</typeStatus>
|
||
differs from the
|
||
<typeStatus id="54BBEAC7FFA216570775F8D97056F8AF" box="[770,873,1850,1876]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
|
||
in the following features: postmentals bordered posteriorly by 3 chin shields; supralabials 11, 9 to midpoint of orbit (L), 8(R); infralabials 9 (L&R); dorsal tubercles in 16 rows at midbody; no precloacal or femoral pores; subdigital lamellae: (left manus) 46777, (right manus) 47776, (left pes) 47 776, (right pes) 47776.
|
||
<typeStatus id="54BBEAC7FFA316560602FE7771E3FE55" box="[629,732,404,430]" pageId="15" pageNumber="16" type="paratype">Paratype</typeStatus>
|
||
similar in color to
|
||
<typeStatus id="54BBEAC7FFA316560795FE777773FE55" box="[994,1100,404,430]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
|
||
, but more boldly patterned with four complete dark bands on original portion of postpygal tail.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A07EEFFA31656057FFDEF7109FC9D" pageId="15" pageNumber="16" type="distribution">
|
||
<paragraph id="8BBF5465FFA31656057FFDEF71BDFDDE" blockId="15.[264,1323,524,870]" box="[264,642,524,549]" pageId="15" pageNumber="16">
|
||
<emphasis id="B9748877FFA31656057FFDEF71BDFDDE" box="[264,642,524,549]" italics="true" pageId="15" pageNumber="16">Distribution and natural history</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BBF5465FFA31656054FFDD77109FC9D" blockId="15.[264,1323,524,870]" pageId="15" pageNumber="16">
|
||
At present known only from the
|
||
<typeStatus id="54BBEAC7FFA3165606A1FDD77036FDB5" box="[726,777,564,590]" pageId="15" pageNumber="16">type</typeStatus>
|
||
locality in the relatively hilly region of the Département des Collines in central
|
||
<collectingCountry id="F31714F5FFA3165606CFFDBF71C1FD8D" box="[696,766,604,630]" name="Benin" pageId="15" pageNumber="16">Bénin</collectingCountry>
|
||
(
|
||
<figureCitation id="133B48E0FFA316560778FDBF7067FD8D" box="[783,856,604,630]" captionStart="FIGURE 6" captionStartId="15.[264,368,1323,1347]" captionTargetBox="[267,1319,926,1298]" captionTargetId="figure@15.[267,1319,926,1298]" captionTargetPageId="15" captionText="FIGURE 6. Type locality of Hemidactylus beninensis sp. nov. in the Collines de DassaZoumè Département des Collines, central Bénin." httpUri="https://zenodo.org/record/273471/files/figure.png" pageId="15" pageNumber="16">Fig. 6</figureCitation>
|
||
). The area lies adjacent to the Nigerian border and it is likely that this species occurs in that country as well. A juvenile
|
||
<taxonomicName id="4C002FE6FFA31656057FFD4F7119FD3E" box="[264,550,684,709]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="angulatus">
|
||
<emphasis id="B9748877FFA31656057FFD4F7119FD3E" box="[264,550,684,709]" italics="true" pageId="15" pageNumber="16">Hemidactylus angulatus</emphasis>
|
||
</taxonomicName>
|
||
was collected at the same site as the
|
||
<typeStatus id="54BBEAC7FFA316560792FD4F771CFD3D" box="[997,1059,684,710]" pageId="15" pageNumber="16">types</typeStatus>
|
||
of
|
||
<taxonomicName id="4C002FE6FFA31656003BFD4F77D3FD3E" box="[1100,1260,684,709]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="beninensis">
|
||
<emphasis id="B9748877FFA31656003BFD4F77D3FD3E" box="[1100,1260,684,709]" italics="true" pageId="15" pageNumber="16">H. beninensis</emphasis>
|
||
</taxonomicName>
|
||
. The
|
||
<typeStatus id="54BBEAC7FFA31656057FFD377251FD15" box="[264,366,724,750]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
|
||
exhibits damage to the skin of the dorsum of the head (
|
||
<figureCitation id="133B48E0FFA31656007FFD377771FD15" box="[1032,1102,724,750]" captionStart="FIGURE 1" captionStartId="2.[264,368,1356,1380]" captionTargetBox="[633,960,710,1355]" captionTargetId="figure@2.[620,979,700,1360]" captionTargetPageId="2" captionText="FIGURE 1. Map of Bénin showing the localities referred to in the text. Coordinates are degrees north and east. Large font — neighboring countries, small font — départements. Numbered localities: 1 = Chutes de Koudou, Parc National du W du Bénin, 2 = Parc National de la Pendjari, 3 = Koussokoinga (= Koussoukoingou), 4 = Bassila, 5 = Manigri, 6 = Diho (= Idiho), 7 = Agougon (= Agouagon), 8 = Collines de DassaZoumè, 9 = Kétou, 10 = Abomey, 11 = Lokoli, 12 = Niaouli, 13 = Attogon, 14 = Ouidah, 15 = Godomé, 16 = Grand Popo, 17 = Porto Novo." httpUri="https://zenodo.org/record/273465/files/figure.png" pageId="15" pageNumber="16">Fig. 1</figureCitation>
|
||
). Such damage is indicative of mechanically weak skin, which is associated with regional integumentary loss, an escape mechanism employed against certain predators by a variety of geckos (
|
||
<bibRefCitation id="EF912994FFA316560567FCAF72E1FC9D" author="Bauer" box="[272,478,844,870]" pageId="15" pageNumber="18" refString="Bauer, A. M., Russell, A. P. & Shadwick, R. E. (1989) Mechanical properties and morphological correlates of fragile skin in gekkonid lizards. Journal of Experimental Biology, 145, 79 - 102." type="journal article" year="1989">
|
||
Bauer
|
||
<emphasis id="B9748877FFA316560528FCAF72A5FC9E" box="[351,410,844,869]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
|
||
1989
|
||
</bibRefCitation>
|
||
, 1993).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |