treatments-xml/data/03/A9/24/03A92416AB47B23DFF74FB06CCE4F4F7.xml
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<document id="D13395F0060968522528078F349BC5DE" ID-CLB-Dataset="31470" ID-DOI="10.11646/zootaxa.4350.1.5" ID-GBIF-Dataset="fba6f08d-2e56-47f6-81d2-dac9346dd00b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1050941" ID-ZooBank="D225AE6E-BE98-40FD-8706-E7699FFFB3C5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510814393772" checkinUser="plazi" docAuthor="Stubbs, Alexander L., Karin, Benjamin R., Arifin, Umilaela, Iskandar, Djoko T., Arida, Evy, Reilly, Sean B., Bloch, Luke M., Kusnadi, Agus &amp; Mcguire, Jimmy A." docDate="2017" docId="03A92416AB47B23DFF74FB06CCE4F4F7" docLanguage="en" docName="zootaxa.4350.1.5.pdf" docOrigin="Zootaxa 4350 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepidodactylus pantai Stubbs &amp; Karin &amp; Arifin &amp; Iskandar &amp; Arida &amp; Reilly &amp; Bloch &amp; Kusnadi &amp; Mcguire 2017, new species" docType="treatment" docVersion="7" lastPageNumber="99" masterDocId="FF905C6EAB43B235FFE3FFE1CD65F366" masterDocTitle="A new species of Lepidodactylus (Reptilia: Squamata: Gekkonidae) from the Kei Islands, Maluku, Indonesia" masterLastPageNumber="105" masterPageNumber="91" pageNumber="95" updateTime="1698497448523" updateUser="ExternalLinkService">
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<mods:title id="B1C4669E1948A9A597803171135E1C81">A new species of Lepidodactylus (Reptilia: Squamata: Gekkonidae) from the Kei Islands, Maluku, Indonesia</mods:title>
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<mods:namePart id="44DC80EC25730E84DE7EDEAEA0FC47FD">Stubbs, Alexander L.</mods:namePart>
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<mods:namePart id="A4FA4AE55837C41D903A27B1999C2B2A">Karin, Benjamin R.</mods:namePart>
</mods:name>
<mods:name id="C5F06974C4C1BCF4171099C48B11E53C" type="personal">
<mods:role id="6D23E0E84AA9BF233F2E7F8CF4B92C2C">
<mods:roleTerm id="782656DCFA4F84F6A651CB4ADA968BCB">Author</mods:roleTerm>
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<mods:namePart id="7BF27EFDCF01388810DAC2769C68DAB1">Arifin, Umilaela</mods:namePart>
</mods:name>
<mods:name id="DC78D6BCC0CD3D4568E3427E9DF348EC" type="personal">
<mods:role id="5B9A3D474622B020A81C27ABABB79ED9">
<mods:roleTerm id="CE0AE6FFF2C01771976FE565E71E823A">Author</mods:roleTerm>
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<mods:namePart id="47A32DD0569A27B5E6B345568E8287D5">Iskandar, Djoko T.</mods:namePart>
</mods:name>
<mods:name id="DC41C9F41CC836945E04B13966A79037" type="personal">
<mods:role id="05211123AF9FF1F3D1796146E47F6E63">
<mods:roleTerm id="D654F438548434E70F4F479177501F86">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="72D995BFC72460DF37D196C77C9EEB06">Arida, Evy</mods:namePart>
</mods:name>
<mods:name id="994D7DFED9107557CD805127786D9A6C" type="personal">
<mods:role id="B663A3707FFC3CD5534D629C25B89913">
<mods:roleTerm id="A581B8BB0A114A257DEA674CD42D3397">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2B397395E5152CFED480661F519B5F6D">Reilly, Sean B.</mods:namePart>
</mods:name>
<mods:name id="407CCA38825D39831C5074F44E9E6B21" type="personal">
<mods:role id="78BC773C66C57A58120A728AC761B818">
<mods:roleTerm id="285E6E07632FE936C2EAC06E7A9D26E5">Author</mods:roleTerm>
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<mods:namePart id="742E4C2C472FD6364ED82D57E2212F83">Bloch, Luke M.</mods:namePart>
</mods:name>
<mods:name id="1DB23E5C1CF87A41FC210EC37187B708" type="personal">
<mods:role id="B39E41D022B20D514C91DED318EE949B">
<mods:roleTerm id="C8B3254942B85D8D32F491AA2ACA5FCF">Author</mods:roleTerm>
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<mods:namePart id="F6CE2F07C17AD04E9F443BCACD3492A2">Kusnadi, Agus</mods:namePart>
</mods:name>
<mods:name id="BF06D76134F9EA9E571F9136104441BD" type="personal">
<mods:role id="CB0199D59F1BFAD3773C3B5D9C5EADCA">
<mods:roleTerm id="AEED78BCB150413F09EF0210AC7AB8E8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0EC971FCD541021DD61049FAB1D6DB95">Mcguire, Jimmy A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="F4E6CD60937F3F3FE8C3796C8F9B8CA0">text</mods:typeOfResource>
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<mods:titleInfo id="20C8960D5520932CCA522F90C866884C">
<mods:title id="9450E5D0A608B00059F11651399A582D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2F47FA7218EC3511039602078DD3569D">
<mods:date id="AF26DD7A79DC417B3705EE153345DC8E">2017</mods:date>
<mods:detail id="7CFA21BD4E8FCE3A096EA67AF69AB7F1" type="pubDate">
<mods:number id="E2FF03DFF3898748338D8AE3784361D1">2017-11-16</mods:number>
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<mods:number id="9A1DA3775D658E62A994021B81676355">4350</mods:number>
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<mods:number id="9DC994B798AC4AB1E60AEC2E205C1F11">1</mods:number>
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<mods:start id="C5CF18E2A9C29638183EFF957FF7AC52">91</mods:start>
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<mods:classification id="AFE04FC00F829FCF549190E20CD17636">journal article</mods:classification>
<mods:identifier id="8224FBCEA5ED63CCC441EE1C2C101843" type="CLB-Dataset">31470</mods:identifier>
<mods:identifier id="63F5C93BD2A6B259DF538901033C92F2" type="DOI">10.11646/zootaxa.4350.1.5</mods:identifier>
<mods:identifier id="0A412168D1800A435CF1003FFE57307A" type="GBIF-Dataset">fba6f08d-2e56-47f6-81d2-dac9346dd00b</mods:identifier>
<mods:identifier id="0A9EF3DB7A407D8434DFA2999F8AB0D7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B91FBB4449BD07A9ED5781D3F1965509" type="Zenodo-Dep">1050941</mods:identifier>
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<treatment id="03A92416AB47B23DFF74FB06CCE4F4F7" ID-DOI="http://doi.org/10.5281/zenodo.6030121" ID-GBIF-Taxon="137307884" ID-Zenodo-Dep="6030121" LSID="urn:lsid:plazi:treatment:03A92416AB47B23DFF74FB06CCE4F4F7" httpUri="http://treatment.plazi.org/id/03A92416AB47B23DFF74FB06CCE4F4F7" lastPageId="8" lastPageNumber="99" pageId="4" pageNumber="95">
<subSubSection id="C31AC68BAB47B231FF74FB06CC58F645" pageId="4" pageNumber="95" type="nomenclature">
<paragraph id="8BBF9500AB47B231FF74FB06CF5AF664" blockId="4.[151,575,1255,1315]" box="[151,575,1255,1282]" pageId="4" pageNumber="95">
<heading id="D0F7226CAB47B231FF74FB06CF5AF664" bold="true" box="[151,575,1255,1282]" fontSize="11" level="1" pageId="4" pageNumber="95" reason="1">
<emphasis id="B9744912AB47B231FF74FB06CF5AF664" bold="true" box="[151,575,1255,1282]" pageId="4" pageNumber="95">
<taxonomicName id="4C00EE83AB47B231FF74FB06CCC4F667" authority="Stubbs &amp; Karin &amp; Arifin &amp; Iskandar &amp; Arida &amp; Reilly &amp; Bloch &amp; Kusnadi &amp; Mcguire, 2017" authorityName="Stubbs &amp; Karin &amp; Arifin &amp; Iskandar &amp; Arida &amp; Reilly &amp; Bloch &amp; Kusnadi &amp; Mcguire" authorityYear="2017" box="[151,417,1255,1281]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="95" phylum="Chordata" rank="species" species="pantai" status="sp. nov.">
<emphasis id="B9744912AB47B231FF74FB06CCC4F667" bold="true" box="[151,417,1255,1281]" italics="true" pageId="4" pageNumber="95">Lepidodactylus pantai</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A247F469AB47B231FE4CFB09CF5AF664" box="[431,575,1256,1282]" pageId="4" pageNumber="95" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBF9500AB47B231FF74FAEBCC58F645" blockId="4.[151,575,1255,1315]" box="[151,317,1290,1315]" pageId="4" pageNumber="95">(Figures 2,3,4)</paragraph>
</subSubSection>
<subSubSection id="C31AC68BAB47B231FF74FAB2C9EEF5C9" pageId="4" pageNumber="95" type="materials_examined">
<paragraph id="8BBF9500AB47B231FF74FAB2CCD5F69D" blockId="4.[151,1436,1362,2035]" pageId="4" pageNumber="95">
<materialsCitation id="3B689F5DAB47B231FF74FAB2CCD5F69D" ID-GBIF-Occurrence="1700783923" collectingDate="2016-10" collectionCode="MZB" country="Indonesia" county="Museum of Vertebrate Zoology" latitude="-5.646671" location="Field" longLatPrecision="1" longitude="132.6383" municipality="Museum Zoologicum Bogoriense" pageId="4" pageNumber="95" specimenCode="MVZ 273691" specimenCount="1" stateProvince="Maluku" typeStatus="holotype">
<emphasis id="B9744912AB47B231FF74FAB2CC6EF60D" bold="true" box="[151,267,1363,1387]" pageId="4" pageNumber="95">
<typeStatus id="54BB2BA2AB47B231FF74FAB2CC63F60D" box="[151,262,1363,1387]" pageId="4" pageNumber="95" type="holotype">Holotype</typeStatus>
.
</emphasis>
(Museum Zoologicum Bogoriense.Lace.14062,
<location id="8EDFC3DBAB47B231FCDFFAB3CE13F60D" LSID="urn:lsid:plazi:treatment:03A92416AB47B23DFF74FB06CCE4F4F7:8EDFC3DBAB47B231FCDFFAB3CE13F60D" box="[828,886,1362,1387]" country="Indonesia" county="Museum of Vertebrate Zoology" latitude="-5.646671" longLatPrecision="1" longitude="132.6383" municipality="Museum Zoologicum Bogoriense" name="Field" pageId="4" pageNumber="95" stateProvince="Maluku">Field</location>
number ALS 534)
<location id="8EDFC3DBAB47B231FBBCFAB2C9E4F60D" LSID="urn:lsid:plazi:treatment:03A92416AB47B23DFF74FB06CCE4F4F7:8EDFC3DBAB47B231FBBCFAB2C9E4F60D" box="[1119,1153,1363,1387]" country="Indonesia" county="Museum of Vertebrate Zoology" latitude="-5.646671" longLatPrecision="1" longitude="132.6383" municipality="Museum Zoologicum Bogoriense" name="An" pageId="4" pageNumber="95" stateProvince="Maluku">An</location>
adult male collected by ALS, BRK, and UA from beachside rocks at
<location id="8EDFC3DBAB47B231FD7EFA97CE1EF6E9" LSID="urn:lsid:plazi:treatment:03A92416AB47B23DFF74FB06CCE4F4F7:8EDFC3DBAB47B231FD7EFA97CE1EF6E9" box="[669,891,1398,1423]" country="Indonesia" county="Museum of Vertebrate Zoology" latitude="-5.646671" longLatPrecision="1" longitude="132.6383" municipality="Museum Zoologicum Bogoriense" name="Pasir Panjan Beach" pageId="4" pageNumber="95" stateProvince="Maluku">Pasir Panjan Beach</location>
,
<location id="8EDFC3DBAB47B231FC6BFA96C94FF6E9" LSID="urn:lsid:plazi:treatment:03A92416AB47B23DFF74FB06CCE4F4F7:8EDFC3DBAB47B231FC6BFA96C94FF6E9" box="[904,1066,1398,1423]" country="Indonesia" county="Museum of Vertebrate Zoology" latitude="-5.646671" longLatPrecision="1" longitude="132.6383" municipality="Museum Zoologicum Bogoriense" name="Desa Ohoililir" pageId="4" pageNumber="95" stateProvince="Maluku">Desa Ohoililir</location>
,
<location id="8EDFC3DBAB47B231FBD4FA97C9C2F6E9" LSID="urn:lsid:plazi:treatment:03A92416AB47B23DFF74FB06CCE4F4F7:8EDFC3DBAB47B231FBD4FA97C9C2F6E9" box="[1079,1191,1398,1423]" country="Indonesia" county="Museum of Vertebrate Zoology" latitude="-5.646671" longLatPrecision="1" longitude="132.6383" municipality="Museum Zoologicum Bogoriense" name="Kei Kecil" pageId="4" pageNumber="95" stateProvince="Maluku">Kei Kecil</location>
,
<collectingRegion id="49C45BE2AB47B231FB57FA97C86BF6E9" box="[1204,1294,1398,1423]" country="Indonesia" name="Maluku Utara" pageId="4" pageNumber="95">Maluku</collectingRegion>
,
<collectingCountry id="F317D590AB47B231FAF9FA97C8E2F6E9" box="[1306,1415,1398,1423]" name="Indonesia" pageId="4" pageNumber="95">Indonesia</collectingCountry>
a few hours after sunset at
<geoCoordinate id="EE34F3C7AB47B231FE57FA7BCF24F6D5" box="[436,577,1434,1459]" direction="south" orientation="latitude" pageId="4" pageNumber="95" precision="1" value="-5.646671">5.646671° S</geoCoordinate>
,
<geoCoordinate id="EE34F3C7AB47B231FDACFA7BCF9DF6D4" box="[591,760,1434,1459]" direction="east" orientation="longitude" pageId="4" pageNumber="95" precision="1" value="132.6383">132.638312° E</geoCoordinate>
(WGS84) on
<collectingDate id="EFFA4A28AB47B231FC7AFA7BC97CF6D5" box="[921,1049,1434,1459]" pageId="4" pageNumber="95" value="2016-10">
<date id="FFBEB3C0AB47B231FC7AFA7BC97CF6D5" box="[921,1049,1434,1459]" pageId="4" pageNumber="95" value="2016-10">16 October</date>
</collectingDate>
, 2011.
<location id="8EDFC3DBAB47B231FB88FA7BC9C2F6D5" LSID="urn:lsid:plazi:treatment:03A92416AB47B23DFF74FB06CCE4F4F7:8EDFC3DBAB47B231FB88FA7BC9C2F6D5" box="[1131,1191,1434,1459]" country="Indonesia" county="Museum of Vertebrate Zoology" latitude="-5.646671" longLatPrecision="1" longitude="132.6383" municipality="Museum Zoologicum Bogoriense" name="Liver" pageId="4" pageNumber="95" stateProvince="Maluku">Liver</location>
tissue is preserved in duplicate in RNA-Later at
<collectingMunicipality id="6BDB0F7AAB47B231FE03FA5ECE03F6B1" box="[480,870,1470,1495]" pageId="4" pageNumber="95">Museum Zoologicum Bogoriense</collectingMunicipality>
(
<collectionCode id="ED110DC5AB47B231FC9EFA5FCED8F6B1" box="[893,957,1470,1495]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="4" pageNumber="95">MZB</collectionCode>
.Lace.14062) and
<collectingCounty id="62DEED8CAB47B231FB70FA5ECD93F69D" pageId="4" pageNumber="95">Museum of Vertebrate Zoology</collectingCounty>
(
<specimenCode id="DBA63D7BAB47B231FEE7FA02CCC1F69D" box="[260,420,1506,1531]" collectionCode="MVZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34777" name="Museum of Vertebrate Zoology, University of California Berkeley" pageId="4" pageNumber="95">MVZ 273691</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BBF9500AB47B231FF24F9E6C9EEF5C9" blockId="4.[151,1436,1362,2035]" pageId="4" pageNumber="95">
<materialsCitation id="3B689F5DAB47B231FF24F9E6CE3DF525" ID-GBIF-Occurrence="1700783924" collectionCode="MVZ" collectorName="Paratypes. A" pageId="4" pageNumber="95" specimenCount="2" typeStatus="paratype">
<collectorName id="26F5F0D6AB47B231FF24F9E6CC07F579" box="[199,354,1543,1567]" pageId="4" pageNumber="95">
<emphasis id="B9744912AB47B231FF24F9E6CC23F579" bold="true" box="[199,326,1543,1567]" pageId="4" pageNumber="95">
<typeStatus id="54BB2BA2AB47B231FF24F9E6CC24F579" box="[199,321,1543,1567]" pageId="4" pageNumber="95" type="paratype">Paratypes</typeStatus>
.
</emphasis>
A
</collectorName>
series of ten additional specimens were collected at the same locality and time as the
<typeStatus id="54BB2BA2AB47B231FAD9F9E7C8F9F579" box="[1338,1436,1542,1567]" pageId="4" pageNumber="95" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED110DC5AB47B231FF7DF9CACDBBF525" box="[158,222,1579,1603]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34777" name="Museum of Vertebrate Zoology, University of California Berkeley" pageId="4" pageNumber="95">MVZ</collectionCode>
tissue number and field number in parenthesis)
</materialsCitation>
: MZB.Lace.14064 (MVZ 273686; ALS 502), MZB.Lace.14065 (MVZ 273687; ALS 505), MZB.Lace.14066 (MVZ 273688; ALS 530), MZB.Lace.14067 (MVZ 273689; ALS 531), MZB.Lace.14068 (MVZ 273690; ALS 533), MVZ 273692 (ALS 501), MVZ 273693 (ALS 503), MVZ 273694 (ALS 504), MVZ 273695 (ALS 532), MVZ 273696 (ALS 535).
</paragraph>
</subSubSection>
<subSubSection id="C31AC68BAB47B231FF24F95BCEA8F4E1" pageId="4" pageNumber="95" type="diagnosis">
<paragraph id="8BBF9500AB47B231FF24F95BCEA8F4E1" blockId="4.[151,1436,1362,2035]" pageId="4" pageNumber="95">
<emphasis id="B9744912AB47B231FF24F95BCC24F5B5" bold="true" box="[199,321,1722,1747]" pageId="4" pageNumber="95">Diagnosis.</emphasis>
A moderate-sized bisexual species of
<taxonomicName id="4C00EE83AB47B231FD1AF95ACEC1F5B5" box="[761,932,1723,1747]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="95" phylum="Chordata" rank="genus">
<emphasis id="B9744912AB47B231FD1AF95ACEC1F5B5" box="[761,932,1723,1747]" italics="true" pageId="4" pageNumber="95">Lepidodactylus</emphasis>
</taxonomicName>
, SVL 36.940.5 (mean = 38.3) mm for five adult males and 32.040.5 (mean = 37.4) for five adult females, distinguished from other species by the following combination of characters: 113 rows of scales around the midbody; subdigital scansors 1012 on toe IV, and 79 on toe I; terminal scansor is divided on digits II through V on both the fingers and toes; 3 scansors on 4th toe divided or deeply notched; interdigital webbing small, less than 1/5th the 4th toe length; 1824 pores in precloacal and femoral regions of male. Tail nearly cylindrical without lateral serrations.
</paragraph>
</subSubSection>
<subSubSection id="C31AC68BAB47B230FF24F873C87AF44E" lastPageId="5" lastPageNumber="96" pageId="4" pageNumber="95" type="description">
<paragraph id="8BBF9500AB47B230FF24F873CE49F39E" blockId="4.[151,1436,1362,2035]" lastBlockId="5.[151,1437,151,2013]" lastPageId="5" lastPageNumber="96" pageId="4" pageNumber="95">
<emphasis id="B9744912AB47B231FF24F873CC8BF4CD" bold="true" box="[199,494,1938,1963]" pageId="4" pageNumber="95">
Description of
<typeStatus id="54BB2BA2AB47B231FE62F872CC8FF4CD" box="[385,490,1939,1963]" pageId="4" pageNumber="95" type="holotype">holotype</typeStatus>
.
</emphasis>
MZB.Lace.14062 (measurements in mm, after preservation). Snoutvent length 40.57; head length 11.74; head width 7.56; head height 4.48; jaw length left/right 6.06/6.34; snouteye length 4.54; nariseye length 3.13; naris circular, approximately 0.4×0.5; orbit diameter 2.46; eyeear length 3.38; snout width 1.78; ear opening length×width 0.62×0.41; interorbital width 4.00; snoutforelimb length 14.13; axillagroin distance 20.05; length of hind limb 15.13 (75.5 % of axillagroin distance); length of forelimb 10.95; crus length 6.41; tail length 24 (entire); tail width 4.34; tail depth 3.88.
</paragraph>
<paragraph id="8BBF9500AB46B230FF24FEE5C9E0F2B6" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="96">Snout tapered, rounded at tip; three scales touching rostral between left and right nares; supranasals separated by three scales in contact with rostral; rostral entering nares, broader than high, 2.05×0.89 (width about 2.3 times height); no rostral cleft; nares bordered by five scales: three nasals, one rostral, and one supralabial; 3435 interorbital scales; 11 left and 11 right supralabials, 10th below center of eye; 10 left and 10 right infralabials; mental scale distinct, triangular, its anterior width nearly equal to midline length (0.69×.0.72); Mental is bordered posteriorly by two enlarged primary postmentals, each in contact with the first infralabial.</paragraph>
<paragraph id="8BBF9500AB46B230FF24FE3DCF7EF05E" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="96">
Body slightly depressed; 113 rows of scales (average of 3 counts: 109, 114, 116) around midbody, grading into granular scales on lower lateral surfaces; dorsal and lateral scales granular, without enlarged tubercles; ventral scales almost flat, hexagonal, and 23 times larger than dorsal scales; limbs well developed; digits moderately dilated, undersurface (
<figureCitation id="133B8985AB46B230FE73FDA6CCB7F106" box="[400,466,583,608]" captionStart="FIGURE 3" captionStartId="7.[151,250,1249,1271]" captionTargetBox="[151,1436,193,1228]" captionTargetId="figure@7.[151,1436,193,1228]" captionTargetPageId="7" captionText="FIGURE 3. Plantar view of the right foot of the holotype (MZB. Lace. 14069). Note the 3 divided terminal scansors on toes 1 - 4." httpUri="https://zenodo.org/record/1051486/files/figure.png" pageId="5" pageNumber="96">Fig. 3</figureCitation>
) bearing left/right scansors as follow: fingers—
<date id="FFBEB3C0AB46B230FC13FDA9C96BF106" box="[1008,1038,584,608]" pageId="5" pageNumber="96">I 9</date>
/9, II 9/
<date id="FFBEB3C0AB46B230FB81FDA9C9ADF106" box="[1122,1224,584,608]" pageId="5" pageNumber="96" value="2010-03-12">10, III 12</date>
/
<date id="FFBEB3C0AB46B230FB31FDA9C85FF106" box="[1234,1338,584,608]" pageId="5" pageNumber="96" value="2012-04-13">12, IV 13</date>
/
<date id="FFBEB3C0AB46B230FAA7FDA9C8F6F106" box="[1348,1427,584,608]" pageId="5" pageNumber="96" value="2013-05-09">13, V 9</date>
/ 9; toes—I 10/(not intact), II 11/
<date id="FFBEB3C0AB46B230FDE5FD8DCF14F1E2" box="[518,625,620,644]" pageId="5" pageNumber="96" value="2011-03-14">11, III 14</date>
/
<date id="FFBEB3C0AB46B230FD98FD8DCF8DF1E2" box="[635,744,620,644]" pageId="5" pageNumber="96" value="2014-04-12">14, IV 12</date>
/
<date id="FFBEB3C0AB46B230FD11FD8DCE2DF1E2" box="[754,840,620,644]" pageId="5" pageNumber="96" value="2012-05-09">12, V 9</date>
/9; distal three scansors, including the terminal one, divided on all digits except the first on fingers and toes; first digit with complete terminal and two divided subterminal scansors; all digits except first with 3 undivided or deeply notched scansors, including the terminal; all digits except first clawed; compressed claw-bearing phalanges arising from distal margin of the dilated part and extending only a short distance beyond; slight webbing 1/6th of the length of the 4th toe; digits elongate and slender, toe pads slightly enlarged.
</paragraph>
<paragraph id="8BBF9500AB46B230FF24FCA4C850F08B" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="96">Twenty five enlarged precloacal and femoral scales; two post-cloacal spurs on each side of vent; tail entire, subcylindrical throughout length, gradually tapering to a blunt tip; lateral margins without spines or skin flanges; tail constricted at base posterior to vent; scales on tail annulate, squarish or rectangular, ventral scales about twice as large as dorsal scales; base of tail distinctly swollen by hemipenes; hemipenes everted, forked with small, flared scales on the midbody of each forked end, transitioning into even smaller distal scales about half the size.</paragraph>
<paragraph id="8BBF9500AB46B230FF24FC16CE6DF7A3" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="96">
<emphasis id="B9744912AB46B230FF24FC16CCAEF776" bold="true" box="[199,459,1015,1040]" pageId="5" pageNumber="96">Color in preservative.</emphasis>
Color of
<typeStatus id="54BB2BA2AB46B230FDDBFC16CFFFF776" box="[568,666,1015,1040]" pageId="5" pageNumber="96" type="holotype">holotype</typeStatus>
after about three years in ethanol is similar to coloration in life (
<figureCitation id="133B8985AB46B230FA8CFC16CDCDF752" captionStart="FIGURE 2" captionStartId="6.[151,250,2000,2022]" captionTargetBox="[320,1227,1353,1935]" captionTargetId="figure@6.[320,1227,1353,1935]" captionTargetPageId="6" captionText="FIGURE 2. Photo in life of Lepidodacylus pantai sp. nov. from the type locality. (Photo by LMB, 2014)." httpUri="https://zenodo.org/record/1051484/files/figure.png" pageId="5" pageNumber="96">Fig. 2</figureCitation>
). Overall dorsal coloration is pale grey with some lateral streaks of darker pigment. Two pairs of dark spots are clearly visible on the dorsal surface of tail. Venter of body pale cream in color with no distinct pigmentation. Ventral surface of tail slightly darker posteriorly. Slight dark spotting on the dorsal surface of the head. A brown orbital stripe extends from the nostril to the anterior insertion of the forelimb. Three elongated spots on dorsal surface of body between the insertions of the forelimbs.
</paragraph>
<paragraph id="8BBF9500AB46B230FF24FB2ECFF5F6A6" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="96">
<emphasis id="B9744912AB46B230FF24FB2ECC5AF78E" bold="true" box="[199,319,1231,1256]" pageId="5" pageNumber="96">Variation.</emphasis>
The
<typeStatus id="54BB2BA2AB46B230FE9CFB30CCCAF78F" box="[383,431,1233,1257]" pageId="5" pageNumber="96">type</typeStatus>
series varies in SVL (20.640.5; n=11), number of precloacal pores in males (1725; n=5), number of scansors on Toe IV (1012, n=11), number of lamellae on Toe I (79, n=11), number of supralabials (1114, n =11), number of infralabials (911, n=11), head width as a proportion of SVL (1821%, n=11), and number of scale rows around the midbody (108127, n=11). Nearly all specimens have two cloacal spurs, however two specimens have an additional cloacal spur on one side of the body. All specimens have three divided terminal scansors. Both males and females of the
<typeStatus id="54BB2BA2AB46B230FCC5FA64CE33F6FB" box="[806,854,1413,1437]" pageId="5" pageNumber="96">type</typeStatus>
series possess enlarged, pale-white, endolymphatic sacs, which are more pronounced in females.
</paragraph>
<paragraph id="8BBF9500AB46B230FF24FA2DCF50F5FE" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="96">
Coloration is similar among the
<typeStatus id="54BB2BA2AB46B230FDCEFA2CCF38F683" box="[557,605,1485,1509]" pageId="5" pageNumber="96">type</typeStatus>
series (
<figureCitation id="133B8985AB46B230FD51FA2DCF93F682" box="[690,758,1484,1509]" captionStart="FIGURE 4" captionStartId="8.[151,250,1162,1184]" captionTargetBox="[151,1436,215,1098]" captionTargetId="figure@8.[151,1436,215,1098]" captionTargetPageId="8" captionText="FIGURE 4. Dorsal color pattern variation in the type series. Specimen tags overlaid with museum accession numbers." httpUri="https://zenodo.org/record/1051488/files/figure.png" pageId="5" pageNumber="96">Fig. 4</figureCitation>
), however patterning is somewhat variable. Some specimens possess distinct dark chevron patterning along the dorsal surface of the body. All members of the
<typeStatus id="54BB2BA2AB46B230FB0BFA10C87DF56F" box="[1256,1304,1521,1545]" pageId="5" pageNumber="96">type</typeStatus>
series have three elongate dark spots on the dorsal surface of the body between the insertions of the forelimbs. The lateral surfaces of some specimens have darker brown pigmentation, with a pale cream dorsal band running from the temporal region posterior to the base of the tail. The orbital stripe is variable in conspicuousness, though all specimens possess it to some degree.
</paragraph>
<paragraph id="8BBF9500AB46B230FF24F945C87AF44E" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="96">
Coloration in life (
<figureCitation id="133B8985AB46B230FE7FF945CC85F5DA" box="[412,480,1700,1725]" captionStart="FIGURE 2" captionStartId="6.[151,250,2000,2022]" captionTargetBox="[320,1227,1353,1935]" captionTargetId="figure@6.[320,1227,1353,1935]" captionTargetPageId="6" captionText="FIGURE 2. Photo in life of Lepidodacylus pantai sp. nov. from the type locality. (Photo by LMB, 2014)." httpUri="https://zenodo.org/record/1051484/files/figure.png" pageId="5" pageNumber="96">Fig. 2</figureCitation>
) is very similar to that in preservative (
<figureCitation id="133B8985AB46B230FC45F945CE89F5DA" box="[934,1004,1700,1725]" captionStart="FIGURE 4" captionStartId="8.[151,250,1162,1184]" captionTargetBox="[151,1436,215,1098]" captionTargetId="figure@8.[151,1436,215,1098]" captionTargetPageId="8" captionText="FIGURE 4. Dorsal color pattern variation in the type series. Specimen tags overlaid with museum accession numbers." httpUri="https://zenodo.org/record/1051488/files/figure.png" pageId="5" pageNumber="96">Fig. 4</figureCitation>
), however some of the darker regions also contain reddish pigment that has been lost in preservation. As with many geckos, these individuals change their level of pigmentation with time of day and light environment.
<figureCitation id="133B8985AB46B230FC52F90DC976F462" box="[945,1043,1772,1797]" captionStart="FIGURE 2" captionStartId="6.[151,250,2000,2022]" captionTargetBox="[320,1227,1353,1935]" captionTargetId="figure@6.[320,1227,1353,1935]" captionTargetPageId="6" captionText="FIGURE 2. Photo in life of Lepidodacylus pantai sp. nov. from the type locality. (Photo by LMB, 2014)." httpUri="https://zenodo.org/record/1051484/files/figure.png" pageId="5" pageNumber="96">Figure 2</figureCitation>
shows a color photograph of this species
<emphasis id="B9744912AB46B230FF0CF8F1CC51F441" box="[239,308,1808,1831]" italics="true" pageId="5" pageNumber="96">in situ</emphasis>
(specimen uncollected) from the
<typeStatus id="54BB2BA2AB46B230FD4EF8F0CFB8F44F" box="[685,733,1809,1833]" pageId="5" pageNumber="96">type</typeStatus>
locality taken on a more recent expedition in 2014.
</paragraph>
</subSubSection>
<subSubSection id="C31AC68BAB46B232FF24F8D2C9E8F4D9" lastPageId="7" lastPageNumber="98" pageId="5" pageNumber="96" type="distribution">
<paragraph id="8BBF9500AB46B230FF24F8D2CFF2F4BB" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="96">
<emphasis id="B9744912AB46B230FF24F8D2CC05F42A" bold="true" box="[199,352,1843,1868]" pageId="5" pageNumber="96">Distribution.</emphasis>
The
<typeStatus id="54BB2BA2AB46B230FE4BF8D4CCBDF42B" box="[424,472,1845,1869]" pageId="5" pageNumber="96">type</typeStatus>
series are all from Kei Kecil, however we recently collected another series of specimens from Kur Island (~
<quantity id="4CF838E5AB46B230FE1AF8B9CF22F416" box="[505,583,1879,1904]" metricMagnitude="4" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="96" unit="km" value="80.0">80 km</quantity>
to the northwest) that appears morphologically similar, however further analysis is needed to determine the level of divergence between these two populations. Our phylogeny also indicates that specimens from
<collectingCountry id="F317D590AB46B230FE0CF841CF48F4DF" box="[495,557,1952,1977]" name="Palau" pageId="5" pageNumber="96">Palau</collectingCountry>
are remarkably similar genetically but we refrain from assigning members of that population to this species at present time.
</paragraph>
<paragraph id="8BBF9500AB45B233FF24FF79C999F3B3" blockId="6.[151,1436,151,213]" pageId="6" pageNumber="97">
<emphasis id="B9744912AB45B233FF24FF79CCEDF3D6" bold="true" box="[199,392,151,176]" pageId="6" pageNumber="97">Natural history.</emphasis>
We have experience with this species only on two islands, Kei Kecil and Kur. Specimens were associated with exposed intertidal limestone rock formations in both cases (see
<figureCitation id="133B8985AB45B233FBACFF5DC9F4F3B2" box="[1103,1169,188,213]" captionStart="FIGURE 1" captionStartId="6.[151,250,1250,1272]" captionTargetBox="[151,1436,265,1229]" captionTargetId="figure@6.[151,1436,265,1229]" captionTargetPageId="6" captionText="FIGURE 1. Habitat of L. pantai sp. nov. at the type locality on Kei Kecil. Specimens were found exclusively in association with limestone rocks in the intertidal zone." httpUri="https://zenodo.org/record/1051482/files/figure.png" pageId="6" pageNumber="97">Fig. 1</figureCitation>
; habitat).
</paragraph>
<caption id="DF7FC588AB45B233FF74FB03CF2DF671" httpUri="https://zenodo.org/record/1051482/files/figure.png" pageId="6" pageNumber="97" targetBox="[151,1436,265,1229]" targetPageId="6">
<paragraph id="8BBF9500AB45B233FF74FB03CF2DF671" blockId="6.[151,1435,1250,1303]" pageId="6" pageNumber="97">
<emphasis id="B9744912AB45B233FF74FB03CC71F79E" bold="true" box="[151,276,1250,1272]" pageId="6" pageNumber="97">FIGURE 1.</emphasis>
Habitat of
<taxonomicName id="4C00EE83AB45B233FE68FB02CC8DF79E" authority="Stubbs &amp; Karin &amp; Arifin &amp; Iskandar &amp; Arida &amp; Reilly &amp; Bloch &amp; Kusnadi &amp; Mcguire, 2017" authorityName="Stubbs &amp; Karin &amp; Arifin &amp; Iskandar &amp; Arida &amp; Reilly &amp; Bloch &amp; Kusnadi &amp; Mcguire" authorityYear="2017" box="[395,488,1251,1272]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="97" phylum="Chordata" rank="species" species="pantai" status="sp. nov.">
<emphasis id="B9744912AB45B233FE68FB02CC8DF79E" box="[395,488,1251,1272]" italics="true" pageId="6" pageNumber="97">L. pantai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247F469AB45B233FE0CFB03CF24F79E" box="[495,577,1250,1272]" pageId="6" pageNumber="97" rank="species">
<emphasis id="B9744912AB45B233FE0CFB03CF24F79E" bold="true" box="[495,577,1250,1272]" pageId="6" pageNumber="97">sp. nov.</emphasis>
</taxonomicNameLabel>
at the type locality on Kei Kecil. Specimens were found exclusively in association with limestone rocks in the intertidal zone.
</paragraph>
</caption>
<caption id="DF7FC588AB45B233FF74F831C9A0F480" box="[151,1221,2000,2022]" httpUri="https://zenodo.org/record/1051484/files/figure.png" pageId="6" pageNumber="97" targetBox="[320,1227,1353,1935]" targetPageId="6">
<paragraph id="8BBF9500AB45B233FF74F831C9A0F480" blockId="6.[151,1221,2000,2022]" box="[151,1221,2000,2022]" pageId="6" pageNumber="97">
<emphasis id="B9744912AB45B233FF74F831CC77F480" bold="true" box="[151,274,2000,2022]" pageId="6" pageNumber="97">FIGURE 2.</emphasis>
Photo in life of
<taxonomicName id="4C00EE83AB45B233FE5BF831CFF6F480" authority="Stubbs &amp; Karin &amp; Arifin &amp; Iskandar &amp; Arida &amp; Reilly &amp; Bloch &amp; Kusnadi &amp; Mcguire, 2017" authorityName="Stubbs &amp; Karin &amp; Arifin &amp; Iskandar &amp; Arida &amp; Reilly &amp; Bloch &amp; Kusnadi &amp; Mcguire" authorityYear="2017" box="[440,659,2000,2022]" class="Reptilia" family="Gekkonidae" genus="Lepidodacylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="97" phylum="Chordata" rank="species" species="pantai" status="sp. nov.">
<emphasis id="B9744912AB45B233FE5BF831CFF6F480" box="[440,659,2000,2022]" italics="true" pageId="6" pageNumber="97">Lepidodacylus pantai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247F469AB45B233FD7AF831CF8CF480" box="[665,745,2000,2022]" pageId="6" pageNumber="97" rank="species">
<emphasis id="B9744912AB45B233FD7AF831CF8CF480" bold="true" box="[665,745,2000,2022]" pageId="6" pageNumber="97">sp. nov.</emphasis>
</taxonomicNameLabel>
from the type locality. (Photo by LMB, 2014).
</paragraph>
</caption>
<caption id="DF7FC588AB44B232FF74FB00C8F9F791" box="[151,1436,1249,1271]" httpUri="https://zenodo.org/record/1051486/files/figure.png" pageId="7" pageNumber="98" targetBox="[151,1436,193,1228]" targetPageId="7">
<paragraph id="8BBF9500AB44B232FF74FB00C8F9F791" blockId="7.[151,1436,1249,1271]" box="[151,1436,1249,1271]" pageId="7" pageNumber="98">
<emphasis id="B9744912AB44B232FF74FB00CC77F791" bold="true" box="[151,274,1249,1271]" pageId="7" pageNumber="98">FIGURE 3.</emphasis>
Plantar view of the right foot of the holotype (MZB.Lace.14069). Note the 3 divided terminal scansors on toes 1-4.
</paragraph>
</caption>
<paragraph id="8BBF9500AB44B232FF24FAFFCC7AF5A5" blockId="7.[151,1437,1310,1984]" pageId="7" pageNumber="98">
All specimens were found at night on beachside limestone rocks (
<figureCitation id="133B8985AB44B232FC49FAFFCE8FF651" box="[938,1002,1310,1335]" captionStart="FIGURE 1" captionStartId="6.[151,250,1250,1272]" captionTargetBox="[151,1436,265,1229]" captionTargetId="figure@6.[151,1436,265,1229]" captionTargetPageId="6" captionText="FIGURE 1. Habitat of L. pantai sp. nov. at the type locality on Kei Kecil. Specimens were found exclusively in association with limestone rocks in the intertidal zone." httpUri="https://zenodo.org/record/1051482/files/figure.png" pageId="7" pageNumber="98">Fig. 1</figureCitation>
) or in mangroves within meters of the high tide line. Despite nearly a month of intensive nightly collecting effort (targeting geckos) on Kei Kecil we were unable to find this species in the adjacent disturbed forest. While on Kei Kecil, we resided in a small house approximately 30 meters from the high tide line and collected many other species of geckos on a small wooden shed but
<taxonomicName id="4C00EE83AB44B232FF1CFA51CC03F6A1" authority="Stubbs &amp; Karin &amp; Arifin &amp; Iskandar &amp; Arida &amp; Reilly &amp; Bloch &amp; Kusnadi &amp; Mcguire, 2017" authorityName="Stubbs &amp; Karin &amp; Arifin &amp; Iskandar &amp; Arida &amp; Reilly &amp; Bloch &amp; Kusnadi &amp; Mcguire" authorityYear="2017" box="[255,358,1456,1479]" class="Reptilia" family="Gekkonidae" genus="Lepidodacylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="98" phylum="Chordata" rank="species" species="pantai" status="sp. nov.">
<emphasis id="B9744912AB44B232FF1CFA51CC03F6A1" box="[255,358,1456,1479]" italics="true" pageId="7" pageNumber="98">L. pantai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247F469AB44B232FE8DFA4ECCACF6A1" box="[366,457,1455,1479]" pageId="7" pageNumber="98" rank="species">
<emphasis id="B9744912AB44B232FE8DFA4ECCACF6A1" bold="true" box="[366,457,1455,1479]" pageId="7" pageNumber="98">sp. nov.</emphasis>
</taxonomicNameLabel>
were never encountered on this structure. This species was discovered accidentally at the end of our stay on Kei Kecil while surveying for laticaudine sea snakes in the intertidal zone. We think this shows the potential importance of sampling intertidal areas at night, something rarely done by herpetologists. All
<specimenCount id="9D065E89AB44B232FF74F9FACC55F555" box="[151,304,1562,1587]" pageId="7" pageNumber="98" type="generic">11 specimens</specimenCount>
in the
<typeStatus id="54BB2BA2AB44B232FE67F9FACCD1F555" box="[388,436,1563,1587]" pageId="7" pageNumber="98">type</typeStatus>
series were collected over the span of less than one hour, and subsequent visits to Kei Kecil confirmed that this species is locally abundant, occasionally in densities of up to 1 individual per square meter. Terrestrial hermit crabs and marine isopods were also abundant on the same rocks and we believe this resource subsidy of food from the marine environment likely contribute to the locally high densities observed in this species.
</paragraph>
<paragraph id="8BBF9500AB44B232FF24F92EC9E8F4D9" blockId="7.[151,1437,1310,1984]" pageId="7" pageNumber="98">
On Kur island, only
<quantity id="4CF838E5AB44B232FE20F92ECF75F581" box="[451,528,1742,1767]" metricMagnitude="4" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="98" unit="km" value="80.0">80 km</quantity>
northwest of Kei Kecil, morphologically similar geckos were also found in association with the intertidal zone and limestone rocks, but they were also numerous in a small patch of mangroves. The roots of these mangroves were submerged during high tide, but specimens were found at an even higher density on mangroves than on exposed limestone rocks (though the small mangroves were themselves growing among limestone rocks). When associated with mangroves, this species appeared to be very abundant close to the water, but trees slightly above the high-tide line were inhabited almost exclusively by another undescribed species of
<taxonomicName id="4C00EE83AB44B232FE78F846CF23F4D9" box="[411,582,1959,1983]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="98" phylum="Chordata" rank="genus">
<emphasis id="B9744912AB44B232FE78F846CF23F4D9" box="[411,582,1959,1983]" italics="true" pageId="7" pageNumber="98">Lepidodactylus</emphasis>
</taxonomicName>
and members of the gekkonid genus
<taxonomicName id="4C00EE83AB44B232FC0AF846C9E3F4D9" authorityName="Gray" authorityYear="1827" box="[1001,1158,1959,1983]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="98" phylum="Chordata" rank="genus">
<emphasis id="B9744912AB44B232FC0AF846C9E3F4D9" box="[1001,1158,1959,1983]" italics="true" pageId="7" pageNumber="98">Cyrtodactylus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DF7FC588AB4BB23DFF74FB6BC82CF7C6" box="[151,1353,1162,1184]" httpUri="https://zenodo.org/record/1051488/files/figure.png" pageId="8" pageNumber="99" targetBox="[151,1436,215,1098]" targetPageId="8">
<paragraph id="8BBF9500AB4BB23DFF74FB6BC82CF7C6" blockId="8.[151,1353,1162,1184]" box="[151,1353,1162,1184]" pageId="8" pageNumber="99">
<emphasis id="B9744912AB4BB23DFF74FB6BCC77F7C6" bold="true" box="[151,274,1162,1184]" pageId="8" pageNumber="99">FIGURE 4.</emphasis>
Dorsal color pattern variation in the type series. Specimen tags overlaid with museum accession numbers.
</paragraph>
</caption>
<subSubSection id="C31AC68BAB4BB23DFF24FB2CCCE4F4F7" pageId="8" pageNumber="99" type="etymology">
<paragraph id="8BBF9500AB4BB23DFF24FB2CCF1CF637" blockId="8.[151,1437,1228,1937]" pageId="8" pageNumber="99">
<emphasis id="B9744912AB4BB23DFF24FB2CCC2BF783" bold="true" box="[199,334,1229,1253]" pageId="8" pageNumber="99">Etymology.</emphasis>
The species epithet,
<taxonomicName id="4C00EE83AB4BB23DFDA7FB2FCFEEF783" authorityName="Stubbs &amp; Karin &amp; Arifin &amp; Iskandar &amp; Arida &amp; Reilly &amp; Bloch &amp; Kusnadi &amp; Mcguire" authorityYear="2017" box="[580,651,1230,1253]" class="Reptilia" family="Gekkonidae" genus="Lepidodacylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="pantai">
<emphasis id="B9744912AB4BB23DFDA7FB2FCFEEF783" box="[580,651,1230,1253]" italics="true" pageId="8" pageNumber="99">pantai</emphasis>
</taxonomicName>
, is the word for beach in the Indonesian national language (Bahasa
<collectingCountry id="F317D590AB4BB23DFF74FB11CC6CF66F" box="[151,265,1264,1289]" name="Indonesia" pageId="8" pageNumber="99">Indonesia</collectingCountry>
). It reflects the habitat in which the new species was discovered—a seemingly obligate association with the seashore. All specimens were found within
<quantity id="4CF838E5AB4BB23DFD48FAF4CFB3F64B" box="[683,726,1301,1325]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="99" unit="m" value="2.0">2 m</quantity>
of the high tide line. We suggest “Beach Scaly-toed Gecko” as the English common name for this species.
</paragraph>
<paragraph id="8BBF9500AB4BB23DFF24FABDC9E9F687" blockId="8.[151,1437,1228,1937]" pageId="8" pageNumber="99">
<emphasis id="B9744912AB4BB23DFF24FABDCC08F613" bold="true" box="[199,365,1372,1397]" pageId="8" pageNumber="99">Comparisons.</emphasis>
The new species is the only member of the genus with divided terminal 4th toe scansors and a cylindrical tail without lateral serrations.
<bibRefCitation id="EF91E8F1AB4BB23DFD99FA60CEF7F6FF" author="Brown" box="[634,914,1408,1433]" pageId="8" pageNumber="99" refString="Brown, W. &amp; Parker, F. (1977) Lizards of the genus Lepidodactylus (Gekkonidae) from the Indo-Australian Archipelago and the islands of the Pacific, with descriptions of new species. Proceedings of the California Academy of Science, 41, 253 - 265." type="journal article" year="1977">Brown &amp; Parker (1977)</bibRefCitation>
divided the genus
<taxonomicName id="4C00EE83AB4BB23DFB94FA60C847F6FF" box="[1143,1314,1409,1433]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="genus">
<emphasis id="B9744912AB4BB23DFB94FA60C847F6FF" box="[1143,1314,1409,1433]" italics="true" pageId="8" pageNumber="99">Lepidodactylus</emphasis>
</taxonomicName>
into three groups but
<taxonomicName id="4C00EE83AB4BB23DFEFEFA47CCE2F6DB" authority="Stubbs &amp; Karin &amp; Arifin &amp; Iskandar &amp; Arida &amp; Reilly &amp; Bloch &amp; Kusnadi &amp; Mcguire, 2017" authorityName="Stubbs &amp; Karin &amp; Arifin &amp; Iskandar &amp; Arida &amp; Reilly &amp; Bloch &amp; Kusnadi &amp; Mcguire" authorityYear="2017" box="[285,391,1446,1469]" class="Reptilia" family="Gekkonidae" genus="Lepidodacylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="pantai" status="sp. nov.">
<emphasis id="B9744912AB4BB23DFEFEFA47CCE2F6DB" box="[285,391,1446,1469]" italics="true" pageId="8" pageNumber="99">L. pantai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247F469AB4BB23DFE71FA44CC95F6DB" box="[402,496,1445,1469]" pageId="8" pageNumber="99" rank="species">
<emphasis id="B9744912AB4BB23DFE71FA44CC95F6DB" bold="true" box="[402,496,1445,1469]" pageId="8" pageNumber="99">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from all of the previously recognized groups in having divided terminal scansors on toes 25 and a tail that is fully cylindrical without any fringes or compression.
</paragraph>
<paragraph id="8BBF9500AB4BB23DFF24FA0CCCE4F4F7" blockId="8.[151,1437,1228,1937]" pageId="8" pageNumber="99">
The presence of divided scansors distinguishes
<taxonomicName id="4C00EE83AB4BB23DFD34FA0FCE59F563" authority="Stubbs &amp; Karin &amp; Arifin &amp; Iskandar &amp; Arida &amp; Reilly &amp; Bloch &amp; Kusnadi &amp; Mcguire, 2017" authorityName="Stubbs &amp; Karin &amp; Arifin &amp; Iskandar &amp; Arida &amp; Reilly &amp; Bloch &amp; Kusnadi &amp; Mcguire" authorityYear="2017" box="[727,828,1518,1541]" class="Reptilia" family="Gekkonidae" genus="Lepidodacylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="pantai" status="sp. nov.">
<emphasis id="B9744912AB4BB23DFD34FA0FCE59F563" box="[727,828,1518,1541]" italics="true" pageId="8" pageNumber="99">L. pantai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247F469AB4BB23DFCA1FA0CCEFEF563" box="[834,923,1517,1541]" pageId="8" pageNumber="99" rank="species">
<emphasis id="B9744912AB4BB23DFCA1FA0CCEFEF563" bold="true" box="[834,923,1517,1541]" pageId="8" pageNumber="99">sp. nov.</emphasis>
</taxonomicNameLabel>
from all Group I
<taxonomicName id="4C00EE83AB4BB23DFBBCFA0CC86FF563" box="[1119,1290,1517,1541]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="genus">
<emphasis id="B9744912AB4BB23DFBBCFA0CC86FF563" box="[1119,1290,1517,1541]" italics="true" pageId="8" pageNumber="99">Lepidodactylus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C00EE83AB4BB23DFAF9FA0FC812F563" baseAuthorityName="Boulenger" baseAuthorityYear="1888" box="[1306,1399,1517,1541]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="listeri">
<emphasis id="B9744912AB4BB23DFAF9FA0FC812F563" box="[1306,1399,1517,1541]" italics="true" pageId="8" pageNumber="99">L. listeri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00EE83AB4BB23DFA66FA0FCD8BF54E" authorityName="Brown &amp; Parker" authorityYear="1977" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="magnus">
<emphasis id="B9744912AB4BB23DFA66FA0FCD8BF54E" italics="true" pageId="8" pageNumber="99">L. magnus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00EE83AB4BB23DFF1FF9F0CC3AF54E" authorityName="Schmidt" authorityYear="1923" box="[252,351,1553,1576]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="manni">
<emphasis id="B9744912AB4BB23DFF1FF9F0CC3AF54E" box="[252,351,1553,1576]" italics="true" pageId="8" pageNumber="99">L. manni</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00EE83AB4BB23DFE8EF9F0CCBDF54F" authorityName="Brown &amp; Parker" authorityYear="1977" box="[365,472,1553,1577]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="mutahi">
<emphasis id="B9744912AB4BB23DFE8EF9F0CCBDF54F" box="[365,472,1553,1577]" italics="true" pageId="8" pageNumber="99">L. mutahi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00EE83AB4BB23DFE06F9F0CF5CF54E" baseAuthorityName="Kopstein" baseAuthorityYear="1926" box="[485,569,1553,1576]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="oorti">
<emphasis id="B9744912AB4BB23DFE06F9F0CF5CF54E" box="[485,569,1553,1576]" italics="true" pageId="8" pageNumber="99">L. oorti</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00EE83AB4BB23DFDA4F9F0CFAAF54F" authorityName="Brown &amp; Parker" authorityYear="1977" box="[583,719,1553,1577]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="orientalis">
<emphasis id="B9744912AB4BB23DFDA4F9F0CFAAF54F" box="[583,719,1553,1577]" italics="true" pageId="8" pageNumber="99">L. orientalis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00EE83AB4BB23DFD3EF9F0CE31F54F" baseAuthorityName="Boulenger" baseAuthorityYear="1885" box="[733,852,1553,1577]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="B9744912AB4BB23DFD3EF9F0CE31F54F" box="[733,852,1553,1577]" italics="true" pageId="8" pageNumber="99">L. pumilus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00EE83AB4BB23DFC82F9F0CEA8F54F" authorityName="Pernetta &amp; Black" authorityYear="1983" box="[865,973,1553,1577]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="browni">
<emphasis id="B9744912AB4BB23DFC82F9F0CEA8F54F" box="[865,973,1553,1577]" italics="true" pageId="8" pageNumber="99">L. browni</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00EE83AB4BB23DFC38F9F0C93CF54E" authorityName="Gibbons &amp; Brown" authorityYear="1988" box="[987,1113,1553,1576]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="euaensis">
<emphasis id="B9744912AB4BB23DFC38F9F0C93CF54E" box="[987,1113,1553,1576]" italics="true" pageId="8" pageNumber="99">L. euaensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C00EE83AB4BB23DFB76F9F0C825F54F" authorityName="Brown, McCoy &amp; Rodda" authorityYear="1992" box="[1173,1344,1553,1577]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="flaviocularis">
<emphasis id="B9744912AB4BB23DFB76F9F0C825F54F" box="[1173,1344,1553,1577]" italics="true" pageId="8" pageNumber="99">L. flaviocularis</emphasis>
</taxonomicName>
), which have no divided scansors (
<bibRefCitation id="EF91E8F1AB4BB23DFE3CF9D4CF93F52B" author="Brown" box="[479,758,1589,1614]" pageId="8" pageNumber="99" refString="Brown, W. &amp; Parker, F. (1977) Lizards of the genus Lepidodactylus (Gekkonidae) from the Indo-Australian Archipelago and the islands of the Pacific, with descriptions of new species. Proceedings of the California Academy of Science, 41, 253 - 265." type="journal article" year="1977">Brown &amp; Parker 1977</bibRefCitation>
). The presence of divided terminal scansors on toe IV distinguishes
<taxonomicName id="4C00EE83AB4BB23DFED7F9B8CCF9F516" authority="Stubbs &amp; Karin &amp; Arifin &amp; Iskandar &amp; Arida &amp; Reilly &amp; Bloch &amp; Kusnadi &amp; Mcguire, 2017" authorityName="Stubbs &amp; Karin &amp; Arifin &amp; Iskandar &amp; Arida &amp; Reilly &amp; Bloch &amp; Kusnadi &amp; Mcguire" authorityYear="2017" box="[308,412,1625,1648]" class="Reptilia" family="Gekkonidae" genus="Lepidodacylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="pantai" status="sp. nov.">
<emphasis id="B9744912AB4BB23DFED7F9B8CCF9F516" box="[308,412,1625,1648]" italics="true" pageId="8" pageNumber="99">L. pantai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247F469AB4BB23DFE4BF9B8CF60F517" box="[424,517,1625,1649]" pageId="8" pageNumber="99" rank="species">
<emphasis id="B9744912AB4BB23DFE4BF9B8CF60F517" bold="true" box="[424,517,1625,1649]" pageId="8" pageNumber="99">sp. nov.</emphasis>
</taxonomicNameLabel>
from all Group II
<taxonomicName id="4C00EE83AB4BB23DFD0AF9B8CEF1F517" box="[745,916,1625,1649]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="genus">
<emphasis id="B9744912AB4BB23DFD0AF9B8CEF1F517" box="[745,916,1625,1649]" italics="true" pageId="8" pageNumber="99">Lepidodactylus</emphasis>
</taxonomicName>
which have undivided terminal scansors (
<taxonomicName id="4C00EE83AB4BB23DFA66F9B8CC61F5F3" authorityName="Brown &amp; Parker" authorityYear="1977" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="gardeneri">
<emphasis id="B9744912AB4BB23DFA66F9B8CC61F5F3" italics="true" pageId="8" pageNumber="99">L. gardeneri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00EE83AB4BB23DFEF9F99FCCE9F5F3" authorityName="Boulenger" authorityYear="1884" box="[282,396,1662,1685]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="guppyi">
<emphasis id="B9744912AB4BB23DFEF9F99FCCE9F5F3" box="[282,396,1662,1685]" italics="true" pageId="8" pageNumber="99">L. guppyi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00EE83AB4BB23DFE41F99FCF31F5F3" authorityName="Brown &amp; Parker" authorityYear="1977" box="[418,596,1662,1685]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="novaeguineae">
<emphasis id="B9744912AB4BB23DFE41F99FCF31F5F3" box="[418,596,1662,1685]" italics="true" pageId="8" pageNumber="99">L.novaeguineae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00EE83AB4BB23DFD89F99FCF81F5F3" authorityName="Boulenger" authorityYear="1885" box="[618,740,1661,1685]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="pulcher">
<emphasis id="B9744912AB4BB23DFD89F99FCF81F5F3" box="[618,740,1661,1685]" italics="true" pageId="8" pageNumber="99">L. pulcher</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00EE83AB4BB23DFD19F99FCE0BF5F3" authorityName="Brown &amp; Tanner" authorityYear="1949" box="[762,878,1661,1685]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="shebae">
<emphasis id="B9744912AB4BB23DFD19F99FCE0BF5F3" box="[762,878,1661,1685]" italics="true" pageId="8" pageNumber="99">L. shebae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00EE83AB4BB23DFC67F99FCE8FF5F3" authorityName="Ineich" authorityYear="2008" box="[900,1002,1661,1685]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="buleli">
<emphasis id="B9744912AB4BB23DFC67F99FCE8FF5F3" box="[900,1002,1661,1685]" italics="true" pageId="8" pageNumber="99">L. buleli</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00EE83AB4BB23DFC1CF99FC9CCF5F3" authorityName="Darevsky" authorityYear="1964" box="[1023,1193,1661,1685]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="B9744912AB4BB23DFC1CF99FC9CCF5F3" box="[1023,1193,1661,1685]" italics="true" pageId="8" pageNumber="99">L. intermedius</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00EE83AB4BB23DFB5CF99FC80AF5F3" authorityName="Mertens" authorityYear="1929" box="[1215,1391,1661,1685]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="lombocensis">
<emphasis id="B9744912AB4BB23DFB5CF99FC80AF5F3" box="[1215,1391,1661,1685]" italics="true" pageId="8" pageNumber="99">L. lombocensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00EE83AB4BB23DFA66F99FCC6BF5DF" authorityName="Ota, Fisher &amp; Ineich" authorityYear="1995" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="paurolepis">
<emphasis id="B9744912AB4BB23DFA66F99FCC6BF5DF" italics="true" pageId="8" pageNumber="99">L. paurolepis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00EE83AB4BB23DFEC3F940CCA3F5DE" authorityName="Ota, Fisher, Ineich, Case, Radtkey &amp; Zug" authorityYear="1998" box="[288,454,1697,1720]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="vanatuensis">
<emphasis id="B9744912AB4BB23DFEC3F940CCA3F5DE" box="[288,454,1697,1720]" italics="true" pageId="8" pageNumber="99">L. vanatuensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00EE83AB4BB23DFE3BF940CF16F5DF" authorityName="Buden" authorityYear="2007" box="[472,627,1697,1721]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="oligoporus">
<emphasis id="B9744912AB4BB23DFE3BF940CF16F5DF" box="[472,627,1697,1721]" italics="true" pageId="8" pageNumber="99">L. oligoporus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00EE83AB4BB23DFD65F940CE72F5DF" authorityName="Zug, Watling, Alefaio, Alefaio &amp; Ludescher" authorityYear="2003" box="[646,791,1697,1721]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="tepukapili">
<emphasis id="B9744912AB4BB23DFD65F940CE72F5DF" box="[646,791,1697,1721]" italics="true" pageId="8" pageNumber="99">L. tepukapili</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C00EE83AB4BB23DFCBDF940CE9EF5DE" authorityName="Ota &amp; Hikida Ota &amp; Hikida" authorityYear="1988" box="[862,1019,1697,1720]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="ranauensis">
<emphasis id="B9744912AB4BB23DFCBDF940CE9EF5DE" box="[862,1019,1697,1720]" italics="true" pageId="8" pageNumber="99">L. ranauensis</emphasis>
</taxonomicName>
). The presence of a cylindrical tail without lateral serrations distinguishes
<taxonomicName id="4C00EE83AB4BB23DFDB1F927CFD2F5BB" authority="Stubbs &amp; Karin &amp; Arifin &amp; Iskandar &amp; Arida &amp; Reilly &amp; Bloch &amp; Kusnadi &amp; Mcguire, 2017" authorityName="Stubbs &amp; Karin &amp; Arifin &amp; Iskandar &amp; Arida &amp; Reilly &amp; Bloch &amp; Kusnadi &amp; Mcguire" authorityYear="2017" box="[594,695,1734,1757]" class="Reptilia" family="Gekkonidae" genus="Lepidodacylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="pantai" status="sp. nov.">
<emphasis id="B9744912AB4BB23DFDB1F927CFD2F5BB" box="[594,695,1734,1757]" italics="true" pageId="8" pageNumber="99">L. pantai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247F469AB4BB23DFD23F924CE7EF5BB" box="[704,795,1733,1757]" pageId="8" pageNumber="99" rank="species">
<emphasis id="B9744912AB4BB23DFD23F924CE7EF5BB" bold="true" box="[704,795,1733,1757]" pageId="8" pageNumber="99">sp. nov.</emphasis>
</taxonomicNameLabel>
from Group III
<taxonomicName id="4C00EE83AB4BB23DFC37F924C91AF5BB" box="[980,1151,1733,1757]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="genus">
<emphasis id="B9744912AB4BB23DFC37F924C91AF5BB" box="[980,1151,1733,1757]" italics="true" pageId="8" pageNumber="99">Lepidodactylus</emphasis>
</taxonomicName>
that have depressed tails with lateral serrations (
<taxonomicName id="4C00EE83AB4BB23DFE4CF908CF4BF466" baseAuthorityName="Peters" baseAuthorityYear="1867" box="[431,558,1769,1792]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="moestus">
<emphasis id="B9744912AB4BB23DFE4CF908CF4BF466" box="[431,558,1769,1792]" italics="true" pageId="8" pageNumber="99">L. moestus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00EE83AB4BB23DFDA7F908CFA0F467" baseAuthorityName="Dumeril &amp; Bibron" baseAuthorityYear="1836" box="[580,709,1769,1793]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="lugubris">
<emphasis id="B9744912AB4BB23DFDA7F908CFA0F467" box="[580,709,1769,1793]" italics="true" pageId="8" pageNumber="99">L. lugubris</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00EE83AB4BB23DFD38F908CE17F467" authorityName="Boulenger" authorityYear="1887" box="[731,882,1769,1793]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="woodfordi">
<emphasis id="B9744912AB4BB23DFD38F908CE17F467" box="[731,882,1769,1793]" italics="true" pageId="8" pageNumber="99">L. woodfordi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00EE83AB4BB23DFC64F908CE84F466" authorityName="Ota" authorityYear="1987" box="[903,993,1769,1792]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="yami">
<emphasis id="B9744912AB4BB23DFC64F908CE84F466" box="[903,993,1769,1792]" italics="true" pageId="8" pageNumber="99">L. yami</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00EE83AB4BB23DFC14F908C9D0F467" authorityName="Taylor" authorityYear="1915" box="[1015,1205,1769,1793]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="aureolineatus">
<emphasis id="B9744912AB4BB23DFC14F908C9D0F467" box="[1015,1205,1769,1793]" italics="true" pageId="8" pageNumber="99">L. aureolineatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00EE83AB4BB23DFB29F908C815F467" authorityName="Ota &amp; Crombie" authorityYear="1989" box="[1226,1392,1769,1793]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="balioburius">
<emphasis id="B9744912AB4BB23DFB29F908C815F467" box="[1226,1392,1769,1793]" italics="true" pageId="8" pageNumber="99">L. balioburius</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00EE83AB4BB23DFA66F908CC67F443" authorityName="Taylor" authorityYear="1917" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="christiani">
<emphasis id="B9744912AB4BB23DFA66F908CC67F443" italics="true" pageId="8" pageNumber="99">L. christiani</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00EE83AB4BB23DFEF1F8EFCC13F443" authorityName="Taylor" authorityYear="1923" box="[274,374,1805,1829]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="herrei">
<emphasis id="B9744912AB4BB23DFEF1F8EFCC13F443" box="[274,374,1805,1829]" italics="true" pageId="8" pageNumber="99">L. herrei</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C00EE83AB4BB23DFE5BF8EFCF04F443" authorityName="Stejneger" authorityYear="1905" box="[440,609,1805,1829]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="planicaudus">
<emphasis id="B9744912AB4BB23DFE5BF8EFCF04F443" box="[440,609,1805,1829]" italics="true" pageId="8" pageNumber="99">L. planicaudus</emphasis>
</taxonomicName>
). Lateral serrations are sometimes absent on
<taxonomicName id="4C00EE83AB4BB23DFB8FF8EFC82FF443" authority="(Ota 1987)" authorityName="Ota" authorityYear="1987" baseAuthorityName="Ota" baseAuthorityYear="1987" box="[1132,1354,1805,1829]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="yami">
<emphasis id="B9744912AB4BB23DFB8FF8EFC9A7F443" box="[1132,1218,1806,1829]" italics="true" pageId="8" pageNumber="99">L. yami</emphasis>
(
<bibRefCitation id="EF91E8F1AB4BB23DFB30F8ECC827F443" author="Ota" box="[1235,1346,1805,1829]" pageId="8" pageNumber="99" refString="Ota, H. (1987) A new species of Lepidodactylus (Gekkonidae: Reptilia) from Lanyu Island Taiwan. Copeia, 1987, 164 - 169. https: // doi. org / 10.2307 / 1446048" type="journal article" year="1987">Ota 1987</bibRefCitation>
)
</taxonomicName>
, but
<taxonomicName id="4C00EE83AB4BB23DFA66F8EFCDBAF42E" authority="Stubbs &amp; Karin &amp; Arifin &amp; Iskandar &amp; Arida &amp; Reilly &amp; Bloch &amp; Kusnadi &amp; Mcguire, 2017" authorityName="Stubbs &amp; Karin &amp; Arifin &amp; Iskandar &amp; Arida &amp; Reilly &amp; Bloch &amp; Kusnadi &amp; Mcguire" authorityYear="2017" class="Reptilia" family="Gekkonidae" genus="Lepidodacylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="pantai" status="sp. nov.">
<emphasis id="B9744912AB4BB23DFA66F8EFCDBAF42E" italics="true" pageId="8" pageNumber="99">L. pantai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247F469AB4BB23DFF06F8D0CC5AF42F" box="[229,319,1841,1865]" pageId="8" pageNumber="99" rank="species">
<emphasis id="B9744912AB4BB23DFF06F8D0CC5AF42F" bold="true" box="[229,319,1841,1865]" pageId="8" pageNumber="99">sp. nov.</emphasis>
</taxonomicNameLabel>
can be further distinguished from
<taxonomicName id="4C00EE83AB4BB23DFD20F8D0CE73F42E" baseAuthorityName="Ota" baseAuthorityYear="1987" box="[707,790,1841,1864]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="yami">
<emphasis id="B9744912AB4BB23DFD20F8D0CE73F42E" box="[707,790,1841,1864]" italics="true" pageId="8" pageNumber="99">L. yami</emphasis>
</taxonomicName>
by the number of scansors on Toe IV (
<quantity id="4CF838E5AB4BB23DFB2FF8D0C855F42F" box="[1228,1328,1840,1865]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="3.048" metricValueMin="2.54" pageId="8" pageNumber="99" unit="in" value="11.0" valueMax="12.0" valueMin="10.0">1012 in</quantity>
<taxonomicName id="4C00EE83AB4BB23DFAD4F8D0C8F9F42E" authority="Stubbs &amp; Karin &amp; Arifin &amp; Iskandar &amp; Arida &amp; Reilly &amp; Bloch &amp; Kusnadi &amp; Mcguire, 2017" authorityName="Stubbs &amp; Karin &amp; Arifin &amp; Iskandar &amp; Arida &amp; Reilly &amp; Bloch &amp; Kusnadi &amp; Mcguire" authorityYear="2017" box="[1335,1436,1841,1864]" class="Reptilia" family="Gekkonidae" genus="Lepidodacylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="pantai" status="sp. nov.">
<emphasis id="B9744912AB4BB23DFAD4F8D0C8F9F42E" box="[1335,1436,1841,1864]" italics="true" pageId="8" pageNumber="99">L. pantai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247F469AB4BB23DFF74F8B4CD97F40B" box="[151,242,1877,1901]" pageId="8" pageNumber="99" rank="species">
<emphasis id="B9744912AB4BB23DFF74F8B4CD97F40B" bold="true" box="[151,242,1877,1901]" pageId="8" pageNumber="99">sp. nov.</emphasis>
</taxonomicNameLabel>
<emphasis id="B9744912AB4BB23DFF1FF8B7CC26F40B" box="[252,323,1878,1901]" italics="true" pageId="8" pageNumber="99">versus</emphasis>
<quantity id="4CF838E5AB4BB23DFEAFF8B4CCD4F408" box="[332,433,1877,1902]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" metricValueMax="3.81" metricValueMin="3.302" pageId="8" pageNumber="99" unit="in" value="14.0" valueMax="15.0" valueMin="13.0">1315 in</quantity>
<taxonomicName id="4C00EE83AB4BB23DFE59F8B7CF75F40B" baseAuthorityName="Ota" baseAuthorityYear="1987" box="[442,528,1878,1901]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="yami">
<emphasis id="B9744912AB4BB23DFE59F8B7CF75F40B" box="[442,528,1878,1901]" italics="true" pageId="8" pageNumber="99">L. yami</emphasis>
</taxonomicName>
) and the number of midbody scale rows (
<quantity id="4CF838E5AB4BB23DFC17F8B4C913F408" box="[1012,1142,1877,1902]" metricMagnitude="0" metricUnit="m" metricValue="2.9844999999999997" metricValueMax="3.2257999999999996" metricValueMin="2.7432" pageId="8" pageNumber="99" unit="in" value="117.5" valueMax="127.0" valueMin="108.0">108127 in</quantity>
<taxonomicName id="4C00EE83AB4BB23DFB9CF8B7C983F40B" authority="Stubbs &amp; Karin &amp; Arifin &amp; Iskandar &amp; Arida &amp; Reilly &amp; Bloch &amp; Kusnadi &amp; Mcguire, 2017" authorityName="Stubbs &amp; Karin &amp; Arifin &amp; Iskandar &amp; Arida &amp; Reilly &amp; Bloch &amp; Kusnadi &amp; Mcguire" authorityYear="2017" box="[1151,1254,1878,1901]" class="Reptilia" family="Gekkonidae" genus="Lepidodacylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="pantai" status="sp. nov.">
<emphasis id="B9744912AB4BB23DFB9CF8B7C983F40B" box="[1151,1254,1878,1901]" italics="true" pageId="8" pageNumber="99">L. pantai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247F469AB4BB23DFB0CF8B4C82EF40A" box="[1263,1355,1877,1901]" pageId="8" pageNumber="99" rank="species">
<emphasis id="B9744912AB4BB23DFB0CF8B4C82EF40A" bold="true" box="[1263,1355,1877,1901]" pageId="8" pageNumber="99">sp. nov.</emphasis>
</taxonomicNameLabel>
<emphasis id="B9744912AB4BB23DFAB7F8B7C8FEF40B" box="[1364,1435,1878,1901]" italics="true" pageId="8" pageNumber="99">versus</emphasis>
<quantity id="4CF838E5AB4BB23DFF74F899CC73F4F7" box="[151,278,1912,1937]" metricMagnitude="0" metricUnit="m" metricValue="3.7211" metricValueMax="3.7592" metricValueMin="3.683" pageId="8" pageNumber="99" unit="in" value="146.5" valueMax="148.0" valueMin="145.0">145148 in</quantity>
<taxonomicName id="4C00EE83AB4BB23DFEFFF898CC15F4F6" baseAuthorityName="Ota" baseAuthorityYear="1987" box="[284,368,1913,1936]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="99" phylum="Chordata" rank="species" species="yami">
<emphasis id="B9744912AB4BB23DFEFFF898CC15F4F6" box="[284,368,1913,1936]" italics="true" pageId="8" pageNumber="99">L. yami</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>