309 lines
40 KiB
XML
309 lines
40 KiB
XML
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<mods:title id="B8ADDB03C05BB93BF16C2AA2476426EF">Revision of the Chinese endemic subgenus Tschitscherinea Berg, with descriptions of three new species (Coleoptera, Carabidae)</mods:title>
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<mods:namePart id="F16E7434821182016C28D551E8DF87F6">Yin, Wen-Qi</mods:namePart>
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<mods:affiliation id="E83093EE122C74E7090E430C76A2F0F9">College of Forestry, Beijing Forestry University, Beijing, 100083, P. R. China</mods:affiliation>
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<mods:namePart id="50E07D65F9CAF375460897EDF9AA9645">Chen, Jia-Heng</mods:namePart>
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<mods:affiliation id="C54A14F4A735C46ADE8024491E1E4730">College of Forestry, Beijing Forestry University, Beijing, 100083, P. R. China</mods:affiliation>
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<mods:namePart id="6EBCDBFE033ECD09898BF0C897377056">Shi, Hong-Liang</mods:namePart>
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<mods:affiliation id="A02C8074FAFD53C0D151787887712E4D">College of Forestry, Beijing Forestry University, Beijing, 100083, P. R. China</mods:affiliation>
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<mods:date id="19596668D4AAFA638B19207FCFB96657">2024</mods:date>
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<mods:number id="7842ED720B7967870123D262AF146D6F">2024-07-15</mods:number>
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<treatment id="03A6DA14FFFD8D71289D7DBDFE1FF4F4" ID-DOI="http://doi.org/10.5281/zenodo.12742734" ID-Zenodo-Dep="12742734" LSID="urn:lsid:plazi:treatment:03A6DA14FFFD8D71289D7DBDFE1FF4F4" httpUri="http://treatment.plazi.org/id/03A6DA14FFFD8D71289D7DBDFE1FF4F4" lastPageId="6" lastPageNumber="49" pageId="5" pageNumber="48">
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<paragraph id="8BB06B02FFFD8D72289D7DBDFD16F638" blockId="5.[151,751,1518,1617]" box="[151,751,1518,1545]" pageId="5" pageNumber="48">
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<heading id="D0F8DC6EFFFD8D72289D7DBDFD16F638" bold="true" box="[151,751,1518,1545]" fontSize="11" level="1" pageId="5" pageNumber="48" reason="1">
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<taxonomicName id="4C0F1081FFFD8D72289D7DBDFD73F638" authorityName="Yin & Chen & Shi" authorityYear="2024" box="[151,650,1518,1545]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="spectabilis" status="sp. nov." subGenus="Tschitscherinea">
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<emphasis id="B97BB710FFFD8D72289D7DBDFED4F638" bold="true" box="[151,301,1518,1545]" italics="true" pageId="5" pageNumber="48">Pterostichus</emphasis>
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(
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<emphasis id="B97BB710FFFD8D7229347DBDFE05F638" bold="true" box="[318,508,1518,1545]" italics="true" pageId="5" pageNumber="48">Tschitscherinea</emphasis>
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)
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<emphasis id="B97BB710FFFD8D722A077DBDFD73F638" bold="true" box="[525,650,1518,1545]" italics="true" pageId="5" pageNumber="48">spectabilis</emphasis>
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<taxonomicNameLabel id="A2480A6BFFFD8D722A987DBCFD16F638" box="[658,751,1519,1545]" pageId="5" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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<subSubSection id="C3153889FFFD8D72289D7E40FDB4F661" pageId="5" pageNumber="48" type="description">
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<paragraph id="8BB06B02FFFD8D72289D7E40FE51F61C" blockId="5.[151,751,1518,1617]" box="[151,424,1555,1581]" pageId="5" pageNumber="48">
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<figureCitation id="13347787FFFD8D72289D7E40FEFBF61C" box="[151,258,1555,1581]" captionStart="FIGURES 1–6" captionStartId="13.[152,270,1795,1819]" captionTargetBox="[158,1429,187,1766]" captionTargetId="figure-18@13.[151,1436,181,1771]" captionTargetPageId="13" captionText="FIGURES 1–6. Habitus of Pterostichus (Tschitscherinea) spp. (1) P. spissipedes sp. nov., holotype. (2) P. spissipedes sp. nov., a male paratype from Jianchuan. (3) P. spissipedes sp. nov., a male paratype from Deqin. (4) P. spectabilis sp. nov., holotype. (5) P. jiuzhaiicus sp. nov., holotype. (6) P. filum (Tschitschérine), a female from Wolong. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742726" httpUri="https://zenodo.org/record/12742726/files/figure.png" pageId="5" pageNumber="48">Figures 4</figureCitation>
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,
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<figureCitation id="13347787FFFD8D7229047E40FED5F61C" box="[270,300,1555,1581]" captionStart="FIGURES 13–18. 13–14" captionStartId="15.[152,270,1767,1791]" captionTargetBox="[158,1431,181,1736]" captionTargetId="figure-18@15.[151,1436,181,1743]" captionTargetPageId="15" captionText="FIGURES 13–18. 13–14. Habitus and labels of P. sterbai Jedlička: (13) holotype. (14) a male paratype. Scale bars: 2 mm. 15–18. Male genitalia of Pterostichus (Tschitscherinea) spp., left lateral view, dorsal view of median lobe of aedeagus, and right paramere. (15) P. sterbai Jedlička, Pe-Yen-Tsin. (16) P. spissipedes sp. nov., holotype. (17) P. spissipedes sp. nov., Deqin, paratype. (18) P. spectabilis sp. nov., holotype. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742732" httpUri="https://zenodo.org/record/12742732/files/figure.png" pageId="5" pageNumber="48">18</figureCitation>
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,
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<figureCitation id="13347787FFFD8D7229327E40FE51F61C" box="[312,424,1555,1581]" captionStart="FIGURES 35–47. 35–43" captionStartId="19.[152,270,1487,1511]" captionTargetBox="[158,1436,181,1457]" captionTargetId="figure-18@19.[151,1436,181,1463]" captionTargetPageId="19" captionText="FIGURES 35–47. 35–43. Metatarsi of Pterostichus (Tschitscherinea) spp. (35) P. spissipedes sp. nov., male. (36) P. spissipedes sp. nov., female. (37) P. spectabilis sp. nov., male. (38) P. spectabilis sp. nov., female. (39) P. mulensis Sciaky, male. (40) P. mulensis Sciaky, female. (41) P. farkaci Sciaky, male. (42) P. krali Sciaky, male. (43) P. sterbai Jedlička, male. Scale bar: 1 mm. 44–47. Elytral microsculpture of Pterostichus (Tschitscherinea) spp. (44) isodiametric, P. specillatus sp. nov., holotype. (45) transverse-isodiametric, P. mulensis Sciaky, a male from Muli. (46) isodiametric, P. farkaci Sciaky, a male from Tianchi, Zhongdian. (47) transverse-linear, P. krali Sciaky, a male from Haba Snow Mountain, Zhongdian. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742746" httpUri="https://zenodo.org/record/12742746/files/figure.png" pageId="5" pageNumber="48">37–38, 44</figureCitation>
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Chinese vernacular name:
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<emphasis id="B97BB710FFFD8D7229CB7E64FDB4F661" bold="true" box="[449,589,1591,1616]" pageId="5" pageNumber="48">îdzā±Ƥ</emphasis>
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locality.
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, male (
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,
|
||
<date id="FFB14DC2FFFD8D722A207EB7FD4CF6CF" box="[554,693,1764,1790]" pageId="5" pageNumber="48" value="2009-07-19">
|
||
<collectingDate id="EFF5B42AFFFD8D722A207EB7FD4CF6CF" box="[554,693,1764,1790]" pageId="5" pageNumber="48" value="2009-07-19">2009.VII.19</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26FA0ED4FFFD8D722ACA7EB7FCE6F6CE" box="[704,799,1764,1791]" pageId="5" pageNumber="48">Shi H.L.</collectorName>
|
||
coll.,
|
||
<collectingMethod id="524E1315FFFD8D722B697EB7FC25F6CE" box="[867,988,1764,1791]" pageId="5" pageNumber="48">Pitfall trap</collectingMethod>
|
||
,
|
||
<location id="8ED03DD9FFFD8D722BEC7EB6FB62F6CF" LSID="urn:lsid:plazi:treatment:03A6DA14FFFD8D71289D7DBDFE1FF4F4:8ED03DD9FFFD8D722BEC7EB6FB62F6CF" box="[998,1179,1764,1791]" country="China" county="Lijiang" latitude="27.63205" longLatPrecision="1" longitude="100.81781" municipality="Ninglang" name="Inst. of Zoology" pageId="5" pageNumber="48" stateProvince="Yunnan">Inst. of Zoology</location>
|
||
, CAS.”, “
|
||
<typeStatus id="54B4D5A0FFFD8D722D027EB7FA65F6CF" box="[1288,1436,1764,1790]" pageId="5" pageNumber="48" type="holotype">HOLOTYPE</typeStatus>
|
||
of
|
||
<taxonomicName id="4C0F1081FFFD8D7228BF7F5BFD7FF713" authorityName="Yin & Chen & Shi" authorityYear="2024" box="[181,646,1800,1826]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="spectabilis" status="sp. nov." subGenus="Tschitscherinea">
|
||
<emphasis id="B97BB710FFFD8D7228BF7F5BFEB9F713" box="[181,320,1800,1826]" italics="true" pageId="5" pageNumber="48">Pterostichus</emphasis>
|
||
(
|
||
<emphasis id="B97BB710FFFD8D7229457F5BFE07F713" box="[335,510,1800,1826]" italics="true" pageId="5" pageNumber="48">Tschitscherinea</emphasis>
|
||
)
|
||
<emphasis id="B97BB710FFFD8D722A047F5BFD7FF713" box="[526,646,1800,1826]" italics="true" pageId="5" pageNumber="48">spectabilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97BB710FFFD8D722A867F5BFD1CF713" bold="true" box="[652,741,1800,1826]" pageId="5" pageNumber="48">
|
||
<taxonomicNameLabel id="A2480A6BFFFD8D722A867F5BFD1CF713" box="[652,741,1800,1826]" pageId="5" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
det.
|
||
<determinerName id="7E6C87C6FFFD8D722B137F5BFCB8F713" box="[793,833,1800,1826]" pageId="5" pageNumber="48">Yin</determinerName>
|
||
& Shi, 2024” [red label]
|
||
</materialsCitation>
|
||
.
|
||
<emphasis id="B97BB710FFFD8D722C687F5BFB23F713" bold="true" box="[1122,1242,1800,1826]" pageId="5" pageNumber="48">
|
||
<typeStatus id="54B4D5A0FFFD8D722C687F5BFB23F713" box="[1122,1242,1800,1826]" pageId="5" pageNumber="48" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
(a total of
|
||
<specimenCount id="9D09A08BFFFD8D722D587F5BFA65F713" box="[1362,1436,1800,1826]" count="1" pageId="5" pageNumber="48" type="male">1 male</specimenCount>
|
||
and
|
||
<specimenCount id="9D09A08BFFFD8D7228C27F7FFED6F777" box="[200,303,1836,1862]" count="1" pageId="5" pageNumber="48" type="female">1 female</specimenCount>
|
||
),
|
||
<materialsCitation id="3B67615FFFFD8D7229497F7FFABAF776" box="[323,1347,1836,1863]" collectingDate="2009-07-17" collectionCode="IZAS" collectorName="Shi H. L." country="China" county="Lijiang" determinerName="Yin" elevation="3367" latitude="27.63205" location="Luguhu" longLatPrecision="1" longitude="100.81781" municipality="Ninglang" pageId="5" pageNumber="48" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
|
||
<specimenCount id="9D09A08BFFFD8D7229497F7FFE5CF777" box="[323,421,1836,1862]" count="1" pageId="5" pageNumber="48" type="female">1 female</specimenCount>
|
||
(
|
||
<collectionCode id="ED1EF3C7FFFD8D7229BC7F7FFE02F777" box="[438,507,1836,1862]" country="0" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="5" pageNumber="48">IZAS</collectionCode>
|
||
): the same data as holotype but date
|
||
<date id="FFB14DC2FFFD8D722B947F7FFBD0F777" box="[926,1065,1836,1862]" pageId="5" pageNumber="48" value="2009-07-17">
|
||
<collectingDate id="EFF5B42AFFFD8D722B947F7FFBD0F777" box="[926,1065,1836,1862]" pageId="5" pageNumber="48" value="2009-07-17">2009.VII.17</collectingDate>
|
||
</date>
|
||
, and labeled as
|
||
<typeStatus id="54B4D5A0FFFD8D722CD57F7EFABAF776" box="[1247,1347,1837,1863]" pageId="5" pageNumber="48" type="paratype">paratype</typeStatus>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B67615FFFFD8D722D5A7F7FFB06F783" collectingDate="2018-07-23" collectionCode="IZAS" country="China" county="Yanyuan County" determinerName="Yin" elevation="3468" latitude="27.5721" location="Shangluoshuidong" longLatPrecision="6" longitude="101.212" municipality="Mianbu Town" pageId="5" pageNumber="48" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan" typeStatus="paratype">
|
||
<specimenCount id="9D09A08BFFFD8D722D5A7F7FFA65F777" box="[1360,1436,1836,1862]" count="1" pageId="5" pageNumber="48" type="male">1 male</specimenCount>
|
||
(
|
||
<collectionCode id="ED1EF3C7FFFD8D7228957F03FF1DF75B" box="[159,228,1872,1898]" country="0" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="5" pageNumber="48">IZAS</collectionCode>
|
||
): “
|
||
<collectingCountry id="F3182B92FFFD8D7229087F03FE99F75B" box="[258,352,1872,1898]" name="China" pageId="5" pageNumber="48">CHINA</collectingCountry>
|
||
,
|
||
<collectingRegion id="49CBA5E0FFFD8D7229667F03FE30F75B" box="[364,457,1872,1898]" country="China" name="Sichuan" pageId="5" pageNumber="48">Sichuan</collectingRegion>
|
||
,
|
||
<collectingCounty id="62D1138EFFFD8D7229DF7F02FD6AF75B" box="[469,659,1872,1899]" pageId="5" pageNumber="48">Yanyuan County</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BD4F178FFFD8D722A957F03FCB9F75A" box="[671,832,1872,1899]" pageId="5" pageNumber="48">Mianbu Town</collectingMunicipality>
|
||
,
|
||
<location id="8ED03DD9FFFD8D722B467F03FBE6F75B" LSID="urn:lsid:plazi:treatment:03A6DA14FFFD8D71289D7DBDFE1FF4F4:8ED03DD9FFFD8D722B467F03FBE6F75B" box="[844,1055,1872,1898]" country="China" county="Yanyuan County" latitude="27.5721" longLatPrecision="6" longitude="101.212" municipality="Mianbu Town" name="Shangluoshuidong" pageId="5" pageNumber="48" stateProvince="Sichuan">Shangluoshuidong</location>
|
||
, mixed forest,
|
||
<quantity id="4CF7C6E7FFFD8D722CC37F03FAE3F75B" box="[1225,1306,1872,1898]" metricMagnitude="3" metricUnit="m" metricValue="3.468" pageId="5" pageNumber="48" unit="m" value="3468.0">
|
||
<elevation id="00228C31FFFD8D722CC37F03FAE3F75B" box="[1225,1306,1872,1898]" metricMagnitude="3" metricUnit="m" metricValue="3.468" pageId="5" pageNumber="48" unit="m" value="3468.0">3468m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="EE3B0DC5FFFD8D722D2C7F03FA61F75A" box="[1318,1432,1872,1899]" degrees="27.5721" direction="north" orientation="latitude" pageId="5" pageNumber="48" precision="5" value="27.5721">N27.5721</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3B0DC5FFFD8D72289D7F27FEEEF7BF" box="[151,279,1908,1934]" degrees="101.2120" direction="east" orientation="longitude" pageId="5" pageNumber="48" precision="5" value="101.212">E101.2120</geoCoordinate>
|
||
”, “
|
||
<date id="FFB14DC2FFFD8D72293C7F27FE3BF7BF" box="[310,450,1908,1934]" pageId="5" pageNumber="48" value="2018-07-23">
|
||
<collectingDate id="EFF5B42AFFFD8D72293C7F27FE3BF7BF" box="[310,450,1908,1934]" pageId="5" pageNumber="48" value="2018-07-23">2018.VII.23</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMethod id="524E1315FFFD8D7229C77F27FDBCF7BE" box="[461,581,1908,1935]" pageId="5" pageNumber="48">pitfall trap</collectingMethod>
|
||
. Shi H.L., Yan W.F., Zhu P.Z. & Jiang Z.Y. lgt., IZAS & BJFU exp. of 2018”, “
|
||
<typeStatus id="54B4D5A0FFFD8D7228AA7FCBFED7F783" box="[160,302,1944,1970]" pageId="5" pageNumber="48" type="paratype">PARATYPE</typeStatus>
|
||
of
|
||
<taxonomicName id="4C0F1081FFFD8D72295E7FCBFCDFF783" authorityName="Yin & Chen & Shi" authorityYear="2024" box="[340,806,1944,1970]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="spectabilis" status="sp. nov." subGenus="Tschitscherinea">
|
||
<emphasis id="B97BB710FFFD8D72295E7FCBFE26F783" box="[340,479,1944,1970]" italics="true" pageId="5" pageNumber="48">Pterostichus</emphasis>
|
||
(
|
||
<emphasis id="B97BB710FFFD8D7229E57FCBFD67F783" box="[495,670,1944,1970]" italics="true" pageId="5" pageNumber="48">Tschitscherinea</emphasis>
|
||
)
|
||
<emphasis id="B97BB710FFFD8D722AA47FCBFCDFF783" box="[686,806,1944,1970]" italics="true" pageId="5" pageNumber="48">spectabilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97BB710FFFD8D722B277FCBFC7EF783" bold="true" box="[813,903,1944,1970]" pageId="5" pageNumber="48">
|
||
<taxonomicNameLabel id="A2480A6BFFFD8D722B277FCBFC7EF783" box="[813,903,1944,1970]" pageId="5" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
det.
|
||
<determinerName id="7E6C87C6FFFD8D722BB77FCBFC1CF783" box="[957,997,1944,1970]" pageId="5" pageNumber="48">Yin</determinerName>
|
||
& Shi, 2024” [red label]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3153889FFFD8D7128CD7FEFFE17F154" lastPageId="6" lastPageNumber="49" pageId="5" pageNumber="48" type="diagnosis">
|
||
<paragraph id="8BB06B02FFFD8D7128CD7FEFFDB0F12C" blockId="5.[151,1436,1656,2043]" lastBlockId="6.[151,1437,151,1221]" lastPageId="6" lastPageNumber="49" pageId="5" pageNumber="48">
|
||
<emphasis id="B97BB710FFFD8D7228CD7FEFFEBBF7E7" bold="true" box="[199,322,1980,2006]" pageId="5" pageNumber="48">Diagnosis.</emphasis>
|
||
Very large and macrocephalic species, much larger than other species of the subgenus, BL =
|
||
<quantity id="4CF7C6E7FFFD8D722D577FEFFF06F7CA" metricMagnitude="-2" metricUnit="m" metricValue="1.515" metricValueMax="1.55" metricValueMin="1.48" pageId="5" pageNumber="48" unit="mm" value="15.15" valueMax="15.5" valueMin="14.8">14.8– 15.5 mm</quantity>
|
||
(versus 9.0–
|
||
<quantity id="4CF7C6E7FFFD8D7229927FB3FE06F7CA" box="[408,511,2016,2043]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="5" pageNumber="48" unit="mm" value="12.5">12.5 mm</quantity>
|
||
in other species); maxim width of head at outer margin of tempora; pronotum subquadrate; basal foveae distinctly ridged between lateral margin and outer groove. Elytra with isodiametric microsculpture (
|
||
<figureCitation id="13347787FFFE8D71294578E8FE5DF0E4" box="[335,420,187,213]" captionStart="FIGURES 35–47. 35–43" captionStartId="19.[152,270,1487,1511]" captionTargetBox="[158,1436,181,1457]" captionTargetId="figure-18@19.[151,1436,181,1463]" captionTargetPageId="19" captionText="FIGURES 35–47. 35–43. Metatarsi of Pterostichus (Tschitscherinea) spp. (35) P. spissipedes sp. nov., male. (36) P. spissipedes sp. nov., female. (37) P. spectabilis sp. nov., male. (38) P. spectabilis sp. nov., female. (39) P. mulensis Sciaky, male. (40) P. mulensis Sciaky, female. (41) P. farkaci Sciaky, male. (42) P. krali Sciaky, male. (43) P. sterbai Jedlička, male. Scale bar: 1 mm. 44–47. Elytral microsculpture of Pterostichus (Tschitscherinea) spp. (44) isodiametric, P. specillatus sp. nov., holotype. (45) transverse-isodiametric, P. mulensis Sciaky, a male from Muli. (46) isodiametric, P. farkaci Sciaky, a male from Tianchi, Zhongdian. (47) transverse-linear, P. krali Sciaky, a male from Haba Snow Mountain, Zhongdian. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742746" httpUri="https://zenodo.org/record/12742746/files/figure.png" pageId="6" pageNumber="49">Fig. 44</figureCitation>
|
||
); parascutellar pore absent; interval 3 with two setigerous pores; metatarsomere thickened in both sexes (
|
||
<figureCitation id="13347787FFFE8D71294B788CFE3EF0C8" box="[321,455,223,249]" captionStart="FIGURES 35–47. 35–43" captionStartId="19.[152,270,1487,1511]" captionTargetBox="[158,1436,181,1457]" captionTargetId="figure-18@19.[151,1436,181,1463]" captionTargetPageId="19" captionText="FIGURES 35–47. 35–43. Metatarsi of Pterostichus (Tschitscherinea) spp. (35) P. spissipedes sp. nov., male. (36) P. spissipedes sp. nov., female. (37) P. spectabilis sp. nov., male. (38) P. spectabilis sp. nov., female. (39) P. mulensis Sciaky, male. (40) P. mulensis Sciaky, female. (41) P. farkaci Sciaky, male. (42) P. krali Sciaky, male. (43) P. sterbai Jedlička, male. Scale bar: 1 mm. 44–47. Elytral microsculpture of Pterostichus (Tschitscherinea) spp. (44) isodiametric, P. specillatus sp. nov., holotype. (45) transverse-isodiametric, P. mulensis Sciaky, a male from Muli. (46) isodiametric, P. farkaci Sciaky, a male from Tianchi, Zhongdian. (47) transverse-linear, P. krali Sciaky, a male from Haba Snow Mountain, Zhongdian. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742746" httpUri="https://zenodo.org/record/12742746/files/figure.png" pageId="6" pageNumber="49">Figs 37–38</figureCitation>
|
||
), with outer-lateral ridge on metatarsomere 1; right paramere falcate, not carinate on dorsal surface, apex rounded (
|
||
<figureCitation id="13347787FFFE8D7129E27950FDC5F12C" box="[488,572,259,285]" captionStart="FIGURES 13–18. 13–14" captionStartId="15.[152,270,1767,1791]" captionTargetBox="[158,1431,181,1736]" captionTargetId="figure-18@15.[151,1436,181,1743]" captionTargetPageId="15" captionText="FIGURES 13–18. 13–14. Habitus and labels of P. sterbai Jedlička: (13) holotype. (14) a male paratype. Scale bars: 2 mm. 15–18. Male genitalia of Pterostichus (Tschitscherinea) spp., left lateral view, dorsal view of median lobe of aedeagus, and right paramere. (15) P. sterbai Jedlička, Pe-Yen-Tsin. (16) P. spissipedes sp. nov., holotype. (17) P. spissipedes sp. nov., Deqin, paratype. (18) P. spectabilis sp. nov., holotype. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742732" httpUri="https://zenodo.org/record/12742732/files/figure.png" pageId="6" pageNumber="49">Fig. 18</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB06B02FFFE8D7128CD7974FE17F154" blockId="6.[151,1437,151,1221]" pageId="6" pageNumber="49">This remarkable new species can be readily distinguished from all other related species by the very large size and macrocephalic body form.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3153889FFFE8D7128CD793DFACDF420" pageId="6" pageNumber="49" type="description">
|
||
<paragraph id="8BB06B02FFFE8D7128CD793DFACDF420" blockId="6.[151,1437,151,1221]" pageId="6" pageNumber="49">
|
||
<emphasis id="B97BB710FFFE8D7128CD793DFEA0F1B9" bold="true" box="[199,345,366,392]" pageId="6" pageNumber="49">Description.</emphasis>
|
||
BL =
|
||
<quantity id="4CF7C6E7FFFE8D712997793DFDB8F1B8" box="[413,577,366,393]" metricMagnitude="-2" metricUnit="m" metricValue="1.515" metricValueMax="1.55" metricValueMin="1.48" pageId="6" pageNumber="49" unit="mm" value="15.15" valueMax="15.5" valueMin="14.8">14.8–15.5 mm</quantity>
|
||
, BW =
|
||
<quantity id="4CF7C6E7FFFE8D712A99793CFCE2F1B8" box="[659,795,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="4.699999999999999" metricValueMax="4.8" metricValueMin="4.6" pageId="6" pageNumber="49" unit="mm" value="4.699999999999999" valueMax="4.8" valueMin="4.6">4.6–4.8 mm</quantity>
|
||
, dorsal surface and appendages dark reddish brown. Head very large and strongly thickened, slightly narrower than maxim width of pronotum, HW/PW = 0.78–0.81; frons without puncture; antennae short, hardly reaching elytra base; eyes very small, tempora strongly swollen behind eyes, length greater than eyes, eyL/tpL = 0.56–0.65, maxim width of head at outer margin of tempora. Pronotum subquadrate, PW/PL = 1.13–1.17, PBW/PW = 0.71–0.74, widest near anterior fourth; anterior margin slightly wider than posterior margin, PAW/PBW = 1.04–1.07; anterior angles narrowly rounded, apex weakly projected; lateral margins subparallel, nearly straight before posterior angles; posterior angles rectangular, apex with a faint denticle. Basal foveae shallow, with a few punctures; inner and outer grooves faintly defined, area between them depressed; outer groove about two-thirds length of inner one; area between lateral margin and outer groove distinctly ridged. Pronotum disc with fine transverse wrinkles aside median line; median line shallow. Elytra oblong and parallel-sided, BW/EL = 0.62–0.65, with isodiametric microsculpture in both sexes (
|
||
<figureCitation id="13347787FFFE8D712BF67A84FBABF2C0" box="[1020,1106,727,753]" captionStart="FIGURES 35–47. 35–43" captionStartId="19.[152,270,1487,1511]" captionTargetBox="[158,1436,181,1457]" captionTargetId="figure-18@19.[151,1436,181,1463]" captionTargetPageId="19" captionText="FIGURES 35–47. 35–43. Metatarsi of Pterostichus (Tschitscherinea) spp. (35) P. spissipedes sp. nov., male. (36) P. spissipedes sp. nov., female. (37) P. spectabilis sp. nov., male. (38) P. spectabilis sp. nov., female. (39) P. mulensis Sciaky, male. (40) P. mulensis Sciaky, female. (41) P. farkaci Sciaky, male. (42) P. krali Sciaky, male. (43) P. sterbai Jedlička, male. Scale bar: 1 mm. 44–47. Elytral microsculpture of Pterostichus (Tschitscherinea) spp. (44) isodiametric, P. specillatus sp. nov., holotype. (45) transverse-isodiametric, P. mulensis Sciaky, a male from Muli. (46) isodiametric, P. farkaci Sciaky, a male from Tianchi, Zhongdian. (47) transverse-linear, P. krali Sciaky, a male from Haba Snow Mountain, Zhongdian. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742746" httpUri="https://zenodo.org/record/12742746/files/figure.png" pageId="6" pageNumber="49">Fig. 44</figureCitation>
|
||
); parascutellar stria joined to stria 1, angular base of stria 1 relatively long, stria 1 entire; parascutellar pore absent; humerus with small tooth, slightly projected out; elytral interval 3 with two setigerous pores, both adjacent to stria 2, the first one near middle, the second one near apical fifth. Male sternite VII without secondary sexual modification; metafemora slightly thickened in males; metatarsus in a stout form in both sexes, more thickened in males (mt1 length/width =
|
||
<quantity id="4CF7C6E7FFFE8D712D517B34FA65F3B0" box="[1371,1436,871,897]" metricMagnitude="-2" metricUnit="m" metricValue="4.318" pageId="6" pageNumber="49" unit="in" value="1.7">1.7 in</quantity>
|
||
males, =
|
||
<quantity id="4CF7C6E7FFFE8D7128F37BD8FECEF394" box="[249,311,907,933]" metricMagnitude="-2" metricUnit="m" metricValue="5.334" pageId="6" pageNumber="49" unit="in" value="2.1">2.1 in</quantity>
|
||
females,
|
||
<figureCitation id="13347787FFFE8D7129947BD8FDE7F394" box="[414,542,907,933]" captionStart="FIGURES 35–47. 35–43" captionStartId="19.[152,270,1487,1511]" captionTargetBox="[158,1436,181,1457]" captionTargetId="figure-18@19.[151,1436,181,1463]" captionTargetPageId="19" captionText="FIGURES 35–47. 35–43. Metatarsi of Pterostichus (Tschitscherinea) spp. (35) P. spissipedes sp. nov., male. (36) P. spissipedes sp. nov., female. (37) P. spectabilis sp. nov., male. (38) P. spectabilis sp. nov., female. (39) P. mulensis Sciaky, male. (40) P. mulensis Sciaky, female. (41) P. farkaci Sciaky, male. (42) P. krali Sciaky, male. (43) P. sterbai Jedlička, male. Scale bar: 1 mm. 44–47. Elytral microsculpture of Pterostichus (Tschitscherinea) spp. (44) isodiametric, P. specillatus sp. nov., holotype. (45) transverse-isodiametric, P. mulensis Sciaky, a male from Muli. (46) isodiametric, P. farkaci Sciaky, a male from Tianchi, Zhongdian. (47) transverse-linear, P. krali Sciaky, a male from Haba Snow Mountain, Zhongdian. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742746" httpUri="https://zenodo.org/record/12742746/files/figure.png" pageId="6" pageNumber="49">Figs 37–38</figureCitation>
|
||
); metatarsomere 1 with outer-lateral ridge. Median lobe of aedeagus small (length about
|
||
<quantity id="4CF7C6E7FFFE8D7128D67BFCFEBDF3F8" box="[220,324,943,969]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="49" unit="mm" value="1.7">1.70 mm</quantity>
|
||
), apex not bent ventrally in lateral view; apical lamella short and wide, apex rounded, not twisted to right; right paramere falcate, elongated and curved, apical portion thickened, not carinate on dorsal surface, cross-section ovate at middle, apex rounded (
|
||
<figureCitation id="13347787FFFE8D712A457BA4FD5BF420" box="[591,674,1015,1041]" captionStart="FIGURES 13–18. 13–14" captionStartId="15.[152,270,1767,1791]" captionTargetBox="[158,1431,181,1736]" captionTargetId="figure-18@15.[151,1436,181,1743]" captionTargetPageId="15" captionText="FIGURES 13–18. 13–14. Habitus and labels of P. sterbai Jedlička: (13) holotype. (14) a male paratype. Scale bars: 2 mm. 15–18. Male genitalia of Pterostichus (Tschitscherinea) spp., left lateral view, dorsal view of median lobe of aedeagus, and right paramere. (15) P. sterbai Jedlička, Pe-Yen-Tsin. (16) P. spissipedes sp. nov., holotype. (17) P. spissipedes sp. nov., Deqin, paratype. (18) P. spectabilis sp. nov., holotype. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742732" httpUri="https://zenodo.org/record/12742732/files/figure.png" pageId="6" pageNumber="49">Fig. 18</figureCitation>
|
||
). Endophallus not studied due to restricted male materials.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3153889FFFE8D7128CD7C49FC15F468" pageId="6" pageNumber="49" type="distribution">
|
||
<paragraph id="8BB06B02FFFE8D7128CD7C49FC15F468" blockId="6.[151,1437,151,1221]" pageId="6" pageNumber="49">
|
||
<emphasis id="B97BB710FFFE8D7128CD7C49FE98F405" bold="true" box="[199,353,1050,1076]" pageId="6" pageNumber="49">Distribution.</emphasis>
|
||
This species is known from two localities on the border of
|
||
<collectingRegion id="49CBA5E0FFFE8D712BEF7C48FBC6F404" box="[997,1087,1051,1077]" country="China" name="Sichuan" pageId="6" pageNumber="49">Sichuan</collectingRegion>
|
||
and
|
||
<collectingRegion id="49CBA5E0FFFE8D712C787C48FB31F404" box="[1138,1224,1051,1077]" country="China" name="Yunnan" pageId="6" pageNumber="49">Yunnan</collectingRegion>
|
||
provinces (
|
||
<collectingCountry id="F3182B92FFFE8D712D4D7C48FA68F404" box="[1351,1425,1051,1077]" name="China" pageId="6" pageNumber="49">China</collectingCountry>
|
||
): Luguhu, Ninglang county and Mianbu pass, Yanyuan county. (
|
||
<figureCitation id="13347787FFFE8D712B597C6CFC5FF468" box="[851,934,1086,1113]" captionStart="FIGURES 54–57. 54" captionStartId="21.[152,270,1525,1549]" captionTargetBox="[151,1435,181,1494]" captionTargetId="figure-18@21.[151,1435,181,1501]" captionTargetPageId="21" captionText="FIGURES 54–57. 54. Confirmed distribution of the Pterostichus (Tschitscherinea) spp. in China: Red P. spissipedes sp. nov. Blue P. spectabilis sp. nov. Black P. jiuzhaiicus sp. nov. Pink P. filum (Tschitschérine). Purple P. mulensis Sciaky. Green P. fakarci Sciaky. Yellow P. krali Sciaky. Cyan P. sterbai Jedlička. 55–57. Endophallus of Pterostichus spp., right lateral view and left lateral view: (55) Pterostichus (Cryobius) jaechi Krischenhofer, Changbaishan, Jilin. (56) Pterostichus (Morphohaptoderus) janatai Sciaky & Wrase, Pingheliang, Shaanxi. (57) Pterostichus (Morphohaptoderus) dundai Sciaky, Houzhenzi, Shaanxi. Scale bars = 0.5 mm, a (55), b (56–57)." figureDoi="http://doi.org/10.5281/zenodo.12742754" httpUri="https://zenodo.org/record/12742754/files/figure.png" pageId="6" pageNumber="49">Fig. 54</figureCitation>
|
||
, blue)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3153889FFFE8D7128CD7C30FE1FF4F4" pageId="6" pageNumber="49" type="etymology">
|
||
<paragraph id="8BB06B02FFFE8D7128CD7C30FE1FF4F4" blockId="6.[151,1437,151,1221]" pageId="6" pageNumber="49">
|
||
<emphasis id="B97BB710FFFE8D7128CD7C30FEB7F44C" bold="true" box="[199,334,1123,1149]" pageId="6" pageNumber="49">Etymology.</emphasis>
|
||
The scientific name of the new species is derived from Latin, which means spectacular, referring the new species is remarkable among all
|
||
<taxonomicName id="4C0F1081FFFE8D712A577CD4FCC6F490" box="[605,831,1159,1185]" pageId="6" pageNumber="49">
|
||
<emphasis id="B97BB710FFFE8D712A577CD4FD11F490" box="[605,744,1159,1185]" italics="true" pageId="6" pageNumber="49">Pterostichus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
in southwest
|
||
<collectingCountry id="F3182B92FFFE8D712BDD7CD4FBE3F490" box="[983,1050,1159,1185]" name="China" pageId="6" pageNumber="49">China</collectingCountry>
|
||
for its special macrocephalic body form and relatively large size.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |