204 lines
24 KiB
XML
204 lines
24 KiB
XML
<document id="BEA4093843B842ED616E602894BEC0A8" ID-DOI="10.11646/zootaxa.5481.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="12742724" ID-ZooBank="FEC2B59D-D79F-4F9C-B082-D5BAABE1A453" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721033477194" checkinUser="plazi" docAuthor="Yin, Wen-Qi, Chen, Jia-Heng & Shi, Hong-Liang" docDate="2024" docId="03A6DA14FFF48D7B289D7AA9FB5FF773" docLanguage="en" docName="zootaxa.5481.1.3.pdf" docOrigin="Zootaxa 5481 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5481.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pterostichus (Morphohaptoderus) parvicollis Sciaky & Wrase 1997" docType="treatment" docVersion="2" lastPageNumber="55" masterDocId="FF9FA26CFFF88D77280A7853FFF9F031" masterDocTitle="Revision of the Chinese endemic subgenus Tschitscherinea Berg, with descriptions of three new species (Coleoptera, Carabidae)" masterLastPageNumber="65" masterPageNumber="43" pageNumber="55" updateTime="1721034011096" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9496655BE09AB5D4542120EAEF8DFA68">Revision of the Chinese endemic subgenus Tschitscherinea Berg, with descriptions of three new species (Coleoptera, Carabidae)</mods:title>
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<mods:namePart id="EBACA91D3F4DAF415371AEE9BB678187">Yin, Wen-Qi</mods:namePart>
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<mods:namePart id="C7BA4D0F58993FD3646DF587D2EB0BF2">Chen, Jia-Heng</mods:namePart>
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<mods:affiliation id="8D521943036838BFBEFF608CDD34B36B">College of Forestry, Beijing Forestry University, Beijing, 100083, P. R. China</mods:affiliation>
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<mods:affiliation id="8AFFA494D9D3809DDC1EBB759855E986">College of Forestry, Beijing Forestry University, Beijing, 100083, P. R. China</mods:affiliation>
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<mods:date id="E243C629F46B02C83DAB0A36C10422E7">2024</mods:date>
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<taxonomicName id="4C0F1081FFF48D7B289D7AA9FC2BF325" ID-CoL="4Q3Y3" authority="Sciaky & Wrase, 1997" authorityName="Sciaky & Wrase" authorityYear="1997" box="[151,978,762,789]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="55" phylum="Arthropoda" rank="species" species="parvicollis" subGenus="Morphohaptoderus">
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<emphasis id="B97BB710FFF48D7B289D7AA9FC2BF325" bold="true" box="[151,978,762,789]" pageId="12" pageNumber="55">
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<emphasis id="B97BB710FFF48D7B289D7AA9FED4F324" bold="true" box="[151,301,762,789]" italics="true" pageId="12" pageNumber="55">Pterostichus</emphasis>
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(
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<emphasis id="B97BB710FFF48D7B29347AA9FDDEF324" bold="true" box="[318,551,762,789]" italics="true" pageId="12" pageNumber="55">Morphohaptoderus</emphasis>
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)
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<emphasis id="B97BB710FFF48D7B2A327AA9FD4CF324" bold="true" box="[568,693,762,789]" italics="true" pageId="12" pageNumber="55">parvicollis</emphasis>
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<bibRefCitation id="EF9E16F3FFF48D7B2AB77AA9FC2BF325" author="Sciaky, R. & Wrase, D. W." box="[701,978,762,789]" pageId="12" pageNumber="55" pagination="1087 - 1139" refId="ref15565" refString="Sciaky, R. & Wrase, D. W. (1997) Twenty-nine new taxa of Pterostichinae from Shaanxi (Coleoptera, Carabidae). Linzer Biologische Beitrage, 29, 1087 - 1139." type="journal article" year="1997">Sciaky & Wrase, 1997</bibRefCitation>
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<subSubSection id="C3153889FFF48D7B289D7B4CFB19F3AB" pageId="12" pageNumber="55" type="description">
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<paragraph id="8BB06B02FFF48D7B289D7B4CFE75F308" blockId="12.[151,978,762,861]" box="[151,396,799,825]" pageId="12" pageNumber="55">
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<figureCitation id="13347787FFF48D7B289D7B4CFE75F308" box="[151,396,799,825]" captionStart="FIGURES 31–34" captionStartId="18.[152,270,1908,1932]" captionTargetBox="[184,1403,188,1878]" captionTargetId="figure-36@18.[177,1409,181,1885]" captionTargetPageId="18" captionText="FIGURES 31–34. habitus and male genitalia of P. parvicollis Sciaky & Wrase. (31) habitus of holotype. (32–34) a male specimen from Pingheliang, Shaanxi. (32) habitus. (33) male genitalia, left lateral view, dorsal view of median lobe of aedeagus, and right paramere. (34) endophallus. Abbreviations: gp gonopore, gpp gonopore piece, la left apical lobe, ra right apical lobe. Scale bars: 2 mm (31–32), 0.5 mm (33–34)." figureDoi="http://doi.org/10.5281/zenodo.12742744" httpUri="https://zenodo.org/record/12742744/files/figure.png" pageId="12" pageNumber="55">Figures 31, 32, 33, 34</figureCitation>
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Chinese vernacular name:
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<emphasis id="B97BB710FFF48D7B29CB7B10FD90F36C" bold="true" box="[449,617,835,861]" pageId="12" pageNumber="55">小ḃdzā±Ƥ</emphasis>
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<bibRefCitation id="EF9E16F3FFF48D7B289D7BD1FE4EF3AB" author="Sciaky, R. & Wrase, D. W." box="[151,439,898,922]" pageId="12" pageNumber="55" pagination="1087 - 1139" refId="ref15565" refString="Sciaky, R. & Wrase, D. W. (1997) Twenty-nine new taxa of Pterostichinae from Shaanxi (Coleoptera, Carabidae). Linzer Biologische Beitrage, 29, 1087 - 1139." type="journal article" year="1997">Sciaky & Wrase 1997: 1101</bibRefCitation>
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. (
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<typeStatus id="54B4D5A0FFF48D7B29CD7BD1FDD8F3AB" box="[455,545,898,922]" pageId="12" pageNumber="55" type="holotype">holotype</typeStatus>
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in
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; type locality:
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<collectingRegion id="49CBA5E0FFF48D7B2B017BD1FC98F3AB" box="[779,865,898,922]" country="China" name="Shaanxi" pageId="12" pageNumber="55">Shaanxi</collectingRegion>
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:
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,
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,
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)
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.
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<subSubSection id="C3153889FFF48D7B289D7B9BFECBF4A7" pageId="12" pageNumber="55" type="materials_examined">
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<paragraph id="8BB06B02FFF48D7B289D7B9BFECBF4A7" blockId="12.[151,1437,968,1859]" pageId="12" pageNumber="55">
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<emphasis id="B97BB710FFF48D7B289D7B9BFE7BF3D3" bold="true" box="[151,386,968,994]" pageId="12" pageNumber="55">Material examined.</emphasis>
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<materialsCitation id="3B67615FFFF48D7B29807B9BFF23F47F" collectionCode="CRS" country="China" elevation="1650" location="Qin Ling Shan" pageId="12" pageNumber="55" specimenCount="2" stateProvince="Shaanxi" typeStatus="holotype">
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<typeStatus id="54B4D5A0FFF48D7B29807B9BFE0BF3D3" box="[394,498,968,994]" pageId="12" pageNumber="55" type="holotype">Holotype</typeStatus>
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of
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<taxonomicName id="4C0F1081FFF48D7B2A107B9AFC8BF3D2" authority="Sciaky & Wrase" authorityName="Sciaky & Wrase" box="[538,882,968,995]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="55" phylum="Arthropoda" rank="species" species="parvicollis">
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<emphasis id="B97BB710FFF48D7B2A107B9AFD56F3D3" box="[538,687,968,994]" italics="true" pageId="12" pageNumber="55">P. parvicollis</emphasis>
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Sciaky & Wrase
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</taxonomicName>
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, female (
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<collectionCode id="ED1EF3C7FFF48D7B2BD17B9BFBEDF3D3" box="[987,1044,968,994]" pageId="12" pageNumber="55">CRS</collectionCode>
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,
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<figureCitation id="13347787FFF48D7B2C2A7B9BFB8CF3D3" box="[1056,1141,968,994]" captionStart="FIGURES 31–34" captionStartId="18.[152,270,1908,1932]" captionTargetBox="[184,1403,188,1878]" captionTargetId="figure-36@18.[177,1409,181,1885]" captionTargetPageId="18" captionText="FIGURES 31–34. habitus and male genitalia of P. parvicollis Sciaky & Wrase. (31) habitus of holotype. (32–34) a male specimen from Pingheliang, Shaanxi. (32) habitus. (33) male genitalia, left lateral view, dorsal view of median lobe of aedeagus, and right paramere. (34) endophallus. Abbreviations: gp gonopore, gpp gonopore piece, la left apical lobe, ra right apical lobe. Scale bars: 2 mm (31–32), 0.5 mm (33–34)." figureDoi="http://doi.org/10.5281/zenodo.12742744" httpUri="https://zenodo.org/record/12742744/files/figure.png" pageId="12" pageNumber="55">Fig. 31</figureCitation>
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): “
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<collectingCountry id="F3182B92FFF48D7B2C9E7B9BFB17F3D3" box="[1172,1262,968,994]" name="China" pageId="12" pageNumber="55">CHINA</collectingCountry>
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(
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<collectingRegion id="49CBA5E0FFF48D7B2CF47B9BFAA7F3D3" box="[1278,1374,968,994]" country="China" name="Shaanxi" pageId="12" pageNumber="55">Shaanxi</collectingRegion>
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),
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<location id="8ED03DD9FFF48D7B2D787B9BFEF4F437" LSID="urn:lsid:plazi:treatment:03A6DA14FFF48D7B289D7AA9FB5FF773:8ED03DD9FFF48D7B2D787B9BFEF4F437" country="China" name="Qin Ling Shan" pageId="12" pageNumber="55" stateProvince="Shaanxi">Qin Ling Shan</location>
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/ 107.56E, 33.45N, autoroute km 93 S Zhouzhi,
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<locationDeviation id="9A027220FFF48D7B2B4E7BBFFBFEF437" box="[836,1031,1004,1030]" location="Xian" pageId="12" pageNumber="55">
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SW Xian
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</locationDeviation>
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, mount. forest,
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<quantity id="4CF7C6E7FFF48D7B2CB67BBFFAF0F436" box="[1212,1289,1004,1031]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="12" pageNumber="55" unit="m" value="1650.0">
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<elevation id="00228C31FFF48D7B2CB67BBFFAF0F436" box="[1212,1289,1004,1031]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="12" pageNumber="55" unit="m" value="1650.0">1650m</elevation>
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</quantity>
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/ 1.–2.IX.95 Schülke”; “
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<typeStatus id="54B4D5A0FFF48D7B29167C43FE3BF41B" box="[284,450,1040,1066]" pageId="12" pageNumber="55" type="holotype">HOLOTYPUS</typeStatus>
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<taxonomicName id="4C0F1081FFF48D7B29C67C43FC34F41B" authorityName="Yin & Chen & Shi" authorityYear="2024" box="[460,973,1040,1066]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="55" phylum="Arthropoda" rank="species" species="parvicollis" status="sp. nov." subGenus="Morphohaptoderus">
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<emphasis id="B97BB710FFF48D7B29C67C43FDAEF41B" box="[460,599,1040,1066]" italics="true" pageId="12" pageNumber="55">Pterostichus</emphasis>
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(
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<emphasis id="B97BB710FFF48D7B2A607C43FCBBF41B" box="[618,834,1040,1066]" italics="true" pageId="12" pageNumber="55">Morphohaptoderus</emphasis>
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)
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<emphasis id="B97BB710FFF48D7B2B5F7C43FC34F41B" box="[853,973,1040,1066]" italics="true" pageId="12" pageNumber="55">parvicollis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2480A6BFFF48D7B2BDD7C42FBEEF41A" box="[983,1047,1041,1067]" pageId="12" pageNumber="55" rank="species">sp. n.</taxonomicNameLabel>
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, Sciaky & Wrase des. 1997” [red label]
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</materialsCitation>
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;
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<materialsCitation id="3B67615FFFF48D7B28E37C66FED7F4A7" collectingDate="2017-07-07" collectionCode="CBFU" collectorName="Gao T." country="China" elevation="2016" latitude="33.4718" location="Non-type" longLatPrecision="6" longitude="108.4961" pageId="12" pageNumber="55" specimenCount="1" specimenCount-male="1" stateProvince="Shaanxi">
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<location id="8ED03DD9FFF48D7B28E37C66FEAAF47E" LSID="urn:lsid:plazi:treatment:03A6DA14FFF48D7B289D7AA9FB5FF773:8ED03DD9FFF48D7B28E37C66FEAAF47E" box="[233,339,1077,1103]" country="China" latitude="33.4718" longLatPrecision="6" longitude="108.4961" name="Non-type" pageId="12" pageNumber="55" stateProvince="Shaanxi">Non-type</location>
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material:
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<specimenCount id="9D09A08BFFF48D7B29C07C67FDE1F47F" box="[458,536,1076,1102]" count="1" pageId="12" pageNumber="55" type="male">1 male</specimenCount>
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(
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<collectionCode id="ED1EF3C7FFF48D7B2A207C67FD83F47F" box="[554,634,1076,1102]" pageId="12" pageNumber="55">CBFU</collectionCode>
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): “
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<collectingCountry id="F3182B92FFF48D7B2A907C67FD01F47F" box="[666,760,1076,1102]" name="China" pageId="12" pageNumber="55">CHINA</collectingCountry>
|
||
,
|
||
<collectingRegion id="49CBA5E0FFF48D7B2B0C7C67FC5FF47E" box="[774,934,1076,1103]" country="China" name="Shaanxi" pageId="12" pageNumber="55">Shaanxi prov.</collectingRegion>
|
||
, Ningshan county, Pingheliang pass; mixed forest;
|
||
<quantity id="4CF7C6E7FFF48D7B28E27C0BFEC0F443" box="[232,313,1112,1138]" metricMagnitude="3" metricUnit="m" metricValue="2.016" pageId="12" pageNumber="55" unit="m" value="2016.0">
|
||
<elevation id="00228C31FFF48D7B28E27C0BFEC0F443" box="[232,313,1112,1138]" metricMagnitude="3" metricUnit="m" metricValue="2.016" pageId="12" pageNumber="55" unit="m" value="2016.0">2016m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="EE3B0DC5FFF48D7B294C7C0BFE41F443" box="[326,440,1112,1138]" degrees="33.4718" direction="north" orientation="latitude" pageId="12" pageNumber="55" precision="5" value="33.4718">N33.4718</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3B0DC5FFF48D7B29CC7C0BFDBCF443" box="[454,581,1112,1138]" degrees="108.4961" direction="east" orientation="longitude" pageId="12" pageNumber="55" precision="5" value="108.4961">E108.4961</geoCoordinate>
|
||
”, “
|
||
<date id="FFB14DC2FFF48D7B2A6D7C0BFD1FF442" box="[615,742,1112,1139]" pageId="12" pageNumber="55" value="2017-07-07">
|
||
<collectingDate id="EFF5B42AFFF48D7B2A6D7C0BFD1FF442" box="[615,742,1112,1139]" pageId="12" pageNumber="55" value="2017-07-07">2017.VII.7</collectingDate>
|
||
</date>
|
||
, under stone, Shi H.L., Lu Z.B., Zhu P.Z.,
|
||
<collectorName id="26FA0ED4FFF48D7B2CD27C0BFADFF442" box="[1240,1318,1112,1139]" pageId="12" pageNumber="55">Gao T.</collectorName>
|
||
leg., Exp. BJFU 2017.”
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3153889FFF48D7B28CD7CF3FD33F72F" pageId="12" pageNumber="55" type="description">
|
||
<paragraph id="8BB06B02FFF48D7B28CD7CF3FD33F72F" blockId="12.[151,1437,968,1859]" pageId="12" pageNumber="55">
|
||
<emphasis id="B97BB710FFF48D7B28CD7CF3FEA0F48B" bold="true" box="[199,345,1184,1210]" pageId="12" pageNumber="55">Description.</emphasis>
|
||
BL = 8.0–
|
||
<quantity id="4CF7C6E7FFF48D7B29DB7CF3FDD0F48A" box="[465,553,1184,1211]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="12" pageNumber="55" unit="mm" value="8.4">8.4 mm</quantity>
|
||
, BW =
|
||
<quantity id="4CF7C6E7FFF48D7B2A757CF3FCF0F48A" box="[639,777,1184,1211]" metricMagnitude="-3" metricUnit="m" metricValue="2.8499999999999996" metricValueMax="2.9" metricValueMin="2.8" pageId="12" pageNumber="55" unit="mm" value="2.8499999999999996" valueMax="2.9" valueMin="2.8">2.8–2.9 mm</quantity>
|
||
. Head slightly thickened, frons without puncture; antennae slightly exceeding elytra base; eyes large, tempora slightly swollen behind eyes, shorter than eyes, eyL/tpL = 1.75– 1.80, maxim width of head at outer margin of eyes, HW/PW = 0.61–0.65; submentum with one seta on each side. Pronotum round-rectangular, PW/PL = 1.07–1.10, PBW/PW = 0.80–0.82, widest near anterior third; anterior margin slightly narrower than posterior margin, PAW/PBW =0.83–0.87; lateral margins slightly arched near middle, almost straight before posterior angles; posterior angles rectangular, with a faint denticle. Basal foveae finely punctate, inner groove shallowly defined, about one-third length of pronotum, outer groove indistinguishable. Elytra oblong and parallel-sided, BW/EL = 0.59–0.61, with transverse microsculpture; parascutellar stria joined to stria 1, angular base of stria 1 very short or absent; parascutellar pore present; elytra humeral tooth small, apex sharp, slightly projected out. Interval 3 with one setigerous pore adjacent to stria 2, near posterior third. Male sternite VII without secondary sexual modification. In males, nearly full length of the inner margin of meso- and metatibiae crenulate; metatarsus slightly thickened in males (mt1 length/width = 2.8), slender as other
|
||
<taxonomicName id="4C0F1081FFF48D7B2C437E7FFAD8F677" box="[1097,1313,1580,1606]" pageId="12" pageNumber="55" rank="subGenus" subGenus="Morphohaptoderus">
|
||
<emphasis id="B97BB710FFF48D7B2C437E7FFAD8F677" box="[1097,1313,1580,1606]" italics="true" pageId="12" pageNumber="55">Morphohaptoderus</emphasis>
|
||
</taxonomicName>
|
||
species in females (mt1 length/width = 3.5); metatarsomeres 1–3 with distinct outer-lateral ridge; fifth tarsomere setose ventrally. Median lobe of aedeagus small (length =
|
||
<quantity id="4CF7C6E7FFF48D7B2AD77E27FCBFF6BF" box="[733,838,1652,1679]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="12" pageNumber="55" unit="mm" value="1.15">1.15 mm</quantity>
|
||
), apex abruptly curved at apical third in lateral view; apical lamella short and wide, apex rounded, slightly twisted to right; right paramere very short, apical portion round (
|
||
<figureCitation id="13347787FFF48D7B28957EEFFF0BF6E7" box="[159,242,1724,1750]" captionStart="FIGURES 31–34" captionStartId="18.[152,270,1908,1932]" captionTargetBox="[184,1403,188,1878]" captionTargetId="figure-36@18.[177,1409,181,1885]" captionTargetPageId="18" captionText="FIGURES 31–34. habitus and male genitalia of P. parvicollis Sciaky & Wrase. (31) habitus of holotype. (32–34) a male specimen from Pingheliang, Shaanxi. (32) habitus. (33) male genitalia, left lateral view, dorsal view of median lobe of aedeagus, and right paramere. (34) endophallus. Abbreviations: gp gonopore, gpp gonopore piece, la left apical lobe, ra right apical lobe. Scale bars: 2 mm (31–32), 0.5 mm (33–34)." figureDoi="http://doi.org/10.5281/zenodo.12742744" httpUri="https://zenodo.org/record/12742744/files/figure.png" pageId="12" pageNumber="55">Fig. 33</figureCitation>
|
||
). Endophallus extended to ventral side of aedeagus, gonopore opened to base of aedeagus, right apical lobe (ra) cordate, left apical lobe (la) spherical, larger than ra, both lobes not sclerotized but with fine scales; a strongly sclerotized piece present near gonopore. (
|
||
<figureCitation id="13347787FFF48D7B2A637F57FD44F72F" box="[617,701,1796,1822]" captionStart="FIGURES 31–34" captionStartId="18.[152,270,1908,1932]" captionTargetBox="[184,1403,188,1878]" captionTargetId="figure-36@18.[177,1409,181,1885]" captionTargetPageId="18" captionText="FIGURES 31–34. habitus and male genitalia of P. parvicollis Sciaky & Wrase. (31) habitus of holotype. (32–34) a male specimen from Pingheliang, Shaanxi. (32) habitus. (33) male genitalia, left lateral view, dorsal view of median lobe of aedeagus, and right paramere. (34) endophallus. Abbreviations: gp gonopore, gpp gonopore piece, la left apical lobe, ra right apical lobe. Scale bars: 2 mm (31–32), 0.5 mm (33–34)." figureDoi="http://doi.org/10.5281/zenodo.12742744" httpUri="https://zenodo.org/record/12742744/files/figure.png" pageId="12" pageNumber="55">Fig. 34</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3153889FFF48D7B28CD7F7BFB5FF773" box="[199,1190,1832,1859]" pageId="12" pageNumber="55" type="distribution">
|
||
<paragraph id="8BB06B02FFF48D7B28CD7F7BFB5FF773" blockId="12.[151,1437,968,1859]" box="[199,1190,1832,1859]" pageId="12" pageNumber="55">
|
||
<emphasis id="B97BB710FFF48D7B28CD7F7BFE98F773" bold="true" box="[199,353,1832,1858]" pageId="12" pageNumber="55">Distribution.</emphasis>
|
||
Only known from two localities on Qinling Mountains,
|
||
<collectingRegion id="49CBA5E0FFF48D7B2BD07F7BFB5BF773" box="[986,1186,1832,1858]" country="China" name="Shaanxi" pageId="12" pageNumber="55">Shaanxi Province</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |