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<document id="442531AE1B6207B5F9C1A2567DCC7666" ID-DOI="10.11646/zootaxa.5481.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="12742724" ID-ZooBank="FEC2B59D-D79F-4F9C-B082-D5BAABE1A453" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1721033477194" checkinUser="plazi" docAuthor="Yin, Wen-Qi, Chen, Jia-Heng &amp; Shi, Hong-Liang" docDate="2024" docId="03A6DA14FFF08D7F289D78C4FC6AF3AB" docLanguage="en" docName="zootaxa.5481.1.3.pdf" docOrigin="Zootaxa 5481 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5481.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pterostichus (Tschitscherinea) filum" docType="treatment" docVersion="3" lastPageNumber="51" masterDocId="FF9FA26CFFF88D77280A7853FFF9F031" masterDocTitle="Revision of the Chinese endemic subgenus Tschitscherinea Berg, with descriptions of three new species (Coleoptera, Carabidae)" masterLastPageNumber="65" masterPageNumber="43" pageNumber="51" updateTime="1721135514183" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title id="0371AE45B81A7B344E10DAF4DA697D80">Revision of the Chinese endemic subgenus Tschitscherinea Berg, with descriptions of three new species (Coleoptera, Carabidae)</mods:title>
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<mods:namePart id="A1CC3E791B21E7441970EB0379D4F2CE">Yin, Wen-Qi</mods:namePart>
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<mods:namePart id="BE490B48739178A4051C0E1DF135C736">Chen, Jia-Heng</mods:namePart>
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<mods:namePart id="64A2540A824E8357E91696787AA73A1C">Shi, Hong-Liang</mods:namePart>
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<mods:affiliation id="2F11AB0C10EFE2B82E25FA48408042BB">College of Forestry, Beijing Forestry University, Beijing, 100083, P. R. China</mods:affiliation>
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<subSubSection id="C3153889FFF08D7F289D78C4FC9FF080" box="[151,870,150,178]" pageId="8" pageNumber="51" type="nomenclature">
<paragraph id="8BB06B02FFF08D7F289D78C4FC9FF080" blockId="8.[151,870,150,250]" box="[151,870,150,178]" pageId="8" pageNumber="51">
<heading id="D0F8DC6EFFF08D7F289D78C4FC9FF080" bold="true" box="[151,870,150,178]" fontSize="11" level="1" pageId="8" pageNumber="51" reason="1">
<taxonomicName id="4C0F1081FFF08D7F289D78C4FC9FF080" ID-CoL="78LTV" authority="(Tschitscherine, 1897)" baseAuthorityName="Tschitscherine" baseAuthorityYear="1897" box="[151,870,150,178]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="51" phylum="Arthropoda" rank="species" species="filum" subGenus="Tschitscherinea">
<emphasis id="B97BB710FFF08D7F289D78C4FC9FF080" bold="true" box="[151,870,150,178]" pageId="8" pageNumber="51">
<emphasis id="B97BB710FFF08D7F289D78C4FED4F083" bold="true" box="[151,301,151,178]" italics="true" pageId="8" pageNumber="51">Pterostichus</emphasis>
(
<emphasis id="B97BB710FFF08D7F293478C4FE05F083" bold="true" box="[318,508,151,178]" italics="true" pageId="8" pageNumber="51">Tschitscherinea</emphasis>
)
<emphasis id="B97BB710FFF08D7F2A0778C4FDB7F083" bold="true" box="[525,590,151,178]" italics="true" pageId="8" pageNumber="51">filum</emphasis>
(
<bibRefCitation id="EF9E16F3FFF08D7F2A5778C5FCA7F080" author="Tschitscherine, T." box="[605,862,150,177]" pageId="8" pageNumber="51" pagination="45 - 75" refId="ref15733" refString="Tschitscherine, T. (1897) Carabiques nouveaux ou peu connus. L'Abeille, 29, 45 - 75." type="journal article" year="1897">Tschitschérine, 1897</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3153889FFF08D7F289D78E8FF0CF0E4" box="[151,245,187,213]" pageId="8" pageNumber="51" type="description">
<paragraph id="8BB06B02FFF08D7F289D78E8FF0CF0E4" blockId="8.[151,870,150,250]" box="[151,245,187,213]" pageId="8" pageNumber="51">
<figureCitation id="13347787FFF08D7F289D78E8FF0CF0E4" box="[151,245,187,213]" captionStart="FIGURES 16" captionStartId="13.[152,270,1795,1819]" captionTargetBox="[158,1429,187,1766]" captionTargetId="figure-18@13.[151,1436,181,1771]" captionTargetPageId="13" captionText="FIGURES 16. Habitus of Pterostichus (Tschitscherinea) spp. (1) P. spissipedes sp. nov., holotype. (2) P. spissipedes sp. nov., a male paratype from Jianchuan. (3) P. spissipedes sp. nov., a male paratype from Deqin. (4) P. spectabilis sp. nov., holotype. (5) P. jiuzhaiicus sp. nov., holotype. (6) P. filum (Tschitschérine), a female from Wolong. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742726" httpUri="https://zenodo.org/record/12742726/files/figure.png" pageId="8" pageNumber="51">Figure 6</figureCitation>
</paragraph>
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<subSubSection id="C3153889FFF08D7F289D788CFDB4F0C8" box="[151,589,223,250]" pageId="8" pageNumber="51" type="vernacular_names">
<paragraph id="8BB06B02FFF08D7F289D788CFDB4F0C8" blockId="8.[151,870,150,250]" box="[151,589,223,250]" pageId="8" pageNumber="51">
Chinese vernacular name:
<emphasis id="B97BB710FFF08D7F29CB78B3FDB4F0C8" bold="true" box="[449,589,224,249]" pageId="8" pageNumber="51">ũdzā±Ƥ</emphasis>
</paragraph>
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<subSubSection id="C3153889FFF08D7F289D7977FDEDF16B" pageId="8" pageNumber="51" type="reference_group">
<paragraph id="8BB06B02FFF08D7F289D7977FDEDF16B" blockId="8.[151,1437,292,346]" pageId="8" pageNumber="51">
<bibRefCitation id="EF9E16F3FFF08D7F289D7977FE7FF10D" author="Tschitscherine, T." box="[151,390,292,316]" pageId="8" pageNumber="51" pagination="45 - 75" refId="ref15733" refString="Tschitscherine, T. (1897) Carabiques nouveaux ou peu connus. L'Abeille, 29, 45 - 75." type="journal article" year="1897">Tschitschérine 1897: 70</bibRefCitation>
(Original:
<taxonomicName id="4C0F1081FFF08D7F29FF7977FDBFF10A" box="[501,582,292,315]" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B97BB710FFF08D7F29FF7977FDBFF10A" box="[501,582,292,315]" italics="true" pageId="8" pageNumber="51">Feronia</emphasis>
</taxonomicName>
: subgen.
<taxonomicName id="4C0F1081FFF08D7F2AAF7977FD01F10A" authorityName="Tschitscherine" authorityYear="1897" box="[677,760,292,315]" class="Gastropoda" family="Lophospiridae" genus="Eunema" kingdom="Animalia" pageId="8" pageNumber="51" phylum="Mollusca" rank="genus">
<emphasis id="B97BB710FFF08D7F2AAF7977FD01F10A" box="[677,760,292,315]" italics="true" pageId="8" pageNumber="51">Eunema</emphasis>
</taxonomicName>
;
<typeStatus id="54B4D5A0FFF08D7F2B0E7977FCA4F10D" box="[772,861,292,316]" pageId="8" pageNumber="51" type="holotype">holotype</typeStatus>
should be in MNHN, but probably missing; type locality: “Mou-Pin”);
<bibRefCitation id="EF9E16F3FFF08D7F29447911FDE9F16B" author="Jedlicka, A." box="[334,528,322,346]" pageId="8" pageNumber="51" pagination="177 - 346" refId="ref15409" refString="Jedlicka, A. (1962) Monographie des Tribus Pterostichini aus Osatasien (Pterostichini, Trigonotomi, Myadi) (Coleoptera- Carabidae). Abhandlungen und Berichte aus dem Staatlichen Museum fur Tierkunde in Dresden, 26, 177 - 346." type="journal article" year="1962">Jedlička 1962: 228</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3153889FFF08D7F289D79DBFBABF1F7" pageId="8" pageNumber="51" type="materials_examined">
<paragraph id="8BB06B02FFF08D7F289D79DBFBABF1F7" blockId="8.[151,1437,392,923]" pageId="8" pageNumber="51">
<emphasis id="B97BB710FFF08D7F289D79DBFE7DF193" bold="true" box="[151,388,392,418]" pageId="8" pageNumber="51">Material examined.</emphasis>
<materialsCitation id="3B67615FFFF08D7F298479DAFBABF1F7" collectingDate="2004-07-22" collectingDateMax="2004-07-25" collectingDateMin="2004-07-22" collectionCode="IZAS" collectorName="Xiaodong Yu" county="Wolong" elevation="2395" location="Institute of Zoology." municipality="Wulidun" pageId="8" pageNumber="51" specimenCount="1" specimenCount-female="1" stateProvince="Sichuan">
Non-type material:
<specimenCount id="9D09A08BFFF08D7F2A7B79DAFD2CF193" box="[625,725,392,419]" count="1" pageId="8" pageNumber="51" type="female">1 female</specimenCount>
(
<collectionCode id="ED1EF3C7FFF08D7F2AE279DAFCD4F192" box="[744,813,393,419]" country="0" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="8" pageNumber="51">IZAS</collectionCode>
): “
<collectingRegion id="49CBA5E0FFF08D7F2B4779DBFC55F193" box="[845,940,392,418]" country="China" name="Sichuan" pageId="8" pageNumber="51">Sichuan</collectingRegion>
,
<collectingCounty id="62D1138EFFF08D7F2BB079DBFBECF193" box="[954,1045,392,418]" pageId="8" pageNumber="51">Wolong</collectingCounty>
,
<collectingMunicipality id="6BD4F178FFF08D7F2C2979DBFB7EF193" box="[1059,1159,392,418]" pageId="8" pageNumber="51">Wulidun</collectingMunicipality>
,
<quantity id="4CF7C6E7FFF08D7F2C9F79DBFB1FF192" box="[1173,1254,392,419]" metricMagnitude="3" metricUnit="m" metricValue="2.395" pageId="8" pageNumber="51" unit="m" value="2395.0">
<elevation id="00228C31FFF08D7F2C9F79DBFB1FF192" box="[1173,1254,392,419]" metricMagnitude="3" metricUnit="m" metricValue="2.395" pageId="8" pageNumber="51" unit="m" value="2395.0">2395m</elevation>
</quantity>
, Artificial larch forest,
<collectingMethod id="524E1315FFF08D7F28EF79FFFEA7F1F6" box="[229,350,428,455]" pageId="8" pageNumber="51">pitfall trap</collectingMethod>
.
<date id="FFB14DC2FFF08D7F296279FEFDE7F1F6" box="[360,542,428,455]" pageId="8" pageNumber="51" value="2004-07-22" valueMax="2004-07-25" valueMin="2004-07-22">
<collectingDate id="EFF5B42AFFF08D7F296279FEFDE7F1F6" box="[360,542,428,455]" pageId="8" pageNumber="51" value="2004-07-22" valueMax="2004-07-25" valueMin="2004-07-22">2004.VII.2225</collectingDate>
</date>
,
<collectorName id="26FA0ED4FFF08D7F2A2379FFFD44F1F6" box="[553,701,428,455]" pageId="8" pageNumber="51">Xiaodong Yu</collectorName>
leg. RG324,
<location id="8ED03DD9FFF08D7F2B6B79FFFBB0F1F7" LSID="urn:lsid:plazi:treatment:03A6DA14FFF08D7F289D78C4FC6AF3AB:8ED03DD9FFF08D7F2B6B79FFFBB0F1F7" box="[865,1097,428,454]" county="Wolong" municipality="Wulidun" name="Institute of Zoology." pageId="8" pageNumber="51" stateProvince="Sichuan">Institute of Zoology.</location>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3153889FFF08D7F28CD7983FE1CF363" pageId="8" pageNumber="51" type="diagnosis">
<paragraph id="8BB06B02FFF08D7F28CD7983FEB6F203" blockId="8.[151,1437,392,923]" pageId="8" pageNumber="51">
<emphasis id="B97BB710FFF08D7F28CD7983FEBBF1DB" bold="true" box="[199,322,464,490]" pageId="8" pageNumber="51">Diagnosis.</emphasis>
This species is different from all the other members of this subgenus by the elytral interval 3 with only one setigerous pore. It is most similar to
<taxonomicName id="4C0F1081FFF08D7F2A9179A6FCD6F23F" authorityName="Yin &amp; Chen &amp; Shi" authorityYear="2024" box="[667,815,500,526]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="51" phylum="Arthropoda" rank="species" species="jiuzhaiicus" status="sp. nov.">
<emphasis id="B97BB710FFF08D7F2A9179A6FCD6F23F" box="[667,815,500,526]" italics="true" pageId="8" pageNumber="51">P. jiuzhaiicus</emphasis>
</taxonomicName>
<emphasis id="B97BB710FFF08D7F2B3D79A6FC68F23E" bold="true" box="[823,913,501,527]" pageId="8" pageNumber="51">
<taxonomicNameLabel id="A2480A6BFFF08D7F2B3D79A6FC68F23E" box="[823,913,501,527]" pageId="8" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Their comparison was already provided under the new species.
</paragraph>
<paragraph id="8BB06B02FFF08D7F28CD7A6FFE1CF363" blockId="8.[151,1437,392,923]" pageId="8" pageNumber="51">
<emphasis id="B97BB710FFF08D7F28CD7A6FFDEAF267" bold="true" box="[199,531,572,598]" pageId="8" pageNumber="51">Supplementary description.</emphasis>
BL =
<quantity id="4CF7C6E7FFF08D7F2A517A6FFD4AF266" box="[603,691,572,599]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="8" pageNumber="51" unit="mm" value="9.5">9.5 mm</quantity>
, BW =
<quantity id="4CF7C6E7FFF08D7F2B007A6EFC9AF266" box="[778,867,573,599]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="8" pageNumber="51" unit="mm" value="3.2">3.2 mm</quantity>
; eyes very small, tempora strongly swollen behind eyes, length greater than eyes, eyL/tpL = 0.52, maxim width of head at outer margin of tempora, HW/PW = 0.75. Pronotum parallel-sided, PW/PL = 1.03, PBW/PW = 0.83; anterior margin slightly wider than posterior margin, PAW/PBW = 1.04; lateral margins straight before posterior angles, posterior angles rectangular, not denticulate. Basal foveae densely punctate, without outer groove, inner groove faintly depressed. Elytra with transverse microsculpture; parascutellar pore absent; humerus with small stout tooth, faintly projected out; interval 3 with one setigerous pore near anterior third, adjacent to stria 2. Metatarsus slender in females, segment 1 with shallow outer-lateral ridge; males unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3153889FFF08D7F28CD7B0FFC6AF3AB" pageId="8" pageNumber="51" type="distribution">
<paragraph id="8BB06B02FFF08D7F28CD7B0FFC6AF3AB" blockId="8.[151,1437,392,923]" pageId="8" pageNumber="51">
<emphasis id="B97BB710FFF08D7F28CD7B0FFE98F347" bold="true" box="[199,353,860,886]" pageId="8" pageNumber="51">Distribution.</emphasis>
This species is known only from two close localities in Western
<collectingRegion id="49CBA5E0FFF08D7F2C2C7B0FFB7AF347" box="[1062,1155,860,886]" country="China" name="Sichuan" pageId="8" pageNumber="51">Sichuan</collectingRegion>
, the
<typeStatus id="54B4D5A0FFF08D7F2CBE7B0EFB1DF346" box="[1204,1252,861,887]" pageId="8" pageNumber="51">type</typeStatus>
locality Moupin in Baoxing county, and Wolong in Wenchuan county. (
<figureCitation id="13347787FFF08D7F2AF07BD3FCB5F3AA" box="[762,844,896,923]" captionStart="FIGURES 5457. 54" captionStartId="21.[152,270,1525,1549]" captionTargetBox="[151,1435,181,1494]" captionTargetId="figure-18@21.[151,1435,181,1501]" captionTargetPageId="21" captionText="FIGURES 5457. 54. Confirmed distribution of the Pterostichus (Tschitscherinea) spp. in China: Red P. spissipedes sp. nov. Blue P. spectabilis sp. nov. Black P. jiuzhaiicus sp. nov. Pink P. filum (Tschitschérine). Purple P. mulensis Sciaky. Green P. fakarci Sciaky. Yellow P. krali Sciaky. Cyan P. sterbai Jedlička. 5557. Endophallus of Pterostichus spp., right lateral view and left lateral view: (55) Pterostichus (Cryobius) jaechi Krischenhofer, Changbaishan, Jilin. (56) Pterostichus (Morphohaptoderus) janatai Sciaky &amp; Wrase, Pingheliang, Shaanxi. (57) Pterostichus (Morphohaptoderus) dundai Sciaky, Houzhenzi, Shaanxi. Scale bars = 0.5 mm, a (55), b (5657)." figureDoi="http://doi.org/10.5281/zenodo.12742754" httpUri="https://zenodo.org/record/12742754/files/figure.png" pageId="8" pageNumber="51">Fig. 54</figureCitation>
, pink)
</paragraph>
</subSubSection>
</treatment>
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