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<document id="BD90B6670439D4DFBEFDF4094ABBD5A1" ID-DOI="10.26107/RBZ-2020-0053" ID-ISSN="2345-7600" ID-Zenodo-Dep="4577302" ID-ZooBank="65CDBE40-FEA7-4F06-A30F-6F970E63AE6C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614785065864" checkinUser="carolina" docAuthor="Chen, Zhuo, Liu, Yingqi &amp; Cai, Wanzhi" docDate="2020" docId="039C879BCF2CC1388F99A970B5896E1A" docLanguage="en" docName="RafflesBZool.68.369-378.pdf" docOrigin="Raffles Bulletin of Zoology 68" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Enoplocephala maculata Chen &amp; Liu &amp; Cai 2020, new species" docType="treatment" docUuid="CFB12690-CEBC-4E65-874F-05C0C8858E7D" docUuidSource="ZooBank" docVersion="4" lastPageNumber="376" masterDocId="FFA5FFE3CF28C13F8C37AE55B458662F" masterDocTitle="Notes on the genus Enoplocephala Miller (Hemiptera: Reduviidae: Stenopodainae), with the description of a new species from Borneo" masterLastPageNumber="378" masterPageNumber="369" pageNumber="373" updateTime="1698878663914" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="5C916BF328A17DEEE7A665178C2D7984">Notes on the genus Enoplocephala Miller (Hemiptera: Reduviidae: Stenopodainae), with the description of a new species from Borneo</mods:title>
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<mods:namePart id="A87709455F4A936981CADEC457725AD1">Chen, Zhuo</mods:namePart>
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<mods:namePart id="5F05B8F85F84DF177A71A5A3E974C862">Liu, Yingqi</mods:namePart>
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<mods:namePart id="D3B4B652DAC9C3719E83E51807A7EACB">Cai, Wanzhi</mods:namePart>
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<treatment id="039C879BCF2CC1388F99A970B5896E1A" LSID="urn:lsid:zoobank.org:act:CFB12690-CEBC-4E65-874F-05C0C8858E7D" httpUri="http://treatment.plazi.org/id/039C879BCF2CC1388F99A970B5896E1A" lastPageId="7" lastPageNumber="376" pageId="4" pageNumber="373">
<subSubSection id="C32F6506CF2CC13B8F99A970B1006112" box="[942,1368,1829,1853]" pageId="4" pageNumber="373" type="nomenclature">
<paragraph id="8B8A368DCF2CC13B8F99A970B1006112" blockId="4.[942,1368,1829,1885]" box="[942,1368,1829,1853]" pageId="4" pageNumber="373">
<heading id="D0C281E1CF2CC13B8F99A970B1006112" bold="true" box="[942,1368,1829,1853]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="373" reason="2">
<emphasis id="B941EA9FCF2CC13B8F99A970B1006112" bold="true" box="[942,1368,1829,1853]" pageId="4" pageNumber="373">
<taxonomicName id="4C354D0ECF2CC13B8F99A970B09B6112" authority="Chen &amp; Liu &amp; Cai, 2020" authorityName="Chen &amp; Liu &amp; Cai" authorityYear="2020" box="[942,1219,1829,1853]" class="Insecta" family="Reduviidae" genus="Enoplocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="species" species="maculata" status="sp. nov.">
<emphasis id="B941EA9FCF2CC13B8F99A970B09B6112" bold="true" box="[942,1219,1829,1853]" italics="true" pageId="4" pageNumber="373">Enoplocephala maculata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27257E4CF2CC13B88E5A970B1006112" box="[1234,1368,1829,1853]" pageId="4" pageNumber="373" rank="species">new species</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C32F6506CF2CC13B8877A910B09D6172" box="[1088,1221,1861,1885]" pageId="4" pageNumber="373" type="description">
<paragraph id="8B8A368DCF2CC13B8877A910B09D6172" blockId="4.[942,1368,1829,1885]" box="[1088,1221,1861,1885]" pageId="4" pageNumber="373">
(
<figureCitation id="130E2A08CF2CC13B887FA910B0E56172" box="[1096,1213,1861,1885]" captionStart-0="Figs" captionStart-1="Figs" captionStart-2="Figs" captionStart-3="Fig" captionStartId-0="4.[136,180,1147,1169]" captionStartId-1="5.[113,157,1658,1680]" captionStartId-2="6.[135,179,1710,1732]" captionStartId-3="8.[135,170,997,1019]" captionTargetBox-0="[111,1500,172,1145]" captionTargetBox-1="[110,1456,188,1625]" captionTargetBox-2="[140,1473,186,1691]" captionTargetBox-3="[135,1473,189,982]" captionTargetId-0="figure-417@4.[111,1500,172,1145]" captionTargetId-1="figure-9@5.[96,1469,172,1658]" captionTargetId-2="figure-8@6.[128,1518,159,1735]" captionTargetId-3="figure-14@8.[135,1473,189,982]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="8" captionText-0="Figs. 911. Enoplocephala maculata, new species, holotype, male, habitus. 9, dorsal view; 10, lateral view; 11, ventral view. Scale bar = 5.00 mm." captionText-1="Figs. 1218. Enoplocephala maculata, new species, holotype, male. 1214, anterior part of body; 15, head; 16, fore wing; 17, 18, posterior part of body. 12, 16, 17, dorsal view; 13, lateral view; 14, 18, ventral view; 15, front view. Scale bars for 1214, 17, 18 = 2.00 mm; 15 = 1.00 mm; 16 = 3.00 mm." captionText-2="Figs. 1928. Enoplocephala maculata, new species, holotype, male. 19, abdominal segment VIII; 2022, pygophore; 2325, paramere; 2628, phallus. 19, 27, ventral view; 20, 26, dorsal view; 21, 28, lateral view; 22, caudal view. Scale bars for 19, 2328 = 0.50 mm; 2022 = 1.00 mm." captionText-3="Fig. 29. Known distribution of Enoplocephala species." figureDoi-0="http://doi.org/10.5281/zenodo.4577310" figureDoi-1="http://doi.org/10.5281/zenodo.4577312" figureDoi-2="http://doi.org/10.5281/zenodo.4577314" figureDoi-3="http://doi.org/10.5281/zenodo.4577316" httpUri-0="https://zenodo.org/record/4577310/files/figure.png" httpUri-1="https://zenodo.org/record/4577312/files/figure.png" httpUri-2="https://zenodo.org/record/4577314/files/figure.png" httpUri-3="https://zenodo.org/record/4577316/files/figure.png" pageId="4" pageNumber="373">Figs. 929</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32F6506CF2CC13A8F74A9D0B5E76157" lastPageId="5" lastPageNumber="374" pageId="4" pageNumber="373" type="diagnosis">
<paragraph id="8B8A368DCF2CC13A8F74A9D0B5E76157" blockId="4.[834,1475,1925,2109]" lastBlockId="5.[113,754,1792,1913]" lastPageId="5" lastPageNumber="374" pageId="4" pageNumber="373">
<emphasis id="B941EA9FCF2CC13B8F74A9D0B7E061B2" bold="true" box="[835,952,1925,1949]" pageId="4" pageNumber="373">Diagnosis.</emphasis>
Body
<quantity id="4CCD9B68CF2CC13B8832A9D0B03F61B2" box="[1029,1127,1925,1949]" metricMagnitude="-2" metricUnit="m" metricValue="1.76" pageId="4" pageNumber="373" unit="mm" value="17.6">17.6 mm</quantity>
in length, brown, mottled with yellowish brown spots and markings; head distinctly shorter than pronotum, about 1.3 times longer than width across eyes; process of antenniferous tubercle relatively short, blunt, nearly horizontal; spines of anterolateral angles of pronotum curved, nearly horizontal; humeral angles protruding posterolaterally; fore femur with two tiny denticles at base of ventral surface; area between Cu and claval suture on fore wing not occupied by blackish patch; blackish patch between M and Cu acuminate.
</paragraph>
</subSubSection>
<caption id="DF4A6605CF2DC13A8C46A82FB5D960E6" ID-DOI="http://doi.org/10.5281/zenodo.4577312" ID-Zenodo-Dep="4577312" httpUri="https://zenodo.org/record/4577312/files/figure.png" pageId="5" pageNumber="374" startId="5.[113,157,1658,1680]" targetBox="[110,1456,188,1625]" targetPageId="5">
<paragraph id="8B8A368DCF2DC13A8C46A82FB5D960E6" blockId="5.[113,1452,1657,1737]" pageId="5" pageNumber="374">
Figs. 1218.
<taxonomicName id="4C354D0ECF2DC13A8CD8A82FB58760A0" authority="Chen &amp; Liu &amp; Cai, 2020" authorityName="Chen &amp; Liu &amp; Cai" authorityYear="2020" box="[239,479,1658,1679]" class="Insecta" family="Reduviidae" genus="Enoplocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="maculata" status="sp. nov.">
<emphasis id="B941EA9FCF2DC13A8CD8A82FB58760A0" box="[239,479,1658,1679]" italics="true" pageId="5" pageNumber="374">Enoplocephala maculata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27257E4CF2DC13A8DDBA82FB63A60BF" box="[492,610,1658,1680]" pageId="5" pageNumber="374" rank="species">new species</taxonomicNameLabel>
, holotype, male. 1214, anterior part of body; 15, head; 16, fore wing; 17, 18, posterior part of body. 12, 16, 17, dorsal view; 13, lateral view; 14, 18, ventral view; 15, front view. Scale bars for 1214, 17, 18 = 2.00 mm; 15 = 1.00 mm; 16 = 3.00 mm.
</paragraph>
</caption>
<subSubSection id="C32F6506CF2DC1388C46A9F5B6F4609A" lastPageId="7" lastPageNumber="376" pageId="5" pageNumber="374" type="description">
<paragraph id="8B8A368DCF2DC1398C46A9F5B7B16E02" blockId="5.[113,754,1952,2105]" lastBlockId="6.[835,1476,1845,2093]" lastPageId="6" lastPageNumber="375" pageId="5" pageNumber="374">
<emphasis id="B941EA9FCF2DC13A8C46A9F5B4A46197" bold="true" box="[113,252,1952,1976]" pageId="5" pageNumber="374">Description.</emphasis>
Macropterous male (
<figureCitation id="130E2A08CF2DC13A8DECA9F5B6166196" box="[475,590,1952,1977]" captionStart="Figs" captionStartId="4.[136,180,1147,1169]" captionTargetBox="[111,1500,172,1145]" captionTargetId="figure-417@4.[111,1500,172,1145]" captionTargetPageId="4" captionText="Figs. 911. Enoplocephala maculata, new species, holotype, male, habitus. 9, dorsal view; 10, lateral view; 11, ventral view. Scale bar = 5.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577310" httpUri="https://zenodo.org/record/4577310/files/figure.png" pageId="5" pageNumber="374">Figs. 911</figureCitation>
).
<emphasis id="B941EA9FCF2DC13A8E68A9F5B6B76197" bold="true" box="[607,751,1952,1976]" pageId="5" pageNumber="374">Colouration.</emphasis>
Body brown. Head with glabrous areas of anteocular part, ventrolateral and ventral surfaces darker; dorsum of head with several blackish markings; antennal segments I and II yellowish brown, segment I mottled with black on basal third and apical third, and with two narrow, incomplete, blackish annuli medially (
<figureCitation id="130E2A08CF2DC13A8FECA975B07B6116" box="[987,1059,1824,1849]" captionStart="Figs" captionStartId="4.[136,180,1147,1169]" captionTargetBox="[111,1500,172,1145]" captionTargetId="figure-417@4.[111,1500,172,1145]" captionTargetPageId="4" captionText="Figs. 911. Enoplocephala maculata, new species, holotype, male, habitus. 9, dorsal view; 10, lateral view; 11, ventral view. Scale bar = 5.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577310" httpUri="https://zenodo.org/record/4577310/files/figure.png" pageId="5" pageNumber="374">Figs. 9</figureCitation>
,
<figureCitation id="130E2A08CF2DC13A881AA974B0116116" box="[1069,1097,1825,1849]" captionStart="Figs" captionStartId="5.[113,157,1658,1680]" captionTargetBox="[110,1456,188,1625]" captionTargetId="figure-9@5.[96,1469,172,1658]" captionTargetPageId="5" captionText="Figs. 1218. Enoplocephala maculata, new species, holotype, male. 1214, anterior part of body; 15, head; 16, fore wing; 17, 18, posterior part of body. 12, 16, 17, dorsal view; 13, lateral view; 14, 18, ventral view; 15, front view. Scale bars for 1214, 17, 18 = 2.00 mm; 15 = 1.00 mm; 16 = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577312" httpUri="https://zenodo.org/record/4577312/files/figure.png" pageId="5" pageNumber="374">13</figureCitation>
), segment II black on base, medial third, and apex (
<figureCitation id="130E2A08CF2DC13A8FD5A915B0686176" box="[994,1072,1856,1881]" captionStart="Figs" captionStartId="4.[136,180,1147,1169]" captionTargetBox="[111,1500,172,1145]" captionTargetId="figure-417@4.[111,1500,172,1145]" captionTargetPageId="4" captionText="Figs. 911. Enoplocephala maculata, new species, holotype, male, habitus. 9, dorsal view; 10, lateral view; 11, ventral view. Scale bar = 5.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577310" httpUri="https://zenodo.org/record/4577310/files/figure.png" pageId="5" pageNumber="374">Figs. 9</figureCitation>
,
<figureCitation id="130E2A08CF2DC13A8808A914B0046176" box="[1087,1116,1857,1881]" captionStart="Figs" captionStartId="5.[113,157,1658,1680]" captionTargetBox="[110,1456,188,1625]" captionTargetId="figure-9@5.[96,1469,172,1658]" captionTargetPageId="5" captionText="Figs. 1218. Enoplocephala maculata, new species, holotype, male. 1214, anterior part of body; 15, head; 16, fore wing; 17, 18, posterior part of body. 12, 16, 17, dorsal view; 13, lateral view; 14, 18, ventral view; 15, front view. Scale bars for 1214, 17, 18 = 2.00 mm; 15 = 1.00 mm; 16 = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577312" httpUri="https://zenodo.org/record/4577312/files/figure.png" pageId="5" pageNumber="374">13</figureCitation>
); segments III and IV greyish brown; labium yellowish brown, with basal spots and apical half (except apex) of segment II black, segment III mottled with black, segment IV darker (
<figureCitation id="130E2A08CF2DC13A88ADA9F5B0B66196" box="[1178,1262,1952,1977]" captionStart="Figs" captionStartId="5.[113,157,1658,1680]" captionTargetBox="[110,1456,188,1625]" captionTargetId="figure-9@5.[96,1469,172,1658]" captionTargetPageId="5" captionText="Figs. 1218. Enoplocephala maculata, new species, holotype, male. 1214, anterior part of body; 15, head; 16, fore wing; 17, 18, posterior part of body. 12, 16, 17, dorsal view; 13, lateral view; 14, 18, ventral view; 15, front view. Scale bars for 1214, 17, 18 = 2.00 mm; 15 = 1.00 mm; 16 = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577312" httpUri="https://zenodo.org/record/4577312/files/figure.png" pageId="5" pageNumber="374">Fig. 13</figureCitation>
). Pronotum with spines of anterolateral angles, medial carinae and spines of humeral angles yellowish brown; propleuron and prosternum darker. Apical process of scutellum yellowish brown. Legs yellowish brown; coxae dark brown; each trochanter with a dark brown spot at base; basal, medial, and subapical parts of all femora each with an irregular, incomplete, blackish annulus (
<figureCitation id="130E2A08CF2EC1398CC3A920B52E61A2" box="[244,374,1909,1933]" captionStart="Figs" captionStartId="4.[136,180,1147,1169]" captionTargetBox="[111,1500,172,1145]" captionTargetId="figure-417@4.[111,1500,172,1145]" captionTargetPageId="4" captionText="Figs. 911. Enoplocephala maculata, new species, holotype, male, habitus. 9, dorsal view; 10, lateral view; 11, ventral view. Scale bar = 5.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577310" httpUri="https://zenodo.org/record/4577310/files/figure.png" pageId="6" pageNumber="375">Figs. 9, 11</figureCitation>
); each tibia with a narrow, basal, blackish annulus and a broad, subbasal, blackish annulus (
<figureCitation id="130E2A08CF2EC1398CA7A9E0B55761E2" box="[144,271,1973,1997]" captionStart="Figs" captionStartId="4.[136,180,1147,1169]" captionTargetBox="[111,1500,172,1145]" captionTargetId="figure-417@4.[111,1500,172,1145]" captionTargetPageId="4" captionText="Figs. 911. Enoplocephala maculata, new species, holotype, male, habitus. 9, dorsal view; 10, lateral view; 11, ventral view. Scale bar = 5.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577310" httpUri="https://zenodo.org/record/4577310/files/figure.png" pageId="6" pageNumber="375">Figs. 9, 11</figureCitation>
); apical two-thirds of fore tibia, apical half of mid tibia, apex of hind tibia darker; tarsomeres I and II brown, tarsomere III and claws darker. Fore wing with blackish patches in corium, clavus and membrane similar with that of
<taxonomicName id="4C354D0ECF2EC1398FF8A960B03A6162" authorityName="Miller" authorityYear="1940" box="[975,1122,1845,1869]" class="Insecta" family="Reduviidae" genus="Enoplocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="perakensis">
<emphasis id="B941EA9FCF2EC1398FF8A960B03A6162" box="[975,1122,1845,1869]" italics="true" pageId="6" pageNumber="375">E. perakensis</emphasis>
</taxonomicName>
, but without patch between Cu and claval suture, and patches in outer cell of membrane much smaller; blackish patch between M and Cu acuminate (
<figureCitation id="130E2A08CF2EC1398F7BA9C0B7C76182" box="[844,927,1941,1965]" captionStart="Figs" captionStartId="4.[136,180,1147,1169]" captionTargetBox="[111,1500,172,1145]" captionTargetId="figure-417@4.[111,1500,172,1145]" captionTargetPageId="4" captionText="Figs. 911. Enoplocephala maculata, new species, holotype, male, habitus. 9, dorsal view; 10, lateral view; 11, ventral view. Scale bar = 5.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577310" httpUri="https://zenodo.org/record/4577310/files/figure.png" pageId="6" pageNumber="375">Figs. 9</figureCitation>
,
<figureCitation id="130E2A08CF2EC1398F99A9C0B7956182" box="[942,973,1941,1965]" captionStart="Figs" captionStartId="5.[113,157,1658,1680]" captionTargetBox="[110,1456,188,1625]" captionTargetId="figure-9@5.[96,1469,172,1658]" captionTargetPageId="5" captionText="Figs. 1218. Enoplocephala maculata, new species, holotype, male. 1214, anterior part of body; 15, head; 16, fore wing; 17, 18, posterior part of body. 12, 16, 17, dorsal view; 13, lateral view; 14, 18, ventral view; 15, front view. Scale bars for 1214, 17, 18 = 2.00 mm; 15 = 1.00 mm; 16 = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577312" httpUri="https://zenodo.org/record/4577312/files/figure.png" pageId="6" pageNumber="375">16</figureCitation>
). Anterolateral and posterolateral angles of each abdominal segment yellowish brown (
<figureCitation id="130E2A08CF2EC139892EA9E0B13161E2" box="[1305,1385,1973,1997]" captionStart="Figs" captionStartId="5.[113,157,1658,1680]" captionTargetBox="[110,1456,188,1625]" captionTargetId="figure-9@5.[96,1469,172,1658]" captionTargetPageId="5" captionText="Figs. 1218. Enoplocephala maculata, new species, holotype, male. 1214, anterior part of body; 15, head; 16, fore wing; 17, 18, posterior part of body. 12, 16, 17, dorsal view; 13, lateral view; 14, 18, ventral view; 15, front view. Scale bars for 1214, 17, 18 = 2.00 mm; 15 = 1.00 mm; 16 = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577312" httpUri="https://zenodo.org/record/4577312/files/figure.png" pageId="6" pageNumber="375">Fig. 17</figureCitation>
); lateral margins of each segment with a reddish spot medially (
<figureCitation id="130E2A08CF2EC13989AFA980B7396E22" captionStart="Figs" captionStartId="5.[113,157,1658,1680]" captionTargetBox="[110,1456,188,1625]" captionTargetId="figure-9@5.[96,1469,172,1658]" captionTargetPageId="5" captionText="Figs. 1218. Enoplocephala maculata, new species, holotype, male. 1214, anterior part of body; 15, head; 16, fore wing; 17, 18, posterior part of body. 12, 16, 17, dorsal view; 13, lateral view; 14, 18, ventral view; 15, front view. Scale bars for 1214, 17, 18 = 2.00 mm; 15 = 1.00 mm; 16 = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577312" httpUri="https://zenodo.org/record/4577312/files/figure.png" pageId="6" pageNumber="375">Fig. 17</figureCitation>
); ventral surface of abdomen mottled with several light spots (
<figureCitation id="130E2A08CF2EC1398FBCA640B7846E02" box="[907,988,2069,2093]" captionStart="Figs" captionStartId="5.[113,157,1658,1680]" captionTargetBox="[110,1456,188,1625]" captionTargetId="figure-9@5.[96,1469,172,1658]" captionTargetPageId="5" captionText="Figs. 1218. Enoplocephala maculata, new species, holotype, male. 1214, anterior part of body; 15, head; 16, fore wing; 17, 18, posterior part of body. 12, 16, 17, dorsal view; 13, lateral view; 14, 18, ventral view; 15, front view. Scale bars for 1214, 17, 18 = 2.00 mm; 15 = 1.00 mm; 16 = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577312" httpUri="https://zenodo.org/record/4577312/files/figure.png" pageId="6" pageNumber="375">Fig. 18</figureCitation>
).
</paragraph>
<caption id="DF4A6605CF2EC1398CB0A8FBB56760D2" ID-DOI="http://doi.org/10.5281/zenodo.4577314" ID-Zenodo-Dep="4577314" httpUri="https://zenodo.org/record/4577314/files/figure.png" pageId="6" pageNumber="375" startId="6.[135,179,1710,1732]" targetBox="[140,1473,186,1691]" targetPageId="6">
<paragraph id="8B8A368DCF2EC1398CB0A8FBB56760D2" blockId="6.[135,1474,1710,1789]" pageId="6" pageNumber="375">
Figs. 1928.
<taxonomicName id="4C354D0ECF2EC1398D3DA8FBB5A560EC" authority="Chen &amp; Liu &amp; Cai, 2020" authorityName="Chen &amp; Liu &amp; Cai" authorityYear="2020" box="[266,509,1710,1731]" class="Insecta" family="Reduviidae" genus="Enoplocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="maculata" status="sp. nov.">
<emphasis id="B941EA9FCF2EC1398D3DA8FBB5A560EC" box="[266,509,1710,1731]" italics="true" pageId="6" pageNumber="375">Enoplocephala maculata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27257E4CF2EC1398E3BA8FBB6DC60EB" box="[524,644,1710,1732]" pageId="6" pageNumber="375" rank="species">new species</taxonomicNameLabel>
, holotype, male. 19, abdominal segment VIII; 2022, pygophore; 2325, paramere; 2628, phallus. 19, 27, ventral view; 20, 26, dorsal view; 21, 28, lateral view; 22, caudal view. Scale bars for 19, 2328 = 0.50 mm; 2022 = 1.00 mm.
</paragraph>
</caption>
<paragraph id="8B8A368DCF2FC1388C46AEE8B6AA64DA" blockId="7.[112,755,189,757]" pageId="7" pageNumber="376">
<emphasis id="B941EA9FCF2FC1388C46AEE8B4BC66FA" bold="true" box="[113,228,189,213]" pageId="7" pageNumber="376">Structure.</emphasis>
Body shape and vestiture as in generic description. Head (
<figureCitation id="130E2A08CF2FC1388C80AE88B56F66DA" box="[183,311,221,245]" captionStart="Figs" captionStartId="5.[113,157,1658,1680]" captionTargetBox="[110,1456,188,1625]" captionTargetId="figure-9@5.[96,1469,172,1658]" captionTargetPageId="5" captionText="Figs. 1218. Enoplocephala maculata, new species, holotype, male. 1214, anterior part of body; 15, head; 16, fore wing; 17, 18, posterior part of body. 12, 16, 17, dorsal view; 13, lateral view; 14, 18, ventral view; 15, front view. Scale bars for 1214, 17, 18 = 2.00 mm; 15 = 1.00 mm; 16 = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577312" httpUri="https://zenodo.org/record/4577312/files/figure.png" pageId="7" pageNumber="376">Figs. 1214</figureCitation>
) 1.29 times as long as width across eyes, distinctly shorter than pronotum; ventral margin of anteocular part strongly concave in lateral view (
<figureCitation id="130E2A08CF2FC1388E39AF48B607671A" box="[526,607,285,309]" captionStart="Figs" captionStartId="5.[113,157,1658,1680]" captionTargetBox="[110,1456,188,1625]" captionTargetId="figure-9@5.[96,1469,172,1658]" captionTargetPageId="5" captionText="Figs. 1218. Enoplocephala maculata, new species, holotype, male. 1214, anterior part of body; 15, head; 16, fore wing; 17, 18, posterior part of body. 12, 16, 17, dorsal view; 13, lateral view; 14, 18, ventral view; 15, front view. Scale bars for 1214, 17, 18 = 2.00 mm; 15 = 1.00 mm; 16 = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577312" httpUri="https://zenodo.org/record/4577312/files/figure.png" pageId="7" pageNumber="376">Fig. 13</figureCitation>
); mandibular plates produced anteriorly, nearly parallel to each other; spinelike process of antenniferous tubercle relatively short, nearly horizontal, apically blunt (
<figureCitation id="130E2A08CF2FC1388DBFAF28B65067BA" box="[392,520,381,405]" captionStart="Figs" captionStartId="5.[113,157,1658,1680]" captionTargetBox="[110,1456,188,1625]" captionTargetId="figure-9@5.[96,1469,172,1658]" captionTargetPageId="5" captionText="Figs. 1218. Enoplocephala maculata, new species, holotype, male. 1214, anterior part of body; 15, head; 16, fore wing; 17, 18, posterior part of body. 12, 16, 17, dorsal view; 13, lateral view; 14, 18, ventral view; 15, front view. Scale bars for 1214, 17, 18 = 2.00 mm; 15 = 1.00 mm; 16 = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577312" httpUri="https://zenodo.org/record/4577312/files/figure.png" pageId="7" pageNumber="376">Figs. 12, 13</figureCitation>
). Pronotum (
<figureCitation id="130E2A08CF2FC1388EA5AF28B4D4679A" captionStart="Figs" captionStartId="5.[113,157,1658,1680]" captionTargetBox="[110,1456,188,1625]" captionTargetId="figure-9@5.[96,1469,172,1658]" captionTargetPageId="5" captionText="Figs. 1218. Enoplocephala maculata, new species, holotype, male. 1214, anterior part of body; 15, head; 16, fore wing; 17, 18, posterior part of body. 12, 16, 17, dorsal view; 13, lateral view; 14, 18, ventral view; 15, front view. Scale bars for 1214, 17, 18 = 2.00 mm; 15 = 1.00 mm; 16 = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577312" httpUri="https://zenodo.org/record/4577312/files/figure.png" pageId="7" pageNumber="376">Figs. 12, 13</figureCitation>
) 1.41 times wider than its length; spines of anterolateral angles curved outwardly, nearly horizontal; tubercles on lateral pronotal margins blunt, wart-like; humeral angles spine-shaped, strongly protruding posterolaterally (
<figureCitation id="130E2A08CF2FC1388EA4AFA8B6BA643A" box="[659,738,509,533]" captionStart="Figs" captionStartId="5.[113,157,1658,1680]" captionTargetBox="[110,1456,188,1625]" captionTargetId="figure-9@5.[96,1469,172,1658]" captionTargetPageId="5" captionText="Figs. 1218. Enoplocephala maculata, new species, holotype, male. 1214, anterior part of body; 15, head; 16, fore wing; 17, 18, posterior part of body. 12, 16, 17, dorsal view; 13, lateral view; 14, 18, ventral view; 15, front view. Scale bars for 1214, 17, 18 = 2.00 mm; 15 = 1.00 mm; 16 = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577312" httpUri="https://zenodo.org/record/4577312/files/figure.png" pageId="7" pageNumber="376">Fig. 12</figureCitation>
); posterior pronotal margin nearly straight. Scutellum (to apex of apical process) 1.06 times as broad as its basal width, lateral margins convex, base of apical process distinctly constricted (
<figureCitation id="130E2A08CF2FC1388CC8AC28B50E64BA" box="[255,342,637,661]" captionStart="Figs" captionStartId="5.[113,157,1658,1680]" captionTargetBox="[110,1456,188,1625]" captionTargetId="figure-9@5.[96,1469,172,1658]" captionTargetPageId="5" captionText="Figs. 1218. Enoplocephala maculata, new species, holotype, male. 1214, anterior part of body; 15, head; 16, fore wing; 17, 18, posterior part of body. 12, 16, 17, dorsal view; 13, lateral view; 14, 18, ventral view; 15, front view. Scale bars for 1214, 17, 18 = 2.00 mm; 15 = 1.00 mm; 16 = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577312" httpUri="https://zenodo.org/record/4577312/files/figure.png" pageId="7" pageNumber="376">Fig. 12</figureCitation>
). Ventral surface of fore trochanter with one (right) or three (left) small denticles; fore femur with two tiny denticles at base of ventral surface (
<figureCitation id="130E2A08CF2FC1388EA6ACE8B4E264DA" captionStart="Figs" captionStartId="5.[113,157,1658,1680]" captionTargetBox="[110,1456,188,1625]" captionTargetId="figure-9@5.[96,1469,172,1658]" captionTargetPageId="5" captionText="Figs. 1218. Enoplocephala maculata, new species, holotype, male. 1214, anterior part of body; 15, head; 16, fore wing; 17, 18, posterior part of body. 12, 16, 17, dorsal view; 13, lateral view; 14, 18, ventral view; 15, front view. Scale bars for 1214, 17, 18 = 2.00 mm; 15 = 1.00 mm; 16 = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577312" httpUri="https://zenodo.org/record/4577312/files/figure.png" pageId="7" pageNumber="376">Figs. 12, 14, 15</figureCitation>
). Abdomen 1.61 times as long as its greatest width.
</paragraph>
<paragraph id="8B8A368DCF2FC1388C46AD48B6BF629A" blockId="7.[112,753,797,1205]" pageId="7" pageNumber="376">
Male genitalia. Pygophore (
<figureCitation id="130E2A08CF2FC1388D94AD48B67E651A" box="[419,550,797,821]" captionStart="Figs" captionStartId="6.[135,179,1710,1732]" captionTargetBox="[140,1473,186,1691]" captionTargetId="figure-8@6.[128,1518,159,1735]" captionTargetPageId="6" captionText="Figs. 1928. Enoplocephala maculata, new species, holotype, male. 19, abdominal segment VIII; 2022, pygophore; 2325, paramere; 2628, phallus. 19, 27, ventral view; 20, 26, dorsal view; 21, 28, lateral view; 22, caudal view. Scale bars for 19, 2328 = 0.50 mm; 2022 = 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577314" httpUri="https://zenodo.org/record/4577314/files/figure.png" pageId="7" pageNumber="376">Figs. 2022</figureCitation>
) oblong, posterior part of pygophore expanded, posterior margin slightly concave at middle; median process (
<figureCitation id="130E2A08CF2FC1388E3DAD08B604655A" box="[522,604,861,885]" captionStart="Figs" captionStartId="6.[135,179,1710,1732]" captionTargetBox="[140,1473,186,1691]" captionTargetId="figure-8@6.[128,1518,159,1735]" captionTargetPageId="6" captionText="Figs. 1928. Enoplocephala maculata, new species, holotype, male. 19, abdominal segment VIII; 2022, pygophore; 2325, paramere; 2628, phallus. 19, 27, ventral view; 20, 26, dorsal view; 21, 28, lateral view; 22, caudal view. Scale bars for 19, 2328 = 0.50 mm; 2022 = 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577314" httpUri="https://zenodo.org/record/4577314/files/figure.png" pageId="7" pageNumber="376">Fig. 22</figureCitation>
) short, stout, apically rounded. Parameres (
<figureCitation id="130E2A08CF2FC1388D8BAD28B61965BA" box="[444,577,893,917]" captionStart="Figs" captionStartId="6.[135,179,1710,1732]" captionTargetBox="[140,1473,186,1691]" captionTargetId="figure-8@6.[128,1518,159,1735]" captionTargetPageId="6" captionText="Figs. 1928. Enoplocephala maculata, new species, holotype, male. 19, abdominal segment VIII; 2022, pygophore; 2325, paramere; 2628, phallus. 19, 27, ventral view; 20, 26, dorsal view; 21, 28, lateral view; 22, caudal view. Scale bars for 19, 2328 = 0.50 mm; 2022 = 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577314" httpUri="https://zenodo.org/record/4577314/files/figure.png" pageId="7" pageNumber="376">Figs. 2325</figureCitation>
) angularly bent near middle, with apical half covered with dense setae; subapical process lamellose. Phallus as shown in
<figureCitation id="130E2A08CF2FC1388E8EADE8B4EF65DA" captionStart="Figs" captionStartId="6.[135,179,1710,1732]" captionTargetBox="[140,1473,186,1691]" captionTargetId="figure-8@6.[128,1518,159,1735]" captionTargetPageId="6" captionText="Figs. 1928. Enoplocephala maculata, new species, holotype, male. 19, abdominal segment VIII; 2022, pygophore; 2325, paramere; 2628, phallus. 19, 27, ventral view; 20, 26, dorsal view; 21, 28, lateral view; 22, caudal view. Scale bars for 19, 2328 = 0.50 mm; 2022 = 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577314" httpUri="https://zenodo.org/record/4577314/files/figure.png" pageId="7" pageNumber="376">Figs. 2628</figureCitation>
; articulatory apparatus thick; basal plates relatively narrow, basal plate bridge thin and short, with two pairs of capitate processes; pedicel thin, slightly longer than basal plates; phallotheca elongate oval; dorsal sclerotised plate (
<figureCitation id="130E2A08CF2FC1388C4EAA08B4A3625A" box="[121,251,1117,1141]" captionStart="Figs" captionStartId="6.[135,179,1710,1732]" captionTargetBox="[140,1473,186,1691]" captionTargetId="figure-8@6.[128,1518,159,1735]" captionTargetPageId="6" captionText="Figs. 1928. Enoplocephala maculata, new species, holotype, male. 19, abdominal segment VIII; 2022, pygophore; 2325, paramere; 2628, phallus. 19, 27, ventral view; 20, 26, dorsal view; 21, 28, lateral view; 22, caudal view. Scale bars for 19, 2328 = 0.50 mm; 2022 = 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577314" httpUri="https://zenodo.org/record/4577314/files/figure.png" pageId="7" pageNumber="376">Figs. 26, 27</figureCitation>
) wrinkled at apical half and slightly incised at apex; struts (
<figureCitation id="130E2A08CF2FC1388CCFAA28B52E62BA" box="[248,374,1149,1173]" captionStart="Figs" captionStartId="6.[135,179,1710,1732]" captionTargetBox="[140,1473,186,1691]" captionTargetId="figure-8@6.[128,1518,159,1735]" captionTargetPageId="6" captionText="Figs. 1928. Enoplocephala maculata, new species, holotype, male. 19, abdominal segment VIII; 2022, pygophore; 2325, paramere; 2628, phallus. 19, 27, ventral view; 20, 26, dorsal view; 21, 28, lateral view; 22, caudal view. Scale bars for 19, 2328 = 0.50 mm; 2022 = 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577314" httpUri="https://zenodo.org/record/4577314/files/figure.png" pageId="7" pageNumber="376">Figs. 2628</figureCitation>
) nearly parallel to each other, apical portion converged and fused with dorsal sclerotised plate.
</paragraph>
<paragraph id="8B8A368DCF2FC1388C46AA88B6F4609A" blockId="7.[113,753,1245,1717]" pageId="7" pageNumber="376">
<emphasis id="B941EA9FCF2FC1388C46AA88B54062DA" bold="true" box="[113,280,1245,1269]" pageId="7" pageNumber="376">Measurements</emphasis>
[in mm, male (n = 1,
<typeStatus id="548E882FCF2FC1388E3EAA88B63662DA" box="[521,622,1245,1269]" pageId="7" pageNumber="376" type="holotype">holotype</typeStatus>
)]. Length of body (to apex of abdomen) = 17.60; length of head = 2.70; length of anteocular part = 1.35; length of postocular part = 0.55; width across eyes = 2.10; interocular space = 0.90; interocellar space = 0.40; length of antennal segments IIV = 3.15, 3.85, 0.80, 0.90; length of labial segments IIIV = 1.70, 0.70, 0.50; length of pronotum = 3.20; length of anterior pronotal lobe = 1.20; length of posterior pronotal lobe = 2.00; width of anterior pronotal lobe = 2.25; width of posterior pronotal lobe = 4.50; median length of scutellum = 1.85; basal width of scutellum = 1.75; length of fore femur, tibia, tarsus = 4.55, 5.25, 0.95; length of mid femur, tibia, tarsus = 5.15, 5.80, 0.95; length of hind femur, tibia, tarsus = 7.15, 10.20, 1.15; length of fore wing = 11.75; length of abdomen = 9.90; greatest width of abdomen = 6.15.
</paragraph>
</subSubSection>
<subSubSection id="C32F6506CF2FC1388C46A888B6FE615A" pageId="7" pageNumber="376" type="materials_examined">
<paragraph id="8B8A368DCF2FC1388C46A888B6FE615A" blockId="7.[112,754,1757,1909]" pageId="7" pageNumber="376">
<emphasis id="B941EA9FCF2FC1388C46A888B54F60DA" bold="true" box="[113,279,1757,1781]" pageId="7" pageNumber="376">Type material.</emphasis>
<materialsCitation id="3B5D3CD0CF2FC1388D2CA888B6FE615A" collectingDate="2017-02" collectionCode="CAU, CHEN" collectorName="Liu Yingqi" country="Malaysia" location="Borneo Jungle Girl Camp" pageId="7" pageNumber="376" specimenCount="2" stateProvince="Sabah" typeStatus="holotype">
<typeStatus id="548E882FCF2FC1388D2CA888B52660DA" box="[283,382,1757,1781]" pageId="7" pageNumber="376" type="holotype">Holotype</typeStatus>
(male): “
<collectingCountry id="F322761DCF2FC1388DE9A888B61C60DA" box="[478,580,1757,1781]" name="Malaysia" pageId="7" pageNumber="376">Malaysia</collectingCountry>
,
<collectingRegion id="49F1F86FCF2FC1388E79A888B6D760DA" box="[590,655,1757,1781]" country="Malaysia" name="Sabah" pageId="7" pageNumber="376">Sabah</collectingRegion>
\
<location id="8EEA6056CF2FC1388E97A888B564613A" LSID="urn:lsid:plazi:treatment:039C879BCF2CC1388F99A970B5896E1A:8EEA6056CF2FC1388E97A888B564613A" country="Malaysia" name="Borneo Jungle Girl Camp" pageId="7" pageNumber="376" stateProvince="Sabah">Borneo Jungle Girl Camp</location>
\
<date id="FF8B104DCF2FC1388D6EA8A8B5EE613A" box="[345,438,1789,1813]" bridgedPair="-" pageId="7" pageNumber="376" value="2017-02">
<collectingDate id="EFCFE9A5CF2FC1388D6EA8A8B5EE613A" box="[345,438,1789,1813]" bridgedPair="-" pageId="7" pageNumber="376" value="2017-02">2017-II-</collectingDate>
</date>
[printed] 4 [handwritten] \ leg.
<collectorName id="26C0535BCF2FC1388C96A948B541611A" box="[161,281,1821,1845]" pageId="7" pageNumber="376">Liu Yingqi</collectorName>
\ Ent. Mus.
<collectionCode id="ED24AE48CF2FC1388D92A948B5B9611A" box="[421,481,1821,1845]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="7" pageNumber="376">CAU</collectionCode>
. Beijing”; “
<typeStatus id="548E882FCF2FC1388E56A948B6A8611A" box="[609,752,1821,1845]" pageId="7" pageNumber="376" type="holotype">HOLOTYPE</typeStatus>
[printed] \
<taxonomicName id="4C354D0ECF2FC1388CD9A968B5D7617A" authorityName="Miller" authorityYear="1940" box="[238,399,1853,1877]" class="Insecta" family="Reduviidae" genus="Enoplocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="376" phylum="Arthropoda" rank="genus">Enoplocephala</taxonomicName>
[handwritten] \
<taxonomicName id="4C354D0ECF2FC1388E7EA968B6F3617A" authority="Chen &amp; Liu &amp; Cai, 2020" authorityName="Chen &amp; Liu &amp; Cai" authorityYear="2020" box="[585,683,1853,1877]" class="Insecta" family="Reduviidae" genus="Enoplocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="376" phylum="Arthropoda" rank="species" species="maculata" status="sp. nov.">maculata</taxonomicName>
<taxonomicNameLabel id="A27257E4CF2FC1388E82A968B6AA617B" box="[693,754,1853,1876]" pageId="7" pageNumber="376" rank="species">sp. n.</taxonomicNameLabel>
[handwritten] \ Det.
<collectionCode id="ED24AE48CF2FC1388D66A908B5C2615A" box="[337,410,1885,1909]" pageId="7" pageNumber="376">CHEN</collectionCode>
Zhuo [printed]” (
<collectionCode id="ED24AE48CF2FC1388E6AA908B6C3615A" box="[605,667,1885,1909]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="7" pageNumber="376">CAU</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32F6506CF2FC1388C46A9C8B5D761DA" pageId="7" pageNumber="376" type="etymology">
<paragraph id="8B8A368DCF2FC1388C46A9C8B5D761DA" blockId="7.[112,752,1949,2037]" pageId="7" pageNumber="376">
<emphasis id="B941EA9FCF2FC1388C46A9C8B4A1619A" bold="true" box="[113,249,1949,1973]" pageId="7" pageNumber="376">Etymology.</emphasis>
The specific epithet is derived from Latin
<emphasis id="B941EA9FCF2FC1388C4DA9E8B4B061FA" box="[122,232,1981,2005]" italics="true" pageId="7" pageNumber="376">maculatus</emphasis>
(meaning spotted), referring to the mottled body colour of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C32F6506CF2FC1388C46A648B5896E1A" box="[113,465,2077,2101]" pageId="7" pageNumber="376" type="distribution">
<paragraph id="8B8A368DCF2FC1388C46A648B5896E1A" blockId="7.[113,465,2077,2101]" box="[113,465,2077,2101]" pageId="7" pageNumber="376">
<emphasis id="B941EA9FCF2FC1388C46A648B55C6E1A" bold="true" box="[113,260,2077,2101]" pageId="7" pageNumber="376">Distribution.</emphasis>
<collectingCountry id="F322761DCF2FC1388D3BA648B5366E1A" box="[268,366,2077,2101]" name="Malaysia" pageId="7" pageNumber="376">Malaysia</collectingCountry>
(
<collectingRegion id="49F1F86FCF2FC1388D49A648B59D6E1A" box="[382,453,2077,2101]" country="Malaysia" name="Sabah" pageId="7" pageNumber="376">Sabah</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>