235 lines
36 KiB
XML
235 lines
36 KiB
XML
<document id="7ABAE92AC5EDC73A05FEFFBC373EDD3C" ID-DOI="10.26107/RBZ-2020-0053" ID-ISSN="2345-7600" ID-Zenodo-Dep="4577302" ID-ZooBank="65CDBE40-FEA7-4F06-A30F-6F970E63AE6C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614785065864" checkinUser="carolina" docAuthor="Chen, Zhuo, Liu, Yingqi & Cai, Wanzhi" docDate="2020" docId="039C879BCF2BC13B8CE2A9CEB00460D2" docLanguage="en" docName="RafflesBZool.68.369-378.pdf" docOrigin="Raffles Bulletin of Zoology 68" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Enoplocephala perakensis Miller 1940" docType="treatment" docVersion="3" lastPageNumber="373" masterDocId="FFA5FFE3CF28C13F8C37AE55B458662F" masterDocTitle="Notes on the genus Enoplocephala Miller (Hemiptera: Reduviidae: Stenopodainae), with the description of a new species from Borneo" masterLastPageNumber="378" masterPageNumber="369" pageNumber="372" updateTime="1698878663914" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="BB9BDC38B9FD4453F288E63C95AFE491">Notes on the genus Enoplocephala Miller (Hemiptera: Reduviidae: Stenopodainae), with the description of a new species from Borneo</mods:title>
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<mods:namePart id="509A2F8FA9B3BA2692CE41CAB9B018E2">Chen, Zhuo</mods:namePart>
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<mods:namePart id="5FB1E1CD9AF37192A7376471C9A97261">Liu, Yingqi</mods:namePart>
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<mods:namePart id="B4326714F25D5306358AB131A79DB5BB">Cai, Wanzhi</mods:namePart>
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<mods:title id="EC87B0703D5E162C686B439587F242E0">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="C51C3AAA87B3772679790915711A747F">2020</mods:date>
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<mods:identifier id="0648F34248D7DDA1EE54C64956126038" type="DOI">10.26107/RBZ-2020-0053</mods:identifier>
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<paragraph id="8B8A368DCF2BC13C8CE2A9CEB6D6619C" blockId="3.[213,654,1947,2004]" box="[213,654,1947,1971]" pageId="3" pageNumber="372">
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<heading id="D0C281E1CF2BC13C8CE2A9CEB6D6619C" bold="true" box="[213,654,1947,1971]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="372" reason="2">
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<taxonomicName id="4C354D0ECF2BC13C8CE2A9CEB6D6619C" ID-CoL="39WKC" authority="Miller, 1940" authorityName="Miller" authorityYear="1940" box="[213,654,1947,1971]" class="Insecta" family="Reduviidae" genus="Enoplocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="perakensis">
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<emphasis id="B941EA9FCF2BC13C8CE2A9CEB6D6619C" bold="true" box="[213,654,1947,1971]" pageId="3" pageNumber="372">
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<emphasis id="B941EA9FCF2BC13C8CE2A9CEB5A0619C" bold="true" box="[213,504,1947,1971]" italics="true" pageId="3" pageNumber="372">Enoplocephala perakensis</emphasis>
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<bibRefCitation id="EFA44B7CCF2BC13C8E34A9CEB6D6619C" author="Miller NCE" box="[515,654,1947,1971]" pageId="3" pageNumber="372" pagination="1 - 115" refId="ref7497" refString="Miller NCE (1940) New genera and species of Malaysian Reduviidae (Hemiptera: Reduviidae). Canadian Journal of Arthropod (Part I). Journal of the Federated Malay States Museums, 18: Identification, 26: 1 - 115. doi: 10.3752 / cjai. 2014.26." type="journal article" year="1940">Miller, 1940</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32F6506CF2BC13C8D57A9EEB65A61FB" box="[352,514,1979,2004]" pageId="3" pageNumber="372" type="description">
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<paragraph id="8B8A368DCF2BC13C8D57A9EEB65A61FB" blockId="3.[213,654,1947,2004]" box="[352,514,1979,2004]" pageId="3" pageNumber="372">
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(
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<figureCitation id="130E2A08CF2BC13C8D5FA9EEB58861FB" box="[360,464,1979,2004]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="1.[114,159,1015,1037]" captionStartId-1="2.[135,179,1605,1627]" captionTargetBox-0="[97,1475,170,1066]" captionTargetBox-1="[137,1470,203,1571]" captionTargetId-0="figure-427@1.[97,1475,170,1048]" captionTargetId-1="figure-7@2.[117,1499,170,1609]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Figs. 1–4. Enoplocephala perakensis Miller, 1940, holotype, male, habitus. 1, dorsal view; 2, lateral view; 3, ventral view; 4, labels. Scale bar of 1–3 = 5.00 mm." captionText-1="Figs. 5–8. Enoplocephala perakensis Miller, 1940, holotype, male. 5, 6, anterior part of body; 7, body habitus; 8, posterior part of body. 5, 8, dorsal view; 6, lateral view; 7, ventral view. Scale bars = 3.00 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4577304" figureDoi-1="http://doi.org/10.5281/zenodo.4577308" httpUri-0="https://zenodo.org/record/4577304/files/figure.png" httpUri-1="https://zenodo.org/record/4577308/files/figure.png" pageId="3" pageNumber="372">Figs. 1–8</figureCitation>
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,
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<figureCitation id="130E2A08CF2BC13C8DE9A9E9B5A261FB" box="[478,506,1980,2004]" captionStart="Fig" captionStartId="8.[135,170,997,1019]" captionTargetBox="[135,1473,189,982]" captionTargetId="figure-14@8.[135,1473,189,982]" captionTargetPageId="8" captionText="Fig. 29. Known distribution of Enoplocephala species." figureDoi="http://doi.org/10.5281/zenodo.4577316" httpUri="https://zenodo.org/record/4577316/files/figure.png" pageId="3" pageNumber="372">29</figureCitation>
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)
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</paragraph>
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<subSubSection id="C32F6506CF2BC13C8C46A9AFB6766E20" box="[113,558,2042,2064]" pageId="3" pageNumber="372" type="reference_group">
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<paragraph id="8B8A368DCF2BC13C8C46A9AFB6766E20" blockId="3.[113,558,2042,2064]" box="[113,558,2042,2064]" pageId="3" pageNumber="372">
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<treatmentCitation id="0A94109CCF2BC13C8C46A9AFB6726E20" author="Miller NCE" box="[113,554,2042,2064]" page="441" pageId="3" pageNumber="372" year="1940">
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<taxonomicName id="4C354D0ECF2BC13C8C46A9AFB6726E20" ID-CoL="39WKC" authority="Miller, 1940: 441" authorityName="Miller" authorityPageNumber="441" authorityYear="1940" box="[113,554,2042,2064]" class="Insecta" family="Reduviidae" genus="Enoplocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="perakensis">
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<emphasis id="B941EA9FCF2BC13C8C46A9AFB5296E20" box="[113,369,2042,2063]" italics="true" pageId="3" pageNumber="372">Enoplocephala perakensis</emphasis>
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<bibRefCitation id="EFA44B7CCF2BC13C8D4CA9AFB6726E20" author="Miller NCE" box="[379,554,2042,2064]" pageId="3" pageNumber="372" pagination="1 - 115" refId="ref7497" refString="Miller NCE (1940) New genera and species of Malaysian Reduviidae (Hemiptera: Reduviidae). Canadian Journal of Arthropod (Part I). Journal of the Federated Malay States Museums, 18: Identification, 26: 1 - 115. doi: 10.3752 / cjai. 2014.26." type="journal article" year="1940">Miller, 1940: 441</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<subSubSection id="C32F6506CF2BC13C8F1AAEE8B0A666FA" box="[813,1278,189,213]" pageId="3" pageNumber="372" type="materials_examined">
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<paragraph id="8B8A368DCF2BC13C8F1AAEE8B0A666FA" blockId="3.[813,1278,189,213]" box="[813,1278,189,213]" pageId="3" pageNumber="372">
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<materialsCitation id="3B5D3CD0CF2BC13C8F1AAEE8B0A666FA" box="[813,1278,189,213]" collectionCode="BMNH" country="Malaysia" location="Perak" pageId="3" pageNumber="372" specimenCount="1" stateProvince="Perak" typeStatus="holotype">
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<emphasis id="B941EA9FCF2BC13C8F1AAEE8B7C366FA" bold="true" box="[813,923,189,213]" pageId="3" pageNumber="372">
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<typeStatus id="548E882FCF2BC13C8F1AAEE8B7CF66FA" box="[813,919,189,213]" pageId="3" pageNumber="372" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Male.
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<collectingCountry id="F322761DCF2BC13C8FDDAEE8B01666FA" box="[1002,1102,189,213]" name="Malaysia" pageId="3" pageNumber="372">Malaysia</collectingCountry>
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,
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<collectingRegion id="49F1F86FCF2BC13C886CAEE8B0C366FA" box="[1115,1179,189,213]" country="Malaysia" name="Perak" pageId="3" pageNumber="372">Perak</collectingRegion>
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,
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<collectionCode id="ED24AE48CF2BC13C8890AEEBB0A266FA" box="[1191,1274,190,213]" pageId="3" pageNumber="372">BMNH</collectionCode>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32F6506CF2BC13C8F1AAEA8B0D367DA" pageId="3" pageNumber="372" type="diagnosis">
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<paragraph id="8B8A368DCF2BC13C8F1AAEA8B0D367DA" blockId="3.[811,1451,253,501]" pageId="3" pageNumber="372">
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<emphasis id="B941EA9FCF2BC13C8F1AAEA8B7F9673A" bold="true" box="[813,929,253,277]" pageId="3" pageNumber="372">Diagnosis.</emphasis>
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Body
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<quantity id="4CCD9B68CF2BC13C8FD1AEA8B068673A" box="[998,1072,253,277]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="372" unit="mm" value="18.0">18 mm</quantity>
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in length, dark brown, mottled with yellowish brown spots and markings; head slightly shorter than pronotum, about 1.4 times longer than width across eyes; process of antenniferous tubercle long, acute, suberect; spines of anterolateral angles of pronotum straight, suberect; humeral angles protruding laterally; fore wing with blackish patch occupying area between Cu and claval suture; blackish patch between M and Cu broad.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32F6506CF2BC13B8F1AAC48B63C6052" lastPageId="4" lastPageNumber="373" pageId="3" pageNumber="372" type="description">
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<paragraph id="8B8A368DCF2BC13C8F1AAC48B047639A" blockId="3.[811,1454,541,1461]" pageId="3" pageNumber="372">
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<emphasis id="B941EA9FCF2BC13C8F1AAC48B796641A" bold="true" box="[813,974,541,565]" pageId="3" pageNumber="372">Redescription.</emphasis>
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Macropterous male (
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<figureCitation id="130E2A08CF2BC13C889EAC48B154641A" box="[1193,1292,541,565]" captionStart="Figs" captionStartId="1.[114,159,1015,1037]" captionTargetBox="[97,1475,170,1066]" captionTargetId="figure-427@1.[97,1475,170,1048]" captionTargetPageId="1" captionText="Figs. 1–4. Enoplocephala perakensis Miller, 1940, holotype, male, habitus. 1, dorsal view; 2, lateral view; 3, ventral view; 4, labels. Scale bar of 1–3 = 5.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577304" httpUri="https://zenodo.org/record/4577304/files/figure.png" pageId="3" pageNumber="372">Figs. 1–3</figureCitation>
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).
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<emphasis id="B941EA9FCF2BC13C892BAC48B1F2641A" bold="true" box="[1308,1450,541,565]" pageId="3" pageNumber="372">Colouration.</emphasis>
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Body dark brown. Head with ventrolateral and ventral surfaces darker; ocelli yellowish brown; antennal segments I and II brown, segment I mottled with black on basal third and apical third, and with two narrow, incomplete, blackish annuli medially (
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<figureCitation id="130E2A08CF2BC13C8869ACE8B0FE64FA" box="[1118,1190,701,725]" captionStart="Figs" captionStartId="2.[135,179,1605,1627]" captionTargetBox="[137,1470,203,1571]" captionTargetId="figure-7@2.[117,1499,170,1609]" captionTargetPageId="2" captionText="Figs. 5–8. Enoplocephala perakensis Miller, 1940, holotype, male. 5, 6, anterior part of body; 7, body habitus; 8, posterior part of body. 5, 8, dorsal view; 6, lateral view; 7, ventral view. Scale bars = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577308" httpUri="https://zenodo.org/record/4577308/files/figure.png" pageId="3" pageNumber="372">Fig. 5</figureCitation>
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), segment II black on base, medial third, and apex (
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<figureCitation id="130E2A08CF2BC13C884FAC88B0E564DA" box="[1144,1213,733,757]" captionStart="Figs" captionStartId="2.[135,179,1605,1627]" captionTargetBox="[137,1470,203,1571]" captionTargetId="figure-7@2.[117,1499,170,1609]" captionTargetPageId="2" captionText="Figs. 5–8. Enoplocephala perakensis Miller, 1940, holotype, male. 5, 6, anterior part of body; 7, body habitus; 8, posterior part of body. 5, 8, dorsal view; 6, lateral view; 7, ventral view. Scale bars = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577308" httpUri="https://zenodo.org/record/4577308/files/figure.png" pageId="3" pageNumber="372">Fig. 6</figureCitation>
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), segment III greyish brown; labium brown, with dorsal and lateral surfaces of apical half of segment II and apical third of segment III black (
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<figureCitation id="130E2A08CF2BC13C8F4FAD68B7BE657A" box="[888,998,829,853]" captionStart="Figs" captionStartId="2.[135,179,1605,1627]" captionTargetBox="[137,1470,203,1571]" captionTargetId="figure-7@2.[117,1499,170,1609]" captionTargetPageId="2" captionText="Figs. 5–8. Enoplocephala perakensis Miller, 1940, holotype, male. 5, 6, anterior part of body; 7, body habitus; 8, posterior part of body. 5, 8, dorsal view; 6, lateral view; 7, ventral view. Scale bars = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577308" httpUri="https://zenodo.org/record/4577308/files/figure.png" pageId="3" pageNumber="372">Figs. 6, 7</figureCitation>
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). Pronotum with apical half of spines of anterolateral angles, tubercles on lateral margins, submedian carinae, and spines of humeral angles lighter (
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<figureCitation id="130E2A08CF2BC13C8928AD28B13965BA" box="[1311,1377,893,917]" captionStart="Figs" captionStartId="2.[135,179,1605,1627]" captionTargetBox="[137,1470,203,1571]" captionTargetId="figure-7@2.[117,1499,170,1609]" captionTargetPageId="2" captionText="Figs. 5–8. Enoplocephala perakensis Miller, 1940, holotype, male. 5, 6, anterior part of body; 7, body habitus; 8, posterior part of body. 5, 8, dorsal view; 6, lateral view; 7, ventral view. Scale bars = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577308" httpUri="https://zenodo.org/record/4577308/files/figure.png" pageId="3" pageNumber="372">Fig. 5</figureCitation>
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). Legs brown; basal, medial, and subapical parts of all femora each with a blackish, irregular, incomplete annulus (
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<figureCitation id="130E2A08CF2BC13C8903ADE8B1C565FA" box="[1332,1437,957,981]" captionStart="Figs" captionStartId="1.[114,159,1015,1037]" captionTargetBox="[97,1475,170,1066]" captionTargetId="figure-427@1.[97,1475,170,1048]" captionTargetPageId="1" captionText="Figs. 1–4. Enoplocephala perakensis Miller, 1940, holotype, male, habitus. 1, dorsal view; 2, lateral view; 3, ventral view; 4, labels. Scale bar of 1–3 = 5.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577304" httpUri="https://zenodo.org/record/4577304/files/figure.png" pageId="3" pageNumber="372">Figs. 1–3</figureCitation>
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); basal and subapical parts of fore and mid tibiae each with a blackish annulus (
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<figureCitation id="130E2A08CF2BC13C8826ADA8B026623A" box="[1041,1150,1021,1045]" captionStart="Figs" captionStartId="1.[114,159,1015,1037]" captionTargetBox="[97,1475,170,1066]" captionTargetId="figure-427@1.[97,1475,170,1048]" captionTargetPageId="1" captionText="Figs. 1–4. Enoplocephala perakensis Miller, 1940, holotype, male, habitus. 1, dorsal view; 2, lateral view; 3, ventral view; 4, labels. Scale bar of 1–3 = 5.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577304" httpUri="https://zenodo.org/record/4577304/files/figure.png" pageId="3" pageNumber="372">Figs. 1–3</figureCitation>
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). Fore wing with blackish patches in corium occupying whole area of cubital cell and areas between R+M and Cu (paler), M and Cu, Cu and claval suture, and R and M+Cu (
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<figureCitation id="130E2A08CF2BC13C889EAA08B0B7625A" box="[1193,1263,1117,1141]" captionStart="Figs" captionStartId="2.[135,179,1605,1627]" captionTargetBox="[137,1470,203,1571]" captionTargetId="figure-7@2.[117,1499,170,1609]" captionTargetPageId="2" captionText="Figs. 5–8. Enoplocephala perakensis Miller, 1940, holotype, male. 5, 6, anterior part of body; 7, body habitus; 8, posterior part of body. 5, 8, dorsal view; 6, lateral view; 7, ventral view. Scale bars = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577308" httpUri="https://zenodo.org/record/4577308/files/figure.png" pageId="3" pageNumber="372">Fig. 8</figureCitation>
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); blackish patch between M and Cu relatively broad (
|
||
<figureCitation id="130E2A08CF2BC13C888CAA28B15D62BA" box="[1211,1285,1149,1173]" captionStart="Figs" captionStartId="1.[114,159,1015,1037]" captionTargetBox="[97,1475,170,1066]" captionTargetId="figure-427@1.[97,1475,170,1048]" captionTargetPageId="1" captionText="Figs. 1–4. Enoplocephala perakensis Miller, 1940, holotype, male, habitus. 1, dorsal view; 2, lateral view; 3, ventral view; 4, labels. Scale bar of 1–3 = 5.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577304" httpUri="https://zenodo.org/record/4577304/files/figure.png" pageId="3" pageNumber="372">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="130E2A08CF2BC13C8923AA28B17B62BA" box="[1300,1315,1149,1173]" captionStart="Figs" captionStartId="2.[135,179,1605,1627]" captionTargetBox="[137,1470,203,1571]" captionTargetId="figure-7@2.[117,1499,170,1609]" captionTargetPageId="2" captionText="Figs. 5–8. Enoplocephala perakensis Miller, 1940, holotype, male. 5, 6, anterior part of body; 7, body habitus; 8, posterior part of body. 5, 8, dorsal view; 6, lateral view; 7, ventral view. Scale bars = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577308" httpUri="https://zenodo.org/record/4577308/files/figure.png" pageId="3" pageNumber="372">8</figureCitation>
|
||
); exocorium with an elongate, blackish patch medially; medial border of Pcu with a narrow, elongate, blackish spot; clavus with an elongate, blackish patch along its medial border; membrane with a basal, subtriangular spot and a subapical, irregular patch in outer cell, and a narrow, elongate, blackish spot near base of M in membrane (
|
||
<figureCitation id="130E2A08CF2BC13C88B1AB68B093637A" box="[1158,1227,1341,1365]" captionStart="Figs" captionStartId="2.[135,179,1605,1627]" captionTargetBox="[137,1470,203,1571]" captionTargetId="figure-7@2.[117,1499,170,1609]" captionTargetPageId="2" captionText="Figs. 5–8. Enoplocephala perakensis Miller, 1940, holotype, male. 5, 6, anterior part of body; 7, body habitus; 8, posterior part of body. 5, 8, dorsal view; 6, lateral view; 7, ventral view. Scale bars = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577308" httpUri="https://zenodo.org/record/4577308/files/figure.png" pageId="3" pageNumber="372">Fig. 8</figureCitation>
|
||
). Abdomen mottled with yellowish brown; basal two-thirds of lateral margins of each abdominal segment yellowish brown; posterolateral angles darker (
|
||
<figureCitation id="130E2A08CF2BC13C8FF9ABC8B049639A" box="[974,1041,1437,1461]" captionStart="Figs" captionStartId="2.[135,179,1605,1627]" captionTargetBox="[137,1470,203,1571]" captionTargetId="figure-7@2.[117,1499,170,1609]" captionTargetPageId="2" captionText="Figs. 5–8. Enoplocephala perakensis Miller, 1940, holotype, male. 5, 6, anterior part of body; 7, body habitus; 8, posterior part of body. 5, 8, dorsal view; 6, lateral view; 7, ventral view. Scale bars = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577308" httpUri="https://zenodo.org/record/4577308/files/figure.png" pageId="3" pageNumber="372">Fig. 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8A368DCF2BC13C8F1AAB88B03D615A" blockId="3.[811,1453,1501,1910]" pageId="3" pageNumber="372">
|
||
<emphasis id="B941EA9FCF2BC13C8F1AAB88B7C763DA" bold="true" box="[813,927,1501,1525]" pageId="3" pageNumber="372">Structure.</emphasis>
|
||
Body shape and vestiture as in generic description. Head (
|
||
<figureCitation id="130E2A08CF2BC13C8F42ABA8B784603A" box="[885,988,1533,1557]" captionStart="Figs" captionStartId="2.[135,179,1605,1627]" captionTargetBox="[137,1470,203,1571]" captionTargetId="figure-7@2.[117,1499,170,1609]" captionTargetPageId="2" captionText="Figs. 5–8. Enoplocephala perakensis Miller, 1940, holotype, male. 5, 6, anterior part of body; 7, body habitus; 8, posterior part of body. 5, 8, dorsal view; 6, lateral view; 7, ventral view. Scale bars = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577308" httpUri="https://zenodo.org/record/4577308/files/figure.png" pageId="3" pageNumber="372">Figs. 5–7</figureCitation>
|
||
) 1.43 times as long as width across eyes, slightly shorter than pronotum; ventral margin of anteocular part nearly straight in lateral view (
|
||
<figureCitation id="130E2A08CF2BC13C88FCA868B14B607A" box="[1227,1299,1597,1621]" captionStart="Figs" captionStartId="2.[135,179,1605,1627]" captionTargetBox="[137,1470,203,1571]" captionTargetId="figure-7@2.[117,1499,170,1609]" captionTargetPageId="2" captionText="Figs. 5–8. Enoplocephala perakensis Miller, 1940, holotype, male. 5, 6, anterior part of body; 7, body habitus; 8, posterior part of body. 5, 8, dorsal view; 6, lateral view; 7, ventral view. Scale bars = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577308" httpUri="https://zenodo.org/record/4577308/files/figure.png" pageId="3" pageNumber="372">Fig. 6</figureCitation>
|
||
); mandibular plates slightly produced anterolaterally (
|
||
<figureCitation id="130E2A08CF2BC13C88D1A808B172605A" box="[1254,1322,1629,1653]" captionStart="Figs" captionStartId="2.[135,179,1605,1627]" captionTargetBox="[137,1470,203,1571]" captionTargetId="figure-7@2.[117,1499,170,1609]" captionTargetPageId="2" captionText="Figs. 5–8. Enoplocephala perakensis Miller, 1940, holotype, male. 5, 6, anterior part of body; 7, body habitus; 8, posterior part of body. 5, 8, dorsal view; 6, lateral view; 7, ventral view. Scale bars = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577308" httpUri="https://zenodo.org/record/4577308/files/figure.png" pageId="3" pageNumber="372">Fig. 5</figureCitation>
|
||
); spine-like process of antenniferous tubercle long, suberect, apically acute. Pronotum (
|
||
<figureCitation id="130E2A08CF2BC13C8FDFA8C8B013609A" box="[1000,1099,1693,1717]" captionStart="Figs" captionStartId="2.[135,179,1605,1627]" captionTargetBox="[137,1470,203,1571]" captionTargetId="figure-7@2.[117,1499,170,1609]" captionTargetPageId="2" captionText="Figs. 5–8. Enoplocephala perakensis Miller, 1940, holotype, male. 5, 6, anterior part of body; 7, body habitus; 8, posterior part of body. 5, 8, dorsal view; 6, lateral view; 7, ventral view. Scale bars = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577308" httpUri="https://zenodo.org/record/4577308/files/figure.png" pageId="3" pageNumber="372">Figs. 5, 6</figureCitation>
|
||
) 1.45 times wider than its length; spines of anterolateral angles straight, suberect; tubercles on lateral pronotal margins relatively acute; humeral angles strongly protruding laterally (
|
||
<figureCitation id="130E2A08CF2BC13C88B6A8A8B091613A" box="[1153,1225,1789,1813]" captionStart="Figs" captionStartId="2.[135,179,1605,1627]" captionTargetBox="[137,1470,203,1571]" captionTargetId="figure-7@2.[117,1499,170,1609]" captionTargetPageId="2" captionText="Figs. 5–8. Enoplocephala perakensis Miller, 1940, holotype, male. 5, 6, anterior part of body; 7, body habitus; 8, posterior part of body. 5, 8, dorsal view; 6, lateral view; 7, ventral view. Scale bars = 3.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577308" httpUri="https://zenodo.org/record/4577308/files/figure.png" pageId="3" pageNumber="372">Fig. 5</figureCitation>
|
||
); posterior pronotal margin slightly concave. Ventral surface of fore trochanter and femur without denticle and spine. Abdomen 1.56 times as long as its greatest width.
|
||
</paragraph>
|
||
<paragraph id="8B8A368DCF2BC13C8F1AA9C8B7C461FA" blockId="3.[813,1451,1949,2005]" pageId="3" pageNumber="372">Male genitalia. As in generic description. Phallus not examined.</paragraph>
|
||
<paragraph id="8B8A368DCF2BC13C8F1AA9A8B1F36E1A" blockId="3.[813,1451,2045,2101]" pageId="3" pageNumber="372">
|
||
<emphasis id="B941EA9FCF2BC13C8F1AA9A8B78F6E3A" bold="true" box="[813,983,2045,2069]" pageId="3" pageNumber="372">Measurements</emphasis>
|
||
[in mm, male (n = 1,
|
||
<typeStatus id="548E882FCF2BC13C88EBA9A8B11A6E3A" box="[1244,1346,2045,2069]" pageId="3" pageNumber="372" type="holotype">holotype</typeStatus>
|
||
)]. Length of body (to apex of abdomen) = 18.00; length of head =
|
||
</paragraph>
|
||
<paragraph id="8B8A368DCF2CC13B8E01AE27B04B66A5" blockId="4.[566,1043,114,138]" box="[566,1043,114,138]" pageId="4" pageNumber="373">
|
||
<emphasis id="B941EA9FCF2CC13B8E01AE27B78D66A6" box="[566,981,114,137]" italics="true" pageId="4" pageNumber="373">RAFFLES BULLETIN OF ZOOLOGY</emphasis>
|
||
2020
|
||
</paragraph>
|
||
<caption id="DF4A6605CF2CC13B8CBFAA2EB4A46281" ID-DOI="http://doi.org/10.5281/zenodo.4577310" ID-Zenodo-Dep="4577310" httpUri="https://zenodo.org/record/4577310/files/figure.png" pageId="4" pageNumber="373" startId="4.[136,180,1147,1169]" targetBox="[111,1500,172,1145]" targetPageId="4">
|
||
<paragraph id="8B8A368DCF2CC13B8CBFAA2EB4A46281" blockId="4.[136,1474,1147,1198]" pageId="4" pageNumber="373">
|
||
Figs. 9–11.
|
||
<taxonomicName id="4C354D0ECF2CC13B8CC9AA2EB5A962BF" authority="Chen & Liu & Cai, 2020" authorityName="Chen & Liu & Cai" authorityYear="2020" box="[254,497,1147,1168]" class="Insecta" family="Reduviidae" genus="Enoplocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="species" species="maculata" status="sp. nov.">
|
||
<emphasis id="B941EA9FCF2CC13B8CC9AA2EB5A962BF" box="[254,497,1147,1168]" italics="true" pageId="4" pageNumber="373">Enoplocephala maculata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A27257E4CF2CC13B8DC8AA29B62F62BE" box="[511,631,1147,1169]" pageId="4" pageNumber="373" rank="species">new species</taxonomicNameLabel>
|
||
, holotype, male, habitus. 9, dorsal view; 10, lateral view; 11, ventral view. Scale bar = 5.00 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8A368DCF2CC13B8CBFAAB0B63C6052" blockId="4.[135,776,1253,1662]" pageId="4" pageNumber="373">3.00; length of anteocular part = 1.65; length of postocular part = 0.60; width across eyes = 2.10; interocular space = 0.80; interocellar space = 0.20; length of antennal segments I–IV = 3.30, 3.65, 0.90,?; length of labial segments II–IV = 2.10, 0.85, 0.60; length of pronotum = 3.30; length of anterior pronotal lobe = 1.30; length of posterior pronotal lobe = 2.00; width of anterior pronotal lobe = 2.10; width of posterior pronotal lobe = 4.80; median length of scutellum = 1.60; basal width of scutellum = 1.40; length of fore femur, tibia, tarsus = 6.00, 6.20, 1.10; length of mid femur, tibia, tarsus = 6.30, 7.40, 1.10; length of hind femur, tibia, tarsus = 9.00,?,?; length of fore wing = 12.40; length of abdomen = 10.20; greatest width of abdomen = 6.55.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32F6506CF2CC13B8CBFA8F0B50761D2" pageId="4" pageNumber="373" type="materials_examined">
|
||
<paragraph id="8B8A368DCF2CC13B8CBFA8F0B50761D2" blockId="4.[135,776,1701,2045]" pageId="4" pageNumber="373">
|
||
<emphasis id="B941EA9FCF2CC13B8CBFA8F0B5F26092" bold="true" box="[136,426,1701,1725]" pageId="4" pageNumber="373">Type material examined.</emphasis>
|
||
<typeStatus id="548E882FCF2CC13B8D83A8F0B64F6092" box="[436,535,1701,1725]" pageId="4" pageNumber="373" type="holotype">Holotype</typeStatus>
|
||
(male): red-margined
|
||
<typeStatus id="548E882FCF2CC13B8CBFA890B4BD60F2" box="[136,229,1733,1757]" pageId="4" pageNumber="373" type="holotype">holotype</typeStatus>
|
||
disc; red-margined type disc; “male sex symbol”; “
|
||
<collectingRegion id="49F1F86FCF2CC13B8CA3A8B0B4AB60D2" box="[148,243,1765,1789]" country="Malaysia" name="Perak" pageId="4" pageNumber="373">PERAK</collectingRegion>
|
||
\ DOHERTY”; “Distant Coll. \ 1911-383”; “
|
||
<taxonomicName id="4C354D0ECF2CC13B8CA6A950B5696132" authorityName="Miller" authorityYear="1940" box="[145,305,1797,1821]" class="Insecta" family="Reduviidae" genus="Enoplocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">Enoplocephala</taxonomicName>
|
||
[Miller’s handwriting] \
|
||
<taxonomicName id="4C354D0ECF2CC13B8E03A950B6FC6132" authorityName="Miller" authorityYear="1940" box="[564,676,1797,1821]" class="Insecta" family="Reduviidae" genus="Enoplocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="species" species="perakensis">perakensis</taxonomicName>
|
||
[Miller’s handwriting] \ gen. nov. sp. n. [Miller’s handwriting] \ det. N.C.E.Miller 193 [printed] 8 [Miller’s handwriting]”; “
|
||
<collectionCode id="ED24AE48CF2CC13B8CA6A930B4A06152" box="[145,248,1893,1917]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="4" pageNumber="373" type="Museum">NHMUK</collectionCode>
|
||
013587199”. Pinned specimen, fourth antennal segments of both antennae, tarsus of left fore leg, tibiae and tarsi of left mid leg and both hind legs missing, male genitalia taken off and glued on card pinned with the specimen (
|
||
<collectionCode id="ED24AE48CF2CC13B8CCAA9B0B50B61D2" box="[253,339,2021,2045]" pageId="4" pageNumber="373">BMNH</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32F6506CF2CC13B8CBFA670B5BB6E12" box="[136,483,2085,2109]" pageId="4" pageNumber="373" type="distribution">
|
||
<paragraph id="8B8A368DCF2CC13B8CBFA670B5BB6E12" blockId="4.[136,483,2085,2109]" box="[136,483,2085,2109]" pageId="4" pageNumber="373">
|
||
<emphasis id="B941EA9FCF2CC13B8CBFA670B5426E12" bold="true" box="[136,282,2085,2109]" pageId="4" pageNumber="373">Distribution.</emphasis>
|
||
<collectingCountry id="F322761DCF2CC13B8D14A670B5DD6E12" box="[291,389,2085,2109]" name="Malaysia" pageId="4" pageNumber="373">Malaysia</collectingCountry>
|
||
(
|
||
<collectingRegion id="49F1F86FCF2CC13B8DA2A670B5806E12" box="[405,472,2085,2109]" country="Malaysia" name="Perak" pageId="4" pageNumber="373">Perak</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32F6506CF2CC13B8F74AAB0B00460D2" pageId="4" pageNumber="373" type="discussion">
|
||
<paragraph id="8B8A368DCF2CC13B8F74AAB0B0BD6052" blockId="4.[834,1476,1253,1661]" pageId="4" pageNumber="373">
|
||
<emphasis id="B941EA9FCF2CC13B8F74AAB0B7E862D2" bold="true" box="[835,944,1253,1277]" pageId="4" pageNumber="373">Remarks.</emphasis>
|
||
<taxonomicName id="4C354D0ECF2CC13B8F83AAB0B09E62D2" authorityName="Miller" authorityYear="1940" box="[948,1222,1253,1277]" class="Insecta" family="Reduviidae" genus="Enoplocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="species" species="perakensis">
|
||
<emphasis id="B941EA9FCF2CC13B8F83AAB0B09E62D2" box="[948,1222,1253,1277]" italics="true" pageId="4" pageNumber="373">Enoplocephala perakensis</emphasis>
|
||
</taxonomicName>
|
||
was described by
|
||
<bibRefCitation id="EFA44B7CCF2CC13B8948AAB0B7D36332" author="Miller NCE" pageId="4" pageNumber="373" pagination="1 - 115" refId="ref7497" refString="Miller NCE (1940) New genera and species of Malaysian Reduviidae (Hemiptera: Reduviidae). Canadian Journal of Arthropod (Part I). Journal of the Federated Malay States Museums, 18: Identification, 26: 1 - 115. doi: 10.3752 / cjai. 2014.26." type="journal article" year="1940">Miller (1940)</bibRefCitation>
|
||
based on the
|
||
<typeStatus id="548E882FCF2CC13B8805AB50B0CB6332" box="[1074,1171,1285,1309]" pageId="4" pageNumber="373" type="holotype">holotype</typeStatus>
|
||
from Distant’s collection. After our examination of the
|
||
<typeStatus id="548E882FCF2CC13B88BAAB70B0B56312" box="[1165,1261,1317,1341]" pageId="4" pageNumber="373" type="holotype">holotype</typeStatus>
|
||
, it can be declared that Miller’s illustrations (see
|
||
<bibRefCitation id="EFA44B7CCF2CC13B88ADAB10B1396372" author="Miller NCE" box="[1178,1377,1349,1373]" pageId="4" pageNumber="373" pagination="1 - 115" refId="ref7497" refString="Miller NCE (1940) New genera and species of Malaysian Reduviidae (Hemiptera: Reduviidae). Canadian Journal of Arthropod (Part I). Journal of the Federated Malay States Museums, 18: Identification, 26: 1 - 115. doi: 10.3752 / cjai. 2014.26." type="journal article" year="1940">Miller, 1940: 441</bibRefCitation>
|
||
) are not completely accurate, especially for the blackish patches on the fore wing. The “dissected” male genitalia of the specimen are glued on a card, pinned with the specimen. Morphological information on the male genitalia of this species is obtained from our observation on the specimen, and Miller’s illustration, thus the phallus of this species remains unknown. Data of the specimen labels (
|
||
<figureCitation id="130E2A08CF2CC13B8978A870B1C96012" box="[1359,1425,1573,1597]" captionStart="Figs" captionStartId="1.[114,159,1015,1037]" captionTargetBox="[97,1475,170,1066]" captionTargetId="figure-427@1.[97,1475,170,1048]" captionTargetPageId="1" captionText="Figs. 1–4. Enoplocephala perakensis Miller, 1940, holotype, male, habitus. 1, dorsal view; 2, lateral view; 3, ventral view; 4, labels. Scale bar of 1–3 = 5.00 mm." figureDoi="http://doi.org/10.5281/zenodo.4577304" httpUri="https://zenodo.org/record/4577304/files/figure.png" pageId="4" pageNumber="373">Fig. 4</figureCitation>
|
||
) are scarce, and no additional specimen is available, resulting in our limited knowledge of this species.
|
||
</paragraph>
|
||
<paragraph id="8B8A368DCF2CC13B8F74A8F3B00460D2" blockId="4.[835,1474,1701,1789]" pageId="4" pageNumber="373">
|
||
In his worldwide catalogue of
|
||
<taxonomicName id="4C354D0ECF2CC13B889DA8F0B1746092" box="[1194,1324,1701,1725]" class="Insecta" family="Reduviidae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="family">Reduviidae</taxonomicName>
|
||
, Maldonado- Capriles (1990) documented the
|
||
<typeStatus id="548E882FCF2CC13B8897A893B09660F1" box="[1184,1230,1734,1758]" pageId="4" pageNumber="373">type</typeStatus>
|
||
locality of this species incorrectly as “
|
||
<collectingRegion id="49F1F86FCF2CC13B8FDFA8B0B01660D2" box="[1000,1102,1765,1789]" country="Malaysia" name="Selangor" pageId="4" pageNumber="373">Selangor</collectingRegion>
|
||
”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |