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<document ID-DOI="10.11646/phytotaxa.579.4.6" ID-GBIF-Dataset="4d923433-070a-4feb-8ec7-b1ccb4566d70" ID-ISSN="1179-3163" ID-Zenodo-Dep="7564013" checkinTime="1674543614717" checkinUser="plazi" docAuthor="Pérez-Moreno, Jesús, Martínez-Reyes, Magdalena, Martínezgonzález, César Ramiro, Ramírez-Carbajal, Elisette, Carrera-Martínez, Anaitzi, Fuente, Javier Isaac De La, Olvera-Noriega, Joan Windhoek &amp; Ayala-Vásquez, Olivia" docDate="2023" docId="039C878AA63A723B76E0FF31FC5927EA" docLanguage="en" docName="phytotaxa.579.4.6.pdf" docOrigin="Phytotaxa 579 (4)" docSource="http://dx.doi.org/10.11646/phytotaxa.579.4.6" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.5:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="5" docTitle="Chroogomphus conacytiensis Ayala-Vasquez, Perez-Moreno, Ramirez-Carbajal &amp; Gonzalez-Martinez 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="294" masterDocId="FFA5FFF2A63E723E7668FFC5FF992447" masterDocTitle="Two new species of Chroogomphus (Gomphidiaceae, Boletales) with biocultural importance in the Tlahuica-Pjiekakjoo culture from Central Mexico" masterLastPageNumber="298" masterPageNumber="289" pageNumber="293" updateTime="1674654249919" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title>Two new species of Chroogomphus (Gomphidiaceae, Boletales) with biocultural importance in the Tlahuica-Pjiekakjoo culture from Central Mexico</mods:title>
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<mods:namePart>Pérez-Moreno, Jesús</mods:namePart>
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<mods:namePart>Martínez-Reyes, Magdalena</mods:namePart>
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<mods:affiliation>Colegio de Posgraduados, Campus Montecillo, Edafología, Texcoco, México &amp; martinezreyes 2012 @ gmail. com; https: // orcid. org / 0000 - 0003 - 2352 - 917 X</mods:affiliation>
<mods:nameIdentifier type="email">martinezreyes2012@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Martínezgonzález, César Ramiro</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-0256-0840</mods:nameIdentifier>
<mods:affiliation>Tecnológico Nacional de México. Instituto Tecnológico de Ciudad Victoria. Blvd. Emilio Portes Gil # 1301 Pte. CP 87010, Ciudad Victoria, Tamaulipas, México &amp; cesar. ramiro. mg @ gmail. com; https: // orcid. org / 0000 - 0002 - 0256 - 0840</mods:affiliation>
<mods:nameIdentifier type="email">cesar.ramiro.mg@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ramírez-Carbajal, Elisette</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-5421-589X</mods:nameIdentifier>
<mods:affiliation>Colegio de Posgraduados, Campus Montecillo, Edafología, Texcoco, México &amp; lisrc @ hotmail. com; https: // orcid. org / 0000 - 0001 - 5421 - 589 X</mods:affiliation>
<mods:nameIdentifier type="email">lisrc@hotmail.com</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Carrera-Martínez, Anaitzi</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-8184-0217</mods:nameIdentifier>
<mods:affiliation>Colegio de Posgraduados, Campus Montecillo, Edafología, Texcoco, México &amp; anaitzicm @ gmail. com; https: // orcid. org / 0000 - 0002 - 8184 - 0217</mods:affiliation>
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<mods:namePart>Fuente, Javier Isaac De La</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-4667-1574</mods:nameIdentifier>
<mods:affiliation>Colegio de Posgraduados, Campus Montecillo, Edafología, Texcoco, México &amp; jdelafuenteitcv @ gmail. com; https: // orcid. org / 0000 - 0003 - 4667 - 1574</mods:affiliation>
<mods:nameIdentifier type="email">jdelafuenteitcv@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Olvera-Noriega, Joan Windhoek</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-0892-684X</mods:nameIdentifier>
<mods:affiliation>Colegio de Posgraduados, Campus Montecillo, Edafología, Texcoco, México &amp; joan 8913 @ hotmail. com; https: // orcid. org / 0000 - 0002 - 0892 - 684 X</mods:affiliation>
<mods:nameIdentifier type="email">joan8913@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ayala-Vásquez, Olivia</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-8970-9571</mods:nameIdentifier>
<mods:affiliation>Colegio de Posgraduados, Campus Montecillo, Edafología, Texcoco, México &amp; yootspooj @ gmail. com; https: // orcid. org / 0000 - 0002 - 8970 - 9571</mods:affiliation>
<mods:nameIdentifier type="email">yootspooj@gmail.com</mods:nameIdentifier>
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<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
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<mods:date>2023</mods:date>
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<mods:number>2023-01-24</mods:number>
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<mods:detail type="volume">
<mods:number>579</mods:number>
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<mods:detail type="issue">
<mods:number>4</mods:number>
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<mods:start>289</mods:start>
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<mods:url>http://dx.doi.org/10.11646/phytotaxa.579.4.6</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.579.4.6</mods:identifier>
<mods:identifier type="GBIF-Dataset">4d923433-070a-4feb-8ec7-b1ccb4566d70</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7564013</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7568725" ID-Zenodo-Dep="7568725" LSID="urn:lsid:plazi:treatment:039C878AA63A723B76E0FF31FC5927EA" httpUri="http://treatment.plazi.org/id/039C878AA63A723B76E0FF31FC5927EA" lastPageId="5" lastPageNumber="294" pageId="4" pageNumber="293">
<subSubSection pageId="4" pageNumber="293" type="nomenclature">
<paragraph blockId="4.[136,1452,244,342]" pageId="4" pageNumber="293">
<taxonomicName authority="Ayala-Vasquez, Perez-Moreno, Ramirez-Carbajal &amp; Gonzalez-Martinez" authorityName="Ayala-Vasquez, Perez-Moreno, Ramirez-Carbajal &amp; Gonzalez-Martinez" authorityYear="2023" box="[136,1295,244,271]" class="Agaricomycetes" family="Gomphidiaceae" genus="Chroogomphus" kingdom="Fungi" order="Boletales" pageId="4" pageNumber="293" phylum="Basidiomycota" rank="species" species="conacytiensis" status="sp. nov.">
<emphasis bold="true" box="[136,476,244,271]" italics="true" pageId="4" pageNumber="293">Chroogomphus conacytiensis</emphasis>
Ayala-Vásquez, Pérez-Moreno, Ramírez-Carbajal &amp; González-Martínez
</taxonomicName>
,
<emphasis box="[1306,1391,245,270]" italics="true" pageId="4" pageNumber="293">
<taxonomicNameLabel box="[1306,1391,245,270]" pageId="4" pageNumber="293" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation captionStart="FIGURE 2" captionStartId="4.[136,229,1893,1915]" captionTargetBox="[493,1095,1015,1867]" captionTargetId="figure-272@4.[493,1095,1015,1867]" captionTargetPageId="4" captionText="FIGURE 2. Chroogomphus conacytiensis (MEXU-HO 30431, holotype, MEXU-HO- 30452 paratype). A. General view of basidiomata. B. Basidioma. C. Stipe surface and partial velum. D. Basidiospores. E. Basidium and cystidium. F. Basidia. G. Lamellar trama. HI. Two different cystidia forms. JL Pileipellis. Scale bars: 10 mm (AC); 10µm (D, E, F, H, I, K, L); 100 µm (G,); 25 µm (J)." figureDoi="http://doi.org/10.5281/zenodo.7564017" httpUri="https://zenodo.org/record/7564017/files/figure.png" pageId="4" pageNumber="293">Fig. 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection box="[136,635,320,342]" pageId="4" pageNumber="293" type="multiple">
<paragraph blockId="4.[136,1452,244,342]" box="[136,635,320,342]" pageId="4" pageNumber="293">
Mycobank no. 845567, GenBank:
<accessionNumber box="[468,570,320,342]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/OP580489" pageId="4" pageNumber="293">OP580489</accessionNumber>
(ITS).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="293" type="diagnosis">
<paragraph blockId="4.[136,1452,388,991]" pageId="4" pageNumber="293">
<emphasis bold="true" box="[136,261,388,414]" pageId="4" pageNumber="293">Diagnosis:</emphasis>
—Basidiomata small to medium, pileus surface furfuraceous to velvety, brown, brown-yellowish, brownolivaceus, grey to greyish, sterile margin, when touch reddish-brown to vinaceous. Basidiospores 1316(19) × 67 (8) μm,
<collectionCode box="[241,262,460,486]" country="Ecuador" lsid="urn:lsid:biocol.org:col:14273" name="Universidad Central" pageId="4" pageNumber="293" type="Herbarium">Q</collectionCode>
=2.2, (n=34), ellipsoid, golden yellow to yellowish-yellow on KOH.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="293" type="materials_examined">
<paragraph blockId="4.[136,1452,388,991]" pageId="4" pageNumber="293">
<materialsCitation collectingDate="2021-09-09" collectionCode="MEXU-HO" collectorName="Martinez-Reyes M. &amp; Ayala-Vasquez O &amp; Perez-Moreno J." country="Mexico" elevation="2989" location="Lomas de Teocaltzingo Town" municipality="Ocuilan Municipaly" pageId="4" pageNumber="293" specimenCount="1" typeStatus="holotype">
<emphasis bold="true" box="[189,257,496,522]" pageId="4" pageNumber="293">Type:</emphasis>
<collectingCountry box="[286,398,496,522]" name="Mexico" pageId="4" pageNumber="293">MEXICO</collectingCountry>
, STATE OF MEXICO:
<collectingMunicipality box="[675,903,496,522]" pageId="4" pageNumber="293">Ocuilan Municipaly</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:039C878AA63A723B76E0FF31FC5927EA:8EEA6047A63A723A75FCFE34FB71264C" box="[916,1256,496,523]" country="Mexico" municipality="Ocuilan Municipaly" name="Lomas de Teocaltzingo Town" pageId="4" pageNumber="293">Lomas de Teocaltzingo Town</location>
, mixed
<emphasis italics="true" pageId="4" pageNumber="293">QuercusPinus</emphasis>
forests, 462878N; 2102619W,
<quantity box="[545,631,532,559]" metricMagnitude="3" metricUnit="m" metricValue="2.989" pageId="4" pageNumber="293" unit="m" value="2989.0">
<elevation box="[545,631,532,559]" metricMagnitude="3" metricUnit="m" metricValue="2.989" pageId="4" pageNumber="293" unit="m" value="2989.0">2989 m</elevation>
</quantity>
,
<date box="[641,845,532,558]" pageId="4" pageNumber="293" value="2021-09-09">
<collectingDate box="[641,845,532,558]" pageId="4" pageNumber="293" value="2021-09-09">9 September 2021</collectingDate>
</date>
,
<collectorName box="[855,1074,532,559]" pageId="4" pageNumber="293">Martínez-Reyes M.</collectorName>
,
<collectorName box="[1083,1280,532,558]" pageId="4" pageNumber="293">Ayala-Vásquez O</collectorName>
,
<collectorName pageId="4" pageNumber="293">Pérez-Moreno J.</collectorName>
(
<collectionCode box="[168,302,568,594]" pageId="4" pageNumber="293">MEXU-HO</collectionCode>
30431,
<typeStatus box="[393,496,568,594]" pageId="4" pageNumber="293">holotype</typeStatus>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="294" pageId="4" pageNumber="293" type="description">
<paragraph blockId="4.[136,1452,388,991]" pageId="4" pageNumber="293">
<emphasis bold="true" box="[189,337,604,630]" pageId="4" pageNumber="293">Description:</emphasis>
<taxonomicName box="[365,435,604,630]" class="Magnoliopsida" family="Caricaceae" genus="Pileus" kingdom="Plantae" order="Brassicales" pageId="4" pageNumber="293" phylum="Tracheophyta" rank="genus">
<emphasis box="[365,435,604,630]" italics="true" pageId="4" pageNumber="293">Pileus</emphasis>
</taxonomicName>
<quantity box="[447,572,604,631]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" metricValueMax="3.5" metricValueMin="2.3" pageId="4" pageNumber="293" unit="mm" value="29.0" valueMax="35.0" valueMin="23.0">2335 mm</quantity>
diameter, broadly convex somewhat umbonate when young, hemispherical towards at maturity, pale yellowish-brown (5D85E8), brownish-orange (5C35D3), grey (30D130E1), greyishbrown (5D3), brownish-grey (6D26
<collectionCode box="[558,575,676,702]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="4" pageNumber="293" type="Herbarium">F</collectionCode>
2) to dark brown (6
<collectionCode box="[799,817,676,702]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="4" pageNumber="293" type="Herbarium">F</collectionCode>
8), when touch reddish-brown (8D68E6) to vinaceous, humid surface, viscid when wet, smooth, shining when dry, margin enrolled when young, margin with curtain remnants at mature.
<emphasis box="[253,411,748,774]" italics="true" pageId="4" pageNumber="293">Hymenophore</emphasis>
Lamellae, subdecurrent, distant, orange-greyish (5B45B5) to light-brown (6D46E4), when touch reddish-brown (8D68E6) to vinaceous, lamellae
<quantity box="[757,863,784,811]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="4" pageNumber="293" unit="mm" value="8.5" valueMax="10.0" valueMin="7.0">710 mm</quantity>
width, slightly serrate, irregular lamellulae common, attenuate.
<emphasis box="[252,339,820,846]" italics="true" pageId="4" pageNumber="293">Context</emphasis>
pileus
<quantity box="[423,523,820,846]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="4" pageNumber="293" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">810mm</quantity>
thick, light orange (5
<collectionCode box="[768,791,820,847]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="4" pageNumber="293" type="Herbarium">A</collectionCode>
4), pale orange (5
<collectionCode box="[989,1012,820,847]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="4" pageNumber="293" type="Herbarium">A</collectionCode>
5) to salmon, basal context of the stipe orange (5
<collectionCode box="[243,266,856,883]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="4" pageNumber="293" type="Herbarium">A</collectionCode>
6), immutable to court.
<emphasis box="[534,590,856,882]" italics="true" pageId="4" pageNumber="293">Stipe</emphasis>
23 70 ×
<quantity box="[709,828,856,883]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.4" metricValueMin="0.6" pageId="4" pageNumber="293" unit="mm" value="10.0" valueMax="14.0" valueMin="6.0">6 14 mm</quantity>
, cylindrical, orange-yellow (4
<collectionCode box="[1170,1193,856,882]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="4" pageNumber="293" type="Herbarium">A</collectionCode>
8), orange (5
<collectionCode box="[1335,1358,856,883]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="4" pageNumber="293" type="Herbarium">A</collectionCode>
4), paleorange (5
<collectionCode box="[244,266,892,919]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="4" pageNumber="293" type="Herbarium">A</collectionCode>
35
<collectionCode box="[309,331,892,919]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="4" pageNumber="293" type="Herbarium">A</collectionCode>
4), when cut it becomes greyish-vinaceous (12E7) to deep vinaceous (12
<collectionCode box="[1181,1199,892,918]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="4" pageNumber="293" type="Herbarium">F</collectionCode>
8), sometimes slightly attenuate downwards, fibrillose surface with ring ephemeral remains very close to the apex whiting to pale pink (12
<collectionCode box="[173,196,964,990]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="4" pageNumber="293" type="Herbarium">A</collectionCode>
4).
<emphasis box="[231,437,964,990]" italics="true" pageId="4" pageNumber="293">Chemical reaction</emphasis>
when applying KOH, all basidiomata turn vinaceous.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7564017" ID-Zenodo-Dep="7564017" httpUri="https://zenodo.org/record/7564017/files/figure.png" pageId="4" pageNumber="293" startId="4.[136,229,1893,1915]" targetBox="[493,1095,1015,1867]" targetPageId="4">
<paragraph blockId="4.[136,1453,1893,1987]" pageId="4" pageNumber="293">
<emphasis bold="true" box="[136,248,1893,1915]" pageId="4" pageNumber="293">FIGURE 2</emphasis>
.
<taxonomicName authorityName="Ayala-Vasquez, Perez-Moreno, Ramirez-Carbajal &amp; Gonzalez-Martinez" authorityYear="2023" box="[260,542,1893,1914]" class="Agaricomycetes" family="Gomphidiaceae" genus="Chroogomphus" kingdom="Fungi" order="Boletales" pageId="4" pageNumber="293" phylum="Basidiomycota" rank="species" species="conacytiensis">
<emphasis box="[260,542,1893,1914]" italics="true" pageId="4" pageNumber="293">Chroogomphus conacytiensis</emphasis>
</taxonomicName>
(MEXU-HO 30431, holotype, MEXU-HO- 30452 paratype). A. General view of basidiomata. B. Basidioma. C. Stipe surface and partial velum. D. Basidiospores. E. Basidium and cystidium. F. Basidia. G. Lamellar trama. HI. Two different cystidia forms. JL Pileipellis. Scale bars: 10 mm (AC); 10µm (D, E, F, H, I, K, L); 100 µm (G,); 25 µm (J).
</paragraph>
</caption>
<paragraph blockId="5.[136,1452,158,941]" pageId="5" pageNumber="294">
<emphasis box="[189,347,158,184]" italics="true" pageId="5" pageNumber="294">Basidiospores</emphasis>
(13)1718 20(23) × 56 (7) µm, (
<collectionCode box="[782,800,158,184]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="5" pageNumber="294" type="Herbarium">L</collectionCode>
= 17.89,
<collectionCode box="[903,929,159,185]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="5" pageNumber="294" type="Herbarium">W</collectionCode>
= 6.1,
<collectionCode box="[1006,1026,158,184]" country="Ecuador" lsid="urn:lsid:biocol.org:col:14273" name="Universidad Central" pageId="5" pageNumber="294" type="Herbarium">Q</collectionCode>
= 2.5 2.7 (2.9),
<collectionCode box="[1232,1252,159,185]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="5" pageNumber="294" type="Herbarium">N</collectionCode>
= 65), cylindrical to elliptical, fusiform, smooth, brown-yellowish, thick-walled.
<emphasis box="[840,926,194,220]" italics="true" pageId="5" pageNumber="294">Basidia</emphasis>
(35) 375564 (70) × (7) 1014 (15) µm, clavate, hyaline to pale brown in KOH, brown-chocolate, in Melzers reagent, with long sterigmata (5) 78× 23 µm.
<taxonomicName authorityName="Hubner" authorityYear="1819" box="[188,281,266,292]" class="Insecta" family="Geometridae" genus="Cystidia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="294" phylum="Arthropoda" rank="genus">
<emphasis box="[188,281,266,292]" italics="true" pageId="5" pageNumber="294">Cystidia</emphasis>
</taxonomicName>
(100) 120150 (150) × (15) (12)1721(22) µm, fusoid-capitate, subclavate to subfusiform, thickwalled (≤3), hyaline to brown in KOH, brown-yellowish in Melzers reagent.
<emphasis box="[1006,1182,302,328]" italics="true" pageId="5" pageNumber="294">Lamellar trama</emphasis>
made up of hyphae 46 µm in width, cylindrical, hyaline to brownish in KOH. Pileipellis formed by ixotrichoderm, with terminal cells (14) 254560 (70) × (8) 101520 (30) µm, clavate, fusiform, to ventricose-fusoid, brown- vinaceous, to chocolate pigment but soon dissolved in KOH, with gelatinous wall that disappears with Melzers reagent, amyloid.
<emphasis box="[1324,1452,410,436]" italics="true" pageId="5" pageNumber="294">Stipitipellis</emphasis>
composed of light amyloid, vertical hyphae (4)512 µm in diameter, thick walled (≤2). With clamp connections.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="294" type="etymology">
<paragraph blockId="5.[136,1452,158,941]" pageId="5" pageNumber="294">
<emphasis bold="true" box="[189,328,482,508]" pageId="5" pageNumber="294">Etymology:</emphasis>
—The name is in honor of the Mexican Council of Science and Technology (or CONACYT for its abbreviation in Spanish) which is the public institution responsible for promoting the advancement of scientific research, innovation and technological development, including that of food security and mycocultural heritage, of the country during the last half-century.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="294" type="biology_ecology">
<paragraph blockId="5.[136,1452,158,941]" pageId="5" pageNumber="294">
<emphasis bold="true" box="[189,578,626,652]" pageId="5" pageNumber="294">Habitat, habit, and distribution:</emphasis>
—Solitary to scattered growing under mixed forest to
<emphasis box="[1195,1362,627,653]" italics="true" pageId="5" pageNumber="294">Quercus-Pinus</emphasis>
forests, forming ectomycorrhizal associations with
<taxonomicName box="[617,847,662,688]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="294" phylum="Tracheophyta" rank="species" species="pseudostrobus">
<emphasis box="[617,847,662,688]" italics="true" pageId="5" pageNumber="294">Pinus pseudostrobus</emphasis>
</taxonomicName>
and
<taxonomicName box="[899,1033,662,688]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="294" phylum="Tracheophyta" rank="species" species="hartwegii">
<emphasis box="[899,1033,662,688]" italics="true" pageId="5" pageNumber="294">P. hartwegii</emphasis>
</taxonomicName>
, during July to September at altitudes between
<quantity box="[236,416,698,725]" metricMagnitude="3" metricUnit="m" metricValue="2.95" metricValueMax="3.0" metricValueMin="2.9" pageId="5" pageNumber="294" unit="m" value="2950.0" valueMax="3000.0" valueMin="2900.0">2900 to 3000 m</quantity>
. Only known currently from Central and Southwestern
<collectingRegion box="[1039,1128,698,724]" country="Mexico" name="Mexico" pageId="5" pageNumber="294">Mexico</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="294" type="materials_examined">
<paragraph blockId="5.[136,1452,158,941]" pageId="5" pageNumber="294">
<emphasis bold="true" box="[189,448,734,760]" pageId="5" pageNumber="294">Specimens examined:</emphasis>
<collectingCountry box="[484,597,734,760]" name="Mexico" pageId="5" pageNumber="294">MEXICO</collectingCountry>
,
<collectionCode box="[610,736,734,760]" pageId="5" pageNumber="294">STATE OF</collectionCode>
<collectingCountry box="[744,859,734,760]" name="Mexico" pageId="5" pageNumber="294">MEXICO</collectingCountry>
: Ocuilan Municipaly, Lomas de Teocaltzingo Town, mixed forest, 4664720
<collectionCode box="[405,425,770,796]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="5" pageNumber="294" type="Herbarium">N</collectionCode>
; 2111033
<collectionCode box="[539,565,770,796]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="5" pageNumber="294" type="Herbarium">W</collectionCode>
,
<quantity box="[583,708,770,796]" metricMagnitude="3" metricUnit="m" metricValue="3.3899999999999997" pageId="5" pageNumber="294" unit="m" value="3390.0">3390 masl</quantity>
,
<date box="[725,945,770,796]" pageId="5" pageNumber="294" value="2021-09-09">9 September 2021</date>
, Mycoredes (MEXU-HO 30426
<typeStatus box="[1338,1440,770,796]" pageId="5" pageNumber="294">paratype</typeStatus>
);
<collectingRegion box="[136,260,806,832]" country="Mexico" name="Oaxaca" pageId="5" pageNumber="294">OAXACA</collectingRegion>
: Santa María Tlahuitoltepec Municipaly, Bridge to Zempoaltepetl Town,
<taxonomicName box="[1078,1255,806,832]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="294" phylum="Tracheophyta" rank="species" species="hartwegii">
<emphasis box="[1078,1255,806,832]" italics="true" pageId="5" pageNumber="294">Pinus hartwegii</emphasis>
</taxonomicName>
forest, 462878
<collectionCode box="[1427,1448,806,832]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="5" pageNumber="294" type="Herbarium">N</collectionCode>
; 2102619
<collectionCode box="[234,261,842,868]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="5" pageNumber="294" type="Herbarium">W</collectionCode>
,
<quantity box="[273,394,842,869]" metricMagnitude="3" metricUnit="m" metricValue="3.45" pageId="5" pageNumber="294" unit="m" value="3450.0">3450 masl</quantity>
,
<date box="[406,557,842,869]" pageId="5" pageNumber="294" value="2022-07-15">15 July 2022</date>
, Ayala-Vásquez
<collectionCode box="[746,768,842,868]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="5" pageNumber="294" type="Herbarium">O</collectionCode>
, Martínez-Gonzalez CR., Martínez-Reyes
<collectionCode box="[1253,1283,843,869]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="5" pageNumber="294" type="Herbarium">M</collectionCode>
., (MEXU-HO-30452);
<collectionCode box="[232,359,878,904]" pageId="5" pageNumber="294">STATE OF</collectionCode>
<collectingRegion box="[367,522,878,904]" country="Mexico" name="Tlaxcala" pageId="5" pageNumber="294">TLAXCALA</collectingRegion>
: Nanacamilpa Municipaly, San Felipe
<collectingRegion box="[965,1055,878,904]" country="Mexico" name="Hidalgo" pageId="5" pageNumber="294">Hidalgo</collectingRegion>
Town, El Astillero,
<quantity box="[1289,1410,878,904]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="5" pageNumber="294" unit="m" value="2900.0">2900 masl</quantity>
,
<date pageId="5" pageNumber="294" value="2021-09-15">15 September 2021</date>
, Martínez-Reyes
<collectionCode box="[519,549,915,941]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="5" pageNumber="294" type="Herbarium">M</collectionCode>
., Ayala-Vásquez
<collectionCode box="[740,762,914,940]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="5" pageNumber="294" type="Herbarium">O</collectionCode>
, (MEXU 30451).
</paragraph>
</subSubSection>
</treatment>
</document>