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<document id="303F948B15B49F45C049FBB179F6DC89" ID-CLB-Dataset="22458" ID-DOI="10.5852/ejt.2020.622" ID-GBIF-Dataset="9cb3ea96-0ee4-40b4-8006-19b5fed80b46" ID-Zenodo-Dep="3777282" ID-ZooBank="9CD3B2CD-F072-4994-8CA9-24145D343401" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1588217079841" checkinUser="valdenar" docAuthor="Darby, Michael" docDate="2020" docId="039987A1386DFF4E2B2BFEE5FE40FBEA" docLanguage="en" docName="ejt-622_darby.pdf.imf" docOrigin="European Journal of Taxonomy 622" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cissidium subfaustum Darby 2020, sp. nov." docType="treatment" docUuid="E204C76D-EAD4-4C6E-BDCA-DDDED90DD366" docUuidSource="ZooBank" docVersion="7" lastPageNumber="129" masterDocId="FFA0FFD93812FFCE2949FFEEFF9CFF98" masterDocTitle="A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species" masterLastPageNumber="188" masterPageNumber="1" pageNumber="128" updateTime="1698838121940" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="AEA3EE3C016EC9915EF2BF6C5584BA46" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9F47EF92A1661D40A63825953D6C39BF">
<mods:title id="B3FECED6C165010C133BE4B2262B5627">A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species</mods:title>
</mods:titleInfo>
<mods:name id="C324800180D45580E885AC1718F6B144" type="personal">
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<mods:roleTerm id="111B49202B046E205EF5880EB2328243">Author</mods:roleTerm>
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<mods:namePart id="0025CC90A28296142B9713CB5D6A7B43">Darby, Michael</mods:namePart>
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<mods:title id="E7E6154DBD34DD5488EE68C6C24797DB">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="172D66E0032249452D68B7A7F216043B">
<mods:date id="492642912FCDD1E8EC116373B5ADFCC3">2020</mods:date>
<mods:detail id="7E7FF20799D23639581D4C50327A4934" type="pubDate">
<mods:number id="CFDA4CD3F45B07CCEE67D007D5C37FFB">2020-04-02</mods:number>
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<mods:number id="00090CD8F6D75965E17A6A1976CD34BB">622</mods:number>
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<mods:identifier id="ACB1C5F4EC9859B821FCA2C30CC7719A" type="DOI">10.5852/ejt.2020.622</mods:identifier>
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<treatment id="039987A1386DFF4E2B2BFEE5FE40FBEA" ID-DOI="http://doi.org/10.5281/zenodo.3795642" ID-GBIF-Taxon="163877818" ID-Zenodo-Dep="3795642" LSID="urn:lsid:zoobank.org:act:E204C76D-EAD4-4C6E-BDCA-DDDED90DD366" httpUri="http://treatment.plazi.org/id/039987A1386DFF4E2B2BFEE5FE40FBEA" lastPageId="128" lastPageNumber="129" pageId="127" pageNumber="128">
<subSubSection id="C32A653C386DFFB12B2BFEE5FB42FED0" pageId="127" pageNumber="128" type="nomenclature">
<paragraph id="8B8F36B7386DFFB12B2BFEE5FC4EFEBE" blockId="127.[341,1246,267,364]" box="[610,978,267,294]" pageId="127" pageNumber="128">
<heading id="D0C781DB386DFFB12B2BFEE5FC4EFEBE" box="[610,978,267,294]" centered="true" fontSize="11" level="2" pageId="127" pageNumber="128" reason="2">
<taxonomicName id="4C304D34386DFFB12B2BFEE5FCECFEBE" ID-CoL="8XXRT" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[610,880,267,294]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="127" pageNumber="128" phylum="Arthropoda" rank="species" species="subfaustum" status="sp. nov.">
<emphasis id="B944EAA5386DFFB12B2BFEE5FCECFEBE" bold="true" box="[610,880,267,294]" italics="true" pageId="127" pageNumber="128">Cissidium subfaustum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE386DFFB12A3EFEE2FC4EFEBE" box="[887,978,268,294]" pageId="127" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8B8F36B7386DFFB1281CFEC0FB42FED0" blockId="127.[341,1246,267,364]" box="[341,1246,302,328]" pageId="127" pageNumber="128">
<uri id="FFA13AB5386DFFB1281CFEC0FB42FED0" box="[341,1246,302,328]" pageId="127" pageNumber="128">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
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<subSubSection id="C32A653C386DFFB12BB9FEBCFCDFFEF4" box="[752,835,338,364]" pageId="127" pageNumber="128" type="description">
<paragraph id="8B8F36B7386DFFB12BB9FEBCFCDFFEF4" blockId="127.[341,1246,267,364]" box="[752,835,338,364]" pageId="127" pageNumber="128">
<figureCitation id="130B2A32386DFFB12BB9FEBCFCDFFEF4" box="[752,835,338,364]" captionStart="Fig" captionStartId="127.[189,232,1964,1990]" captionTargetBox="[205,1381,980,1921]" captionTargetId="figure@127.[204,1382,977,1922]" captionTargetPageId="127" captionText="Fig. 81. Cissidium subfaustum sp. nov. A. Habitus. B. Pronotum, × 410. C. Mesoventrite showing median process of collar, mid-keel and keel, × 1280." figureDoi="http://doi.org/10.5281/zenodo.3777446" httpUri="https://zenodo.org/record/3777446/files/figure.png" pageId="127" pageNumber="128">Fig. 81</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C386DFFB129F4FE7DFBA3FE44" box="[189,1087,403,476]" pageId="127" pageNumber="128" type="etymology">
<paragraph id="8B8F36B7386DFFB129F4FE7DFED8FE35" blockId="127.[189,324,403,429]" box="[189,324,403,429]" pageId="127" pageNumber="128">
<heading id="D0C781DB386DFFB129F4FE7DFED8FE35" bold="true" box="[189,324,403,429]" fontSize="11" level="3" pageId="127" pageNumber="128" reason="3">
<emphasis id="B944EAA5386DFFB129F4FE7DFED8FE35" bold="true" box="[189,324,403,429]" pageId="127" pageNumber="128">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B7386DFFB129F4FE2CFBA3FE44" blockId="127.[189,1087,450,476]" box="[189,1087,450,476]" pageId="127" pageNumber="128">
Johnson ms name referring to similarity of the species to
<taxonomicName id="4C304D34386DFFB12A17FE2CFC41FE44" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[862,989,450,476]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="127" pageNumber="128" phylum="Arthropoda" rank="species" species="faustum" status="sp. nov.">
<emphasis id="B944EAA5386DFFB12A17FE2CFC41FE44" box="[862,989,450,476]" italics="true" pageId="127" pageNumber="128">C. faustum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE386DFFB12AADFE2CFBA3FE44" box="[996,1087,450,476]" pageId="127" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C386DFFB129F4FDEDFCE2FD0B" pageId="127" pageNumber="128" type="materials_examined">
<paragraph id="8B8F36B7386DFFB129F4FDEDFE36FD85" blockId="127.[189,426,515,541]" box="[189,426,515,541]" pageId="127" pageNumber="128">
<heading id="D0C781DB386DFFB129F4FDEDFE36FD85" bold="true" box="[189,426,515,541]" fontSize="11" level="3" pageId="127" pageNumber="128" reason="3">
<emphasis id="B944EAA5386DFFB129F4FDEDFE36FD85" bold="true" box="[189,426,515,541]" pageId="127" pageNumber="128">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B583CEA386DFFB129A3FDDCFCE2FD0B" ID-GBIF-Occurrence="2609493526" collectingDate="1959-03-09" collectionCode="FMNH" collectorName="Berlese &amp; H. S. Dybas" country="Panama" elevation="1494" location="Nueva California" municipality="Chirique Prov." pageId="127" pageNumber="128" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<paragraph id="8B8F36B7386DFFB129A3FDDCFEC0FDD4" blockId="127.[189,1399,562,659]" box="[234,348,562,588]" pageId="127" pageNumber="128">
<emphasis id="B944EAA5386DFFB129A3FDDCFEC0FDD4" bold="true" box="[234,348,562,588]" pageId="127" pageNumber="128">
<typeStatus id="548B8815386DFFB129A3FDDCFEC0FDD4" box="[234,348,562,588]" pageId="127" pageNumber="128" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B8F36B7386DFFB129F4FDBBFCE2FD0B" blockId="127.[189,1399,562,659]" pageId="127" pageNumber="128">
<collectingCountry id="F3277627386DFFB129F4FDBBFEA4FDF7" box="[189,312,597,623]" name="Panama" pageId="127" pageNumber="128">PANAMA</collectingCountry>
•
<specimenCount id="9D36FD3E386DFFB12819FDBBFEFBFDF7" box="[336,359,597,623]" pageId="127" pageNumber="128" type="female">♀</specimenCount>
;
<locationDeviation id="9A3D2F95386DFFB1283DFDBBFDE3FDF7" box="[372,639,597,623]" location="Finca Palo Santo" pageId="127" pageNumber="128">W of Finca Palo Santo</locationDeviation>
, nr.
<location id="8EEF606C386DFFB12BE6FDBBFCE0FDF7" LSID="urn:lsid:plazi:treatment:039987A1386DFF4E2B2BFEE5FE40FBEA:8EEF606C386DFFB12BE6FDBBFCE0FDF7" box="[687,892,597,623]" country="Panama" municipality="Chirique Prov." name="Nueva California" pageId="127" pageNumber="128">Nueva California</location>
,
<collectingMunicipality id="6BEBACCD386DFFB12ACEFDBBFBAAFDF7" box="[903,1078,597,623]" pageId="127" pageNumber="128">Chirique Prov.</collectingMunicipality>
;
<elevation id="001DD184386DFFB12D08FDBBFB48FDF7" box="[1089,1236,597,623]" metricMagnitude="3" metricUnit="m" metricValue="1.49352" pageId="127" pageNumber="128" unit="ft" value="4900.0">
<quantity id="4CC89B52386DFFB12D08FDBBFB08FDF7" box="[1089,1172,597,623]" metricMagnitude="3" metricUnit="m" metricValue="1.49352" pageId="127" pageNumber="128" unit="ft" value="4900.0">4900 ft</quantity>
a.s.l.
</elevation>
;
<date id="FF8E1077386DFFB12DA8FDBBFAEEFDE8" box="[1249,1394,597,624]" pageId="127" pageNumber="128" value="1959-03-09">
<collectingDate id="EFCAE99F386DFFB12DA8FDBBFAEEFDE8" box="[1249,1394,597,624]" pageId="127" pageNumber="128" value="1959-03-09">9 Mar. 1959</collectingDate>
</date>
;
<collectorName id="26C55361386DFFB129F4FD96FE89FD0A" box="[189,277,632,658]" pageId="127" pageNumber="128">Berlese</collectorName>
floor litter in damp ravine;
<collectorName id="26C55361386DFFB12B10FD96FD7CFD0A" box="[601,736,632,658]" pageId="127" pageNumber="128">H.S. Dybas</collectorName>
leg.;
<collectionCode id="ED21AE72386DFFB12A6BFD97FCE7FD0B" box="[802,891,633,659]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="127" pageNumber="128" type="Museum">FMNH</collectionCode>
.
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="C32A653C386DFF4E29F4FD57FCF1FC42" lastPageId="128" lastPageNumber="129" pageId="127" pageNumber="128" type="description">
<paragraph id="8B8F36B7386DFFB129F4FD57FED2FD4B" blockId="127.[189,334,697,723]" box="[189,334,697,723]" pageId="127" pageNumber="128">
<heading id="D0C781DB386DFFB129F4FD57FED2FD4B" bold="true" box="[189,334,697,723]" fontSize="11" level="3" pageId="127" pageNumber="128" reason="3">
<emphasis id="B944EAA5386DFFB129F4FD57FED2FD4B" bold="true" box="[189,334,697,723]" pageId="127" pageNumber="128">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B7386DFFB129F4FD06FD36FC9B" blockId="127.[189,682,744,771]" box="[189,682,744,771]" pageId="127" pageNumber="128">
SIZE. Habitus (
<figureCitation id="130B2A32386DFFB1282EFD06FE4FFC9A" box="[359,467,744,770]" captionStart="Fig" captionStartId="127.[189,232,1964,1990]" captionTargetBox="[205,1381,980,1921]" captionTargetId="figure@127.[204,1382,977,1922]" captionTargetPageId="127" captionText="Fig. 81. Cissidium subfaustum sp. nov. A. Habitus. B. Pronotum, × 410. C. Mesoventrite showing median process of collar, mid-keel and keel, × 1280." figureDoi="http://doi.org/10.5281/zenodo.3777446" httpUri="https://zenodo.org/record/3777446/files/figure.png" pageId="127" pageNumber="128">Fig. 81A</figureCitation>
), length
<quantity id="4CC89B52386DFFB12B73FD06FD39FC9B" box="[570,677,744,771]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" pageId="127" pageNumber="128" unit="mm" value="0.71">0.71 mm</quantity>
.
</paragraph>
<paragraph id="8B8F36B7386DFF4E29F4FCC4FEE2FEF7" blockId="127.[189,978,809,836]" lastBlockId="128.[189,1399,267,367]" lastPageId="128" lastPageNumber="129" pageId="127" pageNumber="128">
COLOUR. Dark brown, pubescence, legs and antennae dusky yellow. HEAD. With a shallow foveolate and setose linear depression behind the eyes, width across eyes
<quantity id="4CC89B523892FF4E29F4FEDEFEB7FED2" box="[189,299,304,330]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="128" pageNumber="129" unit="mm" value="0.17">0.17 mm</quantity>
; mentum as
<figureCitation id="130B2A323892FF4E288EFEDEFDBBFED2" box="[455,551,304,330]" captionStart="Fig" captionStartId="5.[189,232,1927,1953]" captionTargetBox="[232,1334,1274,1884]" captionTargetId="figure@5.[232,1334,1274,1887]" captionTargetPageId="5" captionText="Fig. 4. Menta and submenta. A. Cissidium petri Darby, 2015, × 1380. B. Cissidium tigrum Darby, 2015, × 1220. C. Cissidium amboroense Darby, 2015, × 1260. D. Cissidium losbanos sp. nov., × 1100. E. Cissidium triangulum Darby, 2019, × 650. F. Cissidium pilosellum sp. nov., × 1420." figureDoi="http://doi.org/10.5281/zenodo.3777290" httpUri="https://zenodo.org/record/3777290/files/figure.png" pageId="128" pageNumber="129">Fig. 4A</figureCitation>
; antennomeres III–XI, length
<quantity id="4CC89B523892FF4E2ADBFEDEFB9DFED2" box="[914,1025,304,330]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="128" pageNumber="129" unit="mm" value="0.34">0.34 mm</quantity>
, III–IX length
<quantity id="4CC89B523892FF4E2DFEFEDEFABAFED2" box="[1207,1318,304,331]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="128" pageNumber="129" unit="mm" value="0.15">0.15 mm</quantity>
, X–XI length
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.
</paragraph>
<caption id="DF4F663F386DFFB129F4F842FCB9F872" ID-DOI="http://doi.org/10.5281/zenodo.3777446" ID-Zenodo-Dep="3777446" httpUri="https://zenodo.org/record/3777446/files/figure.png" pageId="127" pageNumber="128" startId="127.[189,232,1964,1990]" targetBox="[205,1381,980,1921]" targetPageId="127">
<paragraph id="8B8F36B7386DFFB129F4F842FCB9F872" blockId="127.[189,1398,1964,2026]" pageId="127" pageNumber="128">
<emphasis id="B944EAA5386DFFB129F4F842FE81F85F" bold="true" box="[189,285,1964,1991]" pageId="127" pageNumber="128">Fig. 81.</emphasis>
<taxonomicName id="4C304D34386DFFB12860F842FDACF85E" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[297,560,1964,1990]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="127" pageNumber="128" phylum="Arthropoda" rank="species" species="subfaustum" status="sp. nov.">
<emphasis id="B944EAA5386DFFB12860F842FDACF85E" box="[297,560,1964,1990]" italics="true" pageId="127" pageNumber="128">Cissidium subfaustum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE386DFFB12B72F843FD06F85F" box="[571,666,1965,1991]" pageId="127" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B944EAA5386DFFB12BEFF842FD27F85E" bold="true" box="[678,699,1964,1990]" pageId="127" pageNumber="128">A</emphasis>
. Habitus.
<emphasis id="B944EAA5386DFFB12A75F843FCCCF85F" bold="true" box="[828,848,1965,1991]" pageId="127" pageNumber="128">B</emphasis>
. Pronotum, × 410.
<emphasis id="B944EAA5386DFFB12D0DF842FBC5F85E" bold="true" box="[1092,1113,1964,1990]" pageId="127" pageNumber="128">C</emphasis>
. Mesoventrite showing median process of collar, mid-keel and keel, × 1280.
</paragraph>
</caption>
<paragraph id="8B8F36B73892FF4E29F4FE76FBC8FE62" blockId="128.[189,1398,407,507]" pageId="128" pageNumber="129">
PRONOTUM.
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long,
<quantity id="4CC89B523892FF4E28BAFE79FDC7FE2A" box="[499,603,407,434]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="128" pageNumber="129" unit="mm" value="0.25">0.25 mm</quantity>
wide, pubescent with shallow foveolae in basal half; lateral margins evenly rounded to acute hind angles, narrowly bordered, the borders continuing along the posterior margin to the angulate medial emargination opposite the scutellum (
<figureCitation id="130B2A323892FF4E2A95FE0EFBDBFE62" box="[988,1095,480,506]" captionStart="Fig" captionStartId="127.[189,232,1964,1990]" captionTargetBox="[205,1381,980,1921]" captionTargetId="figure@127.[204,1382,977,1922]" captionTargetPageId="127" captionText="Fig. 81. Cissidium subfaustum sp. nov. A. Habitus. B. Pronotum, × 410. C. Mesoventrite showing median process of collar, mid-keel and keel, × 1280." figureDoi="http://doi.org/10.5281/zenodo.3777446" httpUri="https://zenodo.org/record/3777446/files/figure.png" pageId="128" pageNumber="129">Fig. 81B</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B73892FF4E29F4FDCAFBBBFDA5" blockId="128.[189,1063,547,574]" box="[189,1063,547,574]" pageId="128" pageNumber="129">
ELYTRA.
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long,
<quantity id="4CC89B523892FF4E289BFDCDFDA7FDA6" box="[466,571,547,574]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="128" pageNumber="129" unit="mm" value="0.34">0.34 mm</quantity>
wide, pubescent and shallowly foveolate.
</paragraph>
<paragraph id="8B8F36B73892FF4E29F4FD88FE64FD75" blockId="128.[189,1398,614,749]" pageId="128" pageNumber="129">
MESOVENTRITE. Medial extension of collar sharply pointed; mid-keel anterior angles effaced before reaching collar, posterior angles stopping well short of mesocoxae; keel long, with ± six setae, tapering to a sharp point near the base of the mesocoxae; mesoventral lateral margins effaced, without serrations; humeri toothed (
<figureCitation id="130B2A323892FF4E2836FD3DFE77FD75" box="[383,491,723,749]" captionStart="Fig" captionStartId="127.[189,232,1964,1990]" captionTargetBox="[205,1381,980,1921]" captionTargetId="figure@127.[204,1382,977,1922]" captionTargetPageId="127" captionText="Fig. 81. Cissidium subfaustum sp. nov. A. Habitus. B. Pronotum, × 410. C. Mesoventrite showing median process of collar, mid-keel and keel, × 1280." figureDoi="http://doi.org/10.5281/zenodo.3777446" httpUri="https://zenodo.org/record/3777446/files/figure.png" pageId="128" pageNumber="129">Fig. 81C</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B73892FF4E29F4FCF9FDDFFCCD" blockId="128.[189,1398,790,853]" pageId="128" pageNumber="129">
METAVENTRITE. Sparsely pubescent, length
<quantity id="4CC89B523892FF4E2BE3FCF8FC89FCA9" box="[682,789,790,817]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="128" pageNumber="129" unit="mm" value="0.12">0.12 mm</quantity>
, disc simple, width across spines
<quantity id="4CC89B523892FF4E2DD0FCF8FA9FFCA9" box="[1177,1283,790,817]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="128" pageNumber="129" unit="mm" value="0.08">0.08 mm</quantity>
; posterior margin of mesocoxae not serrate.
</paragraph>
<paragraph id="8B8F36B73892FF4E29F4FC90FE15FC00" blockId="128.[189,393,894,920]" box="[189,393,894,920]" pageId="128" pageNumber="129">WINGS. Apterous.</paragraph>
<paragraph id="8B8F36B73892FF4E29F4FC2EFCF1FC42" blockId="128.[189,877,960,987]" box="[189,877,960,987]" pageId="128" pageNumber="129">GENITALIA. Female spermatheca globular. Male not known.</paragraph>
</subSubSection>
<subSubSection id="C32A653C3892FF4E29F4FBEDFE40FBEA" pageId="128" pageNumber="129" type="discussion">
<paragraph id="8B8F36B73892FF4E29F4FBEDFEB3FB85" blockId="128.[189,303,1027,1053]" box="[189,303,1027,1053]" pageId="128" pageNumber="129">
<heading id="D0C781DB3892FF4E29F4FBEDFEB3FB85" bold="true" box="[189,303,1027,1053]" fontSize="11" level="3" pageId="128" pageNumber="129" reason="3">
<emphasis id="B944EAA53892FF4E29F4FBEDFEB3FB85" bold="true" box="[189,303,1027,1053]" pageId="128" pageNumber="129">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B73892FF4E29F4FBDDFE40FBEA" blockId="128.[189,1398,1075,1138]" pageId="128" pageNumber="129">One of the five Panamanian species in this group. Distinguished by the sharp borders along the posterior margin of the pronotum.</paragraph>
</subSubSection>
</treatment>
</document>
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