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<document id="80563BEEC5071B1774EE088BF53ADA00" ID-CLB-Dataset="22458" ID-DOI="10.5852/ejt.2020.622" ID-GBIF-Dataset="9cb3ea96-0ee4-40b4-8006-19b5fed80b46" ID-Zenodo-Dep="3777282" ID-ZooBank="9CD3B2CD-F072-4994-8CA9-24145D343401" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1588217079841" checkinUser="valdenar" docAuthor="Darby, Michael" docDate="2020" docId="039987A13808FFD22B13FAD2FB03FE62" docLanguage="en" docName="ejt-622_darby.pdf.imf" docOrigin="European Journal of Taxonomy 622" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cissidium bifoveolatum Darby 2020, sp. nov." docType="treatment" docUuid="793E71EC-48C7-4E7E-A407-98F783A8835D" docUuidSource="ZooBank" docVersion="7" lastPageNumber="29" masterDocId="FFA0FFD93812FFCE2949FFEEFF9CFF98" masterDocTitle="A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species" masterLastPageNumber="188" masterPageNumber="1" pageNumber="27" updateTime="1698838121940" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="D36879A5257D5E0AE4BA3FE9E5CB378E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DA915B2DB63CEB089F9F5FA6079AC1CF">
<mods:title id="81B1F565E74ED603D61AB11C29557A61">A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species</mods:title>
</mods:titleInfo>
<mods:name id="A080178183EDFE61755100300CF1D701" type="personal">
<mods:role id="0C7C6923A3F40755D4A22D86C546C79A">
<mods:roleTerm id="FB4189B9F423882AC94C374EBD2395E8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="75EB8EAF584F896810631D373396DFC5">Darby, Michael</mods:namePart>
</mods:name>
<mods:typeOfResource id="57619E5065F35DF00C922894E88A4B4C">text</mods:typeOfResource>
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<mods:title id="D3C1F46887D91B7DD9D452546E4C9B18">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="BFDABEF1112782C72357AD64A34711A6">
<mods:date id="618484D7650F967C4D2E2163111ADC08">2020</mods:date>
<mods:detail id="C01EE70D5FC38E9FFCEA8BD7F70FDBBC" type="pubDate">
<mods:number id="7BA22355584D9C07924E9D6CE7293E4E">2020-04-02</mods:number>
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<mods:detail id="2AAB33EE7AE25FF76E7F5374F872E3CA" type="volume">
<mods:number id="A44D8B8BDF8C6D2AD4756D4D131A51A3">622</mods:number>
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<mods:classification id="8B7A5F19B7437254FD6253309E98AB15">journal article</mods:classification>
<mods:identifier id="4828D6B06BC6996C468440E851F30C3C" type="CLB-Dataset">22458</mods:identifier>
<mods:identifier id="A0C407C00B5FC8D663F097A2DD48163A" type="DOI">10.5852/ejt.2020.622</mods:identifier>
<mods:identifier id="334CAA5216A41710EC046891DE0C2B46" type="GBIF-Dataset">9cb3ea96-0ee4-40b4-8006-19b5fed80b46</mods:identifier>
<mods:identifier id="D674B426D390A3A1C6124960C338BDBE" type="Zenodo-Dep">3777282</mods:identifier>
<mods:identifier id="998E27BD7057704E85AED5F2DFB2B42F" type="ZooBank">9CD3B2CD-F072-4994-8CA9-24145D343401</mods:identifier>
</mods:mods>
<treatment id="039987A13808FFD22B13FAD2FB03FE62" ID-DOI="http://doi.org/10.5281/zenodo.3795516" ID-GBIF-Taxon="163877831" ID-Zenodo-Dep="3795516" LSID="urn:lsid:zoobank.org:act:793E71EC-48C7-4E7E-A407-98F783A8835D" httpUri="http://treatment.plazi.org/id/039987A13808FFD22B13FAD2FB03FE62" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27">
<subSubSection id="C32A653C3808FFD42B13FAD2FB5CFAE4" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="8B8F36B73808FFD42B13FAD2FC46FACF" blockId="26.[371,1216,1340,1440]" box="[602,986,1340,1367]" pageId="26" pageNumber="27">
<heading id="D0C781DB3808FFD42B13FAD2FC46FACF" box="[602,986,1340,1367]" centered="true" fontSize="11" level="2" pageId="26" pageNumber="27" reason="2">
<taxonomicName id="4C304D343808FFD42B13FAD2FCE4FACF" ID-CoL="8XXPG" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[602,888,1340,1367]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="bifoveolatum" status="sp. nov.">
<emphasis id="B944EAA53808FFD42B13FAD2FCE4FACF" bold="true" box="[602,888,1340,1367]" italics="true" pageId="26" pageNumber="27">Cissidium bifoveolatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE3808FFD42A36FAD3FC46FACF" box="[895,986,1341,1367]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8B8F36B73808FFD4283AFA8FFB5CFAE4" blockId="26.[371,1216,1340,1440]" box="[371,1216,1377,1404]" pageId="26" pageNumber="27">
<uri id="FFA13AB53808FFD4283AFA8FFB5CFAE4" box="[371,1216,1377,1404]" pageId="26" pageNumber="27">
urn:lsid:zoobank.org:act:
<uuid id="FF960C623808FFD42BD8FA8FFB5CFAE4" box="[657,1216,1377,1404]" pageId="26" pageNumber="27">793E71EC-48C7-4E7E-A407-98F783A8835D</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C3808FFD42B9CFA68FCC2FA38" box="[725,862,1413,1440]" pageId="26" pageNumber="27" type="description">
<paragraph id="8B8F36B73808FFD42B9CFA68FCC2FA38" blockId="26.[371,1216,1340,1440]" box="[725,862,1413,1440]" pageId="26" pageNumber="27">
<figureCitation id="130B2A323808FFD42B9CFA68FCA9FA38" box="[725,821,1413,1440]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[195,1398,757,1880]" captionTargetId="figure@6.[189,1398,756,1884]" captionTargetPageId="6" captionText="Fig. 5. Figures to show keel shapes. A. Cissidium bifoveolatum sp. nov., × 515. B. Cissidium lucidulum sp. nov., × 780. C. Cissidium scutellaris (Deane, 1931), × 1480. D. Cissidium tigrum Darby, 2015, × 840. E. Cissidium sawadai Darby, 2015, × 680. F. Cissidium lisae Darby, 2015, × 823." figureDoi="http://doi.org/10.5281/zenodo.3777292" httpUri="https://zenodo.org/record/3777292/files/figure.png" pageId="26" pageNumber="27">Figs 5A</figureCitation>
,
<figureCitation id="130B2A323808FFD42A08FA68FCC2FA38" box="[833,862,1414,1440]" captionStart="Fig" captionStartId="27.[189,232,1967,1993]" captionTargetBox="[206,1397,954,1921]" captionTargetId="figure@27.[189,1398,952,1926]" captionTargetPageId="27" captionText="Fig. 17. Cissidium bifoveolatum sp. nov. A. Habitus. B. Pronotum, × 620. C. Mesoventrite showing median process of collar, mid-keel and keel, × 945." figureDoi="http://doi.org/10.5281/zenodo.3777316" httpUri="https://zenodo.org/record/3777316/files/figure.png" pageId="26" pageNumber="27">17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C3808FFD429F4FA27FBB5F98B" box="[189,1065,1481,1555]" pageId="26" pageNumber="27" type="etymology">
<paragraph id="8B8F36B73808FFD429F4FA27FED8FA7B" blockId="26.[189,324,1481,1507]" box="[189,324,1481,1507]" pageId="26" pageNumber="27">
<heading id="D0C781DB3808FFD429F4FA27FED8FA7B" bold="true" box="[189,324,1481,1507]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B944EAA53808FFD429F4FA27FED8FA7B" bold="true" box="[189,324,1481,1507]" pageId="26" pageNumber="27">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B73808FFD429F4FA16FBB5F98B" blockId="26.[189,1065,1528,1555]" box="[189,1065,1528,1555]" pageId="26" pageNumber="27">Johnson ms name, presumably after the two large foveae on the pronotum.</paragraph>
</subSubSection>
<subSubSection id="C32A653C3808FFD429F4F9D5FBF3F8AE" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="8B8F36B73808FFD429F4F9D5FE36F9CD" blockId="26.[189,426,1595,1621]" box="[189,426,1595,1621]" pageId="26" pageNumber="27">
<heading id="D0C781DB3808FFD429F4F9D5FE36F9CD" bold="true" box="[189,426,1595,1621]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B944EAA53808FFD429F4F9D5FE36F9CD" bold="true" box="[189,426,1595,1621]" pageId="26" pageNumber="27">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B583CEA3808FFD429A3F985FC4BF957" ID-GBIF-Occurrence="2609493503" collectingDate="1959-12" collectionCode="MMUE" collectorName="J. Vadon" country="Madagascar" elevation="500" location="Baie d'Antongil" municipality="East" pageId="26" pageNumber="27" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<paragraph id="8B8F36B73808FFD429A3F985FEC0F91D" blockId="26.[189,1398,1643,1743]" box="[234,348,1643,1669]" pageId="26" pageNumber="27">
<emphasis id="B944EAA53808FFD429A3F985FEC0F91D" bold="true" box="[234,348,1643,1669]" pageId="26" pageNumber="27">
<typeStatus id="548B88153808FFD429A3F985FEC0F91D" box="[234,348,1643,1669]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B8F36B73808FFD429F4F97EFC4BF957" blockId="26.[189,1398,1643,1743]" pageId="26" pageNumber="27">
<collectingCountry id="F32776273808FFD429F4F97EFE12F932" box="[189,398,1680,1706]" name="Madagascar" pageId="26" pageNumber="27">MADAGASCAR</collectingCountry>
•
<specimenCount id="9D36FD3E3808FFD428E4F97EFE59F932" box="[429,453,1680,1706]" pageId="26" pageNumber="27" type="male">♂</specimenCount>
;
<collectingMunicipality id="6BEBACCD3808FFD4289CF97EFD96F932" box="[469,522,1680,1706]" pageId="26" pageNumber="27">East</collectingMunicipality>
,
<location id="8EEF606C3808FFD42B53F961FD42F931" LSID="urn:lsid:plazi:treatment:039987A13808FFD22B13FAD2FB03FE62:8EEF606C3808FFD42B53F961FD42F931" box="[538,734,1679,1705]" country="Madagascar" municipality="East" name="Baie d'Antongil" pageId="26" pageNumber="27">Baie d’Antongil</location>
,
<location id="8EEF606C3808FFD42BA2F961FC47F931" LSID="urn:lsid:plazi:treatment:039987A13808FFD22B13FAD2FB03FE62:8EEF606C3808FFD42BA2F961FC47F931" box="[747,987,1679,1705]" country="Madagascar" municipality="East" name="Ambohitsitondroina" pageId="26" pageNumber="27">Ambohitsitondroina</location>
;
<quantity id="4CC89B523808FFD42AA3F961FBABF932" box="[1002,1079,1679,1706]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="26" pageNumber="27" unit="m" value="500.0">
<elevation id="001DD1843808FFD42AA3F961FBABF932" box="[1002,1079,1679,1706]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="26" pageNumber="27" unit="m" value="500.0">500 m</elevation>
</quantity>
a.s.l.; roots of
<taxonomicName id="4C304D343808FFD42DBBF961FAF2F931" box="[1266,1390,1679,1705]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="26" pageNumber="27" phylum="Tracheophyta" rank="genus">
<emphasis id="B944EAA53808FFD42DBBF961FAF2F931" box="[1266,1390,1679,1705]" italics="true" pageId="26" pageNumber="27">Asplenium</emphasis>
</taxonomicName>
;
<date id="FF8E10773808FFD429F4F95BFEA5F957" box="[189,313,1716,1743]" pageId="26" pageNumber="27" value="1959-12">
<collectingDate id="EFCAE99F3808FFD429F4F95BFEA5F957" box="[189,313,1716,1743]" pageId="26" pageNumber="27" value="1959-12">Dec. 1959</collectingDate>
</date>
;
<collectorName id="26C553613808FFD4280CF95BFE36F956" box="[325,426,1716,1743]" pageId="26" pageNumber="27">J. Vadon</collectorName>
leg.; identified as
<taxonomicName id="4C304D343808FFD42BC8F95AFC85F956" authorityName="Motschulsky" authorityYear="1869" box="[641,793,1716,1742]" class="Insecta" family="Ptiliidae" genus="Camptodium" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B944EAA53808FFD42BC8F95AFC85F956" box="[641,793,1716,1742]" italics="true" pageId="26" pageNumber="27">Camptodium</emphasis>
</taxonomicName>
by CJ;
<collectionCode id="ED21AE723808FFD42A3AF95BFC4EF957" box="[883,978,1717,1743]" collectionName="United Kingdom, Manchester, The University, Manchester Museum" country="United Kingdom" httpUri="http://grbio.org/cool/he6q-xsyw" name="Museum of Manchester University" pageId="26" pageNumber="27">MMUE</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="8B8F36B73808FFD429A3F919FEFBF889" blockId="26.[189,1135,1783,1846]" box="[234,359,1783,1809]" pageId="26" pageNumber="27">
<emphasis id="B944EAA53808FFD429A3F919FEFBF889" bold="true" box="[234,359,1783,1809]" pageId="26" pageNumber="27">
<typeStatus id="548B88153808FFD429A3F919FEFBF889" box="[234,359,1783,1809]" pageId="26" pageNumber="27" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B8F36B73808FFD429F4F8F5FBF3F8AE" blockId="26.[189,1135,1783,1846]" box="[189,1135,1819,1846]" pageId="26" pageNumber="27">
<materialsCitation id="3B583CEA3808FFD429F4F8F5FBF7F8AE" ID-GBIF-Occurrence="2609493544" box="[189,1131,1819,1846]" collectingDate="1959-12" collectionCode="MMUE, BMNH" collectorName="J. Vadon" country="Madagascar" elevation="500" location="MADAGASCAR" municipality="East" pageId="26" pageNumber="27" specimenCount="8" typeStatus="paratype">
<collectingCountry id="F32776273808FFD429F4F8F5FE12F8AD" box="[189,398,1819,1845]" name="Madagascar" pageId="26" pageNumber="27">MADAGASCAR</collectingCountry>
•
<specimenCount id="9D36FD3E3808FFD428EFF8F5FE7EF8AE" box="[422,482,1819,1846]" pageId="26" pageNumber="27" type="generic">8 ex.</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED21AE723808FFD42AECF8F2FB9FF8AE" box="[933,1027,1820,1846]" collectionName="United Kingdom, Manchester, The University, Manchester Museum" country="United Kingdom" httpUri="http://grbio.org/cool/he6q-xsyw" name="Museum of Manchester University" pageId="26" pageNumber="27">MMUE</collectionCode>
,
<collectionCode id="ED21AE723808FFD42D46F8F2FBF7F8AE" box="[1039,1131,1820,1846]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="26" pageNumber="27">BMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32A653C3808FFD229F4F8B0FB19FEFF" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27" type="description">
<paragraph id="8B8F36B73808FFD429F4F8B0FED2F8E0" blockId="26.[189,334,1886,1912]" box="[189,334,1886,1912]" pageId="26" pageNumber="27">
<heading id="D0C781DB3808FFD429F4F8B0FED2F8E0" bold="true" box="[189,334,1886,1912]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B944EAA53808FFD429F4F8B0FED2F8E0" bold="true" box="[189,334,1886,1912]" pageId="26" pageNumber="27">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B73808FFD429F4F860FD36F830" blockId="26.[189,682,1934,1960]" box="[189,682,1934,1960]" pageId="26" pageNumber="27">
SIZE. Habitus (
<figureCitation id="130B2A323808FFD4282EF860FE4FF830" box="[359,467,1934,1960]" captionStart="Fig" captionStartId="27.[189,232,1967,1993]" captionTargetBox="[206,1397,954,1921]" captionTargetId="figure@27.[189,1398,952,1926]" captionTargetPageId="27" captionText="Fig. 17. Cissidium bifoveolatum sp. nov. A. Habitus. B. Pronotum, × 620. C. Mesoventrite showing median process of collar, mid-keel and keel, × 945." figureDoi="http://doi.org/10.5281/zenodo.3777316" httpUri="https://zenodo.org/record/3777316/files/figure.png" pageId="26" pageNumber="27">Fig. 17A</figureCitation>
), length
<quantity id="4CC89B523808FFD42B73F860FD39F830" box="[570,677,1934,1960]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" pageId="26" pageNumber="27" unit="mm" value="0.69">0.69 mm</quantity>
.
</paragraph>
<paragraph id="8B8F36B73808FFD529F4F83FFCF0FEF4" blockId="26.[189,1285,2001,2027]" lastBlockId="27.[189,1398,267,364]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
COLOUR. Yellow brown appearing polished, head darker, pubescence, legs and antennae paler. HEAD. With a shallow depression interrupted medially behind the eyes, width across eyes
<quantity id="4CC89B523809FFD52C4AFEE5FAEEFEBE" box="[1283,1394,267,294]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="27" pageNumber="28" unit="mm" value="0.21">0.21 mm</quantity>
; antennomeres III–XI length
<quantity id="4CC89B523809FFD52B5FFEC0FD1AFED1" box="[534,646,302,329]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="27" pageNumber="28" unit="mm" value="0.35">0.35 mm</quantity>
, III–IX length
<quantity id="4CC89B523809FFD52A09FEC1FC2CFED1" box="[832,944,303,329]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="27" pageNumber="28" unit="mm" value="0.21">0.21 mm</quantity>
, X–XI length
<quantity id="4CC89B523809FFD52D2BFEC1FB4DFED1" box="[1122,1233,303,329]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="27" pageNumber="28" unit="mm" value="0.14">0.14 mm</quantity>
, mentum and submentum as
<figureCitation id="130B2A323809FFD52824FEBCFE55FEF4" box="[365,457,338,364]" captionStart="Fig" captionStartId="5.[189,232,1927,1953]" captionTargetBox="[232,1334,1274,1884]" captionTargetId="figure@5.[232,1334,1274,1887]" captionTargetPageId="5" captionText="Fig. 4. Menta and submenta. A. Cissidium petri Darby, 2015, × 1380. B. Cissidium tigrum Darby, 2015, × 1220. C. Cissidium amboroense Darby, 2015, × 1260. D. Cissidium losbanos sp. nov., × 1100. E. Cissidium triangulum Darby, 2019, × 650. F. Cissidium pilosellum sp. nov., × 1420." figureDoi="http://doi.org/10.5281/zenodo.3777290" httpUri="https://zenodo.org/record/3777290/files/figure.png" pageId="27" pageNumber="28">Fig. 4A</figureCitation>
, but with faint circular depressions.
</paragraph>
<paragraph id="8B8F36B73809FFD529F4FE7AFBB8FD8F" blockId="27.[189,1399,403,535]" pageId="27" pageNumber="28">
PRONOTUM. Length
<quantity id="4CC89B523809FFD528D7FE7DFD96FE36" box="[414,522,403,430]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="27" pageNumber="28" unit="mm" value="0.2">0.20 mm</quantity>
, width
<quantity id="4CC89B523809FFD52B16FE7DFD57FE36" box="[607,715,403,430]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="27" pageNumber="28" unit="mm" value="0.27">0.27 mm</quantity>
, pubescence sparse, with four foveae in the posterior third the two medial ones much larger than the two close to the corners; side margins rounded then sinuate to rectangular hind angles, narrowly bordered, the border extending for a short distance along the posterior margin which has a sinuous emargination before the scutellum (
<figureCitation id="130B2A323809FFD52AE2FE13FB8AFD8F" box="[939,1046,509,535]" captionStart="Fig" captionStartId="27.[189,232,1967,1993]" captionTargetBox="[206,1397,954,1921]" captionTargetId="figure@27.[189,1398,952,1926]" captionTargetPageId="27" captionText="Fig. 17. Cissidium bifoveolatum sp. nov. A. Habitus. B. Pronotum, × 620. C. Mesoventrite showing median process of collar, mid-keel and keel, × 945." figureDoi="http://doi.org/10.5281/zenodo.3777316" httpUri="https://zenodo.org/record/3777316/files/figure.png" pageId="27" pageNumber="28">Fig. 17B</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B73809FFD529F4FDD1FB47FDC0" blockId="27.[189,1243,574,601]" box="[189,1243,574,601]" pageId="27" pageNumber="28">
ELYTRA. Length
<quantity id="4CC89B523809FFD52833FDD0FE7AFDC1" box="[378,486,574,601]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="27" pageNumber="28" unit="mm" value="0.45">0.45 mm</quantity>
, width
<quantity id="4CC89B523809FFD52B72FDD0FD3BFDC1" box="[571,679,574,601]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="27" pageNumber="28" unit="mm" value="0.38">0.38 mm</quantity>
, pubescence mainly confined to lateral margins.
</paragraph>
<paragraph id="8B8F36B73809FFD529F4FD6EFE02FCBF" blockId="27.[189,1399,640,807]" pageId="27" pageNumber="28">
MESOVENTRITE. Medial extension of collar narrow, short; mid-keel posterior and anterior corners of approximately same width, the posterior margin reaching anterior margin of the mesocoxal cavities, sharply raised medially to keel; keel narrow, parallel-sided, bearing ¾ setae, terminating between the mesocoxae (
<figureCitation id="130B2A323809FFD5281AFD07FE2AFC9C" box="[339,438,745,772]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[195,1398,757,1880]" captionTargetId="figure@6.[189,1398,756,1884]" captionTargetPageId="6" captionText="Fig. 5. Figures to show keel shapes. A. Cissidium bifoveolatum sp. nov., × 515. B. Cissidium lucidulum sp. nov., × 780. C. Cissidium scutellaris (Deane, 1931), × 1480. D. Cissidium tigrum Darby, 2015, × 840. E. Cissidium sawadai Darby, 2015, × 680. F. Cissidium lisae Darby, 2015, × 823." figureDoi="http://doi.org/10.5281/zenodo.3777292" httpUri="https://zenodo.org/record/3777292/files/figure.png" pageId="27" pageNumber="28">Fig. 5A</figureCitation>
); mesoventral lateral margins loosely serrate in posterior half; humeri strongly toothed (
<figureCitation id="130B2A323809FFD5286CFCE2FE0CFCBF" box="[293,400,780,807]" captionStart="Fig" captionStartId="27.[189,232,1967,1993]" captionTargetBox="[206,1397,954,1921]" captionTargetId="figure@27.[189,1398,952,1926]" captionTargetPageId="27" captionText="Fig. 17. Cissidium bifoveolatum sp. nov. A. Habitus. B. Pronotum, × 620. C. Mesoventrite showing median process of collar, mid-keel and keel, × 945." figureDoi="http://doi.org/10.5281/zenodo.3777316" httpUri="https://zenodo.org/record/3777316/files/figure.png" pageId="27" pageNumber="28">Fig. 17C</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B73809FFD529F4FCA0FD85FC14" blockId="27.[189,1398,846,908]" pageId="27" pageNumber="28">
METAVENTRITE. Length
<quantity id="4CC89B523809FFD52882FCA0FDA4FCF0" box="[459,568,846,872]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="27" pageNumber="28" unit="mm" value="0.14">0.14 mm</quantity>
, width across spines
<quantity id="4CC89B523809FFD52A7DFCA0FC3DFCF0" box="[820,929,846,872]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="27" pageNumber="28" unit="mm" value="0.09">0.09 mm</quantity>
, glabrous, disc plain. Posterior margins of mesocoxal cavities serrate.
</paragraph>
<caption id="DF4F663F3809FFD529F4F841FC8AF875" ID-DOI="http://doi.org/10.5281/zenodo.3777316" ID-Zenodo-Dep="3777316" httpUri="https://zenodo.org/record/3777316/files/figure.png" pageId="27" pageNumber="28" startId="27.[189,232,1967,1993]" targetBox="[206,1397,954,1921]" targetPageId="27">
<paragraph id="8B8F36B73809FFD529F4F841FC8AF875" blockId="27.[189,1398,1967,2029]" pageId="27" pageNumber="28">
<emphasis id="B944EAA53809FFD529F4F841FE80F852" bold="true" box="[189,284,1967,1994]" pageId="27" pageNumber="28">Fig. 17.</emphasis>
<taxonomicName id="4C304D343809FFD5286FF841FDA3F851" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[294,575,1967,1993]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="bifoveolatum" status="sp. nov.">
<emphasis id="B944EAA53809FFD5286FF841FDA3F851" box="[294,575,1967,1993]" italics="true" pageId="27" pageNumber="28">Cissidium bifoveolatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE3809FFD52B00F85EFD3BF852" box="[585,679,1968,1994]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B944EAA53809FFD52BF8F841FD5AF851" bold="true" box="[689,710,1967,1993]" pageId="27" pageNumber="28">A</emphasis>
. Habitus.
<emphasis id="B944EAA53809FFD52A0DF841FCC4F851" bold="true" box="[836,856,1967,1993]" pageId="27" pageNumber="28">B</emphasis>
. Pronotum, × 620.
<emphasis id="B944EAA53809FFD52D0EF841FBC0F851" bold="true" box="[1095,1116,1967,1993]" pageId="27" pageNumber="28">C</emphasis>
. Mesoventrite showing median process of collar, mid-keel and keel, × 945.
</paragraph>
</caption>
<paragraph id="8B8F36B7380EFFD229F4FEE2FE5EFEBE" blockId="28.[189,450,268,294]" box="[189,450,268,294]" pageId="28" pageNumber="29">WINGS. Macropterous.</paragraph>
<paragraph id="8B8F36B7380EFFD229F4FEA3FB19FEFF" blockId="28.[189,1157,333,359]" box="[189,1157,333,359]" pageId="28" pageNumber="29">
GENITALIA. Female spermatheca globular. Male aedeagus as
<figureCitation id="130B2A32380EFFD22A35FEA3FC4DFEFF" box="[892,977,333,359]" captionStart="Fig" captionStartId="7.[189,232,1825,1851]" captionTargetBox="[202,1398,769,1790]" captionTargetId="figure@7.[189,1398,769,1799]" captionTargetPageId="7" captionText="Fig. 6. Figures to show genitalia types. – Spermathecae. A. Cissidium bilineatum sp. nov. B. Cissidium amboroense Darby, 2015. C. Cissidium sawadai Darby, 2015. D. Cissidium arcuatum sp. nov. E. Cissidium murphyi sp. nov. – Aedeagi. F. Cissidium petri Darby, 2015. G. Cissidium clareae sp. nov. H. Cissidium fraternum sp. nov. I. Cissidium tigrum Darby, 2015. J. Cissidium apterum sp. nov. K. Cissidium brachypterum sp. nov. L. Cissidium greensladei sp. nov. M. Cissidium kolombangaricum sp. nov. N. Cissidium mindanao sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3777294" httpUri="https://zenodo.org/record/3777294/files/figure.png" pageId="28" pageNumber="29">Fig. 6L</figureCitation>
but tip blunter.
</paragraph>
</subSubSection>
<subSubSection id="C32A653C380EFFD229F4FE60FB03FE62" pageId="28" pageNumber="29" type="discussion">
<paragraph id="8B8F36B7380EFFD229F4FE60FEB3FE30" blockId="28.[189,303,398,424]" box="[189,303,398,424]" pageId="28" pageNumber="29">
<heading id="D0C781DB380EFFD229F4FE60FEB3FE30" bold="true" box="[189,303,398,424]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B944EAA5380EFFD229F4FE60FEB3FE30" bold="true" box="[189,303,398,424]" pageId="28" pageNumber="29">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B7380EFFD229F4FE53FB03FE62" blockId="28.[189,1399,445,506]" pageId="28" pageNumber="29">
One of the five species of
<taxonomicName id="4C304D34380EFFD228A2FE53FDFCFE4F" box="[491,608,445,471]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B944EAA5380EFFD228A2FE53FDFCFE4F" box="[491,608,445,471]" italics="true" pageId="28" pageNumber="29">Cissidium</emphasis>
</taxonomicName>
recorded from
<collectingCountry id="F3277627380EFFD22A5DFE53FC3EFE4F" box="[788,930,445,471]" name="Madagascar" pageId="28" pageNumber="29">Madagascar</collectingCountry>
in this group, only likely to be confused with
<taxonomicName id="4C304D34380EFFD229B1FE0EFEF5FE62" authorityName="Darby" authorityYear="2013" box="[248,361,480,506]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="banari">
<emphasis id="B944EAA5380EFFD229B1FE0EFEF5FE62" box="[248,361,480,506]" italics="true" pageId="28" pageNumber="29">C. banari</emphasis>
</taxonomicName>
, but distinguished by its smaller size and longer pronotal pubescence.
</paragraph>
</subSubSection>
</treatment>
</document>
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