treatments-xml/data/03/8B/87/038B87F6A5721F6E0AEDB4FCB9CD521B.xml
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<document id="951C8A20B4FF315F1588B1142F013722" ID-CLB-Dataset="21269" ID-DOI="10.11646/zootaxa.4786.1.3" ID-GBIF-Dataset="70d743db-654b-4050-a05f-a5c71b54e14e" ID-ISSN="1175-5326" ID-Zenodo-Dep="3990784" ID-ZooBank="625F197D-9D26-4B00-AF3E-3045586ECB07" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1597795225921" checkinUser="felipe" docAuthor="Salles, F. F., Nieto, C. &amp; Cruz, P. V." docDate="2020" docId="038B87F6A5721F6E0AEDB4FCB9CD521B" docLanguage="en" docName="zootaxa.4786.1.3.pdf.imf" docOrigin="Zootaxa 4786 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rivudiva oonirikoperi Salles &amp; Nieto &amp; Cruz 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="43" masterDocId="FFB2FF8EA5771F680A7AB129B91D5554" masterDocTitle="New species of Rivudiva Lugo-Ortiz &amp; McCafferty (Ephemeroptera: Baetidae) with comments on R. minantenna Lugo-Ortiz &amp; McCafferty and R. trichobasis Lugo-Ortiz &amp; McCafferty" masterLastPageNumber="52" masterPageNumber="37" pageNumber="42" updateTime="1698844779479" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5D08E0F7187C08166B69529149DE67FE">New species of Rivudiva Lugo-Ortiz &amp; McCafferty (Ephemeroptera: Baetidae) with comments on R. minantenna Lugo-Ortiz &amp; McCafferty and R. trichobasis Lugo-Ortiz &amp; McCafferty</mods:title>
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<mods:name id="06673C8FD98FDBE62D810555243F6C8A" type="personal">
<mods:role id="34AD2439BD9807235AEE1B746191F75C">
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<mods:namePart id="D46A57DE59A211CB6BBCE040ED849D79">Salles, F. F.</mods:namePart>
<mods:affiliation id="D85A329BAE08470C13615602C202DF36">Museu de Entomologia, Depto. de Entomologia, Universidade Federal de Viçosa, CEP 36.570 - 900, Viçosa, MG, Brazil. frederico. salles @ ufv. br; https: // orcid. org / 0000 - 0001 - 8331 - 5929</mods:affiliation>
</mods:name>
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<mods:namePart id="F3349CD724A8C5D8E38775369E4718AD">Nieto, C.</mods:namePart>
<mods:affiliation id="956E913EB07840D6E6F03F3EF52C41E2">Instituto de Biodiversidad Neotropical (IBN), CONICET-U. N. T. &amp; Facultad de Ciencias Naturales, Universidad Nacional de Tucumán, San Miguel de Tucumán, 4.000 Tucumán, Argentina.</mods:affiliation>
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<mods:role id="642EAC3746A7AD42BD16259EF46E3BF7">
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</mods:role>
<mods:namePart id="624CA10A764CBB1E583D300006E40B04">Cruz, P. V.</mods:namePart>
<mods:affiliation id="A16B0E19097797CF1A076F5B4BBF481B">Universidade Federal de Rondônia - UNIR, Programa de Pós-Graduação em Ciências Ambientais, Laboratório de Biodiversidade e Conservação - LABICON, CEP: 76940 - 000, Rolim de Moura, Rondônia, Brazil.</mods:affiliation>
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<mods:title id="8D5025F3460A750167191A39895449D5">Zootaxa</mods:title>
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<mods:date id="65CD227186B6229048D64D39DFD7FC28">2020</mods:date>
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<mods:number id="1A55D006856496D610501C99AB423C05">2020-05-29</mods:number>
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<mods:number id="CEDFF18E1326033AE26B665AFA86F6BF">4786</mods:number>
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<treatment id="038B87F6A5721F6E0AEDB4FCB9CD521B" ID-DOI="http://doi.org/10.5281/zenodo.4328820" ID-GBIF-Taxon="167186778" ID-Zenodo-Dep="4328820" LSID="urn:lsid:plazi:treatment:038B87F6A5721F6E0AEDB4FCB9CD521B" httpUri="http://treatment.plazi.org/id/038B87F6A5721F6E0AEDB4FCB9CD521B" lastPageId="6" lastPageNumber="43" pageId="5" pageNumber="42">
<subSubSection id="C338656BA5721F6D0AEDB4FCBB5050BB" box="[151,589,1493,1519]" pageId="5" pageNumber="42" type="nomenclature">
<paragraph id="8B9D36E0A5721F6D0AEDB4FCBB5050BB" blockId="5.[151,589,1493,1519]" box="[151,589,1493,1519]" pageId="5" pageNumber="42">
<heading id="D0D5818CA5721F6D0AEDB4FCBB5050BB" box="[151,589,1493,1519]" level="2" pageId="5" pageNumber="42">
<taxonomicName id="4C224D63A5721F6D0AEDB4FCB8FF50BB" authority="Cruz" authorityName="Salles &amp; Nieto &amp; Cruz" authorityYear="2020" box="[151,482,1493,1519]" class="Insecta" family="Baetidae" genus="Rivudiva" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="oonirikoperi" status="sp. nov.">
<emphasis id="B956EAF2A5721F6D0AEDB4FCB88750BB" box="[151,410,1493,1519]" italics="true" pageId="5" pageNumber="42">Rivudiva oonirikoperi</emphasis>
<emphasis id="B956EAF2A5721F6D0BDBB4FCB8FF50BB" bold="true" box="[417,482,1493,1519]" pageId="5" pageNumber="42">Cruz</emphasis>
</taxonomicName>
<emphasis id="B956EAF2A5721F6D0B98B4FCBB5050BB" bold="true" box="[482,589,1493,1519]" pageId="5" pageNumber="42">
,
<taxonomicNameLabel id="A2655789A5721F6D0B95B4FCBB5050BB" box="[495,589,1493,1519]" pageId="5" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C338656BA5721F6D0AEDB735BBF7532B" pageId="5" pageNumber="42" type="materials_examined">
<paragraph id="8B9D36E0A5721F6D0AEDB735BBF7532B" blockId="5.[151,1437,1564,1664]" pageId="5" pageNumber="42">
<emphasis id="B956EAF2A5721F6D0AEDB735B8655363" bold="true" box="[151,376,1564,1591]" pageId="5" pageNumber="42">Material examined</emphasis>
.
<materialsCitation id="3B4A3CBDA5721F6D0BFCB734BC84530F" ID-GBIF-Occurrence="2832053310" collectingDate="2011-08-26" collectorName="P. V. Cruz &amp; A. S. Fernandes &amp; E. Reis" country="Brazil" county="Km" location="Amazonas" municipality="Sao Gabriel da Cachoeira" pageId="5" pageNumber="42" specimenCount="1" typeStatus="holotype">
<typeStatus id="54998842A5721F6D0BFCB734B8F35363" box="[390,494,1565,1591]" pageId="5" pageNumber="42" type="holotype">Holotype</typeStatus>
(male nymph on slide):
<collectingCountry id="F3357670A5721F6D0881B734BA7E5363" box="[763,867,1565,1591]" name="Brazil" pageId="5" pageNumber="42">BRAZIL</collectingCountry>
,
<location id="8EFD603BA5721F6D0916B734BAFF5363" LSID="urn:lsid:plazi:treatment:038B87F6A5721F6E0AEDB4FCB9CD521B:8EFD603BA5721F6D0916B734BAFF5363" box="[876,994,1565,1591]" country="Brazil" county="Km" municipality="Sao Gabriel da Cachoeira" name="Amazonas" pageId="5" pageNumber="42">Amazonas</location>
state,
<collectingMunicipality id="6BF9AC9AA5721F6D0E53B734BC5A5363" box="[1065,1351,1565,1591]" pageId="5" pageNumber="42">São Gabriel da Cachoeira</collectingMunicipality>
, stream on
<collectionCode id="ED33AE25A5721F6D0AC0B76BB9C25308" box="[186,223,1602,1628]" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection" pageId="5" pageNumber="42">BR</collectionCode>
307
<collectingCounty id="62FC4E6CA5721F6D0B6CB76BB85D5308" box="[278,320,1602,1628]" pageId="5" pageNumber="42">Km</collectingCounty>
13, 00
<superScript id="7C579BA8A5721F6D0BF6B769B889531A" attach="none" box="[396,404,1600,1614]" fontSize="6" pageId="5" pageNumber="42">o</superScript>
0444.6”S/067
<superScript id="7C579BA8A5721F6D0843B769BB5C531A" attach="right" box="[569,577,1600,1614]" fontSize="6" pageId="5" pageNumber="42">o</superScript>
0015.9”W,
<date id="FF9C1020A5721F6D08B3B768BA4F530F" box="[713,850,1601,1627]" pageId="5" pageNumber="42" value="2011-08-26">
<collectingDate id="EFD8E9C8A5721F6D08B3B768BA4F530F" box="[713,850,1601,1627]" pageId="5" pageNumber="42" value="2011-08-26">26.viii.2011</collectingDate>
</date>
,
<collectorName id="26D75336A5721F6D0927B76BBAD5530F" box="[861,968,1601,1628]" pageId="5" pageNumber="42">
P.
<collectionCode id="ED33AE25A5721F6D090AB76BBA985308" box="[880,901,1602,1628]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="42" type="Museum">V</collectionCode>
. Cruz
</collectorName>
,
<collectorName id="26D75336A5721F6D09ABB768BD61530F" box="[977,1148,1601,1627]" pageId="5" pageNumber="42">A.S. Fernandes</collectorName>
and
<collectorName id="26D75336A5721F6D0EC8B76BBC1E530F" box="[1202,1283,1601,1628]" pageId="5" pageNumber="42">E. Reis</collectorName>
cols. (
<collectionCode id="ED33AE25A5721F6D0F36B768BC8F530F" box="[1356,1426,1601,1627]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="5" pageNumber="42">INPA</collectionCode>
)
</materialsCitation>
.
<typeStatus id="54998842A5721F6D0AEDB74FB9E053D4" box="[151,253,1638,1664]" pageId="5" pageNumber="42" type="paratype">Paratype</typeStatus>
: one nymph, same data as
<typeStatus id="54998842A5721F6D0852B74CBB97532B" box="[552,650,1637,1663]" pageId="5" pageNumber="42" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED33AE25A5721F6D08E2B74CBBC3532B" box="[664,734,1637,1663]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="5" pageNumber="42">INPA</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C338656BA5721F6D0AEDB7EDB88E52A3" pageId="5" pageNumber="42" type="diagnosis">
<paragraph id="8B9D36E0A5721F6D0AEDB7EDB80D538A" blockId="5.[151,272,1732,1758]" box="[151,272,1732,1758]" pageId="5" pageNumber="42">
<heading id="D0D5818CA5721F6D0AEDB7EDB80D538A" bold="true" box="[151,272,1732,1758]" fontSize="11" level="3" pageId="5" pageNumber="42">
<emphasis id="B956EAF2A5721F6D0AEDB7EDB80D538A" bold="true" box="[151,272,1732,1758]" pageId="5" pageNumber="42">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D36E0A5721F6D0AEDB62CBC7D5287" blockId="5.[151,1437,1797,2040]" pageId="5" pageNumber="42">
<emphasis id="B956EAF2A5721F6D0AEDB62CB9FB524B" box="[151,230,1797,1823]" italics="true" pageId="5" pageNumber="42">Nymph</emphasis>
. 1) antenna without spine-like setae on scape and pedicel; 2) distal margin of labrum without medial emargination (
<figureCitation id="13192A65A5721F6D0A90B600B8275217" box="[234,314,1833,1859]" captionStart="FIGURES 1116" captionStartId="7.[151,264,1945,1970]" captionTargetBox="[239,1349,202,1903]" captionTargetId="figure@7.[221,1366,181,1920]" captionTargetPageId="7" captionText="FIGURES 1116. Rivudiva oonirikoperi, sp. nov., nymph. (11) Labrum; (12) Left mandible; (13) Right mandible; (14) Hypopharynx; (15) Maxilla; (16) Labium (left = dorsal surface, right = ventral surface)." figureDoi="http://doi.org/10.5281/zenodo.3990790" httpUri="https://zenodo.org/record/3990790/files/figure.png" pageId="5" pageNumber="42">Fig. 11</figureCitation>
); 3) outer margin of mandibular incisors with spine-like process (
<figureCitation id="13192A65A5721F6D0E79B600BD9E5217" box="[1027,1155,1833,1859]" captionStart="FIGURES 1116" captionStartId="7.[151,264,1945,1970]" captionTargetBox="[239,1349,202,1903]" captionTargetId="figure@7.[221,1366,181,1920]" captionTargetPageId="7" captionText="FIGURES 1116. Rivudiva oonirikoperi, sp. nov., nymph. (11) Labrum; (12) Left mandible; (13) Right mandible; (14) Hypopharynx; (15) Maxilla; (16) Labium (left = dorsal surface, right = ventral surface)." figureDoi="http://doi.org/10.5281/zenodo.3990790" httpUri="https://zenodo.org/record/3990790/files/figure.png" pageId="5" pageNumber="42">Figs 1213</figureCitation>
); 4) tuft on apex of lingua formed by short simple setae (
<figureCitation id="13192A65A5721F6D0B96B664BB5D5233" box="[492,576,1869,1895]" captionStart="FIGURES 1116" captionStartId="7.[151,264,1945,1970]" captionTargetBox="[239,1349,202,1903]" captionTargetId="figure@7.[221,1366,181,1920]" captionTargetPageId="7" captionText="FIGURES 1116. Rivudiva oonirikoperi, sp. nov., nymph. (11) Labrum; (12) Left mandible; (13) Right mandible; (14) Hypopharynx; (15) Maxilla; (16) Labium (left = dorsal surface, right = ventral surface)." figureDoi="http://doi.org/10.5281/zenodo.3990790" httpUri="https://zenodo.org/record/3990790/files/figure.png" pageId="5" pageNumber="42">Fig. 14</figureCitation>
); 5) maxillary palp long, 2.1× the length of galea-lacinia (
<figureCitation id="13192A65A5721F6D0EB4B664BC3C5233" box="[1230,1313,1868,1895]" captionStart="FIGURES 1116" captionStartId="7.[151,264,1945,1970]" captionTargetBox="[239,1349,202,1903]" captionTargetId="figure@7.[221,1366,181,1920]" captionTargetPageId="7" captionText="FIGURES 1116. Rivudiva oonirikoperi, sp. nov., nymph. (11) Labrum; (12) Left mandible; (13) Right mandible; (14) Hypopharynx; (15) Maxilla; (16) Labium (left = dorsal surface, right = ventral surface)." figureDoi="http://doi.org/10.5281/zenodo.3990790" httpUri="https://zenodo.org/record/3990790/files/figure.png" pageId="5" pageNumber="42">Fig. 15</figureCitation>
); 6) glossa subelliptical, not expanded at base (
<figureCitation id="13192A65A5721F6D085BB658BB6F52D8" box="[545,626,1905,1932]" captionStart="FIGURES 1116" captionStartId="7.[151,264,1945,1970]" captionTargetBox="[239,1349,202,1903]" captionTargetId="figure@7.[221,1366,181,1920]" captionTargetPageId="7" captionText="FIGURES 1116. Rivudiva oonirikoperi, sp. nov., nymph. (11) Labrum; (12) Left mandible; (13) Right mandible; (14) Hypopharynx; (15) Maxilla; (16) Labium (left = dorsal surface, right = ventral surface)." figureDoi="http://doi.org/10.5281/zenodo.3990790" httpUri="https://zenodo.org/record/3990790/files/figure.png" pageId="5" pageNumber="42">Fig. 16</figureCitation>
); 7) medial protuberance of labial palp segment II strongly projected medially (
<figureCitation id="13192A65A5721F6D0A9FB6BCB82A52E4" box="[229,311,1941,1968]" captionStart="FIGURES 1116" captionStartId="7.[151,264,1945,1970]" captionTargetBox="[239,1349,202,1903]" captionTargetId="figure@7.[221,1366,181,1920]" captionTargetPageId="7" captionText="FIGURES 1116. Rivudiva oonirikoperi, sp. nov., nymph. (11) Labrum; (12) Left mandible; (13) Right mandible; (14) Hypopharynx; (15) Maxilla; (16) Labium (left = dorsal surface, right = ventral surface)." figureDoi="http://doi.org/10.5281/zenodo.3990790" httpUri="https://zenodo.org/record/3990790/files/figure.png" pageId="5" pageNumber="42">Fig. 16</figureCitation>
); 8) hind wing pads absent; 9) inner margin and anterior surface of fore tarsus with robust long spinelike setae (
<figureCitation id="13192A65A5721F6D0B74B690B8425287" box="[270,351,1977,2003]" captionStart="FIGURES 1721" captionStartId="8.[151,264,1945,1970]" captionTargetBox="[192,1396,189,1917]" captionTargetId="figure@8.[192,1396,181,1920]" captionTargetPageId="8" captionText="FIGURES 1721. Rivudiva oonirikoperi, sp. nov., nymph. (17) Fore leg (anterior surface); (18) detail of foreclaw; (19) mid leg (anterior surface); (20) Paraproct; (21) Dorsal view of body." figureDoi="http://doi.org/10.5281/zenodo.3990792" httpUri="https://zenodo.org/record/3990792/files/figure.png" pageId="5" pageNumber="42">Fig. 17</figureCitation>
); 10) paraproct with three marginal spines, posterolateral extension without spines (
<figureCitation id="13192A65A5721F6D0F7AB690BC4E5287" box="[1280,1363,1977,2003]" captionStart="FIGURES 1721" captionStartId="8.[151,264,1945,1970]" captionTargetBox="[192,1396,189,1917]" captionTargetId="figure@8.[192,1396,181,1920]" captionTargetPageId="8" captionText="FIGURES 1721. Rivudiva oonirikoperi, sp. nov., nymph. (17) Fore leg (anterior surface); (18) detail of foreclaw; (19) mid leg (anterior surface); (20) Paraproct; (21) Dorsal view of body." figureDoi="http://doi.org/10.5281/zenodo.3990792" httpUri="https://zenodo.org/record/3990792/files/figure.png" pageId="5" pageNumber="42">Fig. 20</figureCitation>
).
</paragraph>
<paragraph id="8B9D36E0A5721F6D0ABDB6F7B88E52A3" blockId="5.[151,1437,1797,2040]" box="[199,403,2013,2040]" pageId="5" pageNumber="42">
<emphasis id="B956EAF2A5721F6D0ABDB6F7B81252A3" box="[199,271,2014,2039]" italics="true" pageId="5" pageNumber="42">Imago</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C338656BA5711F6E0AEDB1B0B9CD521B" pageId="6" pageNumber="43" type="description">
<paragraph id="8B9D36E0A5711F6E0AEDB1B0B83555E7" blockId="6.[151,296,153,179]" box="[151,296,153,179]" pageId="6" pageNumber="43">
<heading id="D0D5818CA5711F6E0AEDB1B0B83555E7" bold="true" box="[151,296,153,179]" fontSize="11" level="3" pageId="6" pageNumber="43">
<emphasis id="B956EAF2A5711F6E0AEDB1B0B83555E7" bold="true" box="[151,296,153,179]" pageId="6" pageNumber="43">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D36E0A5711F6E0AEDB1C8BBA055A8" blockId="6.[151,1437,225,1871]" box="[151,701,225,252]" pageId="6" pageNumber="43">
Nymph. Lengths: body,
<quantity id="4CDA9B05A5711F6E0BDCB1C8B8E255A8" box="[422,511,225,252]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="6" pageNumber="43" unit="mm" value="3.2">3.2 mm</quantity>
; cercus,
<quantity id="4CDA9B05A5711F6E081AB1C8BBA455A8" box="[608,697,225,252]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="43" unit="mm" value="1.6">1.6 mm</quantity>
.
</paragraph>
<paragraph id="8B9D36E0A5711F6E0ABDB02CBC4F5147" blockId="6.[151,1437,225,1871]" pageId="6" pageNumber="43">
Head. Coloration: Pale yellow. Antenna white. Compound eyes light orange. Antenna. Scape and pedicel without spine-like setae; flagellum with minute spines and thin setae on apex of each segment. Labrum (
<figureCitation id="13192A65A5711F6E0E9BB000BC2C5417" box="[1249,1329,297,323]" captionStart="FIGURES 1116" captionStartId="7.[151,264,1945,1970]" captionTargetBox="[239,1349,202,1903]" captionTargetId="figure@7.[221,1366,181,1920]" captionTargetPageId="7" captionText="FIGURES 1116. Rivudiva oonirikoperi, sp. nov., nymph. (11) Labrum; (12) Left mandible; (13) Right mandible; (14) Hypopharynx; (15) Maxilla; (16) Labium (left = dorsal surface, right = ventral surface)." figureDoi="http://doi.org/10.5281/zenodo.3990790" httpUri="https://zenodo.org/record/3990790/files/figure.png" pageId="6" pageNumber="43">Fig. 11</figureCitation>
). Rectangular, length about 0.5x maximum width; distal margin without medial emargination, anterolateral and distal margin with robust bifid setae; dorsal surface near distal margin with two rows of thin simple setae; ventral surface with anterolateral robust bifid setae. Left mandible (
<figureCitation id="13192A65A5711F6E08DFB0BCBBE554FB" box="[677,760,405,431]" captionStart="FIGURES 1116" captionStartId="7.[151,264,1945,1970]" captionTargetBox="[239,1349,202,1903]" captionTargetId="figure@7.[221,1366,181,1920]" captionTargetPageId="7" captionText="FIGURES 1116. Rivudiva oonirikoperi, sp. nov., nymph. (11) Labrum; (12) Left mandible; (13) Right mandible; (14) Hypopharynx; (15) Maxilla; (16) Labium (left = dorsal surface, right = ventral surface)." figureDoi="http://doi.org/10.5281/zenodo.3990790" httpUri="https://zenodo.org/record/3990790/files/figure.png" pageId="6" pageNumber="43">Fig. 12</figureCitation>
). Incisors cleft in two sets; outer and inner set of incisors respectively with 3 + 3 denticles, outer incisor with spine-like process; prostheca robust and bifid at base, internal lobe slender, external lobe robust; margin between prostheca and mola straight; tuft of spine-like setae at base of mola absent; subtriangular process wide; denticles of mola not constricted; lateral margins convex. Right mandible (
<figureCitation id="13192A65A5711F6E0F15B328B9AB576B" captionStart="FIGURES 1116" captionStartId="7.[151,264,1945,1970]" captionTargetBox="[239,1349,202,1903]" captionTargetId="figure@7.[221,1366,181,1920]" captionTargetPageId="7" captionText="FIGURES 1116. Rivudiva oonirikoperi, sp. nov., nymph. (11) Labrum; (12) Left mandible; (13) Right mandible; (14) Hypopharynx; (15) Maxilla; (16) Labium (left = dorsal surface, right = ventral surface)." figureDoi="http://doi.org/10.5281/zenodo.3990790" httpUri="https://zenodo.org/record/3990790/files/figure.png" pageId="6" pageNumber="43">Fig. 13</figureCitation>
). Incisors cleft in two sets; outer and inner set of incisors respectively with 2 + 2 denticles and an outer spine-like process; prostheca stout, bifurcated at apex, inner lobe longer and pectinate; margin between prostheca and mola straight; tuft of spine-like setae at base of mola present; denticles of mola not constricted; apex of mola with two simple setae; lateral margins convex. Hypopharynx (
<figureCitation id="13192A65A5711F6E088FB3B8BA5657FF" box="[757,843,657,683]" captionStart="FIGURES 1116" captionStartId="7.[151,264,1945,1970]" captionTargetBox="[239,1349,202,1903]" captionTargetId="figure@7.[221,1366,181,1920]" captionTargetPageId="7" captionText="FIGURES 1116. Rivudiva oonirikoperi, sp. nov., nymph. (11) Labrum; (12) Left mandible; (13) Right mandible; (14) Hypopharynx; (15) Maxilla; (16) Labium (left = dorsal surface, right = ventral surface)." figureDoi="http://doi.org/10.5281/zenodo.3990790" httpUri="https://zenodo.org/record/3990790/files/figure.png" pageId="6" pageNumber="43">Fig. 14</figureCitation>
). Lingua longer than superlingua, sub-quadrangular with a distomedial pointed projection covered by tuft of simple setae; superlingua with rounded outer margin; short, thin, simple setae scattered over distal margin of lingua and superlingua. Maxilla (
<figureCitation id="13192A65A5711F6E0E3DB3F0BD8057A0" box="[1095,1181,729,756]" captionStart="FIGURES 1116" captionStartId="7.[151,264,1945,1970]" captionTargetBox="[239,1349,202,1903]" captionTargetId="figure@7.[221,1366,181,1920]" captionTargetPageId="7" captionText="FIGURES 1116. Rivudiva oonirikoperi, sp. nov., nymph. (11) Labrum; (12) Left mandible; (13) Right mandible; (14) Hypopharynx; (15) Maxilla; (16) Labium (left = dorsal surface, right = ventral surface)." figureDoi="http://doi.org/10.5281/zenodo.3990790" httpUri="https://zenodo.org/record/3990790/files/figure.png" pageId="6" pageNumber="43">Fig. 15</figureCitation>
). Maxillary palp long, 2.1× length of galea-lacinia; segment II 0.8× the length of segment I, apex with inner lobe; maxillary palp with thin simple setae scattered over surface; canines slender, dentisetae similar to canines. Labium (
<figureCitation id="13192A65A5711F6E0EEEB208BDFA5668" box="[1172,1255,801,828]" captionStart="FIGURES 1116" captionStartId="7.[151,264,1945,1970]" captionTargetBox="[239,1349,202,1903]" captionTargetId="figure@7.[221,1366,181,1920]" captionTargetPageId="7" captionText="FIGURES 1116. Rivudiva oonirikoperi, sp. nov., nymph. (11) Labrum; (12) Left mandible; (13) Right mandible; (14) Hypopharynx; (15) Maxilla; (16) Labium (left = dorsal surface, right = ventral surface)." figureDoi="http://doi.org/10.5281/zenodo.3990790" httpUri="https://zenodo.org/record/3990790/files/figure.png" pageId="6" pageNumber="43">Fig. 16</figureCitation>
). Glossa sub-elliptical, basally broad, narrowing apically and shorter than paraglossa; inner margin with row of spine-like setae on apical half; ventral surface covered by thin setae. Paraglossa curved inward; apex with two rows of robust and long spine-like setae; outer margin bare; dorsal surface with many long spine-like setae. Labial palp with segment I 0.9× the length of segments II and III combined; medial protuberance of segment II strongly projected medially, outer margin and distomedial protuberance covered by thin, long simple setae; segment III conical, length 1.1× width, covered by thin simple setae on outer margin, ventral surface with robust spine-like setae near outer margin.
</paragraph>
<paragraph id="8B9D36E0A5711F6E0ABDB534BB125044" blockId="6.[151,1437,225,1871]" pageId="6" pageNumber="43">
Thorax. Mesothorax with two large brown marks (
<figureCitation id="13192A65A5711F6E0979B534BA455163" box="[771,856,1053,1079]" captionStart="FIGURES 1721" captionStartId="8.[151,264,1945,1970]" captionTargetBox="[192,1396,189,1917]" captionTargetId="figure@8.[192,1396,181,1920]" captionTargetPageId="8" captionText="FIGURES 1721. Rivudiva oonirikoperi, sp. nov., nymph. (17) Fore leg (anterior surface); (18) detail of foreclaw; (19) mid leg (anterior surface); (20) Paraproct; (21) Dorsal view of body." figureDoi="http://doi.org/10.5281/zenodo.3990792" httpUri="https://zenodo.org/record/3990792/files/figure.png" pageId="6" pageNumber="43">Fig. 21</figureCitation>
). Hind wing pads absent. Foreleg (
<figureCitation id="13192A65A5711F6E0E9EB534BC255163" box="[1252,1336,1053,1079]" captionStart="FIGURES 1721" captionStartId="8.[151,264,1945,1970]" captionTargetBox="[192,1396,189,1917]" captionTargetId="figure@8.[192,1396,181,1920]" captionTargetPageId="8" captionText="FIGURES 1721. Rivudiva oonirikoperi, sp. nov., nymph. (17) Fore leg (anterior surface); (18) detail of foreclaw; (19) mid leg (anterior surface); (20) Paraproct; (21) Dorsal view of body." figureDoi="http://doi.org/10.5281/zenodo.3990792" httpUri="https://zenodo.org/record/3990792/files/figure.png" pageId="6" pageNumber="43">Fig. 17</figureCitation>
). Femur, tibia and tarsus light yellow. Ratio of foreleg 1.1:(1):0.6:0.17. Forefemur. Length about 2.6× maximum width; dorsal margin, anterior and posterior surface covered by robust long spine-like setae. Tibia. Dorsally bare; inner margin and anterior surface with few robust long spine-like setae. Tarsus. Ventral margin and anterior surface with robust long spine-like setae. Tarsal claws (
<figureCitation id="13192A65A5711F6E0852B584BB675193" box="[552,634,1197,1223]" captionStart="FIGURES 1721" captionStartId="8.[151,264,1945,1970]" captionTargetBox="[192,1396,189,1917]" captionTargetId="figure@8.[192,1396,181,1920]" captionTargetPageId="8" captionText="FIGURES 1721. Rivudiva oonirikoperi, sp. nov., nymph. (17) Fore leg (anterior surface); (18) detail of foreclaw; (19) mid leg (anterior surface); (20) Paraproct; (21) Dorsal view of body." figureDoi="http://doi.org/10.5281/zenodo.3990792" httpUri="https://zenodo.org/record/3990792/files/figure.png" pageId="6" pageNumber="43">Fig. 18</figureCitation>
) 0.3× the length of tarsus, with two rows of conical denticles. Mid and hind leg (
<figureCitation id="13192A65A5711F6E0B7EB5F8B84551BF" box="[260,344,1233,1259]" captionStart="FIGURES 1721" captionStartId="8.[151,264,1945,1970]" captionTargetBox="[192,1396,189,1917]" captionTargetId="figure@8.[192,1396,181,1920]" captionTargetPageId="8" captionText="FIGURES 1721. Rivudiva oonirikoperi, sp. nov., nymph. (17) Fore leg (anterior surface); (18) detail of foreclaw; (19) mid leg (anterior surface); (20) Paraproct; (21) Dorsal view of body." figureDoi="http://doi.org/10.5281/zenodo.3990792" httpUri="https://zenodo.org/record/3990792/files/figure.png" pageId="6" pageNumber="43">Fig. 19</figureCitation>
). Anterior surface of tibiae and tarsi without robust long setae; ventral margin of tibiae and tarsi with one row of short blunt setae.
</paragraph>
<paragraph id="8B9D36E0A5711F6E0ABDB430B89D50CB" blockId="6.[151,1437,225,1871]" pageId="6" pageNumber="43">
Abdomen. Terga pale yellow, I and II with two large brown marks (
<figureCitation id="13192A65A5711F6E09C9B430BD1B5067" box="[947,1030,1305,1331]" captionStart="FIGURES 1721" captionStartId="8.[151,264,1945,1970]" captionTargetBox="[192,1396,189,1917]" captionTargetId="figure@8.[192,1396,181,1920]" captionTargetPageId="8" captionText="FIGURES 1721. Rivudiva oonirikoperi, sp. nov., nymph. (17) Fore leg (anterior surface); (18) detail of foreclaw; (19) mid leg (anterior surface); (20) Paraproct; (21) Dorsal view of body." figureDoi="http://doi.org/10.5281/zenodo.3990792" httpUri="https://zenodo.org/record/3990792/files/figure.png" pageId="6" pageNumber="43">Fig. 21</figureCitation>
); surface covered by pores and short spine-like setae as in fig. 37; posterior margin smooth. Sterna white. Gills missing. Paraproct (
<figureCitation id="13192A65A5711F6E0EBDB414BC075003" box="[1223,1306,1341,1367]" captionStart="FIGURES 1721" captionStartId="8.[151,264,1945,1970]" captionTargetBox="[192,1396,189,1917]" captionTargetId="figure@8.[192,1396,181,1920]" captionTargetPageId="8" captionText="FIGURES 1721. Rivudiva oonirikoperi, sp. nov., nymph. (17) Fore leg (anterior surface); (18) detail of foreclaw; (19) mid leg (anterior surface); (20) Paraproct; (21) Dorsal view of body." figureDoi="http://doi.org/10.5281/zenodo.3990792" httpUri="https://zenodo.org/record/3990792/files/figure.png" pageId="6" pageNumber="43">Fig. 20</figureCitation>
) with three marginal spines, posterolateral extension without spines. Cerci with lateral spines on every segment. Terminal filament without spines.
</paragraph>
<paragraph id="8B9D36E0A5711F6E0ABDB483B9D7537B" blockId="6.[151,1437,225,1871]" pageId="6" pageNumber="43">
<emphasis id="B956EAF2A5711F6E0ABDB483B8555090" bold="true" box="[199,328,1450,1476]" pageId="6" pageNumber="43">Etymology</emphasis>
. Õonirikoperi is a word in Baniwa language, from Baniwa indigenous ethnicity, a group of Native American inhabiting the area where the species was collected. This word is attributed to any specimen of
<taxonomicName id="4C224D63A5711F6E0F3EB4E4B9EB5358" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="order">Ephemeroptera</taxonomicName>
. However, its literal meaning is “a being [malevolent] that walk inside the water”. The name is in apposition.
</paragraph>
<paragraph id="8B9D36E0A5711F6E0ABDB710BA7A53B7" blockId="6.[151,1437,225,1871]" pageId="6" pageNumber="43">
<emphasis id="B956EAF2A5711F6E0ABDB710B8555300" bold="true" box="[199,328,1593,1620]" pageId="6" pageNumber="43">Comments</emphasis>
.
<taxonomicName id="4C224D63A5711F6E0B2FB710BBE15307" authority="(Traver, 1943)" baseAuthorityName="Traver" baseAuthorityYear="1943" box="[341,764,1593,1619]" class="Insecta" family="Baetidae" genus="Rivudiva" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="venezuelensis">
<emphasis id="B956EAF2A5711F6E0B2FB710BB4B5307" box="[341,598,1593,1619]" italics="true" pageId="6" pageNumber="43">Rivudiva venezuelensis</emphasis>
(
<bibRefCitation id="EFB34B11A5711F6E081EB710BBE95307" author="Traver J. R." box="[612,756,1593,1619]" pageId="6" pageNumber="43" pagination="79 - 98" refId="ref8083" refString="Traver J. R. (1943) New Venezuelan mayflies. Boletin de Entomologia Venezolana, 2 (2), 79 - 98." type="journal article" year="1943">Traver, 1943</bibRefCitation>
)
</taxonomicName>
was described based solely on male and female imagos from
<collectingCountry id="F3357670A5711F6E0AEDB774B8175323" box="[151,266,1629,1655]" name="Venezuela" pageId="6" pageNumber="43">Venezuela</collectingCountry>
and since then no other material has been assigned to it. Given the absence of hind wings in this species and in
<taxonomicName id="4C224D63A5711F6E0A9DB7ABB88853CF" authority="Salles &amp; Nieto &amp; Cruz, 2020" authorityName="Salles &amp; Nieto &amp; Cruz" authorityYear="2020" box="[231,405,1665,1692]" class="Insecta" family="Baetidae" genus="Rivudiva" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="oonirikoperi" status="sp. nov.">
R.
<emphasis id="B956EAF2A5711F6E0B73B7A8B88853CF" box="[265,405,1665,1691]" italics="true" pageId="6" pageNumber="43">oonirikoperi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2655789A5711F6E0BDFB7ABBB1D53C8" box="[421,512,1666,1692]" pageId="6" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
, there is a possibility that this new species is in fact the nymphal stage, and thus represents a junior synonym of
<taxonomicName id="4C224D63A5711F6E0B89B78FBBB453EB" baseAuthorityName="Traver" baseAuthorityYear="1943" box="[499,681,1701,1727]" class="Insecta" family="Baetidae" genus="Rivudiva" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="venezuelensis">
<emphasis id="B956EAF2A5711F6E0B89B78FBBB453EB" box="[499,681,1701,1727]" italics="true" pageId="6" pageNumber="43">R. venezuelensis</emphasis>
</taxonomicName>
. Obtaining additional material from both species, including nymphs and reared adults, would be the best solution to solve this doubt.
</paragraph>
<paragraph id="8B9D36E0A5711F6E0ABDB7C4BAD65253" blockId="6.[151,1437,225,1871]" box="[199,971,1773,1800]" pageId="6" pageNumber="43">
<emphasis id="B956EAF2A5711F6E0ABDB7C4B844525C" bold="true" box="[199,345,1773,1800]" pageId="6" pageNumber="43">Distribution</emphasis>
.
<collectingCountry id="F3357670A5711F6E0B1DB7C4B8B35253" box="[359,430,1773,1799]" name="Brazil" pageId="6" pageNumber="43">Brazil</collectingCountry>
, Amazonas, São Gabriel da Cachoeira (
<figureCitation id="13192A65A5711F6E0911B7C4BAA35253" box="[875,958,1773,1799]" captionStart="FIGURE 41" captionStartId="13.[151,250,2017,2042]" captionTargetBox="[165,1418,319,1979]" captionTargetId="figure@13.[158,1430,309,1992]" captionTargetPageId="13" captionText="FIGURE 41. Map of South America showing the distribution of the distinct species of Rivudiva." figureDoi="http://doi.org/10.5281/zenodo.3990800" httpUri="https://zenodo.org/record/3990800/files/figure.png" pageId="6" pageNumber="43">Fig. 41</figureCitation>
).
</paragraph>
<paragraph id="8B9D36E0A5711F6E0ABDB638B9CD521B" blockId="6.[151,1437,225,1871]" pageId="6" pageNumber="43">
<emphasis id="B956EAF2A5711F6E0ABDB638B83C5278" bold="true" box="[199,289,1809,1836]" pageId="6" pageNumber="43">Biology</emphasis>
. Nymphs were collected on sand in a small (
<quantity id="4CDA9B05A5711F6E095BB638BA745278" box="[801,873,1809,1836]" metricMagnitude="-1" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="43" unit="cm" value="90.0">90 cm</quantity>
width) and well preserved stream with low water flow.
</paragraph>
</subSubSection>
</treatment>
</document>