419 lines
47 KiB
XML
419 lines
47 KiB
XML
<document id="57B4BEB16B58B42951D8D514AFE66A01" ID-DOI="10.5281/zenodo.193356" ID-GBIF-Dataset="25d23af4-b9c8-47be-b68e-33ae68076464" ID-ISSN="1175-5326" ID-Zenodo-Dep="193356" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460135747041" checkinUser="plazi" docAuthor="Tarasov, Sergey I. & Kabakov, Oleg N." docDate="2010" docId="038B87DEFFD81E38FF02FB253664F937" docLanguage="en" docName="zt02344p028.pdf" docOrigin="Zootaxa 2344" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Onthophagus (Sunenaga) streltsovi Tarasov & Kabakov, sp. n." docType="treatment" docVersion="10" lastPageNumber="21" masterDocId="FFB2FFA6FFD91E3CFF95FFD23424FFCF" masterDocTitle="Two new species of Onthophagus (Coleoptera: Scarabaeidae) from Indochina, with a discussion of some problems with the classification of Serrophorus and similar subgenera" masterLastPageNumber="28" masterPageNumber="17" pageNumber="18" updateTime="1698235207373" updateUser="plazi">
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<mods:mods id="0D651BC332EFE8C012347D2070DF78A0" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="F0FE3E003DF9D86D690AD4F35734218E">
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<mods:title id="8BFCC4FDA2E9332CBB13CD8D4093AD1E">Two new species of Onthophagus (Coleoptera: Scarabaeidae) from Indochina, with a discussion of some problems with the classification of Serrophorus and similar subgenera</mods:title>
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</mods:titleInfo>
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<mods:name id="8BD7F972CA9155BE15F9A03FF65C3594" type="personal">
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<mods:role id="6F7BB1AB22CA2E797FD8EC2B1CFD1F52">
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<mods:roleTerm id="7F337F365EC76C6769756259A418144F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F79EB5680354053CB8C36701BDD9C430">Tarasov, Sergey I.</mods:namePart>
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</mods:name>
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<mods:name id="21814688FD76613AA93370FE58A6483F" type="personal">
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<mods:roleTerm id="54DF8BEF586C673894039B50B381AF8A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BE701683D0FC68CFAFA925A034CC4972">Kabakov, Oleg N.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="E6AA76376C7993FAFD3459EAC70EF796">text</mods:typeOfResource>
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<mods:title id="365D9F99D6E22038B3AF1435E4D0FD36">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="77ACE92F08B3783A9306E34A53ACBA52">
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<mods:date id="A5EC137FD907123BD7DAB11D995FA45A">2010</mods:date>
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<mods:detail id="C83F0A583E6F54DF978FFD89C763C9FD" type="volume">
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<mods:number id="7F8733DE9BD48A84739050832ACE7B0F">2344</mods:number>
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<mods:start id="4B95A8D23AB58395441DE4A10C66CD2F">17</mods:start>
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<mods:classification id="3926BF4013D9699B2150A3319F3E0011">journal article</mods:classification>
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<mods:identifier id="5A15D1624B0A4A1FA94AE3007D507A41" type="DOI">10.5281/zenodo.193356</mods:identifier>
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<mods:identifier id="C3FC7A15EA4AA3BD58DB994682098C02" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="AB61E43B4E9E61661FB8F8C101337A16" type="Zenodo-Dep">193356</mods:identifier>
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<treatment id="038B87DEFFD81E38FF02FB253664F937" ID-DOI="http://doi.org/10.5281/zenodo.5687352" ID-GBIF-Taxon="119383043" ID-Zenodo-Dep="5687352" LSID="urn:lsid:plazi:treatment:038B87DEFFD81E38FF02FB253664F937" httpUri="http://treatment.plazi.org/id/038B87DEFFD81E38FF02FB253664F937" lastPageId="4" lastPageNumber="21" pageId="1" pageNumber="18">
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<subSubSection id="C3386543FFD81E3DFF02FB2535A7FAFB" pageId="1" pageNumber="18" type="nomenclature">
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<paragraph id="8B9D36C8FFD81E3DFF02FB2537B2FADE" blockId="1.[151,918,1271,1332]" box="[151,918,1271,1297]" pageId="1" pageNumber="18">
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<heading id="D0D581A4FFD81E3DFF02FB2537B2FADE" bold="true" box="[151,918,1271,1297]" fontSize="11" level="1" pageId="1" pageNumber="18" reason="1">
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<emphasis id="B956EADAFFD81E3DFF02FB2537B2FADE" bold="true" box="[151,918,1271,1297]" pageId="1" pageNumber="18">
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<taxonomicName id="4C224D4BFFD81E3DFF02FB253763FADE" ID-CoL="49MWK" authority="Tarasov & Kabakov" authorityName="Tarasov & Kabakov" box="[151,839,1271,1297]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="streltsovi" status="sp. nov." subGenus="Sunenaga">
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<emphasis id="B956EADAFFD81E3DFF02FB253518FADE" bold="true" box="[151,316,1271,1297]" italics="true" pageId="1" pageNumber="18">Onthophagus</emphasis>
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(
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<emphasis id="B956EADAFFD81E3DFED9FB2535E2FADE" bold="true" box="[332,454,1271,1297]" italics="true" pageId="1" pageNumber="18">Sunenaga</emphasis>
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)
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<emphasis id="B956EADAFFD81E3DFE42FB253666FADE" bold="true" box="[471,578,1271,1297]" italics="true" pageId="1" pageNumber="18">streltsovi</emphasis>
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||
Tarasov & Kabakov
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26557A1FFD81E3DFCC1FB2537B2FADE" box="[852,918,1271,1297]" pageId="1" pageNumber="18" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9D36C8FFD81E3DFF02FAC835A7FAFB" blockId="1.[151,918,1271,1332]" box="[151,387,1306,1332]" pageId="1" pageNumber="18">
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(
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<figureCitation id="13192A4DFFD81E3DFF0AFAC834CAFAFB" box="[159,238,1306,1332]" captionStart="FIGURE 1" captionStartId="2.[151,257,1711,1735]" captionTargetBox="[152,1414,315,1710]" captionTargetId="figure@2.[151,1436,269,1712]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Onthophagus. a, b— O. streltsovi; c, d— O. nampatensis; a, c—males, holotypes; b—female, paratype; d—small male paratype." httpUri="https://zenodo.org/record/193357/files/figure.png" pageId="1" pageNumber="18">Figs. 1</figureCitation>
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a, b; 2c, e, f)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3386543FFD81E3EFF02FAB7300CFF33" lastPageId="2" lastPageNumber="19" pageId="1" pageNumber="18" type="materials_examined">
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||
<paragraph id="8B9D36C8FFD81E3DFF02FAB73028FAB0" blockId="1.[151,1437,1381,2004]" box="[151,1036,1381,1407]" pageId="1" pageNumber="18">
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||
<emphasis id="B956EADAFFD81E3DFF02FAB735AFFAB0" bold="true" box="[151,395,1381,1407]" pageId="1" pageNumber="18">Material examined.</emphasis>
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||
<emphasis id="B956EADAFFD81E3DFE07FAB735DAFAB1" box="[402,510,1381,1406]" italics="true" pageId="1" pageNumber="18">
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<typeStatus id="5499886AFFD81E3DFE07FAB735DAFAB1" box="[402,510,1381,1406]" pageId="1" pageNumber="18" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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||
(
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<collectionCode id="ED33AE0DFFD81E3DFD98FAB7364CFAB0" box="[525,616,1381,1407]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="1" pageNumber="18">ZMUC</collectionCode>
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), male bearing the following labels:
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</paragraph>
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<paragraph id="8B9D36C8FFD81E3DFF51FA5837B0FA05" blockId="1.[151,1437,1381,2004]" pageId="1" pageNumber="18">
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1. White, printed: S
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<collectingCountry id="F3357658FFD81E3DFE2AFA583635FA6B" box="[447,529,1418,1444]" name="Laos" pageId="1" pageNumber="18">LAOS</collectingCountry>
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, Bolaven plateau B.Thongvai,
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<geoCoordinate id="EE16500FFFD81E3DFC18FA583003FA6B" box="[909,1063,1418,1444]" direction="north" orientation="latitude" pageId="1" pageNumber="18" precision="1" value="15.245684">N15°14.741'</geoCoordinate>
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<geoCoordinate id="EE16500FFFD81E3DFBBAFA5830F2FA6B" box="[1071,1238,1418,1444]" direction="east" orientation="longitude" pageId="1" pageNumber="18" precision="1" value="106.53194">E106°31.916'</geoCoordinate>
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h=
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||
<quantity id="4CDA9B2DFFD81E3DFB68FA5831B8FA6B" box="[1277,1436,1418,1444]" metricMagnitude="3" metricUnit="m" metricValue="1.225" metricValueMax="1.3" metricValueMin="1.15" pageId="1" pageNumber="18" unit="m" value="1225.0" valueMax="1300.0" valueMin="1150.0">1150–1300m</quantity>
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||
dung trap, primary montane forest, 8–
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<date id="FF9C1008FFD81E3DFDC7FA6236FEFA05" box="[594,730,1456,1482]" pageId="1" pageNumber="18" value="2008-06-16">16.06.2008</date>
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||
, leg. S. Tarasov
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||
</paragraph>
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||
<paragraph id="8B9D36C8FFD81E3DFF51FA073133FA20" blockId="1.[151,1437,1381,2004]" box="[196,1303,1493,1519]" pageId="1" pageNumber="18">
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2. Orange, printed:
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<typeStatus id="5499886AFFD81E3DFE3DFA073673FA20" box="[424,599,1493,1519]" pageId="1" pageNumber="18" type="holotype">HOLOTYPUS</typeStatus>
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||
<taxonomicName id="4C224D4BFFD81E3DFDCBFA073087FA20" authority="S. Tarasov & O. Kabakov" authorityName="S. Tarasov & O. Kabakov" box="[606,1187,1493,1519]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="streltsovi">
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||
<emphasis id="B956EADAFFD81E3DFDCBFA073749FA21" box="[606,877,1493,1518]" italics="true" pageId="1" pageNumber="18">Onthophagus streltsovi</emphasis>
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S. Tarasov & O. Kabakov
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||
</taxonomicName>
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||
det. 2009
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||
</paragraph>
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||
<paragraph id="8B9D36C8FFD81E3DFF51FA2931B8F81B" blockId="1.[151,1437,1381,2004]" pageId="1" pageNumber="18">
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<emphasis id="B956EADAFFD81E3DFF51FA293564F9DB" box="[196,320,1531,1556]" italics="true" pageId="1" pageNumber="18">
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<typeStatus id="5499886AFFD81E3DFF51FA293564F9DB" box="[196,320,1531,1556]" pageId="1" pageNumber="18" type="paratype">Paratypes</typeStatus>
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||
</emphasis>
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||
.
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||
<specimenCount id="9D24FD41FFD81E3DFEC4FA28362FF9DB" box="[337,523,1530,1556]" pageId="1" pageNumber="18" type="generic">147 specimens</specimenCount>
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||
, South
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||
<collectingCountry id="F3357658FFD81E3DFDFCFA28368CF9DB" box="[617,680,1530,1556]" name="Laos" pageId="1" pageNumber="18">Laos</collectingCountry>
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||
, Bolaven Plateau, B.Thongvai,
|
||
<geoCoordinate id="EE16500FFFD81E3DFBA4FA2830EAF9DB" box="[1073,1230,1530,1556]" direction="north" orientation="latitude" pageId="1" pageNumber="18" precision="1" value="15.245684">N15°14.741'</geoCoordinate>
|
||
<geoCoordinate id="EE16500FFFD81E3DFB40FA2831A5F9DB" box="[1237,1409,1530,1556]" direction="east" orientation="longitude" pageId="1" pageNumber="18" precision="1" value="106.53194">E106°31.916'</geoCoordinate>
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||
, h
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||
<quantity id="4CDA9B2DFFD81E3DFF02F9F23564F9F5" box="[151,320,1568,1594]" metricMagnitude="3" metricUnit="m" metricValue="1.225" metricValueMax="1.3" metricValueMin="1.15" pageId="1" pageNumber="18" unit="m" value="1225.0" valueMax="1300.0" valueMin="1150.0">1150–1300m</quantity>
|
||
, dung trap (human), primary montane forest, 8–
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||
<date id="FF9C1008FFD81E3DFC3CF9F2301EF9F5" box="[937,1082,1568,1594]" pageId="1" pageNumber="18" value="2008-06-16">16.06.2008</date>
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||
leg. S. Tarasov (
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||
<collectionCode id="ED33AE0DFFD81E3DFA80F9F23153F9F5" box="[1301,1399,1568,1594]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="1" pageNumber="18">ZMUC</collectionCode>
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||
);
|
||
<specimenCount id="9D24FD41FFD81E3DFA18F9F23529F990" pageId="1" pageNumber="18" type="generic">1 specimen</specimenCount>
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||
, South
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||
<collectingCountry id="F3357658FFD81E3DFEFFF997358CF990" box="[362,424,1605,1631]" name="Laos" pageId="1" pageNumber="18">Laos</collectingCountry>
|
||
, Bolaven Plateau, B.Thongvai,
|
||
<geoCoordinate id="EE16500FFFD81E3DFCA5F99737EFF990" box="[816,971,1605,1631]" direction="north" orientation="latitude" pageId="1" pageNumber="18" precision="1" value="15.245684">N15°14.741'</geoCoordinate>
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||
<geoCoordinate id="EE16500FFFD81E3DFC47F9973059F990" box="[978,1149,1605,1631]" direction="east" orientation="longitude" pageId="1" pageNumber="18" precision="1" value="106.53194">E106°31.916'</geoCoordinate>
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, h
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<quantity id="4CDA9B2DFFD81E3DFB34F99730DCF990" box="[1185,1272,1605,1631]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="18" unit="m" value="1300.0">1300m</quantity>
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||
, carrion trap, primary montane forest, 12–
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||
<date id="FF9C1008FFD81E3DFE73F9B8364AF94B" box="[486,622,1642,1668]" pageId="1" pageNumber="18" value="2008-06-16">16.06.2008</date>
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, leg. S. Tarasov (
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||
<collectionCode id="ED33AE0DFFD81E3DFCAFF9B837B1F94B" box="[826,917,1642,1668]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="1" pageNumber="18">ZMUC</collectionCode>
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);
|
||
<specimenCount id="9D24FD41FFD81E3DFC3EF9B83017F94B" box="[939,1075,1642,1668]" pageId="1" pageNumber="18" type="generic">1 specimen</specimenCount>
|
||
, South
|
||
<collectingCountry id="F3357658FFD81E3DFB19F9B830ECF94B" box="[1164,1224,1642,1668]" name="Laos" pageId="1" pageNumber="18">Laos</collectingCountry>
|
||
, Bolaven Plateau, B.Thongvai,
|
||
<geoCoordinate id="EE16500FFFD81E3DFEA5F94235E1F965" box="[304,453,1680,1706]" direction="north" orientation="latitude" pageId="1" pageNumber="18" precision="1" value="15.245684">N15°14.741'</geoCoordinate>
|
||
<geoCoordinate id="EE16500FFFD81E3DFE5EF942364AF965" box="[459,622,1680,1706]" direction="east" orientation="longitude" pageId="1" pageNumber="18" precision="1" value="106.53194">E106°31.916'</geoCoordinate>
|
||
, h
|
||
<quantity id="4CDA9B2DFFD81E3DFD18F94236C4F965" box="[653,736,1680,1706]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="1" pageNumber="18" unit="m" value="1150.0">1150m</quantity>
|
||
, carrion trap, disturb montane forest, 10–
|
||
<date id="FF9C1008FFD81E3DFB2AF9423163F965" box="[1215,1351,1680,1706]" pageId="1" pageNumber="18" value="2008-06-16">16.06.2008</date>
|
||
, leg. S. Tarasov (
|
||
<collectionCode id="ED33AE0DFFD81E3DFE93F9673547F900" box="[262,355,1717,1743]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="1" pageNumber="18">ZMUC</collectionCode>
|
||
);
|
||
<specimenCount id="9D24FD41FFD81E3DFEEDF967363AF900" box="[376,542,1717,1743]" pageId="1" pageNumber="18" type="generic">81 specimens</specimenCount>
|
||
,
|
||
<collectingCountry id="F3357658FFD81E3DFDBEF967364EF900" box="[555,618,1717,1743]" name="Laos" pageId="1" pageNumber="18">Laos</collectingCountry>
|
||
: Champasak province: Bolaven Plateau, Ban Thongvai, old logging road N of the village,
|
||
<quantity id="4CDA9B2DFFD81E3DFE2DF9083637F93B" box="[440,531,1754,1780]" metricMagnitude="3" metricUnit="m" metricValue="1.095" pageId="1" pageNumber="18" unit="m" value="1095.0">1095m</quantity>
|
||
,
|
||
<geoCoordinate id="EE16500FFFD81E3DFDB4F90836E2F93B" box="[545,710,1754,1780]" direction="north" orientation="latitude" pageId="1" pageNumber="18" precision="1" value="15.238133">15°14.288'N</geoCoordinate>
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,
|
||
<geoCoordinate id="EE16500FFFD81E3DFD41F90837A1F93B" box="[724,901,1754,1780]" direction="east" orientation="longitude" pageId="1" pageNumber="18" precision="1" value="106.53152">106°31.891'E</geoCoordinate>
|
||
, selectively logged forest, 8–
|
||
<date id="FF9C1008FFD81E3DFA97F90831BCF93B" box="[1282,1432,1754,1780]" pageId="1" pageNumber="18" value="2008-06-16">16.06.2008</date>
|
||
;
|
||
<collectionCode id="ED33AE0DFFD81E3DFF02F8D234CBF8D5" box="[151,239,1792,1818]" pageId="1" pageNumber="18">FMHD</collectionCode>
|
||
#2008-039, dung trap (human), A. Solodovnikov, M. Thayer & A. Newton (
|
||
<collectionCode id="ED33AE0DFFD81E3DFBE0F8D230F5F8D5" LSID="urn:lsid:biocol.org:col:34795" box="[1141,1233,1792,1818]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="1" pageNumber="18">FMNH</collectionCode>
|
||
);
|
||
<specimenCount id="9D24FD41FFD81E3DFB73F8D231B3F8D5" box="[1254,1431,1792,1818]" pageId="1" pageNumber="18" type="generic">134 specimens</specimenCount>
|
||
,
|
||
<collectingCountry id="F3357658FFD81E3DFF02F8F734FCF8F0" box="[151,216,1829,1855]" name="Laos" pageId="1" pageNumber="18">Laos</collectingCountry>
|
||
: Champasak province: Bolaven Plateau, Ban Thongvai, old logging road N of the village,
|
||
<quantity id="4CDA9B2DFFD81E3DFAD4F8F731B3F8F0" box="[1345,1431,1829,1855]" metricMagnitude="3" metricUnit="m" metricValue="1.17" pageId="1" pageNumber="18" unit="m" value="1170.0">1170m</quantity>
|
||
,
|
||
<geoCoordinate id="EE16500FFFD81E3DFF02F898350BF8AB" box="[151,303,1866,1892]" direction="north" orientation="latitude" pageId="1" pageNumber="18" precision="1" value="15.241567">15°14.494'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE16500FFFD81E3DFEA9F89835C4F8AB" box="[316,480,1866,1892]" direction="east" orientation="longitude" pageId="1" pageNumber="18" precision="1" value="106.53011">106°31.807'E</geoCoordinate>
|
||
, selectively logged forest, 8–
|
||
<date id="FF9C1008FFD81E3DFCAEF89837E1F8AB" box="[827,965,1866,1892]" pageId="1" pageNumber="18" value="2008-06-16">16.06.2008</date>
|
||
;
|
||
<collectionCode id="ED33AE0DFFD81E3DFC47F898300EF8AB" box="[978,1066,1866,1892]" pageId="1" pageNumber="18">FMHD</collectionCode>
|
||
#2008-042, dung trap (human), A. Solodovnikov, M. Thayer & A. Newton (
|
||
<collectionCode id="ED33AE0DFFD81E3DFD26F8A23735F845" LSID="urn:lsid:biocol.org:col:34795" box="[691,785,1904,1930]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="1" pageNumber="18">FMNH</collectionCode>
|
||
);
|
||
<specimenCount id="9D24FD41FFD81E3DFCB2F8A237EAF845" box="[807,974,1904,1930]" pageId="1" pageNumber="18" type="generic">23 specimens</specimenCount>
|
||
,
|
||
<collectingCountry id="F3357658FFD81E3DFC49F8A23038F845" box="[988,1052,1904,1930]" name="Laos" pageId="1" pageNumber="18">Laos</collectingCountry>
|
||
: Champasak province: Bolaven Plateau, Ban Thongvai, old logging road N of the village,
|
||
<quantity id="4CDA9B2DFFD81E3DFCF1F8473798F860" box="[868,956,1941,1967]" metricMagnitude="3" metricUnit="m" metricValue="1.035" pageId="1" pageNumber="18" unit="m" value="1035.0">1035m</quantity>
|
||
,
|
||
<geoCoordinate id="EE16500FFFD81E3DFC5CF8473073F860" box="[969,1111,1941,1967]" direction="north" orientation="latitude" pageId="1" pageNumber="18" precision="9" value="15.232667">15°13.96'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE16500FFFD81E3DFBF6F8473128F860" box="[1123,1292,1941,1967]" direction="east" orientation="longitude" pageId="1" pageNumber="18" precision="1" value="106.52885">106°31.731'E</geoCoordinate>
|
||
, selectively logged forest, 8–
|
||
<date id="FF9C1008FFD81E3DFEFCF86835DEF81B" box="[361,506,1978,2004]" pageId="1" pageNumber="18" value="2008-06-16">16.06.2008</date>
|
||
;
|
||
<collectionCode id="ED33AE0DFFD81E3DFD9DF8683640F81B" box="[520,612,1978,2004]" pageId="1" pageNumber="18">FMHD</collectionCode>
|
||
#2008-036, dung trap (human), A. Solodovnikov, M. Thayer & A.
|
||
</paragraph>
|
||
<paragraph id="8B9D36C8FFDB1E3EFF02FF4A300CFF33" blockId="2.[151,1436,152,252]" pageId="2" pageNumber="19">
|
||
Newton (
|
||
<collectionCode id="ED33AE0DFFDB1E3EFE97FF4A3579FF7D" LSID="urn:lsid:biocol.org:col:34795" box="[258,349,152,178]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="2" pageNumber="19">FMNH</collectionCode>
|
||
);
|
||
<specimenCount id="9D24FD41FFDB1E3EFEE4FF4A3620FF7D" box="[369,516,152,178]" pageId="2" pageNumber="19" type="generic">6 specimens</specimenCount>
|
||
,
|
||
<collectingCountry id="F3357658FFDB1E3EFD9BFF4A3668FF7D" box="[526,588,152,178]" name="Laos" pageId="2" pageNumber="19">Laos</collectingCountry>
|
||
: Champasak province: Bolaven Plateau, Ban Thongvai, old logging road N of the village,
|
||
<quantity id="4CDA9B2DFFDB1E3EFEF7FF6F358DFF18" box="[354,425,189,215]" metricMagnitude="2" metricUnit="m" metricValue="9.95" pageId="2" pageNumber="19" unit="m" value="995.0">995m</quantity>
|
||
,
|
||
<geoCoordinate id="EE16500FFFDB1E3EFE20FF6F366AFF18" box="[437,590,189,215]" direction="north" orientation="latitude" pageId="2" pageNumber="19" precision="1" value="15.22935">15°13.761'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE16500FFFDB1E3EFDCEFF6F36DBFF18" box="[603,767,189,215]" direction="east" orientation="longitude" pageId="2" pageNumber="19" precision="1" value="106.52915">106°31.749'E</geoCoordinate>
|
||
, selectively logged forest, 8–
|
||
<date id="FF9C1008FFDB1E3EFBCEFF6F30C1FF18" box="[1115,1253,189,215]" pageId="2" pageNumber="19" value="2008-06-16">16.06.2008</date>
|
||
;
|
||
<collectionCode id="ED33AE0DFFDB1E3EFB66FF6F316EFF18" box="[1267,1354,189,215]" pageId="2" pageNumber="19">FMHD</collectionCode>
|
||
#2008- 0 33, dung trap (human), A. Solodovnikov, M. Thayer & A. Newton (
|
||
<collectionCode id="ED33AE0DFFDB1E3EFC55FF303038FF33" LSID="urn:lsid:biocol.org:col:34795" box="[960,1052,226,252]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="2" pageNumber="19">FMNH</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5D6640FFDB1E3EFF02F97D3623F929" httpUri="https://zenodo.org/record/193357/files/figure.png" pageId="2" pageNumber="19" targetBox="[152,1414,315,1710]" targetPageId="2">
|
||
<paragraph id="8B9D36C8FFDB1E3EFF02F97D3623F929" blockId="2.[151,1436,1711,1766]" pageId="2" pageNumber="19">
|
||
<emphasis id="B956EADAFFDB1E3EFF02F97D3538F908" bold="true" box="[151,284,1711,1735]" pageId="2" pageNumber="19">FIGURE 1.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="4C224D4BFFDB1E3EFE0AF97D3616F909" box="[415,562,1711,1734]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B956EADAFFDB1E3EFE0AF97D3616F909" box="[415,562,1711,1734]" italics="true" pageId="2" pageNumber="19">Onthophagus</emphasis>
|
||
</taxonomicName>
|
||
. a, b—
|
||
<taxonomicName id="4C224D4BFFDB1E3EFD10F97D372FF909" box="[645,779,1711,1734]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="streltsovi">
|
||
<emphasis id="B956EADAFFDB1E3EFD10F97D372FF909" box="[645,779,1711,1734]" italics="true" pageId="2" pageNumber="19">O. streltsovi</emphasis>
|
||
</taxonomicName>
|
||
; c, d—
|
||
<taxonomicName id="4C224D4BFFDB1E3EFCC8F97D302CF909" box="[861,1032,1711,1734]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="nampatensis">
|
||
<emphasis id="B956EADAFFDB1E3EFCC8F97D302CF909" box="[861,1032,1711,1734]" italics="true" pageId="2" pageNumber="19">O. nampatensis</emphasis>
|
||
</taxonomicName>
|
||
; a, c—males, holotypes; b—female, paratype; d—small male paratype.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3386543FFDB1E38FF51F8C631B8FF33" lastPageId="4" lastPageNumber="21" pageId="2" pageNumber="19" type="description">
|
||
<paragraph id="8B9D36C8FFDB1E3EFF51F8C637F9F80B" blockId="2.[151,1437,1812,1988]" pageId="2" pageNumber="19">
|
||
<emphasis id="B956EADAFFDB1E3EFF51F8C63579F8E1" bold="true" box="[196,349,1812,1838]" pageId="2" pageNumber="19">Description.</emphasis>
|
||
Head and pronotum black with slight green luster, rarely slightly red; elytra yellow-brown, intervals with black, interrupted stripes; rarely stripe pattern weakly expressed; pygidium, metasternum laterally, femur yellow or yellow-black; abdominal sternites yellow usually with black stripes basally; antennal club yellow; elytra with short, black setae turning yellow depending on light angle; pronotum, head, pygidium, abdominal side with longer yellow setae. Length
|
||
<quantity id="4CDA9B2DFFDB1E3EFCC2F87837FEF80B" box="[855,986,1962,1988]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="2" pageNumber="19" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10–12 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9D36C8FFDA1E3FFF51FF4A37D8FEED" blockId="3.[151,1437,152,703]" pageId="3" pageNumber="20">
|
||
<emphasis id="B956EADAFFDA1E3FFF51FF4A3574FF7E" box="[196,336,152,177]" italics="true" pageId="3" pageNumber="20">Male major</emphasis>
|
||
(
|
||
<figureCitation id="13192A4DFFDA1E3FFEF4FF4A3581FF7D" box="[353,421,152,178]" captionStart="FIGURE 1" captionStartId="2.[151,257,1711,1735]" captionTargetBox="[152,1414,315,1710]" captionTargetId="figure@2.[151,1436,269,1712]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Onthophagus. a, b— O. streltsovi; c, d— O. nampatensis; a, c—males, holotypes; b—female, paratype; d—small male paratype." httpUri="https://zenodo.org/record/193357/files/figure.png" pageId="3" pageNumber="20">Fig. 1</figureCitation>
|
||
a). Head rounded; apex of clypeal margin anteriorly reflexed upwards; frontoclypeal ridge absent in larger males and present with slightly raised strip in smaller males; vertex with small hookshaped horn broadened basally and bent backwards; eyes completely divided with genal appendage; clypeus sparsely punctate; antennal scapus anteriorly with slight ridge, not serrate.
|
||
</paragraph>
|
||
<paragraph id="8B9D36C8FFDA1E3FFF51FEFF3014FE78" blockId="3.[151,1437,152,703]" pageId="3" pageNumber="20">Pronotum convex; anterior, lateral and basal sides marginate, postlateral section strongly sinuate; anterolateral angles rounded, anterior part slightly concave in middle behind cephalic horn; disc sparsely covered with fine punctures (separated by 1–2 puncture diameters) becoming denser and larger laterally; anterior part of propleura with ridge almost reaching propleural lateral margin.</paragraph>
|
||
<paragraph id="8B9D36C8FFDA1E3FFF51FE17362FFDE5" blockId="3.[151,1437,152,703]" pageId="3" pageNumber="20">Elytra densely covered with scattered punctures becoming denser on 7th and 8th intervals. Pygidium covered with scattered large punctures separated by 1–2 diameters of a puncture, with longitudinal black strip medially; basal edge marginate.</paragraph>
|
||
<paragraph id="8B9D36C8FFDA1E3FFF51FDE73751FD55" blockId="3.[151,1437,152,703]" pageId="3" pageNumber="20">Protibia with four outer teeth, internal apical angle with large tooth about two times smaller than apical spur; apical spur modified in males: small, acuminate, slightly bent outward; mesotibial and metatibial apex produced into three distinct lobes bearing short and long setae.</paragraph>
|
||
<paragraph id="8B9D36C8FFDA1E3FFF51FD773758FD70" blockId="3.[151,1437,152,703]" box="[196,892,677,703]" pageId="3" pageNumber="20">
|
||
Aedeagus as in
|
||
<figureCitation id="13192A4DFFDA1E3FFEE9FD7735EAFD70" box="[380,462,677,703]" captionStart="FIGURE 2" captionStartId="3.[151,255,1728,1752]" captionTargetBox="[190,1401,755,1690]" captionTargetId="figure@3.[151,1436,713,1730]" captionTargetPageId="3" captionText="FIGURE 2. Elements of genitalia of Onthophagus. a— O. seniculus (fused area of SL and SR indicated by arrow); b— O. anguliceps (separation of SL and SR indicated by arrow); c, e, f— O. streltsovi; d— O. manipurensis; a – d—lamella copulatrix; e—aedeagus, lateral view; f—aedeagus, apical view." httpUri="https://zenodo.org/record/193358/files/figure.png" pageId="3" pageNumber="20">Figs. 2</figureCitation>
|
||
e, f. Lamella copulatrix as in
|
||
<figureCitation id="13192A4DFFDA1E3FFCB6FD77374EFD70" box="[803,874,677,703]" captionStart="FIGURE 2" captionStartId="3.[151,255,1728,1752]" captionTargetBox="[190,1401,755,1690]" captionTargetId="figure@3.[151,1436,713,1730]" captionTargetPageId="3" captionText="FIGURE 2. Elements of genitalia of Onthophagus. a— O. seniculus (fused area of SL and SR indicated by arrow); b— O. anguliceps (separation of SL and SR indicated by arrow); c, e, f— O. streltsovi; d— O. manipurensis; a – d—lamella copulatrix; e—aedeagus, lateral view; f—aedeagus, apical view." httpUri="https://zenodo.org/record/193358/files/figure.png" pageId="3" pageNumber="20">Fig. 2</figureCitation>
|
||
c.
|
||
</paragraph>
|
||
<caption id="DF5D6640FFDA1E3FFF02F9123760F8D8" httpUri="https://zenodo.org/record/193358/files/figure.png" pageId="3" pageNumber="20" targetBox="[190,1401,755,1690]" targetPageId="3">
|
||
<paragraph id="8B9D36C8FFDA1E3FFF02F9123760F8D8" blockId="3.[151,1436,1728,1815]" pageId="3" pageNumber="20">
|
||
<emphasis id="B956EADAFFDA1E3FFF02F912353CF917" bold="true" box="[151,280,1728,1752]" pageId="3" pageNumber="20">FIGURE 2.</emphasis>
|
||
Elements of genitalia of
|
||
<taxonomicName id="4C224D4BFFDA1E3FFDB7F9123696F918" box="[546,690,1728,1751]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B956EADAFFDA1E3FFDB7F9123696F918" box="[546,690,1728,1751]" italics="true" pageId="3" pageNumber="20">Onthophagus</emphasis>
|
||
</taxonomicName>
|
||
. a—
|
||
<taxonomicName id="4C224D4BFFDA1E3FFD71F9123743F918" box="[740,871,1728,1751]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="seniculus">
|
||
<emphasis id="B956EADAFFDA1E3FFD71F9123743F918" box="[740,871,1728,1751]" italics="true" pageId="3" pageNumber="20">O. seniculus</emphasis>
|
||
</taxonomicName>
|
||
(fused area of SL and SR indicated by arrow); b—
|
||
<taxonomicName id="4C224D4BFFDA1E3FFA17F912352BF938" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="anguliceps">
|
||
<emphasis id="B956EADAFFDA1E3FFA17F912352BF938" italics="true" pageId="3" pageNumber="20">O. anguliceps</emphasis>
|
||
</taxonomicName>
|
||
(separation of SL and SR indicated by arrow); c, e, f—
|
||
<taxonomicName id="4C224D4BFFDA1E3FFC15F932302CF938" box="[896,1032,1760,1783]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="streltsovi">
|
||
<emphasis id="B956EADAFFDA1E3FFC15F932302CF938" box="[896,1032,1760,1783]" italics="true" pageId="3" pageNumber="20">O. streltsovi</emphasis>
|
||
</taxonomicName>
|
||
; d—
|
||
<taxonomicName id="4C224D4BFFDA1E3FFBD4F93230DCF938" box="[1089,1272,1760,1783]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="manipurensis">
|
||
<emphasis id="B956EADAFFDA1E3FFBD4F93230DCF938" box="[1089,1272,1760,1783]" italics="true" pageId="3" pageNumber="20">O. manipurensis</emphasis>
|
||
</taxonomicName>
|
||
; a–d—lamella copulatrix; e—aedeagus, lateral view; f—aedeagus, apical view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9D36C8FFDA1E3FFF51F894359FF83A" blockId="3.[151,1437,1862,2037]" pageId="3" pageNumber="20">
|
||
<emphasis id="B956EADAFFDA1E3FFF51F8943538F890" box="[196,284,1862,1887]" italics="true" pageId="3" pageNumber="20">Female</emphasis>
|
||
(
|
||
<figureCitation id="13192A4DFFDA1E3FFEB9F894354AF8AF" box="[300,366,1862,1888]" captionStart="FIGURE 1" captionStartId="2.[151,257,1711,1735]" captionTargetBox="[152,1414,315,1710]" captionTargetId="figure@2.[151,1436,269,1712]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Onthophagus. a, b— O. streltsovi; c, d— O. nampatensis; a, c—males, holotypes; b—female, paratype; d—small male paratype." httpUri="https://zenodo.org/record/193357/files/figure.png" pageId="3" pageNumber="20">Fig. 1</figureCitation>
|
||
b). Female differs from male by clypeus slightly reflexed along anterior margin and incised medially, with each side of the incision lobed; frontoclypeal ridge present; vertex without horn; head and pronotum sparsely covered with large, dense punctures separated by a diameter or less; pronotum less convex; internal apical angle of protibia with small indistinctive tooth; apical spur of protibia larger, more acuminate and slightly bent inward.
|
||
</paragraph>
|
||
<paragraph id="8B9D36C8FFDD1E38FF51FF4A31B8FF33" blockId="4.[151,1437,152,962]" pageId="4" pageNumber="21">
|
||
<emphasis id="B956EADAFFDD1E38FF51FF4A3515FF7E" box="[196,305,152,177]" italics="true" pageId="4" pageNumber="21">Var i at io n</emphasis>
|
||
. Males vary allometrically in the degree of expression of some features of the clypeus, horn, pronotum and punctation. Small males usually have less anteriorly reflexed clypeus; vertex not armed with horn, or, sometimes, only with small tubercle; pronotum more flat, not concave, disc more densely punctured.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3386543FFDD1E38FF51FEDA36E1F9D7" pageId="4" pageNumber="21" type="materials_examined">
|
||
<paragraph id="8B9D36C8FFDD1E38FF51FEDA309AFE13" blockId="4.[151,1437,152,962]" pageId="4" pageNumber="21">
|
||
<emphasis id="B956EADAFFDD1E38FF51FEDA350BFEEE" box="[196,303,264,289]" italics="true" pageId="4" pageNumber="21">
|
||
<typeStatus id="5499886AFFDD1E38FF51FEDA350BFEEE" box="[196,303,264,289]" pageId="4" pageNumber="21" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13192A4DFFDD1E38FEAAFEDA35A5FEED" box="[319,385,264,290]" captionStart="FIGURE 1" captionStartId="2.[151,257,1711,1735]" captionTargetBox="[152,1414,315,1710]" captionTargetId="figure@2.[151,1436,269,1712]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Onthophagus. a, b— O. streltsovi; c, d— O. nampatensis; a, c—males, holotypes; b—female, paratype; d—small male paratype." httpUri="https://zenodo.org/record/193357/files/figure.png" pageId="4" pageNumber="21">Fig. 1</figureCitation>
|
||
a). Major male, length
|
||
<quantity id="4CDA9B2DFFDD1E38FD05FEDA36D8FEED" box="[656,764,264,290]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="4" pageNumber="21" unit="mm" value="11.4">11.4 mm</quantity>
|
||
. Head and pronotum black with slight green luster; elytra yellow-brown, intervals with black, interrupted stripes; pygidium yellow-black; metasternum laterally and femur yellow; abdominal sternites yellow with black stripes basally, antennal club yellow; apex of clypeal margin anteriorly reflexed upwards; vertex with small hook-shaped horn broadened basally and bent backwards; Pronotum convex, slightly concave in middle behind cephalic horn; disc sparsely covered with fine punctures (separated by 1–2 puncture diameters) becoming denser and larger laterally.
|
||
</paragraph>
|
||
<paragraph id="8B9D36C8FFDD1E38FF51FE35373CFC0D" blockId="4.[151,1437,152,962]" pageId="4" pageNumber="21">
|
||
<emphasis id="B956EADAFFDD1E38FF51FE3535FDFDCE" bold="true" box="[196,473,487,513]" pageId="4" pageNumber="21">Differential diagnosis.</emphasis>
|
||
This species is similar to
|
||
<taxonomicName id="4C224D4BFFDD1E38FC87FE3A3018FDCD" authority="Arrow, 1931" authorityName="Arrow" authorityYear="1931" box="[786,1084,488,514]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="digitatus">
|
||
<emphasis id="B956EADAFFDD1E38FC87FE3A3703FDCE" box="[786,807,488,513]" italics="true" pageId="4" pageNumber="21">O</emphasis>
|
||
.
|
||
<emphasis id="B956EADAFFDD1E38FCA3FE3A37BAFDCE" box="[822,926,488,513]" italics="true" pageId="4" pageNumber="21">digitatus</emphasis>
|
||
<bibRefCitation id="EFB34B39FFDD1E38FC33FE3A3018FDCD" author="Arrow" box="[934,1084,488,514]" pageId="4" pageNumber="28" refString="Arrow, G. J. (1931) The Fauna of British India III, Coprinae. Taylor & Francis, London, 428 pp." type="book" year="1931">Arrow, 1931</bibRefCitation>
|
||
</taxonomicName>
|
||
from Sikkim,
|
||
<collectingCountry id="F3357658FFDD1E38FB7EFE3A310CFDCD" box="[1259,1320,488,514]" name="India" pageId="4" pageNumber="21">India</collectingCountry>
|
||
(we have examined the male
|
||
<typeStatus id="5499886AFFDD1E38FE15FDDF35C9FDE8" box="[384,493,525,551]" pageId="4" pageNumber="21" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C224D4BFFDD1E38FD83FDDF3687FDE9" box="[534,675,525,550]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="digitatus">
|
||
<emphasis id="B956EADAFFDD1E38FD83FDDF3687FDE9" box="[534,675,525,550]" italics="true" pageId="4" pageNumber="21">O. digitatus</emphasis>
|
||
</taxonomicName>
|
||
, which is deposited at the
|
||
<collectionCode id="ED33AE0DFFDD1E38FC76FDDF3065FDE8" box="[995,1089,525,551]" pageId="4" pageNumber="21">BMNH</collectionCode>
|
||
). Both species posses similar body shape, especially of the head and pronotum; vertex in males armed with small horn bent backwards; internal apical angle of protibia with large tooth in males; protibial apical spur small, acuminate, slightly bent outward; mesotibial and metatibial apex produced into three distinct lobes; aedeagus and lamella copulatrix similarly shaped. The characters separating two these species are summarized in
|
||
<tableCitation id="C6A00373FFDD1E38FBF9FD7030E3FD73" box="[1132,1223,674,700]" captionStart="TABLE 1" captionStartId="4.[151,239,1010,1034]" captionTargetBox="[151,1366,1061,1426]" captionTargetPageId="4" captionText="TABLE 1. Diagnostic characters separating the new species O. streltsovi from O. digitatus." httpUri="http://table.plazi.org/id/DF5D6640FFDD1E38FF02FC203043FBC6" pageId="4" pageNumber="21" tableUuid="DF5D6640FFDD1E38FF02FC203043FBC6">Table 1</tableCitation>
|
||
. Both species are probably also similar to
|
||
<taxonomicName id="4C224D4BFFDD1E38FE5BFD1A30F1FD2D" authority="Masumoto, Ochi & Hanboonsong, 2008" authorityName="Masumoto, Ochi & Hanboonsong" authorityYear="2008" box="[462,1237,712,738]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="kaengkrachangus">
|
||
<emphasis id="B956EADAFFDD1E38FE5BFD1A36F7FD2E" box="[462,723,712,737]" italics="true" pageId="4" pageNumber="21">O. kaengkrachangus</emphasis>
|
||
Masumoto, Ochi & Hanboonsong, 2008
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F3357658FFDD1E38FAB0FD1A31BDFD2D" box="[1317,1433,712,738]" name="Thailand" pageId="4" pageNumber="21">Thailand</collectingCountry>
|
||
. Unfortunately the
|
||
<typeStatus id="5499886AFFDD1E38FEFAFD3F3589FCC8" box="[367,429,749,775]" pageId="4" pageNumber="21">types</typeStatus>
|
||
of the latter have not been examined, but as we can see from its original description,
|
||
<taxonomicName id="4C224D4BFFDD1E38FF02FCC035ABFCE4" box="[151,399,786,811]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="kaengkrachangus">
|
||
<emphasis id="B956EADAFFDD1E38FF02FCC035ABFCE4" box="[151,399,786,811]" italics="true" pageId="4" pageNumber="21">O. kaengkrachangus</emphasis>
|
||
</taxonomicName>
|
||
shares almost the same combination of characters with
|
||
<taxonomicName id="4C224D4BFFDD1E38FBA3FCC030EEFCE4" box="[1078,1226,786,811]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="streltsovi">
|
||
<emphasis id="B956EADAFFDD1E38FBA3FCC030EEFCE4" box="[1078,1226,786,811]" italics="true" pageId="4" pageNumber="21">O. streltsovi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C224D4BFFDD1E38FA90FCC031B7FCE4" box="[1285,1427,786,811]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="digitatus">
|
||
<emphasis id="B956EADAFFDD1E38FA90FCC031B7FCE4" box="[1285,1427,786,811]" italics="true" pageId="4" pageNumber="21">O. digitatus</emphasis>
|
||
</taxonomicName>
|
||
: head and the rest of body specifically shaped; “protibiae with inner side of terminal edge bluntly projected; terminal spur rather bold”. Also, all three species have similar shape of aedeagus. Only one remarkable character, the male vertex with small horn bent backwards, is not reported for
|
||
<taxonomicName id="4C224D4BFFDD1E38FBD1FC50311FFC54" box="[1092,1339,898,923]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="kaengkrachangus">
|
||
<emphasis id="B956EADAFFDD1E38FBD1FC50311FFC54" box="[1092,1339,898,923]" italics="true" pageId="4" pageNumber="21">O. kaengkrachangus</emphasis>
|
||
</taxonomicName>
|
||
. This is likely due to the lack of major males in the
|
||
<typeStatus id="5499886AFFDD1E38FD01FC7A36E3FC0D" box="[660,711,936,962]" pageId="4" pageNumber="21">type</typeStatus>
|
||
series.
|
||
</paragraph>
|
||
<caption id="DF5D6640FFDD1E38FF02FC203043FBC6" ID-Table-UUID="DF5D6640FFDD1E38FF02FC203043FBC6" box="[151,1127,1010,1034]" httpUri="http://table.plazi.org/id/DF5D6640FFDD1E38FF02FC203043FBC6" pageId="4" pageNumber="21" targetBox="[151,1366,1061,1426]" targetIsTable="true" targetPageId="4">
|
||
<paragraph id="8B9D36C8FFDD1E38FF02FC203043FBC6" blockId="4.[151,1127,1010,1034]" box="[151,1127,1010,1034]" pageId="4" pageNumber="21">
|
||
<emphasis id="B956EADAFFDD1E38FF02FC20352CFBC5" bold="true" box="[151,264,1010,1034]" pageId="4" pageNumber="21">TABLE 1.</emphasis>
|
||
Diagnostic characters separating the new species
|
||
<taxonomicName id="4C224D4BFFDD1E38FC8AFC203785FBC6" box="[799,929,1010,1033]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="streltsovi">
|
||
<emphasis id="B956EADAFFDD1E38FC8AFC203785FBC6" box="[799,929,1010,1033]" italics="true" pageId="4" pageNumber="21">O. streltsovi</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C224D4BFFDD1E38FC77FC203044FBC6" box="[994,1120,1010,1033]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="digitatus">
|
||
<emphasis id="B956EADAFFDD1E38FC77FC203044FBC6" box="[994,1120,1010,1033]" italics="true" pageId="4" pageNumber="21">O. digitatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9D36C8FFDD1E38FF0AFBF730C8FA5D" pageId="4" pageNumber="21">
|
||
<table id="F922C468FFDDE1C3FF02FBF73172FA5D" box="[151,1366,1061,1426]" gridcols="3" gridrows="5" pageId="4" pageNumber="21">
|
||
<tr id="3512348AFFDDE1C3FF02FBF73172FBF3" box="[151,1366,1061,1084]" gridrow="0" pageId="4" pageNumber="21">
|
||
<th id="76C35DF6FFDDE1C3FF02FBF735DFFBF3" box="[151,507,1061,1084]" gridcol="0" gridrow="0" pageId="4" pageNumber="21">Characters</th>
|
||
<th id="76C35DF6FFDDE1C3FD8FFBF7302FFBF3" box="[538,1035,1061,1084]" gridcol="1" gridrow="0" pageId="4" pageNumber="21">
|
||
<taxonomicName id="4C224D4BFFDD1E38FD8FFBF736BFFBF3" box="[538,667,1061,1084]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="streltsovi">
|
||
<emphasis id="B956EADAFFDD1E38FD8FFBF736BFFBF3" box="[538,667,1061,1084]" italics="true" pageId="4" pageNumber="21">O. streltsovi</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="76C35DF6FFDDE1C3FBB4FBF73172FBF3" box="[1057,1366,1061,1084]" gridcol="2" gridrow="0" pageId="4" pageNumber="21">
|
||
<taxonomicName id="4C224D4BFFDD1E38FBB4FBF730BBFBF3" box="[1057,1183,1061,1084]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="digitatus">
|
||
<emphasis id="B956EADAFFDD1E38FBB4FBF730BBFBF3" box="[1057,1183,1061,1084]" italics="true" pageId="4" pageNumber="21">O. digitatus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="3512348AFFDDE1C3FF02FB823172FBA8" box="[151,1366,1104,1127]" gridrow="1" pageId="4" pageNumber="21">
|
||
<th id="76C35DF6FFDDE1C3FF02FB8235DFFBA8" box="[151,507,1104,1127]" gridcol="0" gridrow="1" pageId="4" pageNumber="21">Matasternal posterior depression</th>
|
||
<td id="76C35DF6FFDDE1C3FD8FFB82302FFBA8" box="[538,1035,1104,1127]" gridcol="1" gridrow="1" pageId="4" pageNumber="21">Absent</td>
|
||
<td id="76C35DF6FFDDE1C3FBB4FB823172FBA8" box="[1057,1366,1104,1127]" gridcol="2" gridrow="1" pageId="4" pageNumber="21">Present</td>
|
||
</tr>
|
||
<tr id="3512348AFFDDE1C3FF02FBA83172FB1E" box="[151,1366,1146,1233]" gridrow="2" pageId="4" pageNumber="21">
|
||
<th id="76C35DF6FFDDE1C3FF02FBA835DFFB1E" box="[151,507,1146,1233]" gridcol="0" gridrow="2" pageId="4" pageNumber="21">Coloration</th>
|
||
<td id="76C35DF6FFDDE1C3FD8FFBA8302FFB1E" box="[538,1035,1146,1233]" gridcol="1" gridrow="2" pageId="4" pageNumber="21">Head and pronotum black; elytra yellow- brown, intervals with black interrupted stripes; pygidium, metasternum laterally, abdomen,</td>
|
||
<td id="76C35DF6FFDDE1C3FBB4FBA83172FB1E" box="[1057,1366,1146,1233]" gridcol="2" gridrow="2" pageId="4" pageNumber="21">Black</td>
|
||
</tr>
|
||
<tr id="3512348AFFDDE1C3FF02FB223172FA88" box="[151,1366,1264,1351]" gridrow="3" pageId="4" pageNumber="21">
|
||
<th id="76C35DF6FFDDE1C3FF02FB2235DFFA88" box="[151,507,1264,1351]" gridcol="0" gridrow="3" pageId="4" pageNumber="21">Punctation of pronotum</th>
|
||
<td id="76C35DF6FFDDE1C3FD8FFB22302FFA88" box="[538,1035,1264,1351]" gridcol="1" gridrow="3" pageId="4" pageNumber="21">Disc sparsely covered with fine punctures becoming denser and larger laterally</td>
|
||
<td id="76C35DF6FFDDE1C3FBB4FB223172FA88" box="[1057,1366,1264,1351]" gridcol="2" gridrow="3" pageId="4" pageNumber="21">Disc not punctate, smooth; pronotum covered with large punctures laterally</td>
|
||
</tr>
|
||
<tr id="3512348AFFDDE1C3FF02FA893172FA5D" box="[151,1366,1371,1426]" gridrow="4" pageId="4" pageNumber="21">
|
||
<th id="76C35DF6FFDDE1C3FF02FA8935DFFA5D" box="[151,507,1371,1426]" gridcol="0" gridrow="4" pageId="4" pageNumber="21">Aedeagus</th>
|
||
<td id="76C35DF6FFDDE1C3FD8FFA89302FFA5D" box="[538,1035,1371,1426]" gridcol="1" gridrow="4" pageId="4" pageNumber="21">Parameral preapical section distinctly excavated</td>
|
||
<td id="76C35DF6FFDDE1C3FBB4FA893172FA5D" box="[1057,1366,1371,1426]" gridcol="2" gridrow="4" pageId="4" pageNumber="21">Parameral preapical section without excavation</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8B9D36C8FFDD1E38FF51FA0A36E1F9D7" blockId="4.[151,1437,1496,1784]" pageId="4" pageNumber="21">These three species probably represent a monophyletic group. Reasons for its current taxonomic position are provided in the “Discussion” section below.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3386543FFDD1E38FF51F9F03664F937" pageId="4" pageNumber="21" type="etymology">
|
||
<paragraph id="8B9D36C8FFDD1E38FF51F9F03664F937" blockId="4.[151,1437,1496,1784]" pageId="4" pageNumber="21">
|
||
<emphasis id="B956EADAFFDD1E38FF51F9F03572F9F3" bold="true" box="[196,342,1570,1596]" pageId="4" pageNumber="21">Etymology.</emphasis>
|
||
The new species is named after Dr. Alexey Streltsov, Professor of Kaluga University in
|
||
<collectingCountry id="F3357658FFDD1E38FF02F99A34CDF9AD" box="[151,233,1608,1634]" name="Russia" pageId="4" pageNumber="21">Russia</collectingCountry>
|
||
, to whom we are much obliged for personal help and support during research as well as in everyday life. In particular, it was thanks to Alexey Streltsov that the long field trip to
|
||
<collectingCountry id="F3357658FFDD1E38FB8DF9BC3075F947" box="[1048,1105,1646,1672]" name="Laos" pageId="4" pageNumber="21">Laos</collectingCountry>
|
||
by S.T. was possible.
|
||
<emphasis id="B956EADAFFDD1E38FF51F9403627F963" bold="true" box="[196,515,1682,1708]" pageId="4" pageNumber="21">Distribution and Ecology.</emphasis>
|
||
The species is known from many specimens, but only from the
|
||
<typeStatus id="5499886AFFDD1E38FB78F9413104F962" box="[1261,1312,1683,1709]" pageId="4" pageNumber="21">type</typeStatus>
|
||
locality in Bolaven Plateau, South
|
||
<collectingCountry id="F3357658FFDD1E38FE2FF96A35D0F91D" box="[442,500,1720,1746]" name="Laos" pageId="4" pageNumber="21">Laos</collectingCountry>
|
||
(
|
||
<figureCitation id="13192A4DFFDD1E38FD90F96A3675F91D" box="[517,593,1720,1746]" captionStart="FIGURE 4" captionStartId="7.[151,255,1747,1771]" captionText="FIGURE 4. Distribution map of O. streltsovi and O. nanpatensis." pageId="4" pageNumber="21">Fig. 4</figureCitation>
|
||
). Almost all specimens were collected in montane forest by means of pifall traps baited with human dung.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |