treatments-xml/data/03/8B/87/038B87C6FFC3F10E102EAFB016235425.xml

<document id="4BCCE9BF8DDB82A39C2E1AF9DAF988C3" ID-DOI="10.5281/zenodo.10982041" ID-ISSN="2295-0214" ID-Zenodo-Dep="10982041" ID-ZooBank="C668F461-F8B3-4E76-B261-B2D166D484E8" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1713293055674" checkinUser="felipe" docAuthor="Limbourg, Pol, Dekoninck, Wouter &amp; Seidel, Matthias" docDate="2024" docId="038B87C6FFC3F10E102EAFB016235425" docLanguage="en" docName="BelgJEntomol.145.1-48.pdf" docOrigin="Belgian Journal of Entomology 145" docStyle="DocumentStyle:32E55E25D8523BC97666C7812278ECC1.2:BelgJEntomol.2022-.journal_article.type1" docStyleId="32E55E25D8523BC97666C7812278ECC1" docStyleName="BelgJEntomol.2022-.journal_article.type1" docStyleVersion="2" docTitle="Anomala koulango Limbourg &amp; Dekoninck &amp; Seidel 2024, sp. nov." docType="treatment" docUuid="B451C64F-0E49-4E41-899A-3DC38F180364" docUuidSource="ZooBank" docVersion="1" lastPageNumber="13" masterDocId="FFB2FFBEFFCBF102124AAC12130A5751" masterDocTitle="Contribution to the knowledge of Afrotropical Rutelinae VII (Coleoptera, Scarabaeidae, Rutelinae)" masterLastPageNumber="48" masterPageNumber="1" pageNumber="9" updateTime="1714478230135" updateUser="tatiana">
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<mods:title id="2BD83E9B97647E86A8D733B17247949F">Contribution to the knowledge of Afrotropical Rutelinae VII (Coleoptera, Scarabaeidae, Rutelinae)</mods:title>
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<mods:namePart id="9134D385CE1A1B4371E5EF9DDA40C764">Limbourg, Pol</mods:namePart>
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<mods:namePart id="86B1F173E7AFF8B7E6A53211B79E7858">Dekoninck, Wouter</mods:namePart>
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<mods:namePart id="952E3B7A2D69E14C88611D57AFDA7177">Seidel, Matthias</mods:namePart>
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<mods:title id="B7B313B43AC3DDC191A135C1CF2CF6FC">Belgian Journal of Entomology</mods:title>
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<mods:date id="8B1969BAAE608CE008383C021BC0ADAC">2024</mods:date>
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<mods:number id="6AFDFB82A51BF833F2D849E249AFBB43">2024-02-21</mods:number>
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<treatment id="038B87C6FFC3F10E102EAFB016235425" LSID="urn:lsid:zoobank.org:act:B451C64F-0E49-4E41-899A-3DC38F180364" httpUri="http://treatment.plazi.org/id/038B87C6FFC3F10E102EAFB016235425" lastPageId="12" lastPageNumber="13" pageId="8" pageNumber="9">
<subSubSection id="C338655BFFC3F10A102EAFB010DA54EE" box="[612,976,930,959]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="8B9D36D0FFC3F10A102EAFB010DA54EE" blockId="8.[409,1178,930,1038]" box="[612,976,930,959]" pageId="8" pageNumber="9">
<heading id="D0D581BCFFC3F10A102EAFB010DA54EE" bold="true" box="[612,976,930,959]" centered="true" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
<emphasis id="B956EAC2FFC3F10A102EAFB010DA54EE" bold="true" box="[612,976,930,959]" pageId="8" pageNumber="9">
<taxonomicName id="4C224D53FFC3F10A102EAFB0106B54EE" ID-CoL="C4KZX" authority="Limbourg &amp; Dekoninck &amp; Seidel, 2024" authorityName="Limbourg &amp; Dekoninck &amp; Seidel" authorityYear="2024" box="[612,865,930,959]" class="Insecta" family="Scarabaeidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="koulango" status="sp. nov.">
<emphasis id="B956EAC2FFC3F10A102EAFB0106B54EE" bold="true" box="[612,865,930,959]" italics="true" pageId="8" pageNumber="9">Anomala koulango</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557B9FFC3F10A1123AFB110DA54EE" box="[873,976,931,959]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C338655BFFC3F10A13D3AFDD105B535F" pageId="8" pageNumber="9" type="description">
<paragraph id="8B9D36D0FFC3F10A13D3AFDD179054B7" blockId="8.[409,1178,930,1038]" box="[409,1178,974,998]" pageId="8" pageNumber="9">
<heading id="D0D581BCFFC3F10A13D3AFDD179054B7" box="[409,1178,974,998]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="9" reason="5">
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urn: lsid:zoobank.org:act: 
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</uri>
</heading>
</paragraph>
<paragraph id="8B9D36D0FFC3F10A10A8AFE3105B535F" blockId="8.[409,1178,930,1038]" box="[738,849,1009,1038]" pageId="8" pageNumber="9">
<heading id="D0D581BCFFC3F10A10A8AFE3105B535F" box="[738,849,1009,1038]" centered="true" fontSize="12" level="2" pageId="8" pageNumber="9" reason="7">
<figureCitation id="13192A55FFC3F10A10A8AFE3105B535F" box="[738,849,1009,1038]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="9.[189,228,1958,1982]" captionStartId-1="10.[189,228,1958,1982]" captionTargetBox-0="[204,1357,382,1930]" captionTargetBox-1="[201,1380,421,1931]" captionTargetId-0="figure-79@9.[190,1397,364,1943]" captionTargetId-1="figure-12@10.[188,1400,399,1944]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="Fig. 3. Anomala koulango sp. nov. A-B, holotype ♂ (RBINS) habitus. A, dorsal view. B, lateral view. C-D, paratype ♀ (ZFMK) habitus. C, dorsal view. D, lateral view. E-G, aedeagus. E, lateral view. F, dorsal view. G, ventral view. H, distribution map of Anomala koulango sp. nov. and Anomala dossidea Ohaus, 1925." captionText-1="Fig. 4. Anomala dossidea Ohaus, 1925. A-B, ♂ (RBINS) habitus. A, dorsal view. B, lateral view. C-D, ♀ (RBINS) habitus. C, dorsal view. D, lateral view. E-G, aedeagus. E, lateral view. F, dorsal view. G, ventral view. H, disc of pronotum of Anomala koulango sp. nov. ♂. I, disc of pronotum of Anomala dossidea Ohaus, 1925 ♂." figureDoi-0="http://doi.org/10.5281/zenodo.10982047" figureDoi-1="http://doi.org/10.5281/zenodo.10982049" httpUri-0="https://zenodo.org/record/10982047/files/figure.png" httpUri-1="https://zenodo.org/record/10982049/files/figure.png" pageId="8" pageNumber="9">Figs 3–4</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C338655BFFC3F10A12F7A857173851B5" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="8B9D36D0FFC3F10A12F7A857128D5330" blockId="8.[189,391,1092,1121]" box="[189,391,1092,1121]" pageId="8" pageNumber="9">
<typeStatus id="54998872FFC3F10A12F7A85713F45330" box="[189,254,1093,1121]" pageId="8" pageNumber="9">TYPE</typeStatus>
MATERIAL.
</paragraph>
<paragraph id="8B9D36D0FFC3F10A12F7A86B173851B5" blockId="8.[189,1399,1145,1764]" pageId="8" pageNumber="9">
<materialsCitation id="3B4A3C8DFFC3F10A12F7A86B123353A0" collectingDate="1998-04" collectionCode="RBINS" collectorName="P. Moretto" country="Ivory Coast" elevation="279" latitude="8.5923605" location="Zamou" longLatPrecision="1" longitude="-3.7686112" pageId="8" pageNumber="9" specimenCount="1" specimenCount-male="1" stateProvince="Comoe" typeStatus="holotype">
<collectingCountry id="F3357640FFC3F10A12F7A86B129453C7" box="[189,414,1145,1174]" name="Ivory Coast" pageId="8" pageNumber="9">IVORY COAST</collectingCountry>
. 
<typeStatus id="54998872FFC3F10A13FEA86B112153C7" box="[436,555,1145,1174]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
<specimenCount id="9D24FD59FFC3F10A1075A868115153C7" box="[575,603,1146,1174]" count="1" pageId="8" pageNumber="9" type="male">♂</specimenCount>
: [Coll. I.R.Sc.N.B., CÔTE d’IVOIRE, 
<collectingRegion id="49E6F832FFC3F10A16C3A86B17EF53C7" box="[1161,1253,1145,1174]" country="Ivory Coast" name="Comoe" pageId="8" pageNumber="9">Comoé</collectingRegion>
, 
<location id="8EFD600BFFC3F10A175EA868167853C7" LSID="urn:lsid:plazi:treatment:038B87C6FFC3F10E102EAFB016235425:8EFD600BFFC3F10A175EA868167853C7" box="[1300,1394,1146,1174]" country="Ivory Coast" latitude="8.5923605" longLatPrecision="1" longitude="-3.7686112" name="Zamou" pageId="8" pageNumber="9" stateProvince="Comoe">Zamou</location>
, 
<geoCoordinate id="EE165017FFC3F10A12F7A8B412795392" box="[189,371,1190,1219]" degrees="08" direction="north" minutes="35" orientation="latitude" pageId="8" pageNumber="9" precision="1" seconds="32.5" value="8.5923605">08°35’32.5’’N</geoCoordinate>
<geoCoordinate id="EE165017FFC3F10A1338A8B4114D5392" box="[370,583,1190,1219]" degrees="003" direction="west" minutes="46" orientation="longitude" pageId="8" pageNumber="9" precision="1" seconds="07.0" value="-3.7686112">003°46’07.0’’W</geoCoordinate>
, 
<quantity id="4CDA9B35FFC3F10A1011A8B511AD5395" box="[603,679,1191,1220]" metricMagnitude="2" metricUnit="m" metricValue="2.79" pageId="8" pageNumber="9" unit="m" value="279.0">
<elevation id="000FD1E3FFC3F10A1011A8B511AD5395" box="[603,679,1191,1220]" metricMagnitude="2" metricUnit="m" metricValue="2.79" pageId="8" pageNumber="9" unit="m" value="279.0">279m</elevation>
</quantity>
, 
<date id="FF9C1010FFC3F10A10F6A8B410175392" box="[700,797,1190,1219]" pageId="8" pageNumber="9" value="1998-04">
<collectingDate id="EFD8E9F8FFC3F10A10F6A8B410175392" box="[700,797,1190,1219]" pageId="8" pageNumber="9" value="1998-04">iv.1998</collectingDate>
</date>
, local coll.&gt; 
<collectorName id="26D75306FFC3F10A11A2A8B5177E5392" box="[1000,1140,1191,1219]" pageId="8" pageNumber="9">P. Moretto</collectorName>
leg., I.G.: 31.706] (
<collectionCode id="ED33AE15FFC3F10A128CA8C6122153A0" box="[198,299,1236,1265]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="8" pageNumber="9" type="Museum">RBINS</collectionCode>
);
</materialsCitation>
<typeStatus id="54998872FFC3F10A1302A8C612C053A1" box="[328,458,1236,1264]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
: same data collection data as for 
<typeStatus id="54998872FFC3F10A11D6A8C6171B53A0" box="[924,1041,1236,1265]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
: 
<specimenCount id="9D24FD59FFC3F10A1669A8C6174553A0" box="[1059,1103,1236,1265]" count="1" pageId="8" pageNumber="9" type="male">1♂</specimenCount>
: (
<collectionCode id="ED33AE15FFC3F10A1626A8C617DB53A0" box="[1132,1233,1236,1265]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="8" pageNumber="9" type="Museum">RBINS</collectionCode>
); 
<specimenCount id="9D24FD59FFC3F10A16A7A8C6161353A0" box="[1261,1305,1236,1265]" count="1" pageId="8" pageNumber="9" type="male">1♂</specimenCount>
: [Coll. I.R.Sc.N.B., 
<collectingCountry id="F3357640FFC3F10A132BA910114C524F" box="[353,582,1281,1313]" name="Ivory Coast" pageId="8" pageNumber="9">CÔTE d’IVOIRE</collectingCountry>
, 
<collectingRegion id="49E6F832FFC3F10A1019A91311A5524F" box="[595,687,1281,1310]" country="Ivory Coast" name="Comoe" pageId="8" pageNumber="9">Comoé</collectingRegion>
, Zamou, 
<geoCoordinate id="EE165017FFC3F10A1163A91310D5524F" box="[809,991,1281,1310]" degrees="08" direction="north" minutes="35" orientation="latitude" pageId="8" pageNumber="9" precision="1" seconds="32.5" value="8.5923605">08°35’32.5’’N</geoCoordinate>
<geoCoordinate id="EE165017FFC3F10A1195A91317B9524F" box="[991,1203,1281,1310]" degrees="003" direction="west" minutes="46" orientation="longitude" pageId="8" pageNumber="9" precision="1" seconds="07.0" value="-3.7686112">003°46’07.0’’W</geoCoordinate>
, 
<quantity id="4CDA9B35FFC3F10A168AA9131606524F" box="[1216,1292,1281,1310]" metricMagnitude="2" metricUnit="m" metricValue="2.79" pageId="8" pageNumber="9" unit="m" value="279.0">279m</quantity>
, 
<date id="FF9C1010FFC3F10A1753A9131678524F" box="[1305,1394,1281,1310]" pageId="8" pageNumber="9" value="1998-05">v.1998</date>
, local coll.&gt; P. Moretto leg., I.G.: 31.706] (
<collectionCode id="ED33AE15FFC3F10A10ABA93D104C521D" box="[737,838,1327,1356]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="8" pageNumber="9" type="Museum">RBINS</collectionCode>
); 
<specimenCount id="9D24FD59FFC3F10A1116A93D1082521A" box="[860,904,1327,1356]" count="1" pageId="8" pageNumber="9" type="male">1♂</specimenCount>
: [Coll. I.R.Sc.N.B., 
<collectingCountry id="F3357640FFC3F10A16C6A93D167B521A" box="[1164,1393,1326,1358]" name="Ivory Coast" pageId="8" pageNumber="9">CÔTE d’IVOIRE</collectingCountry>
, 
<collectingRegion id="49E6F832FFC3F10A12F7A94E12135228" box="[189,281,1372,1401]" country="Ivory Coast" name="Comoe" pageId="8" pageNumber="9">Comoé</collectingRegion>
, Zamou, 
<geoCoordinate id="EE165017FFC3F10A13ECA94E11565228" box="[422,604,1372,1401]" degrees="08" direction="north" minutes="35" orientation="latitude" pageId="8" pageNumber="9" precision="1" seconds="32.5" value="8.5923605">08°35’32.5’’N</geoCoordinate>
<geoCoordinate id="EE165017FFC3F10A1016A94E103A5228" box="[604,816,1372,1401]" degrees="003" direction="west" minutes="46" orientation="longitude" pageId="8" pageNumber="9" precision="1" seconds="07.0" value="-3.7686112">003°46’07.0’’W</geoCoordinate>
, 
<quantity id="4CDA9B35FFC3F10A110DA94E10995228" box="[839,915,1372,1401]" metricMagnitude="2" metricUnit="m" metricValue="2.79" pageId="8" pageNumber="9" unit="m" value="279.0">279m</quantity>
, 
<date id="FF9C1010FFC3F10A11E3A94E171F5228" box="[937,1045,1372,1401]" pageId="8" pageNumber="9" value="1998-09">IX.1998</date>
, local coll.&gt; P. Moretto leg., I.G.: 31.706] (
<collectionCode id="ED33AE15FFC3F10A138CA99B112152F7" box="[454,555,1417,1446]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="8" pageNumber="9" type="Museum">RBINS</collectionCode>
); 
<specimenCount id="9D24FD59FFC3F10A100DA99B118152F7" box="[583,651,1417,1446]" count="5" pageId="8" pageNumber="9" type="male">5♂♂</specimenCount>
: [Coll. I.R.Sc.N.B., 
<collectingCountry id="F3357640FFC3F10A11E8A998178452F7" box="[930,1166,1417,1449]" name="Ivory Coast" pageId="8" pageNumber="9">CÔTE d’IVOIRE</collectingCountry>
, 
<collectingRegion id="49E6F832FFC3F10A16EBA99B17F752F7" box="[1185,1277,1417,1446]" country="Ivory Coast" name="Comoe" pageId="8" pageNumber="9">Comoé</collectingRegion>
, Zamou, 
<geoCoordinate id="EE165017FFC3F10A12F7A9A412795282" box="[189,371,1462,1491]" degrees="08" direction="north" minutes="35" orientation="latitude" pageId="8" pageNumber="9" precision="1" seconds="32.5" value="8.5923605">08°35’32.5’’N</geoCoordinate>
<geoCoordinate id="EE165017FFC3F10A1338A9A4114D5282" box="[370,583,1462,1491]" degrees="003" direction="west" minutes="46" orientation="longitude" pageId="8" pageNumber="9" precision="1" seconds="07.0" value="-3.7686112">003°46’07.0’’W</geoCoordinate>
, 
<quantity id="4CDA9B35FFC3F10A101FA9A511AB5285" box="[597,673,1463,1492]" metricMagnitude="2" metricUnit="m" metricValue="2.79" pageId="8" pageNumber="9" unit="m" value="279.0">279m</quantity>
, 
<date id="FF9C1010FFC3F10A10E5A9A510005285" box="[687,778,1463,1492]" pageId="8" pageNumber="9" value="1998-10">x.1998</date>
, local coll.&gt; P. Moretto leg., I.G.: 31.706] (
<specimenCount id="9D24FD59FFC3F10A170DA9A5167B5285" box="[1351,1393,1463,1492]" count="1" pageId="8" pageNumber="9" type="male">1♂</specimenCount>
: 
<collectionCode id="ED33AE15FFC3F10A12F7A9F612165151" box="[189,284,1508,1536]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="8" pageNumber="9">MFNB</collectionCode>
; 
<specimenCount id="9D24FD59FFC3F10A1363A9F612675151" box="[297,365,1508,1537]" count="4" pageId="8" pageNumber="9" type="male">4♂♂</specimenCount>
: 
<collectionCode id="ED33AE15FFC3F10A1331A9F612D55150" box="[379,479,1508,1537]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="8" pageNumber="9" type="Museum">RBINS</collectionCode>
); 
<collectingCountry id="F3357640FFC3F10A13BCA9F611E85150" box="[502,738,1508,1537]" name="Burkina Faso" pageId="8" pageNumber="9">BURKINA FASO</collectingCountry>
. 
<specimenCount id="9D24FD59FFC3F10A10A4A9F6103B5151" box="[750,817,1508,1537]" count="2" pageId="8" pageNumber="9" type="male">2♂♂</specimenCount>
, 
<specimenCount id="9D24FD59FFC3F10A1175A9F610895150" box="[831,899,1508,1537]" count="2" pageId="8" pageNumber="9" type="female">2♀♀</specimenCount>
: [Obervolta, 
<quantity id="4CDA9B35FFC3F10A1666A9F6176F5150" box="[1068,1125,1508,1537]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="9" unit="km" value="4.0">4km</quantity>
NE Dérégoué, 10,79 N 
<geoCoordinate id="EE165017FFC3F10A129EAA031221517F" box="[212,299,1553,1582]" degrees="4,08" direction="west" orientation="longitude" pageId="8" pageNumber="9" precision="555" value="-4.08">4,08W</geoCoordinate>
, Galeriewald am Koba, 2-4.1985, H. Schreiber, B. Basten] [museum KOENIG Bonn, 
<collectingCountry id="F3357640FFC3F10A12F7AA2D1232510D" box="[189,312,1599,1628]" name="Germany" pageId="8" pageNumber="9">Germany</collectingCountry>
] (
<specimenCount id="9D24FD59FFC3F10A131EAA2D1277510D" box="[340,381,1599,1628]" count="1" pageId="8" pageNumber="9" type="male">1♂</specimenCount>
, 
<specimenCount id="9D24FD59FFC3F10A13C4AA2D12DB510D" box="[398,465,1599,1628]" count="2" pageId="8" pageNumber="9" type="female">2♀♀</specimenCount>
: 
<collectionCode id="ED33AE15FFC3F10A13ABAA2D1134510A" box="[481,574,1599,1627]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="8" pageNumber="9" type="Museum">ZFMK</collectionCode>
; 
<specimenCount id="9D24FD59FFC3F10A1006AA2D1172510D" box="[588,632,1599,1628]" count="1" pageId="8" pageNumber="9" type="male">1♂</specimenCount>
: 
<collectionCode id="ED33AE15FFC3F10A10CDAA2D11FC510D" box="[647,758,1599,1628]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="9" type="Museum">NHMW</collectionCode>
); 
<specimenCount id="9D24FD59FFC3F10A1144AA2D1030510D" box="[782,826,1599,1628]" count="1" pageId="8" pageNumber="9" type="male">1♂</specimenCount>
: [
<collectingCountry id="F3357640FFC3F10A111AAA2C10DD510A" box="[848,983,1598,1627]" name="Burkina Faso" pageId="8" pageNumber="9">Obervolta</collectingCountry>
, 
<quantity id="4CDA9B35FFC3F10A11ACAA2C1724510A" box="[998,1070,1598,1627]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="9" unit="km" value="10.0">10km</quantity>
SO Tiefora, Galeriewald am Sinlo, 11.i-19.ii.(19)82, Nagel, Schreiber, Kalmund] [museum KOENIG Bonn, 
<collectingCountry id="F3357640FFC3F10A16BFAA7E167A51D8" box="[1269,1392,1644,1673]" name="Germany" pageId="8" pageNumber="9">Germany</collectingCountry>
] (
<collectionCode id="ED33AE15FFC3F10A128CAA8B122C51E7" box="[198,294,1689,1718]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="8" pageNumber="9" type="Museum">ZFMK</collectionCode>
); 
<specimenCount id="9D24FD59FFC3F10A130CAA8B127B51E7" box="[326,369,1689,1718]" count="1" pageId="8" pageNumber="9" type="female">1♀</specimenCount>
: [
<collectingCountry id="F3357640FFC3F10A13DAAA8B111D51E7" box="[400,535,1689,1718]" name="Burkina Faso" pageId="8" pageNumber="9">Obervolta</collectingCountry>
, 
<quantity id="4CDA9B35FFC3F10A1067AA8B117C51E7" box="[557,630,1689,1718]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="9" unit="km" value="10.0">10km</quantity>
SO Tiefora, Galeriewald am Sinlo, 
<date id="FF9C1010FFC3F10A1634AA8B164A51E7" box="[1150,1344,1689,1718]" pageId="8" pageNumber="9" value="1983-01-10" valueMax="1983-02-25" valueMin="1983-01-10">10.I-25.ii.1983</date>
, H. Schreiber, M. Paulus] [museum KOENIG Bonn, 
<collectingCountry id="F3357640FFC3F10A1178AAD510A751B5" box="[818,941,1735,1764]" name="Germany" pageId="8" pageNumber="9">Germany</collectingCountry>
] (
<collectionCode id="ED33AE15FFC3F10A118EAAD5172E51B5" box="[964,1060,1735,1764]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="8" pageNumber="9" type="Museum">ZFMK</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C338655BFFC3F10912F7AB0812CF5026" lastPageId="11" lastPageNumber="12" pageId="8" pageNumber="9" type="description">
<paragraph id="8B9D36D0FFC3F10A12F7AB0812665066" blockId="8.[189,364,1818,1847]" box="[189,364,1818,1847]" pageId="8" pageNumber="9">DESCRIPTION.</paragraph>
<paragraph id="8B9D36D0FFC3F10A12F7AB5D10E8503A" blockId="8.[189,994,1870,1900]" box="[189,994,1870,1900]" pageId="8" pageNumber="9">
MALE. Measurements: length: 
<quantity id="4CDA9B35FFC3F10A101EAB5C11F3503A" box="[596,761,1870,1899]" metricMagnitude="-2" metricUnit="m" metricValue="1.725" metricValueMax="1.8" metricValueMin="1.65" pageId="8" pageNumber="9" unit="mm" value="17.25" valueMax="18.0" valueMin="16.5">16.5–18 mm</quantity>
, width: 
<quantity id="4CDA9B35FFC3F10A112AAB5D10D4503A" box="[864,990,1871,1900]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="8" pageNumber="9" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">9–10 mm</quantity>
.
</paragraph>
<paragraph id="8B9D36D0FFC3F10A12A9AB9912415F52" blockId="8.[189,1399,1931,2051]" pageId="8" pageNumber="9">
<emphasis id="B956EAC2FFC3F10A12A9AB99122F50F9" box="[227,293,1931,1960]" italics="true" pageId="8" pageNumber="9">Body</emphasis>
: laterally slightly convex, elytra almost parallel, slightly bent; brownish-yellow except head, all margins, mouthparts, tarsi, protibia and metatibia deep brown to black; without metallic reflections.
</paragraph>
<caption id="DF5D6658FFC2F10B12F7ABB417F850AF" ID-DOI="http://doi.org/10.5281/zenodo.10982047" ID-Zenodo-Dep="10982047" httpUri="https://zenodo.org/record/10982047/files/figure.png" pageId="9" pageNumber="10" startId="9.[189,228,1958,1982]" targetBox="[204,1357,382,1930]" targetPageId="9" targetType="figure">
<paragraph id="8B9D36D0FFC2F10B12F7ABB417F850AF" blockId="9.[189,1399,1958,2046]" pageId="9" pageNumber="10">
Fig. 3. 
<taxonomicName id="4C224D53FFC2F10B135AABB412D450EF" authority="Limbourg &amp; Dekoninck &amp; Seidel, 2024" authorityName="Limbourg &amp; Dekoninck &amp; Seidel" authorityYear="2024" box="[272,478,1958,1982]" class="Insecta" family="Scarabaeidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="koulango" status="sp. nov.">
<emphasis id="B956EAC2FFC2F10B135AABB412D450EF" box="[272,478,1958,1982]" italics="true" pageId="9" pageNumber="10">Anomala koulango</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557B9FFC2F10B13A3ABB5114A50EF" box="[489,576,1959,1982]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B956EAC2FFC2F10B1000ABB4117250EF" bold="true" box="[586,632,1958,1982]" pageId="9" pageNumber="10">A -B</emphasis>
, holotype ♂ (RBINS) habitus. 
<emphasis id="B956EAC2FFC2F10B1193ABB410E650EF" bold="true" box="[985,1004,1958,1982]" pageId="9" pageNumber="10">A</emphasis>
, dorsal view. 
<emphasis id="B956EAC2FFC2F10B16C7ABB4179550EF" bold="true" box="[1165,1183,1958,1982]" pageId="9" pageNumber="10">B</emphasis>
, lateral view. 
<emphasis id="B956EAC2FFC2F10B170AABB4167A50EF" bold="true" box="[1344,1392,1958,1982]" pageId="9" pageNumber="10">C -D</emphasis>
, paratype ♀ (ZFMK) habitus. 
<emphasis id="B956EAC2FFC2F10B13B7ABD4111A508F" bold="true" box="[509,528,1990,2014]" pageId="9" pageNumber="10">C</emphasis>
, dorsal view. 
<emphasis id="B956EAC2FFC2F10B10E3ABD411B6508F" bold="true" box="[681,700,1990,2014]" pageId="9" pageNumber="10">D</emphasis>
, lateral view. 
<emphasis id="B956EAC2FFC2F10B111DABD4108C508F" bold="true" box="[855,902,1990,2014]" pageId="9" pageNumber="10">E -G</emphasis>
, aedeagus. 
<emphasis id="B956EAC2FFC2F10B164CABD41712508F" bold="true" box="[1030,1048,1990,2014]" pageId="9" pageNumber="10">E</emphasis>
, lateral view. 
<emphasis id="B956EAC2FFC2F10B16F8ABD417C8508F" bold="true" box="[1202,1218,1990,2014]" pageId="9" pageNumber="10">F</emphasis>
, dorsal view. 
<emphasis id="B956EAC2FFC2F10B1711ABD4167A508F" bold="true" box="[1371,1392,1990,2014]" pageId="9" pageNumber="10">G</emphasis>
, ventral view. 
<emphasis id="B956EAC2FFC2F10B1307ABF4126850AF" bold="true" box="[333,354,2022,2046]" pageId="9" pageNumber="10">H</emphasis>
, distribution map of 
<taxonomicName id="4C224D53FFC2F10B1008ABF4100650AF" authority="Limbourg &amp; Dekoninck &amp; Seidel, 2024" authorityName="Limbourg &amp; Dekoninck &amp; Seidel" authorityYear="2024" box="[578,780,2022,2046]" class="Insecta" family="Scarabaeidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="koulango" status="sp. nov.">
<emphasis id="B956EAC2FFC2F10B1008ABF4100650AF" box="[578,780,2022,2046]" italics="true" pageId="9" pageNumber="10">Anomala koulango</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557B9FFC2F10B1159ABF5106C50AF" box="[787,870,2023,2046]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C224D53FFC2F10B11D3ABF417E450AF" authority="Ohaus, 1925" authorityName="Ohaus" authorityYear="1925" box="[921,1262,2022,2046]" class="Insecta" family="Scarabaeidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="dossidea">
<emphasis id="B956EAC2FFC2F10B11D3ABF4175750AF" box="[921,1117,2022,2046]" italics="true" pageId="9" pageNumber="10">Anomala dossidea</emphasis>
<bibRefCitation id="EFB34B21FFC2F10B162EABF417E450AF" author="OHAUS F." box="[1124,1262,2022,2046]" pageId="9" pageNumber="10" pagination="42 - 80" refId="ref16704" refString="OHAUS F., 1925. - Revision der afrikanischen Anomala - Arten (Col. lamell. Rutelin.). Stettiner Entomologische Zeitung 86: 42 - 80." type="journal article" year="1925">Ohaus, 1925</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF5D6658FFC1F10812F7ABB417F250AC" ID-DOI="http://doi.org/10.5281/zenodo.10982049" ID-Zenodo-Dep="10982049" httpUri="https://zenodo.org/record/10982049/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,228,1958,1982]" targetBox="[201,1380,421,1931]" targetPageId="10" targetType="figure">
<paragraph id="8B9D36D0FFC1F10812F7ABB417F250AC" blockId="10.[189,1398,1957,2046]" pageId="10" pageNumber="11">
Fig. 4. 
<taxonomicName id="4C224D53FFC1F108134DABB4115050EC" authority="Ohaus, 1925" authorityName="Ohaus" authorityYear="1925" box="[263,602,1957,1982]" class="Insecta" family="Scarabaeidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="dossidea">
<emphasis id="B956EAC2FFC1F108134DABB412C050EF" box="[263,458,1958,1982]" italics="true" pageId="10" pageNumber="11">Anomala dossidea</emphasis>
<bibRefCitation id="EFB34B21FFC1F108139AABB4115050EC" author="OHAUS F." box="[464,602,1957,1982]" pageId="10" pageNumber="11" pagination="42 - 80" refId="ref16704" refString="OHAUS F., 1925. - Revision der afrikanischen Anomala - Arten (Col. lamell. Rutelin.). Stettiner Entomologische Zeitung 86: 42 - 80." type="journal article" year="1925">Ohaus, 1925</bibRefCitation>
</taxonomicName>
. 
<emphasis id="B956EAC2FFC1F1081028ABB7119A50EF" bold="true" box="[610,656,1957,1982]" pageId="10" pageNumber="11">A -B</emphasis>
, ♂ (RBINS) habitus. 
<emphasis id="B956EAC2FFC1F108113CABB7108350EC" bold="true" box="[886,905,1957,1981]" pageId="10" pageNumber="11">A</emphasis>
, dorsal view. 
<emphasis id="B956EAC2FFC1F1081651ABB4172750EF" bold="true" box="[1051,1069,1958,1982]" pageId="10" pageNumber="11">B</emphasis>
, lateral view. 
<emphasis id="B956EAC2FFC1F108168AABB717FA50EF" bold="true" box="[1216,1264,1957,1982]" pageId="10" pageNumber="11">C -D</emphasis>
, ♀ (RBINS) habitus. 
<emphasis id="B956EAC2FFC1F108135DABD71220508C" bold="true" box="[279,298,1989,2013]" pageId="10" pageNumber="11">C</emphasis>
, dorsal view. 
<emphasis id="B956EAC2FFC1F10813F4ABD412DB508F" bold="true" box="[446,465,1990,2014]" pageId="10" pageNumber="11">D</emphasis>
, lateral view. 
<emphasis id="B956EAC2FFC1F108102CABD4119C508C" bold="true" box="[614,662,1989,2014]" pageId="10" pageNumber="11">E -G</emphasis>
, aedeagus. 
<emphasis id="B956EAC2FFC1F108115BABD41029508F" bold="true" box="[785,803,1990,2014]" pageId="10" pageNumber="11">E</emphasis>
, lateral view. 
<emphasis id="B956EAC2FFC1F10811F2ABD410C2508F" bold="true" box="[952,968,1990,2014]" pageId="10" pageNumber="11">F</emphasis>
, dorsal view. 
<emphasis id="B956EAC2FFC1F1081616ABD7177B508C" bold="true" box="[1116,1137,1989,2013]" pageId="10" pageNumber="11">G</emphasis>
, ventral view. 
<emphasis id="B956EAC2FFC1F1081747ABD41628508F" bold="true" box="[1293,1314,1990,2014]" pageId="10" pageNumber="11">H</emphasis>
, disc of pronotum of 
<taxonomicName id="4C224D53FFC1F1081302ABF4111850AF" authority="Limbourg &amp; Dekoninck &amp; Seidel, 2024" authorityName="Limbourg &amp; Dekoninck &amp; Seidel" authorityYear="2024" box="[328,530,2022,2046]" class="Insecta" family="Scarabaeidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="koulango" status="sp. nov.">
<emphasis id="B956EAC2FFC1F1081302ABF4111850AF" box="[328,530,2022,2046]" italics="true" pageId="10" pageNumber="11">Anomala koulango</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557B9FFC1F1081053ABF4116650AC" box="[537,620,2022,2045]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
♂. 
<emphasis id="B956EAC2FFC1F10810D9ABF4119750AF" bold="true" box="[659,669,2022,2046]" pageId="10" pageNumber="11">I</emphasis>
, disc of pronotum of 
<taxonomicName id="4C224D53FFC1F10811CFABF417DC50AC" authority="Ohaus, 1925" authorityName="Ohaus" authorityYear="1925" box="[901,1238,2021,2046]" class="Insecta" family="Scarabaeidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="dossidea">
<emphasis id="B956EAC2FFC1F10811CFABF4174250AF" box="[901,1096,2022,2046]" italics="true" pageId="10" pageNumber="11">Anomala dossidea</emphasis>
<bibRefCitation id="EFB34B21FFC1F1081605ABF417DC50AC" author="OHAUS F." box="[1103,1238,2021,2046]" pageId="10" pageNumber="11" pagination="42 - 80" refId="ref16704" refString="OHAUS F., 1925. - Revision der afrikanischen Anomala - Arten (Col. lamell. Rutelin.). Stettiner Entomologische Zeitung 86: 42 - 80." type="journal article" year="1925">Ohaus, 1925</bibRefCitation>
</taxonomicName>
♂.
</paragraph>
</caption>
<paragraph id="8B9D36D0FFC0F10912A9ACC611EC5615" blockId="11.[189,1398,212,324]" pageId="11" pageNumber="12">
<emphasis id="B956EAC2FFC0F10912A9ACC6122257A0" box="[227,296,212,241]" italics="true" pageId="11" pageNumber="12">Head</emphasis>
: clypeus rounded and totally reflexed, aerolate-rugosely punctate, vertex alveolatepunctate, transversal elongate foveolate punctures near eye margins. Antennae with 9 antennomeres, with club as long as funicle.
</paragraph>
<paragraph id="8B9D36D0FFC0F10912A9AD4D106356A9" blockId="11.[189,1399,351,504]" pageId="11" pageNumber="12">
<emphasis id="B956EAC2FFC0F10912A9AD4D1235562D" box="[227,319,351,380]" italics="true" pageId="11" pageNumber="12">Thorax</emphasis>
: pronotum rectangular, wider than long; sides angular in middle; all margins with complete marginal line; anterior angles slightly projecting and reaching posterior margin of eyes; with shallow horseshoe-shaped punctures, almost coalescent on sides; scutellar shield subobtuse, with shallow horseshoe-shaped punctures.
</paragraph>
<paragraph id="8B9D36D0FFC0F10912A9AE06167D55D2" blockId="11.[189,1399,532,643]" pageId="11" pageNumber="12">
<emphasis id="B956EAC2FFC0F10912A9AE0612395560" box="[227,307,532,561]" italics="true" pageId="11" pageNumber="12">Elytra</emphasis>
: longer than wide; epipleura complete; regularly striate-sulcate; primary costae more convex than interstices; interstices with small isolated rounded punctures and microreticulate; striae with close horseshoe-shaped punctures; humeral calli marked, apical calli slightly marked.
</paragraph>
<paragraph id="8B9D36D0FFC0F10912A9AE8D1176545F" blockId="11.[189,1399,671,782]" pageId="11" pageNumber="12">
<emphasis id="B956EAC2FFC0F10912A9AE8D125455ED" box="[227,350,671,700]" italics="true" pageId="11" pageNumber="12">Pygidium</emphasis>
: triangular, apex rounded, marginated along anal plate, not projecting posteroventrally and not surpassing apex of elytra; punctate on disc, transversally, semicircularly punctate on lateral sides.
</paragraph>
<paragraph id="8B9D36D0FFC0F10912A9AF3917C15421" blockId="11.[189,1399,810,880]" pageId="11" pageNumber="12">
<emphasis id="B956EAC2FFC0F10912A9AF39123F5416" box="[227,309,811,839]" italics="true" pageId="11" pageNumber="12">Venter</emphasis>
: lateral part of metaventrite with setigerous punctures, median part almost smooth; abdominal ventrites with small transversal isolated tubercles, imbricate-punctate.
</paragraph>
<paragraph id="8B9D36D0FFC0F10912A9AF9F106C54AA" blockId="11.[189,1399,908,1019]" pageId="11" pageNumber="12">
<emphasis id="B956EAC2FFC0F10912A9AF9F121554F8" box="[227,287,909,937]" italics="true" pageId="11" pageNumber="12">Legs</emphasis>
: protibiae tridentate, inner claw relatively elongate, wide, flattened and cleft, strongly sinuate at base, one third shorter than last tarsomere; mesotarsal outer claw cleft; outer claw of metatarsus slightly longer than inner one and uncleft.
</paragraph>
<paragraph id="8B9D36D0FFC0F10912A9A80511C9522F" blockId="11.[189,1399,1047,1406]" pageId="11" pageNumber="12">
<emphasis id="B956EAC2FFC0F10912A9A80512465362" box="[227,332,1047,1075]" italics="true" pageId="11" pageNumber="12">Setation</emphasis>
: dorsal side almost glabrous except isolated yellowish setae on antennae, lateral margins of pronotum and along the first part of margins of elytra; line of short, close, erected brownish setae inserted in setigerous punctures along margins of inner part of protibia; long separeted, erected brownish setae inserted in setigerous punctures along margins of inner part of meso and metatibia, closer and shorter on inner face; underside with long, erected yellowish setae inserted in setigerous punctures on sides of metaventrite; long, erected, isolated yellowish setae along margin of anal plate; close and long setae on anterior and posterior margins of prosternum; abdominal ventrites with isolated, long, erected brownish-yellow setae inserted in setigerous punctures; pygidium glabrous.
</paragraph>
<paragraph id="8B9D36D0FFC0F10912A9A98811EA5162" blockId="11.[189,1399,1434,1587]" pageId="11" pageNumber="12">
<emphasis id="B956EAC2FFC0F10912A9A988125552E6" box="[227,351,1434,1463]" italics="true" pageId="11" pageNumber="12">Aedeagus</emphasis>
: in lateral view, parameres elongate and ventrally slightly sinuate before the apex; in dorsal view, parallel from base to middle part, laterally slightly curved in the third part, apex strongly diverging, apico-internal angle obtusely subtriangular; in ventral view, apex of median lobe rounded, apex of parameres incurved.
</paragraph>
<paragraph id="8B9D36D0FFC0F10912F7AA5D10FA513A" blockId="11.[189,1008,1614,1644]" box="[189,1008,1614,1644]" pageId="11" pageNumber="12">
FEMALE: Measurements: length: 
<quantity id="4CDA9B35FFC0F1091030AA5D100D513A" box="[634,775,1615,1644]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="11" pageNumber="12" unit="mm" value="17.5" valueMax="18.0" valueMin="17.0">17–18 mm</quantity>
, width: 
<quantity id="4CDA9B35FFC0F1091124AA5D10E6513A" box="[878,1004,1615,1644]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="11" pageNumber="12" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">9–10 mm</quantity>
.
</paragraph>
<paragraph id="8B9D36D0FFC0F10912A9AA951761519C" blockId="11.[189,1398,1671,1741]" pageId="11" pageNumber="12">
Differs from 
<specimenCount id="9D24FD59FFC0F10913C7AA9512AF51F2" box="[397,421,1671,1699]" count="1" pageId="11" pageNumber="12" type="male">♂</specimenCount>
by: abdomen convex; club of antennae slightly shorter; protibiae narrower with apical tooth rounded and longer; large claw of protibia not modified.
</paragraph>
<paragraph id="8B9D36D0FFC0F10912F7AAED12BD504D" blockId="11.[189,439,1791,1820]" box="[189,439,1791,1820]" pageId="11" pageNumber="12">DERIVATIO NOMINIS.</paragraph>
<paragraph id="8B9D36D0FFC0F10912F7AB2212CF5026" blockId="11.[189,1398,1840,1911]" pageId="11" pageNumber="12">
The species epithet is a noun in opposition, referring to the Koulango ethnic group from North- West of 
<collectingCountry id="F3357640FFC0F109136DAB4812CB5026" box="[295,449,1882,1911]" name="Ivory Coast" pageId="11" pageNumber="12">Ivory Coast</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C338655BFFC0F10912F7ABBB11FB50A7" pageId="11" pageNumber="12" type="distribution">
<paragraph id="8B9D36D0FFC0F10912F7ABBB127D5097" blockId="11.[189,375,1961,1990]" box="[189,375,1961,1990]" pageId="11" pageNumber="12">DISTRIBUTION.</paragraph>
<paragraph id="8B9D36D0FFC0F10912F7ABC811FB50A7" blockId="11.[189,753,2009,2039]" box="[189,753,2009,2039]" pageId="11" pageNumber="12">
Known from 
<collectingCountry id="F3357640FFC0F1091323ABC8110A50A6" box="[361,512,2010,2039]" name="Ivory Coast" pageId="11" pageNumber="12">Ivory Coast</collectingCountry>
and 
<collectingCountry id="F3357640FFC0F1091074ABCB11E450A7" box="[574,750,2009,2038]" name="Burkina Faso" pageId="11" pageNumber="12">Burkina Faso</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C338655BFFC7F10E12F7ACC61221561F" pageId="12" pageNumber="13" type="biology_ecology">
<paragraph id="8B9D36D0FFC7F10E12F7ACC6123F57A0" blockId="12.[189,309,212,241]" box="[189,309,212,241]" pageId="12" pageNumber="13">BIOLOGY.</paragraph>
<paragraph id="8B9D36D0FFC7F10E12F7AD141221561F" blockId="12.[189,1398,262,334]" pageId="12" pageNumber="13">The species was collected in the months of January, February, April, May, September and October.</paragraph>
</subSubSection>
<subSubSection id="C338655BFFC7F10E12F7AD9016235425" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="8B9D36D0FFC7F10E12F7AD90110856CE" blockId="12.[189,514,386,415]" box="[189,514,386,415]" pageId="12" pageNumber="13">DIFFERENTIAL DIAGNOSIS.</paragraph>
<paragraph id="8B9D36D0FFC7F10E12F7ADA6163456AA" blockId="12.[189,1398,436,507]" pageId="12" pageNumber="13">
<taxonomicName id="4C224D53FFC7F10E12F7ADA6125E5680" authority="Limbourg &amp; Dekoninck &amp; Seidel, 2024" authorityName="Limbourg &amp; Dekoninck &amp; Seidel" authorityYear="2024" box="[189,340,436,465]" class="Insecta" family="Scarabaeidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="koulango" status="sp. nov.">
<emphasis id="B956EAC2FFC7F10E12F7ADA6125E5680" box="[189,340,436,465]" italics="true" pageId="12" pageNumber="13">A. koulango</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557B9FFC7F10E131CADA612BC5681" box="[342,438,436,464]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
belongs to the homogeneous 
<emphasis id="B956EAC2FFC7F10E116CADA610B65680" box="[806,956,436,465]" italics="true" pageId="12" pageNumber="13">paleopyga–</emphasis>
group, reviewed by 
<bibRefCitation id="EFB34B21FFC7F10E16F8ADA613F556AA" author="MACHATSCHKE J. W." pageId="12" pageNumber="13" pagination="157 - 165" refId="ref16348" refString="MACHATSCHKE J. W., 1964. - Uber Anomala paleopyga (Benderitter, 1923) und ihre nachsten Verwandten (Coleoptera, Melolonthidae, Rutelinae). Reichenbachia, 4 (19): 157 - 165." type="journal article" year="1964">MACHATSCHKE, 1964</bibRefCitation>
, and strongly resembles 
<taxonomicName id="4C224D53FFC7F10E107DADCD107F56AA" authority="Ohaus, 1925" authorityName="Ohaus" authorityYear="1925" box="[567,885,478,507]" class="Insecta" family="Scarabaeidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dossidea">
<emphasis id="B956EAC2FFC7F10E107DADCD11C156AA" box="[567,715,478,507]" italics="true" pageId="12" pageNumber="13">A. dossidea</emphasis>
<bibRefCitation id="EFB34B21FFC7F10E1099ADCC107F56AA" author="OHAUS F." box="[723,885,478,507]" pageId="12" pageNumber="13" pagination="42 - 80" refId="ref16704" refString="OHAUS F., 1925. - Revision der afrikanischen Anomala - Arten (Col. lamell. Rutelin.). Stettiner Entomologische Zeitung 86: 42 - 80." type="journal article" year="1925">Ohaus, 1925</bibRefCitation>
</taxonomicName>
but can be distinguished as follows:
</paragraph>
<paragraph id="8B9D36D0FFC7F10E1281AE0D17F8556D" blockId="12.[203,1266,543,572]" box="[203,1266,543,572]" pageId="12" pageNumber="13">
<taxonomicName id="4C224D53FFC7F10E1281AE0D126C556D" authority="Limbourg &amp; Dekoninck &amp; Seidel, 2024" authorityName="Limbourg &amp; Dekoninck &amp; Seidel" authorityYear="2024" box="[203,358,543,572]" class="Insecta" family="Scarabaeidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="koulango" status="sp. nov.">
<emphasis id="B956EAC2FFC7F10E1281AE0D126C556D" box="[203,358,543,572]" italics="true" pageId="12" pageNumber="13">A. koulango</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557B9FFC7F10E1324AE3212DB556D" box="[366,465,544,572]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="13192A55FFC7F10E13ABAE0D1158556D" box="[481,594,543,572]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="9.[189,228,1958,1982]" captionStartId-1="10.[189,228,1958,1982]" captionTargetBox-0="[204,1357,382,1930]" captionTargetBox-1="[201,1380,421,1931]" captionTargetId-0="figure-79@9.[190,1397,364,1943]" captionTargetId-1="figure-12@10.[188,1400,399,1944]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="Fig. 3. Anomala koulango sp. nov. A-B, holotype ♂ (RBINS) habitus. A, dorsal view. B, lateral view. C-D, paratype ♀ (ZFMK) habitus. C, dorsal view. D, lateral view. E-G, aedeagus. E, lateral view. F, dorsal view. G, ventral view. H, distribution map of Anomala koulango sp. nov. and Anomala dossidea Ohaus, 1925." captionText-1="Fig. 4. Anomala dossidea Ohaus, 1925. A-B, ♂ (RBINS) habitus. A, dorsal view. B, lateral view. C-D, ♀ (RBINS) habitus. C, dorsal view. D, lateral view. E-G, aedeagus. E, lateral view. F, dorsal view. G, ventral view. H, disc of pronotum of Anomala koulango sp. nov. ♂. I, disc of pronotum of Anomala dossidea Ohaus, 1925 ♂." figureDoi-0="http://doi.org/10.5281/zenodo.10982047" figureDoi-1="http://doi.org/10.5281/zenodo.10982049" httpUri-0="https://zenodo.org/record/10982047/files/figure.png" httpUri-1="https://zenodo.org/record/10982049/files/figure.png" pageId="12" pageNumber="13">Figs 3–4</figureCitation>
) 
<taxonomicName id="4C224D53FFC7F10E1162AE0D176F556D" authority="Ohaus, 1925" authorityName="Ohaus" authorityYear="1925" box="[808,1125,543,572]" class="Insecta" family="Scarabaeidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dossidea">
<emphasis id="B956EAC2FFC7F10E1162AE0D10B6556D" box="[808,956,543,572]" italics="true" pageId="12" pageNumber="13">A. dossidea</emphasis>
<bibRefCitation id="EFB34B21FFC7F10E118EAE0D176F556D" author="OHAUS F." box="[964,1125,543,572]" pageId="12" pageNumber="13" pagination="42 - 80" refId="ref16704" refString="OHAUS F., 1925. - Revision der afrikanischen Anomala - Arten (Col. lamell. Rutelin.). Stettiner Entomologische Zeitung 86: 42 - 80." type="journal article" year="1925">Ohaus, 1925</bibRefCitation>
</taxonomicName>
(
<figureCitation id="13192A55FFC7F10E163CAE0D17E2556D" box="[1142,1256,543,572]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="9.[189,228,1958,1982]" captionStartId-1="10.[189,228,1958,1982]" captionTargetBox-0="[204,1357,382,1930]" captionTargetBox-1="[201,1380,421,1931]" captionTargetId-0="figure-79@9.[190,1397,364,1943]" captionTargetId-1="figure-12@10.[188,1400,399,1944]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="Fig. 3. Anomala koulango sp. nov. A-B, holotype ♂ (RBINS) habitus. A, dorsal view. B, lateral view. C-D, paratype ♀ (ZFMK) habitus. C, dorsal view. D, lateral view. E-G, aedeagus. E, lateral view. F, dorsal view. G, ventral view. H, distribution map of Anomala koulango sp. nov. and Anomala dossidea Ohaus, 1925." captionText-1="Fig. 4. Anomala dossidea Ohaus, 1925. A-B, ♂ (RBINS) habitus. A, dorsal view. B, lateral view. C-D, ♀ (RBINS) habitus. C, dorsal view. D, lateral view. E-G, aedeagus. E, lateral view. F, dorsal view. G, ventral view. H, disc of pronotum of Anomala koulango sp. nov. ♂. I, disc of pronotum of Anomala dossidea Ohaus, 1925 ♂." figureDoi-0="http://doi.org/10.5281/zenodo.10982047" figureDoi-1="http://doi.org/10.5281/zenodo.10982049" httpUri-0="https://zenodo.org/record/10982047/files/figure.png" httpUri-1="https://zenodo.org/record/10982049/files/figure.png" pageId="12" pageNumber="13">Figs 3–4</figureCitation>
)
</paragraph>
<paragraph id="8B9D36D0FFC7F10E1281AE7F12715475" blockId="12.[203,775,621,804]" pageId="12" pageNumber="13">
<emphasis id="B956EAC2FFC7F10E1281AE7F124D55DB" box="[203,327,621,650]" italics="true" pageId="12" pageNumber="13">Aedeagus</emphasis>
: parameres in dorsal view, parallel from base to middle part, laterally slightly curved in the third part, apex strongly diverging, apico-internal angle obtusely subtriangular.
</paragraph>
<paragraph id="8B9D36D0FFC7F10E1162AE7F178E5586" blockId="12.[808,1380,621,727]" pageId="12" pageNumber="13">
<emphasis id="B956EAC2FFC7F10E1162AE7F10AE55DB" box="[808,932,621,650]" italics="true" pageId="12" pageNumber="13">Aedeagus</emphasis>
: parameres in dorsal view, parallel from base to apex; apico-internal angle rounded with a deep notch.
</paragraph>
<paragraph id="8B9D36D0FFC7F10E1281AF4A16235425" blockId="12.[203,1321,855,884]" box="[203,1321,855,884]" pageId="12" pageNumber="13">
Punctures of pronotum deeper (
<figureCitation id="13192A55FFC7F10E1012AF4511B45425" box="[600,702,855,884]" captionStart="Fig" captionStartId="10.[189,228,1958,1982]" captionTargetBox="[201,1380,421,1931]" captionTargetId="figure-12@10.[188,1400,399,1944]" captionTargetPageId="10" captionText="Fig. 4. Anomala dossidea Ohaus, 1925. A-B, ♂ (RBINS) habitus. A, dorsal view. B, lateral view. C-D, ♀ (RBINS) habitus. C, dorsal view. D, lateral view. E-G, aedeagus. E, lateral view. F, dorsal view. G, ventral view. H, disc of pronotum of Anomala koulango sp. nov. ♂. I, disc of pronotum of Anomala dossidea Ohaus, 1925 ♂." figureDoi="http://doi.org/10.5281/zenodo.10982049" httpUri="https://zenodo.org/record/10982049/files/figure.png" pageId="12" pageNumber="13">Fig. 4H</figureCitation>
). Punctures of pronotum shallow (
<figureCitation id="13192A55FFC7F10E1688AF4516115425" box="[1218,1307,855,884]" captionStart="Fig" captionStartId="10.[189,228,1958,1982]" captionTargetBox="[201,1380,421,1931]" captionTargetId="figure-12@10.[188,1400,399,1944]" captionTargetPageId="10" captionText="Fig. 4. Anomala dossidea Ohaus, 1925. A-B, ♂ (RBINS) habitus. A, dorsal view. B, lateral view. C-D, ♀ (RBINS) habitus. C, dorsal view. D, lateral view. E-G, aedeagus. E, lateral view. F, dorsal view. G, ventral view. H, disc of pronotum of Anomala koulango sp. nov. ♂. I, disc of pronotum of Anomala dossidea Ohaus, 1925 ♂." figureDoi="http://doi.org/10.5281/zenodo.10982049" httpUri="https://zenodo.org/record/10982049/files/figure.png" pageId="12" pageNumber="13">Fig. 4I</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>