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<document id="DB89EF5D2D971ACCE0F51563E19B459F" ID-DOI="10.11646/zootaxa.3652.3.3" ID-GBIF-Dataset="664a75d3-f878-42de-a8b4-85a359928c94" ID-ISSN="1175-5326" ID-Zenodo-Dep="221749" ID-ZooBank="B3BDA986-3C7B-45B9-8B55-754F1B28815C" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460610058317" checkinUser="plazi" docAuthor="Artal, Pedro, Van Bakel, Barry W. M., Fraaije, René H. B. &amp; Jagt, John W. M." docDate="2013" docId="038B87C6005A3D73FF2EED64FE783F42" docLanguage="en" docName="zt03652p352.pdf" docOrigin="Zootaxa 3652 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Serrablopluma Artal, Bakel, Fraaije &amp; Jagt, 2013, n. gen." docType="treatment" docVersion="8" lastPageNumber="348" masterDocId="FFB2FFBE005B3D76FFB9EB09FFEC3E5E" masterDocTitle="New retroplumid crabs (Crustacea, Brachyura, Retroplumidae Gill, 1894) from the Eocene of Huesca (Aragón, Spain)" masterLastPageNumber="352" masterPageNumber="343" pageNumber="344" updateTime="1698328574862" updateUser="plazi">
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<mods:title id="0BEFEE2F42CE8E5834DF39C3E2297239">New retroplumid crabs (Crustacea, Brachyura, Retroplumidae Gill, 1894) from the Eocene of Huesca (Aragón, Spain)</mods:title>
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<mods:namePart id="CD343D34CAC9F55F1CD1D7EF944969BF">Artal, Pedro</mods:namePart>
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<mods:namePart id="BF1FC12A8C47596F41F8DB9013383289">Van Bakel, Barry W. M.</mods:namePart>
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<mods:namePart id="ED01C73905A1E9EEFCCA1FA168F40382">Fraaije, René H. B.</mods:namePart>
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<mods:namePart id="D5E8A73B15BA989E1F3F364A126E6F61">Jagt, John W. M.</mods:namePart>
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<treatment id="038B87C6005A3D73FF2EED64FE783F42" ID-DOI="http://doi.org/10.5281/zenodo.6159017" ID-GBIF-Taxon="119568245" ID-Zenodo-Dep="6159017" LSID="urn:lsid:plazi:treatment:038B87C6005A3D73FF2EED64FE783F42" httpUri="http://treatment.plazi.org/id/038B87C6005A3D73FF2EED64FE783F42" lastPageId="5" lastPageNumber="348" pageId="1" pageNumber="344">
<subSubSection id="C338655B005A3D77FF2EED64FE4938D9" box="[151,421,1645,1671]" pageId="1" pageNumber="344" type="nomenclature">
<paragraph id="8B9D36D0005A3D77FF2EED64FE4938D9" blockId="1.[151,421,1645,1671]" box="[151,421,1645,1671]" pageId="1" pageNumber="344">
<heading id="D0D581BC005A3D77FF2EED64FE4938D9" bold="true" box="[151,421,1645,1671]" fontSize="11" level="1" pageId="1" pageNumber="344" reason="1">
<emphasis id="B956EAC2005A3D77FF2EED64FE4938D9" bold="true" box="[151,421,1645,1671]" pageId="1" pageNumber="344">
<taxonomicName id="4C224D53005A3D77FF2EED64FEA038D9" ID-CoL="c8990422-c0e4-42ec-bec3-7102385f99c0" box="[151,332,1645,1671]" class="Malacostraca" family="Retroplumidae" genus="Serrablopluma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="344" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B956EAC2005A3D77FF2EED64FEA038D9" bold="true" box="[151,332,1645,1671]" italics="true" pageId="1" pageNumber="344">Serrablopluma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557B9005A3D77FEEAED64FE4938D9" box="[339,421,1645,1671]" pageId="1" pageNumber="344" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C338655B005A3D77FF2EEDBDFD7D3892" box="[151,657,1715,1740]" pageId="1" pageNumber="344" type="materials_examined">
<paragraph id="8B9D36D0005A3D77FF2EEDBDFD7D3892" blockId="1.[151,1437,1715,2028]" box="[151,657,1715,1740]" pageId="1" pageNumber="344">
<emphasis id="B956EAC2005A3D77FF2EEDBDFEC73892" bold="true" box="[151,299,1715,1740]" pageId="1" pageNumber="344">
<typeStatus id="54998872005A3D77FF2EEDBDFF3E3892" box="[151,210,1716,1740]" pageId="1" pageNumber="344">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="4C224D53005A3D77FE80EDBDFDA73892" box="[313,587,1716,1740]" class="Malacostraca" family="Retroplumidae" genus="Serrablopluma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="344" phylum="Arthropoda" rank="species" species="diminuta" status="sp. nov.">
<emphasis id="B956EAC2005A3D77FE80EDBDFDA73892" box="[313,587,1716,1740]" italics="true" pageId="1" pageNumber="344">Serrablopluma diminuta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557B9005A3D77FDEBEDBDFD7D3892" box="[594,657,1716,1740]" pageId="1" pageNumber="344" rank="species">
<emphasis id="B956EAC2005A3D77FDEBEDBDFD7D3892" bold="true" box="[594,657,1716,1740]" pageId="1" pageNumber="344">n. sp.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C338655B005A3D74FF7EEDDEFB663E8A" lastPageId="2" lastPageNumber="345" pageId="1" pageNumber="344" type="diagnosis">
<paragraph id="8B9D36D0005A3D74FF7EEDDEFB663E8A" blockId="1.[151,1437,1715,2028]" lastBlockId="2.[151,1436,151,391]" lastPageId="2" lastPageNumber="345" pageId="1" pageNumber="344">
<emphasis id="B956EAC2005A3D77FF7EEDDEFED638AE" bold="true" box="[199,314,1751,1776]" pageId="1" pageNumber="344">Diagnosis</emphasis>
. Small size carapace, subtrapezoidal in outline, much wider than long, widest anteriorly, maximum width at position of lateral spines; anterior margin wide, broadly sinuous, terminating in fairly widely projecting lateral spines; front conspicuously reduced, narrow; orbits wide, with sub-orbital spine; lateral margins straight, inclined, converging posteriorly; posterior margin wide, broadly convex; dorsal carapace surface flat in both directions, with 3 strongly marked ridges; anterior ridge continuous, intersecting anterior margin; medium ridge interrupted only by gastric grooves; posterior ridge continuous, straight. Thoracic sternum broad; sternites 3, 4 trapezoidal, sternites 57 subrectangular; vulvae large, subcircular, situated below extremity of sutures 5/6; abdomen subtriangular in both sexes; female abdomen much broader than males; thoracic sternites, abdominal somites notably ridged; thoracic sternite 5, abdominal somite 6 with distinct abdominal holding mechanism; chelipeds equal, robust, relatively large; P2P4 long, flattened; P5 conspicuously reduced.
</paragraph>
</subSubSection>
<subSubSection id="C338655B00593D74FF7EEBE9FD8B3F42" pageId="2" pageNumber="345" type="etymology">
<paragraph id="8B9D36D000593D74FF7EEBE9FD8B3F42" blockId="2.[151,1436,151,391]" pageId="2" pageNumber="345">
<emphasis id="B956EAC200593D74FF7EEBE9FEA23EA6" bold="true" box="[199,334,224,248]" pageId="2" pageNumber="345">Etymology.</emphasis>
From Serrablo, the mediaeval name of the area (Pyrenees, Huesca) and the ending
<emphasis id="B956EAC200593D74FABBEBE9FAA43EA6" box="[1282,1352,224,248]" italics="true" pageId="2" pageNumber="345">pluma</emphasis>
, which refers to the main character of the family.
</paragraph>
</subSubSection>
<subSubSection id="C338655B00593D74FF7EEA21FEFE3C49" pageId="2" pageNumber="345" type="discussion">
<paragraph id="8B9D36D000593D74FF7EEA21FB5D3FD9" blockId="2.[151,1436,151,391]" pageId="2" pageNumber="345">
<emphasis id="B956EAC200593D74FF7EEA21FED73F1E" bold="true" box="[199,315,296,320]" pageId="2" pageNumber="345">Remarks.</emphasis>
The new genus differs from all others currently included in the
<taxonomicName id="4C224D5300593D74FBB5EA2EFB5E3F61" box="[1036,1202,295,319]" class="Malacostraca" family="Retroplumidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="345" phylum="Arthropoda" rank="family">Retroplumidae</taxonomicName>
(see De Grave
<emphasis id="B956EAC200593D74FADAEA20FA793F1E" box="[1379,1429,296,320]" italics="true" pageId="2" pageNumber="345">et al</emphasis>
. 2009: 41) in having a wide, broadly arched anterior margin, a conspicuously reduced front and an anterior dorsal ridge that intersects the anterior margin at the position of long, fairly projecting lateral spines.
</paragraph>
<paragraph id="8B9D36D000593D74FF2EEAD2FEFE3C49" blockId="2.[151,504,475,535]" pageId="2" pageNumber="345">
<emphasis id="B956EAC200593D74FF2EEAD2FE143FAA" bold="true" box="[151,504,475,500]" pageId="2" pageNumber="345">
<taxonomicName id="4C224D5300593D74FF2EEAD2FE5F3FAA" box="[151,435,475,500]" class="Malacostraca" family="Retroplumidae" genus="Serrablopluma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="345" phylum="Arthropoda" rank="species" species="diminuta" status="sp. nov.">
<emphasis id="B956EAC200593D74FF2EEAD2FE5F3FAA" bold="true" box="[151,435,475,500]" italics="true" pageId="2" pageNumber="345">Serrablopluma diminuta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557B900593D74FE00EAD5FE143FAA" box="[441,504,476,500]" pageId="2" pageNumber="345" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13192A5500593D74FF26EAF6FEE53C49" box="[159,265,511,535]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[151,250,1855,1878]" captionStartId-1="4.[151,250,1832,1855]" captionTargetBox-0="[164,1436,195,1817]" captionTargetBox-1="[150,1430,193,1814]" captionTargetId-0="figure@3.[164,1449,195,1817]" captionTargetId-1="figure@4.[146,1431,193,1815]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 1. Serrablopluma diminuta n. gen., n. sp., MGSB 75287 (holotype, indeterminate sex); A, dorsal view. MAB k. 3279 (paratype, male); B, ventral view. MGSB 75290 a (paratype, female); C, ventral view. MGSB 75289 (paratype, male); D, ventral view. MGSB 75290 b (paratype, female); E, ventral view. All from the upper Eocene (Priabonian) of Fanlillo (municipality of Yebra de Basa, Huesca, northeast Spain). Scale bars equal 5 mm." captionText-1="FIGURE 2. Serrablopluma diminuta n. gen., n. sp., MGSB 78335 a (paratype, indeterminate sex; anterior carapace margin damaged); A, oblique frontal view, showing orbits; C, dorsal view. MGSB 78334 a (paratype, indeterminate sex; anterior carapace margin damaged); B, oblique frontal view, showing orbits. MGSB 78334 b (paratype, female partial thoracic sternum); D, ventral view showing vulva and reduced sternite 8. MGSB 78335 c (paratype, male); E, posteroventral view, showing reduced P 5. All from the upper Eocene (Priabonian) of Fanlillo (municipality of Yebra de Basa, Huesca, northeast Spain). Scale bars equal 5 mm (A D), 2 mm (E). O, orbit; sos, sub-orbital spine; ls, lateral spine; V, vulva; 4 / 5, 5 / 6, 6 / 7, 7 / 8, thoracic sternal suture 4 / 5, 5 / 6, 6 / 7, 7 / 8; St 4, St 5, St 6, St 7, St 8, thoracic sternites 4, 5, 6, 7, 8; cx 4, cx 5, P 4, P 5 coxae; mP 4, mP 5, P 4, P 5 merus." httpUri-0="https://zenodo.org/record/221750/files/figure.png" httpUri-1="https://zenodo.org/record/221751/files/figure.png" pageId="2" pageNumber="345">Figs. 12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C338655B00593D74FF2EE949FE433C04" box="[151,431,576,602]" pageId="2" pageNumber="345" type="diagnosis">
<paragraph id="8B9D36D000593D74FF2EE949FE433C04" blockId="2.[151,1437,576,2006]" box="[151,431,576,602]" pageId="2" pageNumber="345">
<emphasis id="B956EAC200593D74FF2EE949FEFD3C07" bold="true" box="[151,273,576,601]" pageId="2" pageNumber="345">Diagnosis.</emphasis>
As for genus.
</paragraph>
</subSubSection>
<subSubSection id="C338655B00593D74FF7EE96CFBF33C23" box="[199,1055,613,637]" pageId="2" pageNumber="345" type="etymology">
<paragraph id="8B9D36D000593D74FF7EE96CFBF33C23" blockId="2.[151,1437,576,2006]" box="[199,1055,613,637]" pageId="2" pageNumber="345">
<emphasis id="B956EAC200593D74FF7EE96CFEA33C23" bold="true" box="[199,335,613,637]" pageId="2" pageNumber="345">Etymology.</emphasis>
From the Latin
<emphasis id="B956EAC200593D74FDBCE96CFD9F3C23" box="[517,627,613,637]" italics="true" pageId="2" pageNumber="345">diminutus</emphasis>
, in allusion to the rather reduced size.
</paragraph>
</subSubSection>
<subSubSection id="C338655B00593D74FF7EE980FC4C3D53" pageId="2" pageNumber="345" type="materials_examined">
<paragraph id="8B9D36D000593D74FF7EE980FC4C3D53" blockId="2.[151,1437,576,2006]" pageId="2" pageNumber="345">
<emphasis id="B956EAC200593D74FF7EE980FE503CFC" bold="true" box="[199,444,649,674]" pageId="2" pageNumber="345">Material examined.</emphasis>
MGSB75287,
<typeStatus id="5499887200593D74FDC6E980FD093CFF" box="[639,741,649,673]" pageId="2" pageNumber="345" type="holotype">holotype</typeStatus>
; MGSB75289, MGSB75290a, b, MGSB78334a, b, and MGSB78335a, b, c,
<typeStatus id="5499887200593D74FE3FE9A7FE193C98" box="[390,501,686,710]" pageId="2" pageNumber="345" type="paratype">paratypes</typeStatus>
; MAB k. 32793281, MAB k. 3293, MAB k. 3295,
<typeStatus id="5499887200593D74FBE5E9A7FB273C98" box="[1116,1227,686,710]" pageId="2" pageNumber="345" type="paratype">paratypes</typeStatus>
, all from Fanlillo, municipality of Yebra de Basa, Huesca, and of late Eocene (Priabonian) age. In the
<typeStatus id="5499887200593D74FB3EE9D8FB003CB7" box="[1159,1260,721,745]" pageId="2" pageNumber="345" type="holotype">holotype</typeStatus>
, the maximum carapace width and length (in millimetres) are 12 and 9, respectively.
</paragraph>
</subSubSection>
<subSubSection id="C338655B00593D74FF7EE810FD9F3B2F" pageId="2" pageNumber="345" type="description">
<paragraph id="8B9D36D000593D74FF7EE810FAF63A0F" blockId="2.[151,1437,576,2006]" pageId="2" pageNumber="345">
<emphasis id="B956EAC200593D74FF7EE810FEB53D6C" bold="true" box="[199,345,793,818]" pageId="2" pageNumber="345">Description.</emphasis>
Carapace small (maximum carapace width
<quantity id="4CDA9B3500593D74FCF7E810FC4F3D6F" box="[846,931,793,817]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="345" unit="mm" value="12.0">12 mm</quantity>
); subtrapezoidal in outline; wider than long, widest anteriorly at level of lateral spines; dorsal surface flat in both directions. Anterior margin sinuous, broadly arched, with projecting lateral spines. Front conspicuously reduced, narrow, directed forwards; orbits wide, with sub-orbital spine. Lateral margins straight, inclined, converging posteriorly. Posterior margin convex, wide, only slightly shorter than orbitofrontal margin. Dorsal surface with 3 ridges; anteriormost intersecting anterior margins laterally; median ridge defined by epibranchial swellings, interrupted by gastric grooves; posterior ridge straight, continuous, interrupted only by small indentations of subtle branchiocardiac grooves. Dorsal surface smooth except for horizontal ridges, shallow grooves. Mesogastric region defined by 2 faint swellings, bounded by weak posterior grooves. Cardiac region large, gently swollen, sub-rhomboidal in shape. Intestinal region flat.
</paragraph>
<paragraph id="8B9D36D000593D74FF7EEF54FD9F3B2F" blockId="2.[151,1437,576,2006]" pageId="2" pageNumber="345">Thoracic sternum much broader than long, sternites with subtle ridges; sternites 1,2 fused, small, subtriangular; sternites 3, 4 subtrapezoidal; sternite 4 with pronounced lateral extensions; small, weak indentation between sternites 3, 4; sternites 57 subrectangular, with subtle inclined ridges; sternite 8 conspicuously reduced, oblique; vulvae large, subcircular, situated below extremity of suture 5/6. Male abdomen subtriangular, with segments 35 fused, all segments with pronounced horizontal ridge; female abdomen much broader, all segments separated, with marked ridges. Abdominal holding mechanism with press-button on sternite 5, and lateral extensions of abdominal segment 6. P1 subequal, robust, relatively large, strong, inner portion of dactyli with numerous denticles. P2P4 long, flattened. P5 conspicuously reduced.</paragraph>
</subSubSection>
<subSubSection id="C338655B00593D73FF7EEE74FE783F42" lastPageId="5" lastPageNumber="348" pageId="2" pageNumber="345" type="discussion">
<paragraph id="8B9D36D000593D74FF7EEE74FEA03833" blockId="2.[151,1437,576,2006]" pageId="2" pageNumber="345">
<emphasis id="B956EAC200593D74FF7EEE74FED83BCB" bold="true" box="[199,308,1405,1429]" pageId="2" pageNumber="345">Remarks</emphasis>
.
<taxonomicName id="4C224D5300593D74FEFDEE74FDBB3BCB" box="[324,599,1405,1429]" class="Malacostraca" family="Retroplumidae" genus="Serrablopluma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="345" phylum="Arthropoda" rank="species" species="diminuta" status="gen. nov.">
<emphasis id="B956EAC200593D74FEFDEE74FDBB3BCB" box="[324,599,1405,1429]" italics="true" pageId="2" pageNumber="345">Serrablopluma diminuta</emphasis>
</taxonomicName>
<emphasis id="B956EAC200593D74FDD8EE74FCEE3BCB" bold="true" box="[609,770,1405,1429]" pageId="2" pageNumber="345">
<taxonomicNameLabel id="A26557B900593D74FDD8EE74FD5E3BCB" box="[609,690,1405,1429]" pageId="2" pageNumber="345" rank="genus">n. gen.</taxonomicNameLabel>
, n. sp.
</emphasis>
shows the plesiomorphic characters of
<taxonomicName id="4C224D5300593D74FB70EE74FA833BCB" box="[1225,1391,1405,1429]" class="Malacostraca" family="Retroplumidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="345" phylum="Arthropoda" rank="family">Retroplumidae</taxonomicName>
(de Saint Laurent 1989: 111112), namely a sinuous anterior margin and a narrow front; a set of dorsal transverse ridges; a thoracic sternum that is much broader than long; subtrapezoidal sternites 3, 4, with pronounced lateral extensions in sternite 4 behind a distinct notch in sternite 3; distinctly wide, sub-rectangular sternites 57; a conspicuously reduced, oblique sternite 8; distinct sexual dimorphism for abdomen; P2P4 long, flattened; P5 conspicuously reduced, sub-dorsal. The thoracic sternum is typical of
<taxonomicName id="4C224D5300593D74FC68ED38FB9B3817" box="[977,1143,1585,1609]" class="Malacostraca" family="Retroplumidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="345" phylum="Arthropoda" rank="family">Retroplumidae</taxonomicName>
(see Guinot &amp; Quenette 2005: fig. 29A).
</paragraph>
<paragraph id="8B9D36D000593D74FF7EED70FAAF38A3" blockId="2.[151,1437,576,2006]" pageId="2" pageNumber="345">
Genera that resemble
<taxonomicName id="4C224D5300593D74FE61ED70FD6C38CF" box="[472,640,1657,1681]" class="Malacostraca" family="Retroplumidae" genus="Serrablopluma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="345" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B956EAC200593D74FE61ED70FD6C38CF" box="[472,640,1657,1681]" italics="true" pageId="2" pageNumber="345">Serrablopluma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557B900593D74FD29ED73FD0438CC" box="[656,744,1658,1682]" pageId="2" pageNumber="345" rank="genus">
<emphasis id="B956EAC200593D74FD29ED73FD0438CC" bold="true" box="[656,744,1658,1682]" pageId="2" pageNumber="345">n. gen.</emphasis>
</taxonomicNameLabel>
are
<taxonomicName id="4C224D5300593D74FC92ED70FC5B38CF" box="[811,951,1657,1681]" class="Malacostraca" family="Retroplumidae" genus="Loerentheya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="345" phylum="Arthropoda" rank="genus">
<emphasis id="B956EAC200593D74FC92ED70FC5B38CF" box="[811,951,1657,1681]" italics="true" pageId="2" pageNumber="345">Loerentheya</emphasis>
</taxonomicName>
(see Glaessner 1969: fig. 338, 5) and
<taxonomicName id="4C224D5300593D74FF2EED94FEBC38EB" box="[151,336,1693,1717]" class="Malacostraca" family="Retroplumidae" genus="Loerenthopluma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="345" phylum="Arthropoda" rank="genus">
<emphasis id="B956EAC200593D74FF2EED94FEBC38EB" box="[151,336,1693,1717]" italics="true" pageId="2" pageNumber="345">Loerenthopluma</emphasis>
</taxonomicName>
, both of which have a sub-trapezoidal carapace, with the maximum width anteriorly and a wide anterior margin.
<taxonomicName id="4C224D5300593D74FEEBEDC8FE163887" box="[338,506,1729,1753]" class="Malacostraca" family="Retroplumidae" genus="Serrablopluma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="345" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B956EAC200593D74FEEBEDC8FE163887" box="[338,506,1729,1753]" italics="true" pageId="2" pageNumber="345">Serrablopluma</emphasis>
</taxonomicName>
<emphasis id="B956EAC200593D74FDB8EDCBFDBA3884" bold="true" box="[513,598,1730,1754]" pageId="2" pageNumber="345">
<taxonomicNameLabel id="A26557B900593D74FDB8EDCBFDBC3884" box="[513,592,1730,1754]" pageId="2" pageNumber="345" rank="genus">n. gen.</taxonomicNameLabel>
,
</emphasis>
however, shows distinct characters such as the anterior carapace ridge that intersects the anterior margin, a markedly long lateral spine and lateral margins that are uniformly straight.
</paragraph>
<paragraph id="8B9D36D000593D74FF7EEC00FC37398B" blockId="2.[151,1437,576,2006]" pageId="2" pageNumber="345">
The extinct genera
<taxonomicName id="4C224D5300593D74FE09EC00FDD8397F" box="[432,564,1801,1825]" class="Malacostraca" family="Retroplumidae" genus="Retropluma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="345" phylum="Arthropoda" rank="genus">
<emphasis id="B956EAC200593D74FE09EC00FDD8397F" box="[432,564,1801,1825]" italics="true" pageId="2" pageNumber="345">Retropluma</emphasis>
</taxonomicName>
and
<taxonomicName id="4C224D5300593D74FDCFEC00FCE9397F" box="[630,773,1801,1825]" class="Malacostraca" family="Retroplumidae" genus="Retrocypoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="345" phylum="Arthropoda" rank="genus">
<emphasis id="B956EAC200593D74FDCFEC00FCE9397F" box="[630,773,1801,1825]" italics="true" pageId="2" pageNumber="345">Retrocypoda</emphasis>
</taxonomicName>
differ (see Beschin
<emphasis id="B956EAC200593D74FC46EC03FBDA397F" box="[1023,1078,1801,1825]" italics="true" pageId="2" pageNumber="345">et al</emphasis>
. 1996: figs 4, 5) in having a subrectangular carapace outline, a much narrower anterior margin, lateral margins broadly arched, not diverging posteriorly and an anterior dorsal carapace ridge without lateral spines. The carapace outline is sub-trapezoidal in
<taxonomicName id="4C224D5300593D74FF2EEC7CFED339D3" box="[151,319,1909,1933]" class="Malacostraca" family="Retroplumidae" genus="Serrablopluma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="345" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B956EAC200593D74FF2EEC7CFED339D3" box="[151,319,1909,1933]" italics="true" pageId="2" pageNumber="345">Serrablopluma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557B900593D74FEECEC7FFE5839D0" box="[341,436,1910,1934]" pageId="2" pageNumber="345" rank="genus">
<emphasis id="B956EAC200593D74FEECEC7FFE5839D0" bold="true" box="[341,436,1910,1934]" pageId="2" pageNumber="345">n. gen.</emphasis>
</taxonomicNameLabel>
, the anterior margin conspicuously wide, clearly broader than the posterior margin;moreover, the anterior margins terminate in a long lateral spine intersecting the first anterior dorsal ridge, while lateral margins are straight and continuous and converge posteriorly.
</paragraph>
<caption id="DF5D665800583D75FF2EEC36FCC639EF" httpUri="https://zenodo.org/record/221750/files/figure.png" pageId="3" pageNumber="346" targetBox="[164,1436,195,1817]" targetPageId="3">
<paragraph id="8B9D36D000583D75FF2EEC36FCC639EF" blockId="3.[151,1436,1855,1969]" pageId="3" pageNumber="346">
<emphasis id="B956EAC200583D75FF2EEC36FEE7390B" bold="true" box="[151,267,1855,1878]" pageId="3" pageNumber="346">FIGURE 1</emphasis>
.
<taxonomicName id="4C224D5300583D75FEA1EC36FDFC390B" box="[280,528,1855,1877]" class="Malacostraca" family="Retroplumidae" genus="Serrablopluma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="diminuta" status="gen. nov.">
<emphasis id="B956EAC200583D75FEA1EC36FDFC390B" box="[280,528,1855,1877]" italics="true" pageId="3" pageNumber="346">Serrablopluma diminuta</emphasis>
</taxonomicName>
<emphasis id="B956EAC200583D75FDAFEC49FD70390B" bold="true" box="[534,668,1856,1877]" pageId="3" pageNumber="346">
<taxonomicNameLabel id="A26557B900583D75FDAFEC49FDB3390B" box="[534,607,1856,1877]" pageId="3" pageNumber="346" rank="genus">n. gen.</taxonomicNameLabel>
, n. sp
</emphasis>
., MGSB75287 (holotype, indeterminate sex); A, dorsal view. MAB k. 3279 (paratype, male); B, ventral view. MGSB75290a (paratype, female); C, ventral view. MGSB75289 (paratype, male); D, ventral view. MGSB75290b (paratype, female); E, ventral view. All from the upper Eocene (Priabonian) of Fanlillo (municipality of Yebra de Basa, Huesca, northeast Spain). Scale bars equal 5 mm.
</paragraph>
</caption>
<caption id="DF5D6658005F3D72FF2EEC21FA7039A8" httpUri="https://zenodo.org/record/221751/files/figure.png" pageId="4" pageNumber="347" targetBox="[150,1430,193,1814]" targetPageId="4">
<paragraph id="8B9D36D0005F3D72FF2EEC21FA7039A8" blockId="4.[151,1436,1832,2039]" pageId="4" pageNumber="347">
<emphasis id="B956EAC2005F3D72FF2EEC21FEE33963" bold="true" box="[151,271,1832,1855]" pageId="4" pageNumber="347">FIGURE 2</emphasis>
.
<taxonomicName id="4C224D53005F3D72FEA6EC21FDF63960" box="[287,538,1832,1854]" class="Malacostraca" family="Retroplumidae" genus="Serrablopluma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="347" phylum="Arthropoda" rank="species" species="diminuta" status="gen. nov.">
<emphasis id="B956EAC2005F3D72FEA6EC21FDF63960" box="[287,538,1832,1854]" italics="true" pageId="4" pageNumber="347">Serrablopluma diminuta</emphasis>
</taxonomicName>
<emphasis id="B956EAC2005F3D72FD9DEC20FD5F3960" bold="true" box="[548,691,1833,1854]" pageId="4" pageNumber="347">
<taxonomicNameLabel id="A26557B9005F3D72FD9DEC20FD833960" box="[548,623,1833,1854]" pageId="4" pageNumber="347" rank="genus">n. gen.</taxonomicNameLabel>
, n. sp
</emphasis>
., MGSB78335a (paratype, indeterminate sex; anterior carapace margin damaged); A, oblique frontal view, showing orbits; C, dorsal view. MGSB78334a (paratype, indeterminate sex; anterior carapace margin damaged); B, oblique frontal view, showing orbits. MGSB78334b (paratype, female partial thoracic sternum); D, ventral view showing vulva and reduced sternite 8. MGSB78335c (paratype, male); E, posteroventral view, showing reduced P5. All from the upper Eocene (Priabonian) of Fanlillo (municipality of Yebra de Basa, Huesca, northeast Spain). Scale bars equal 5 mm (AD), 2 mm (E).
<emphasis id="B956EAC2005F3D72FDBBECC8FDF03986" bold="true" box="[514,540,1985,2008]" pageId="4" pageNumber="347">O,</emphasis>
orbit;
<emphasis id="B956EAC2005F3D72FDD9ECCBFD6B3989" bold="true" box="[608,647,1986,2007]" pageId="4" pageNumber="347">sos,</emphasis>
sub-orbital spine;
<emphasis id="B956EAC2005F3D72FCFDECCBFCB73989" bold="true" box="[836,859,1986,2007]" pageId="4" pageNumber="347">ls,</emphasis>
lateral spine;
<emphasis id="B956EAC2005F3D72FC52ECCBFC113989" bold="true" box="[1003,1021,1986,2007]" pageId="4" pageNumber="347">V,</emphasis>
vulva;
<emphasis id="B956EAC2005F3D72FBF5ECC8FB163986" bold="true" box="[1100,1274,1985,2008]" pageId="4" pageNumber="347">4/5, 5/6, 6/7, 7/8,</emphasis>
thoracic sternal suture 4/5, 5/6, 6/7, 7/8;
<emphasis id="B956EAC2005F3D72FE37ECE9FD8239A9" bold="true" box="[398,622,2016,2039]" pageId="4" pageNumber="347">St4, St5, St6, St7, St8,</emphasis>
thoracic sternites 4, 5, 6, 7, 8;
<emphasis id="B956EAC2005F3D72FC1DECE9FC1339A8" bold="true" box="[932,1023,2016,2038]" pageId="4" pageNumber="347">cx4, cx5,</emphasis>
P4, P5 coxae;
<emphasis id="B956EAC2005F3D72FB2CECE9FAE639A8" bold="true" box="[1173,1290,2016,2038]" pageId="4" pageNumber="347">mP4, mP5,</emphasis>
P4, P5 merus.
</paragraph>
</caption>
<paragraph id="8B9D36D0005E3D73FF7EEB9EFE783F42" blockId="5.[151,1436,151,284]" pageId="5" pageNumber="348">
Species of
<taxonomicName id="4C224D53005E3D73FEF3EB91FE003EEE" box="[330,492,152,176]" class="Malacostraca" family="Retroplumidae" genus="Costacopluma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="B956EAC2005E3D73FEF3EB91FE003EEE" box="[330,492,152,176]" italics="true" pageId="5" pageNumber="348">Costacopluma</emphasis>
</taxonomicName>
and
<taxonomicName id="4C224D53005E3D73FD93EB91FD5C3EEE" box="[554,688,152,176]" class="Malacostraca" family="Retroplumidae" genus="Archaeopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="B956EAC2005E3D73FD93EB91FD5C3EEE" box="[554,688,152,176]" italics="true" pageId="5" pageNumber="348">Archaeopus</emphasis>
</taxonomicName>
, which were included in the
<taxonomicName id="4C224D53005E3D73FBB5EB9EFB5E3EF1" box="[1036,1202,151,175]" class="Malacostraca" family="Retroplumidae" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="348" phylum="Arthropoda" rank="family">Retroplumidae</taxonomicName>
by De Grave
<emphasis id="B956EAC2005E3D73FAD9EB90FA793EEE" box="[1376,1429,152,176]" italics="true" pageId="5" pageNumber="348">et al</emphasis>
. (2009: 41), exhibit much more inflated carapaces, with markedly convex dorsal surfaces. The dorsal surface in
<taxonomicName id="4C224D53005E3D73FF2EEBE9FEF13EA6" box="[151,285,224,248]" class="Malacostraca" family="Retroplumidae" genus="Archaeopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="B956EAC2005E3D73FF2EEBE9FEF13EA6" box="[151,285,224,248]" italics="true" pageId="5" pageNumber="348">Archaeopus</emphasis>
</taxonomicName>
has in addition swollen regions (Rathbun 1908: pl. 47, fig. 5), rather than the thin ridges that define
<taxonomicName id="4C224D53005E3D73FF2EEA0DFED33F42" box="[151,319,260,284]" class="Malacostraca" family="Retroplumidae" genus="Serrablopluma" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="348" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B956EAC2005E3D73FF2EEA0DFED33F42" box="[151,319,260,284]" italics="true" pageId="5" pageNumber="348">Serrablopluma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557B9005E3D73FEFFEA0DFE783F42" box="[326,404,260,284]" pageId="5" pageNumber="348" rank="genus">
<emphasis id="B956EAC2005E3D73FEFFEA0DFE613F42" bold="true" box="[326,397,260,284]" pageId="5" pageNumber="348">n. gen</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>