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<document id="B33D03BEC5282E28E004563B8D892182" ID-CLB-Dataset="31638" ID-DOI="10.11646/zootaxa.4343.1.1" ID-GBIF-Dataset="2b60af66-f747-47b2-8e43-2eb333131a4b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1041210" ID-ZooBank="6E46EE12-D51F-48B0-BC66-0EBBAF9FA981" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509696557008" checkinUser="plazi" docAuthor="Salazar-Vallejo, Sergio I." docDate="2017" docId="038B87B63427FFEE1AB7FF57FC5CF8A0" docLanguage="en" docName="zootaxa.4343.1.1.pdf.imf" docOrigin="Zootaxa 4343 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Bradabyssa hartmanae Salazar-Vallejo 2017, n. sp." docType="treatment" docVersion="9" lastPageNumber="67" masterDocId="FFB2FFCE3466FFAC1A20FFC0FFE8FF94" masterDocTitle="Revision of Brada Stimpson, 1853, and Bradabyssa Hartman, 1967 (Annelida, Flabelligeridae)" masterLastPageNumber="98" masterPageNumber="1" pageNumber="66" updateTime="1698480659823" updateUser="ExternalLinkService">
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<mods:title id="6FAF879FB57550BABB80F471B76CDBE4">Revision of Brada Stimpson, 1853, and Bradabyssa Hartman, 1967 (Annelida, Flabelligeridae)</mods:title>
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<mods:name id="8B16FBA1E039633ECB6D9C9EDE2751E9" type="personal">
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<mods:namePart id="108131C025DE919B094B410C2DC02FDD">Salazar-Vallejo, Sergio I.</mods:namePart>
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<mods:title id="9DC580F871240A25D999CCF57B9B4001">Zootaxa</mods:title>
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<mods:part id="47289F68029BF6CA2670A44251055255">
<mods:date id="C632EAF6DA717B51807BBE0E0F4CA3F5">2017</mods:date>
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<mods:number id="D1869EEFEC61D3BFF677545AD502F314">2017-11-03</mods:number>
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<treatment id="038B87B63427FFEE1AB7FF57FC5CF8A0" ID-DOI="http://doi.org/10.5281/zenodo.6051181" ID-GBIF-Taxon="137233543" ID-Zenodo-Dep="6051181" LSID="urn:lsid:plazi:treatment:038B87B63427FFEE1AB7FF57FC5CF8A0" httpUri="http://treatment.plazi.org/id/038B87B63427FFEE1AB7FF57FC5CF8A0" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66">
<subSubSection id="C338652B3427FFED1AB7FF57FEEBFF47" pageId="65" pageNumber="66" type="nomenclature">
<paragraph id="8B9D36A03427FFED1AB7FF57FE1FFF26" blockId="65.[151,503,151,211]" box="[151,503,151,178]" pageId="65" pageNumber="66">
<heading id="D0D581CC3427FFED1AB7FF57FE1FFF26" bold="true" box="[151,503,151,178]" fontSize="11" level="1" pageId="65" pageNumber="66" reason="1">
<emphasis id="B956EAB23427FFED1AB7FF57FE1FFF26" bold="true" box="[151,503,151,178]" pageId="65" pageNumber="66">
<taxonomicName id="4C224D233427FFED1AB7FF57FE46FF25" authority="Salazar-Vallejo, 2017" authorityName="Salazar-Vallejo" authorityYear="2017" box="[151,430,151,177]" class="Polychaeta" family="Flabelligeridae" genus="Bradabyssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="65" pageNumber="66" phylum="Annelida" rank="species" species="hartmanae" status="sp. nov.">
<emphasis id="B956EAB23427FFED1AB7FF57FE46FF25" bold="true" box="[151,430,151,177]" italics="true" pageId="65" pageNumber="66">Bradabyssa hartmanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557C93427FFED1B95FF58FE1FFF26" box="[437,503,152,178]" pageId="65" pageNumber="66" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D36A03427FFED1AB7FF7AFEEBFF47" blockId="65.[151,503,151,211]" box="[151,259,186,211]" pageId="65" pageNumber="66">
<figureCitation id="13192A253427FFED1AB7FF7AFEEBFF47" box="[151,259,186,211]" captionStart="FIGURE 33" captionStartId="65.[151,250,1674,1696]" captionTargetBox="[176,1419,448,1636]" captionTargetId="figure@65.[172,1419,444,1636]" captionTargetPageId="65" captionText="FIGURE 33. Bradabyssa hartmanae n. sp.: A. Holotype (SIORAS 866), dorsal view. B. Same, anterior end, dorsal view. C. Paratype (ECOSUR 185), prostomium with branchiae and palps removed, frontal view (1 4: branchial scars, PS: palp scar). D. Holotype, anterior end, ventral view. E. Paratype (ECOSUR 185), chaetiger 6, left parapodium, anterior view (No: distal notochaetal region, Ne: neurochaetal tip). Scale bars: A: 1 mm, B: 0.25 mm, C: 0.1 mm, D: 0.27 mm, E: 150 µm." httpUri="https://zenodo.org/record/1041276/files/figure.png" pageId="65" pageNumber="66">Figure 33</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C338652B3427FFED1AB7FEC2FD0FF811" pageId="65" pageNumber="66" type="materials_examined">
<paragraph id="8B9D36A03427FFED1AB7FEC2FA74FE13" blockId="65.[151,1436,258,391]" pageId="65" pageNumber="66">
<emphasis id="B956EAB23427FFED1AB7FEC2FED7FE8F" bold="true" box="[151,319,258,283]" pageId="65" pageNumber="66">
<typeStatus id="549988023427FFED1AB7FEC2FF3AFE8F" box="[151,210,258,283]" pageId="65" pageNumber="66">Type</typeStatus>
material
</emphasis>
.
<emphasis id="B956EAB23427FFED1B6FFEC2FC45FE8F" bold="true" box="[335,941,258,283]" pageId="65" pageNumber="66">
<collectingRegion id="49E6F8423427FFED1B6FFEC2FE55FE8F" box="[335,445,258,283]" country="Sri Lanka" name="Southern" pageId="65" pageNumber="66">Southern</collectingRegion>
Atlantic Ocean, South Sandwich Trench
</emphasis>
.
<typeStatus id="549988023427FFED199EFEC2FBCEFE8F" box="[958,1062,258,283]" pageId="65" pageNumber="66" type="holotype">Holotype</typeStatus>
(SIORAS 866), and
<specimenCount id="9D24FD293427FFED1F3AFEC3FA74FE8F" box="[1306,1436,259,283]" pageId="65" pageNumber="66" type="generic" typeStatus="paratypes">8 paratypes</specimenCount>
(SIORAS 866a,
<collectionCode id="ED33AE653427FFED1B71FEE7FE57FEAB" box="[337,447,295,319]" country="Mexico" httpUri="http://grbio.org/cool/dtqq-2t5p" name="El Colegio de la Frontera Sur (Mexico)" pageId="65" pageNumber="66">ECOSUR</collectionCode>
185), R.V. Akademik Kurchatov, Sta. 866 (
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,
<geoCoordinate id="EE1650673427FFED1E3DFEE6FB60FEAA" box="[1053,1160,294,319]" direction="west" orientation="longitude" pageId="65" pageNumber="66" precision="925" value="-26.716667">26°43' W</geoCoordinate>
<geoCoordinate id="EE1650673427FFED1E93FEE6FAFCFEAB" box="[1203,1300,294,319]" direction="south" orientation="latitude" pageId="65" pageNumber="66" precision="925" value="-55.1">55°06' S</geoCoordinate>
,
<geoCoordinate id="EE1650673427FFED1F02FEE6FA78FEAB" box="[1314,1424,294,319]" direction="west" orientation="longitude" pageId="65" pageNumber="66" precision="925" value="-26.666666">26°40' W</geoCoordinate>
),
<quantity id="4CDA9B453427FFED1AB7FE8AFEDBFEF6" box="[151,307,330,355]" metricMagnitude="3" metricUnit="m" metricValue="7.814" metricValueMax="7.934" metricValueMin="7.694" pageId="65" pageNumber="66" unit="m" value="7814.0" valueMax="7934.0" valueMin="7694.0">76947934 m</quantity>
, Sigsbee trawl,
<date id="FF9C10603427FFED1BC3FE8BFD92FEF7" box="[483,634,331,355]" pageId="65" pageNumber="66" value="1971-11-27">27 Nov. 1971</date>
(8 complete
<typeStatus id="549988023427FFED192CFE8BFC92FEF7" box="[780,890,331,355]" pageId="65" pageNumber="66" type="paratype">paratypes</typeStatus>
,
<quantity id="4CDA9B453427FFED19A4FE8BFC07FEF6" box="[900,1007,331,355]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.3" metricValueMin="0.8" pageId="65" pageNumber="66" unit="mm" value="10.5" valueMax="13.0" valueMin="8.0">813 mm</quantity>
long, 1.52.0 mm wide, cephalic cage chaetae
<quantity id="4CDA9B453427FFED1AD2FEAFFEA0FE12" box="[242,328,367,391]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="65" pageNumber="66" unit="mm" value="0.8">0.8 mm</quantity>
long, 2330 chaetigers; gonopodial lobes in chaetiger 5; 812 transverse series of dorsal papillae).
</paragraph>
<caption id="DF5D66283427FFED1AB7F94AFAFAF968" httpUri="https://zenodo.org/record/1041276/files/figure.png" pageId="65" pageNumber="66" targetBox="[176,1419,448,1636]" targetPageId="65">
<paragraph id="8B9D36A03427FFED1AB7F94AFAFAF968" blockId="65.[151,1436,1674,1788]" pageId="65" pageNumber="66">
<emphasis id="B956EAB23427FFED1AB7F94AFEC9F90B" bold="true" box="[151,289,1674,1696]" pageId="65" pageNumber="66">FIGURE 33.</emphasis>
<taxonomicName id="4C224D233427FFED1B08F94AFDFDF934" authority="Salazar-Vallejo, 2017" authorityName="Salazar-Vallejo" authorityYear="2017" box="[296,533,1674,1696]" class="Polychaeta" family="Flabelligeridae" genus="Bradabyssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="65" pageNumber="66" phylum="Annelida" rank="species" species="hartmanae" status="sp. nov.">
<emphasis id="B956EAB23427FFED1B08F94AFDFDF934" box="[296,533,1674,1696]" italics="true" pageId="65" pageNumber="66">Bradabyssa hartmanae</emphasis>
</taxonomicName>
<emphasis id="B956EAB23427FFED183CF94BFDB6F934" bold="true" box="[540,606,1675,1696]" pageId="65" pageNumber="66">
<taxonomicNameLabel id="A26557C93427FFED183CF94BFDB1F934" box="[540,601,1675,1696]" pageId="65" pageNumber="66" rank="species">n. sp.</taxonomicNameLabel>
:
</emphasis>
A. Holotype (SIORAS 866), dorsal view. B. Same, anterior end, dorsal view. C. Paratype (ECOSUR 185), prostomium with branchiae and palps removed, frontal view (14: branchial scars, PS: palp scar). D. Holotype, anterior end, ventral view. E. Paratype (ECOSUR 185), chaetiger 6, left parapodium, anterior view (No: distal notochaetal region, Ne: neurochaetal tip). Scale bars: A: 1 mm, B: 0.25 mm, C: 0.1 mm, D: 0.27 mm, E: 150 µm.
</paragraph>
</caption>
<paragraph id="8B9D36A03427FFED1AE7F8E4FD0FF811" blockId="65.[151,1436,1828,2032]" pageId="65" pageNumber="66">
<materialsCitation id="3B4A3CFD3427FFED1AE7F8E4FD0FF811" ID-GBIF-Occurrence="1697157828" collectingDate="1965-10-15" collectionCode="LACM" collectorName="R. V. Anton Bruun &amp; Cruise II &amp; Sta." country="Peru Chile" elevation="4377" latitude="-8.516666" location="Eastern" longLatPrecision="1301" longitude="-81.666664" pageId="65" pageNumber="66" specimenCode="LACM 9557" specimenCount="1" stateProvince="Eastern">
<emphasis id="B956EAB23427FFED1AE7F8E4FE5EF8A9" bold="true" box="[199,438,1828,1853]" pageId="65" pageNumber="66">Additional material</emphasis>
.
<emphasis id="B956EAB23427FFED1BE7F8E4FD3FF8A9" bold="true" box="[455,727,1828,1853]" pageId="65" pageNumber="66">
<collectingRegion id="49E6F8423427FFED1BE7F8E4FDCCF8A9" box="[455,548,1828,1853]" country="Sri Lanka" name="Eastern" pageId="65" pageNumber="66">Eastern</collectingRegion>
Pacific Ocean
</emphasis>
. One specimen (
<specimenCode id="DB849EDB3427FFED19BEF8E4FBDFF8A9" box="[926,1079,1828,1853]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="65" pageNumber="66">LACM 9557</specimenCode>
), median fragment,
<collectingCountry id="F33576303427FFED1F01F8E5FA73F8A9" box="[1313,1435,1828,1853]" isEnumeration="true" pageId="65" pageNumber="66">
<collectingCountry id="F33576303427FFED1F01F8E5FABDF8A8" box="[1313,1365,1829,1852]" name="Peru" pageId="65" pageNumber="66">Peru</collectingCountry>
<collectingCountry id="F33576303427FFED1F7FF8E4FA73F8A9" box="[1375,1435,1828,1853]" name="Chile" pageId="65" pageNumber="66">Chile</collectingCountry>
</collectingCountry>
Trench Expedition,
<collectorName id="26D753763427FFED1B58F888FDA1F8CB" box="[376,585,1864,1888]" pageId="65" pageNumber="66">R.V. Anton Bruun</collectorName>
,
<collectorName id="26D753763427FFED1876F887FD54F8CB" box="[598,700,1863,1888]" pageId="65" pageNumber="66">Cruise II</collectorName>
,
<collectorName id="26D753763427FFED18EAF888FD1CF8F4" box="[714,756,1864,1888]" pageId="65" pageNumber="66">Sta.</collectorName>
95 (
<geoCoordinate id="EE1650673427FFED190AF888FC64F8F4" box="[810,908,1864,1888]" direction="south" orientation="latitude" pageId="65" pageNumber="66" precision="925" value="-8.516666">08°31' S</geoCoordinate>
,
<geoCoordinate id="EE1650673427FFED19BAF888FBE1F8F4" box="[922,1033,1864,1888]" direction="west" orientation="longitude" pageId="65" pageNumber="66" precision="925" value="-81.666664">81°40' W</geoCoordinate>
),
<quantity id="4CDA9B453427FFED1E3EF888FB55F8CB" box="[1054,1213,1864,1888]" metricMagnitude="3" metricUnit="m" metricValue="4.3774999999999995" metricValueMax="4.423" metricValueMin="4.332" pageId="65" pageNumber="66" unit="m" value="4377.5" valueMax="4423.0" valueMin="4332.0">
<elevation id="000FD1933427FFED1E3EF888FB55F8CB" box="[1054,1213,1864,1888]" metricMagnitude="3" metricUnit="m" metricValue="4.3774999999999995" metricValueMax="4.423" metricValueMin="4.332" pageId="65" pageNumber="66" unit="m" value="4377.5" valueMax="4423.0" valueMin="4332.0">43324423 m</elevation>
</quantity>
, Menzies trawl and beam trawl, in tandem,
<date id="FF9C10603427FFED1BBFF8ACFDD9F811" box="[415,561,1900,1925]" pageId="65" pageNumber="66" value="1965-10-15">
<collectingDate id="EFD8E9883427FFED1BBFF8ACFDD9F811" box="[415,561,1900,1925]" pageId="65" pageNumber="66" value="1965-10-15">15 Oct. 1965</collectingDate>
</date>
(14 chaetigers).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C338652B3427FFEE1AE7F84FFA71FB1C" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66" type="description">
<paragraph id="8B9D36A03427FFEE1AE7F84FFC33FF6C" blockId="65.[151,1436,1828,2032]" lastBlockId="66.[151,1437,151,1844]" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66">
<emphasis id="B956EAB23427FFED1AE7F84FFEB9F83C" bold="true" box="[199,337,1935,1960]" pageId="65" pageNumber="66">Description</emphasis>
.
<typeStatus id="549988023427FFED1B41F84FFE21F83C" box="[353,457,1935,1960]" pageId="65" pageNumber="66" type="holotype">Holotype</typeStatus>
(SIORAS 866) complete, slightly eroded, soft, pale, most chaetae broken (
<figureCitation id="13192A253427FFED1F05F84FFA78F83C" box="[1317,1424,1935,1960]" captionStart="FIGURE 33" captionStartId="65.[151,250,1674,1696]" captionTargetBox="[176,1419,448,1636]" captionTargetId="figure@65.[172,1419,444,1636]" captionTargetPageId="65" captionText="FIGURE 33. Bradabyssa hartmanae n. sp.: A. Holotype (SIORAS 866), dorsal view. B. Same, anterior end, dorsal view. C. Paratype (ECOSUR 185), prostomium with branchiae and palps removed, frontal view (1 4: branchial scars, PS: palp scar). D. Holotype, anterior end, ventral view. E. Paratype (ECOSUR 185), chaetiger 6, left parapodium, anterior view (No: distal notochaetal region, Ne: neurochaetal tip). Scale bars: A: 1 mm, B: 0.25 mm, C: 0.1 mm, D: 0.27 mm, E: 150 µm." httpUri="https://zenodo.org/record/1041276/files/figure.png" pageId="65" pageNumber="66">Fig. 33A</figureCitation>
). Body elongate, slightly fusiform,
<quantity id="4CDA9B453427FFED1839F874FD97F858" box="[537,639,1972,1997]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="65" pageNumber="66" unit="mm" value="10.5">10.5 mm</quantity>
long,
<quantity id="4CDA9B453427FFED18E9F874FCC9F858" box="[713,801,1972,1997]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="65" pageNumber="66" unit="mm" value="1.5">1.5 mm</quantity>
wide, cephalic cage
<quantity id="4CDA9B453427FFED1E33F874FB83F858" box="[1043,1131,1972,1997]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="65" pageNumber="66" unit="mm" value="0.5">0.5 mm</quantity>
long, 28 chaetigers. Tunic papillated; papillae abundant, filiform, slightly capitate, with fine sediment loosely-adhered, forming irregular globular masses, more or less fused to surrounding ones (
<figureCitation id="13192A253424FFEE1936FF57FC94FF24" box="[790,892,151,176]" captionStart="FIGURE 33" captionStartId="65.[151,250,1674,1696]" captionTargetBox="[176,1419,448,1636]" captionTargetId="figure@65.[172,1419,444,1636]" captionTargetPageId="65" captionText="FIGURE 33. Bradabyssa hartmanae n. sp.: A. Holotype (SIORAS 866), dorsal view. B. Same, anterior end, dorsal view. C. Paratype (ECOSUR 185), prostomium with branchiae and palps removed, frontal view (1 4: branchial scars, PS: palp scar). D. Holotype, anterior end, ventral view. E. Paratype (ECOSUR 185), chaetiger 6, left parapodium, anterior view (No: distal notochaetal region, Ne: neurochaetal tip). Scale bars: A: 1 mm, B: 0.25 mm, C: 0.1 mm, D: 0.27 mm, E: 150 µm." httpUri="https://zenodo.org/record/1041276/files/figure.png" pageId="66" pageNumber="67">Fig. 33B</figureCitation>
), individual papillae arranged in 910 transverse series (after sediment removal from
<typeStatus id="549988023424FFEE1816FF7DFD4CFF41" box="[566,676,189,213]" pageId="66" pageNumber="67" type="paratype">paratypes</typeStatus>
, papillae filiform, thicker basally, about 910 transverse series per chaetiger). Posterior end papillae not decreasing in size towards pygidium.
</paragraph>
<paragraph id="8B9D36A03424FFEE1AE7FEC4FC3DFE1C" blockId="66.[151,1437,151,1844]" pageId="66" pageNumber="67">Anterior end features observed in other specimens (ECOSUR 185). Prostomium low cone, pale. Eyes not seen. Caruncle poorly developed, median keel and lateral ridges low, continued posteriorly, reaching posterior margin of branchial membrane. Palps thick, large, massive, very short after contraction, slightly darker dorsally; palp keels rounded, low. Lateral lips well developed; ventral and dorsal lips reduced.</paragraph>
<paragraph id="8B9D36A03424FFEE1AE7FE54FDEEFE60" blockId="66.[151,1437,151,1844]" pageId="66" pageNumber="67">
Branchiae cirriform, sessile on branchial plate, arranged as a discontinuous marginal series with four filaments per side, largest filaments dorsalmost (
<figureCitation id="13192A253424FFEE1872FE77FD54FE44" box="[594,700,439,464]" captionStart="FIGURE 33" captionStartId="65.[151,250,1674,1696]" captionTargetBox="[176,1419,448,1636]" captionTargetId="figure@65.[172,1419,444,1636]" captionTargetPageId="65" captionText="FIGURE 33. Bradabyssa hartmanae n. sp.: A. Holotype (SIORAS 866), dorsal view. B. Same, anterior end, dorsal view. C. Paratype (ECOSUR 185), prostomium with branchiae and palps removed, frontal view (1 4: branchial scars, PS: palp scar). D. Holotype, anterior end, ventral view. E. Paratype (ECOSUR 185), chaetiger 6, left parapodium, anterior view (No: distal notochaetal region, Ne: neurochaetal tip). Scale bars: A: 1 mm, B: 0.25 mm, C: 0.1 mm, D: 0.27 mm, E: 150 µm." httpUri="https://zenodo.org/record/1041276/files/figure.png" pageId="66" pageNumber="67">Fig. 33C</figureCitation>
). Longest filaments about one-third as long as palps. Nephridial lobes in branchial plate not seen.
</paragraph>
<paragraph id="8B9D36A03424FFEE1AE7FDC2FE46FDF2" blockId="66.[151,1437,151,1844]" pageId="66" pageNumber="67">Cephalic cage chaetae as long as 1/20 body length, or about 1/3 body width. Chaetiger 1 involved in the cephalic cage; chaetae arranged in short dorsoventral series; 2 notochaetae and 3 neurochaetae per fascicle; first neuropodia not projected or swollen.</paragraph>
<paragraph id="8B9D36A03424FFEE1AE7FDB1FF30FD61" blockId="66.[151,1437,151,1844]" pageId="66" pageNumber="67">
Anterior dorsal margin of first chaetiger papillated, rounded. Chaetal transition from cephalic cage to body chaetae abrupt; large aristate neurospines present from chaetiger 2. Ventral gonopodial lobes not seen in
<typeStatus id="549988023424FFEE1F1AFD55FA74FD3A" box="[1338,1436,661,686]" pageId="66" pageNumber="67" type="holotype">holotype</typeStatus>
(
<figureCitation id="13192A253424FFEE1ABFFD79FEE0FD46" box="[159,264,697,722]" captionStart="FIGURE 33" captionStartId="65.[151,250,1674,1696]" captionTargetBox="[176,1419,448,1636]" captionTargetId="figure@65.[172,1419,444,1636]" captionTargetPageId="65" captionText="FIGURE 33. Bradabyssa hartmanae n. sp.: A. Holotype (SIORAS 866), dorsal view. B. Same, anterior end, dorsal view. C. Paratype (ECOSUR 185), prostomium with branchiae and palps removed, frontal view (1 4: branchial scars, PS: palp scar). D. Holotype, anterior end, ventral view. E. Paratype (ECOSUR 185), chaetiger 6, left parapodium, anterior view (No: distal notochaetal region, Ne: neurochaetal tip). Scale bars: A: 1 mm, B: 0.25 mm, C: 0.1 mm, D: 0.27 mm, E: 150 µm." httpUri="https://zenodo.org/record/1041276/files/figure.png" pageId="66" pageNumber="67">Fig. 33D</figureCitation>
);
<typeStatus id="549988023424FFEE1B3FFD7AFE62FD46" box="[287,394,698,722]" pageId="66" pageNumber="67" type="paratype">paratypes</typeStatus>
with gonopodial lobes in chaetiger 5, each short, rounded, slightly darker than surrounding areas.
</paragraph>
<paragraph id="8B9D36A03424FFEE1AE7FCC1FAC0FCAA" blockId="66.[151,1437,151,1844]" pageId="66" pageNumber="67">
Parapodia poorly developed, chaetae emerge from body wall (
<figureCitation id="13192A253424FFEE19A0FCC1FC0EFC8E" box="[896,998,769,794]" captionStart="FIGURE 33" captionStartId="65.[151,250,1674,1696]" captionTargetBox="[176,1419,448,1636]" captionTargetId="figure@65.[172,1419,444,1636]" captionTargetPageId="65" captionText="FIGURE 33. Bradabyssa hartmanae n. sp.: A. Holotype (SIORAS 866), dorsal view. B. Same, anterior end, dorsal view. C. Paratype (ECOSUR 185), prostomium with branchiae and palps removed, frontal view (1 4: branchial scars, PS: palp scar). D. Holotype, anterior end, ventral view. E. Paratype (ECOSUR 185), chaetiger 6, left parapodium, anterior view (No: distal notochaetal region, Ne: neurochaetal tip). Scale bars: A: 1 mm, B: 0.25 mm, C: 0.1 mm, D: 0.27 mm, E: 150 µm." httpUri="https://zenodo.org/record/1041276/files/figure.png" pageId="66" pageNumber="67">Fig. 33E</figureCitation>
). Parapodia lateral; median neuropodia ventrolateral. Notopodia and neuropodia close to each other; short, low conical lobes, neuropodia larger.
</paragraph>
<paragraph id="8B9D36A03424FFEE1AE7FC89FEE9FB8F" blockId="66.[151,1437,151,1844]" pageId="66" pageNumber="67">
Median notochaetae in tufts, two (rarely three) per fascicle, all multiarticulate capillaries, articulated distally, articles medium-sized, longer distally, each internally with 89 transverse annulations (
<figureCitation id="13192A253424FFEE1EB3FCADFB13FC12" box="[1171,1275,877,902]" captionStart="FIGURE 33" captionStartId="65.[151,250,1674,1696]" captionTargetBox="[176,1419,448,1636]" captionTargetId="figure@65.[172,1419,444,1636]" captionTargetPageId="65" captionText="FIGURE 33. Bradabyssa hartmanae n. sp.: A. Holotype (SIORAS 866), dorsal view. B. Same, anterior end, dorsal view. C. Paratype (ECOSUR 185), prostomium with branchiae and palps removed, frontal view (1 4: branchial scars, PS: palp scar). D. Holotype, anterior end, ventral view. E. Paratype (ECOSUR 185), chaetiger 6, left parapodium, anterior view (No: distal notochaetal region, Ne: neurochaetal tip). Scale bars: A: 1 mm, B: 0.25 mm, C: 0.1 mm, D: 0.27 mm, E: 150 µm." httpUri="https://zenodo.org/record/1041276/files/figure.png" pageId="66" pageNumber="67">Fig. 33E</figureCitation>
,
<emphasis id="B956EAB23424FFEE1F2BFCAEFAC4FC11" box="[1291,1324,878,901]" italics="true" pageId="66" pageNumber="67">No</emphasis>
); chaetae about as long as 2/3 body width. Neurochaetae multiarticulate capillaries in chaetiger 1; small aristate neurospines from chaetiger 2, larger in median neuropodia, decrease in size posteriorly, mostly 4 neurospines throughout body. Neurospines with very short rings basally and medially, distally hyaline, aristate, curved in median chaetigers (
<figureCitation id="13192A253424FFEE1F4FFC1FFF2EFB8F" captionStart="FIGURE 33" captionStartId="65.[151,250,1674,1696]" captionTargetBox="[176,1419,448,1636]" captionTargetId="figure@65.[172,1419,444,1636]" captionTargetPageId="65" captionText="FIGURE 33. Bradabyssa hartmanae n. sp.: A. Holotype (SIORAS 866), dorsal view. B. Same, anterior end, dorsal view. C. Paratype (ECOSUR 185), prostomium with branchiae and palps removed, frontal view (1 4: branchial scars, PS: palp scar). D. Holotype, anterior end, ventral view. E. Paratype (ECOSUR 185), chaetiger 6, left parapodium, anterior view (No: distal notochaetal region, Ne: neurochaetal tip). Scale bars: A: 1 mm, B: 0.25 mm, C: 0.1 mm, D: 0.27 mm, E: 150 µm." httpUri="https://zenodo.org/record/1041276/files/figure.png" pageId="66" pageNumber="67">Fig. 33E</figureCitation>
,
<emphasis id="B956EAB23424FFEE1AF2FBC4FF19FB8F" box="[210,241,1028,1051]" italics="true" pageId="66" pageNumber="67">Ne</emphasis>
).
</paragraph>
<paragraph id="8B9D36A03424FFEE1AE7FBE7FE3DFBF0" blockId="66.[151,1437,151,1844]" pageId="66" pageNumber="67">
Posterior end tapering into rounded cone; pygidium with anus terminal, pale muscular ring (dark in some
<typeStatus id="549988023424FFEE1AB7FB8BFEE0FBF7" box="[151,264,1099,1123]" pageId="66" pageNumber="67" type="paratype">paratypes</typeStatus>
); anal cirri absent.
</paragraph>
<paragraph id="8B9D36A03424FFEE1AE7FBAFFA71FB1C" blockId="66.[151,1437,151,1844]" box="[199,1433,1135,1160]" pageId="66" pageNumber="67">
<emphasis id="B956EAB23424FFEE1AE7FBAFFEDDFB1C" bold="true" box="[199,309,1135,1160]" pageId="66" pageNumber="67">Variation</emphasis>
.
<typeStatus id="549988023424FFEE1B65FBB0FE5AFB1C" box="[325,434,1136,1160]" pageId="66" pageNumber="67" type="paratype">Paratypes</typeStatus>
<quantity id="4CDA9B453424FFEE1B99FBAFFDCCFB13" box="[441,548,1135,1159]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.3" metricValueMin="0.8" pageId="66" pageNumber="67" unit="mm" value="10.5" valueMax="13.0" valueMin="8.0">813 mm</quantity>
long, 1.52.0 mm wide, cephalic cage chaetae
<quantity id="4CDA9B453424FFEE1E16FBAFFB64FB13" box="[1078,1164,1135,1159]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="66" pageNumber="67" unit="mm" value="0.8">0.8 mm</quantity>
long, 2330 chaetigers.
</paragraph>
</subSubSection>
<subSubSection id="C338652B3424FFEE1AE7FB53FBA9F930" pageId="66" pageNumber="67" type="discussion">
<paragraph id="8B9D36A03424FFEE1AE7FB53FCAEFAF4" blockId="66.[151,1437,151,1844]" pageId="66" pageNumber="67">
<emphasis id="B956EAB23424FFEE1AE7FB53FEDCFB38" bold="true" box="[199,308,1171,1196]" pageId="66" pageNumber="67">Remarks</emphasis>
.
<taxonomicName id="4C224D233424FFEE1B63FB53FDA0FB38" authority="Salazar-Vallejo, 2017" authorityName="Salazar-Vallejo" authorityYear="2017" box="[323,584,1171,1196]" class="Polychaeta" family="Flabelligeridae" genus="Bradabyssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="66" pageNumber="67" phylum="Annelida" rank="species" species="hartmanae" status="sp. nov.">
<emphasis id="B956EAB23424FFEE1B63FB53FDA0FB38" box="[323,584,1171,1196]" italics="true" pageId="66" pageNumber="67">Bradabyssa hartmanae</emphasis>
</taxonomicName>
<emphasis id="B956EAB23424FFEE1870FB53FD67FB38" bold="true" box="[592,655,1171,1196]" pageId="66" pageNumber="67">
<taxonomicNameLabel id="A26557C93424FFEE1870FB53FD67FB38" box="[592,655,1171,1196]" pageId="66" pageNumber="67" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is very similar to
<taxonomicName id="4C224D233424FFEE197FFB54FC20FB38" authority="Salazar-Vallejo, 2017" authorityName="Salazar-Vallejo" authorityYear="2017" box="[863,968,1171,1196]" class="Polychaeta" family="Flabelligeridae" genus="Bradabyssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="66" pageNumber="67" phylum="Annelida" rank="species" species="jirkovi" status="sp. nov.">
<emphasis id="B956EAB23424FFEE197FFB54FC20FB38" box="[863,968,1171,1196]" italics="true" pageId="66" pageNumber="67">B. jirkovi</emphasis>
</taxonomicName>
<emphasis id="B956EAB23424FFEE19F0FB53FBF8FB38" bold="true" box="[976,1040,1171,1196]" pageId="66" pageNumber="67">
<taxonomicNameLabel id="A26557C93424FFEE19F0FB53FBF8FB38" box="[976,1040,1171,1196]" pageId="66" pageNumber="67" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
These abyssal species are the only two species in the genus that possess multiarticulate notochaetae with articles restricted only to their distal half or third. In
<taxonomicName id="4C224D233424FFEE1ADCFB1CFE7EFB60" authorityName="Salazar-Vallejo" authorityYear="2017" box="[252,406,1243,1268]" class="Polychaeta" family="Flabelligeridae" genus="Bradabyssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="66" pageNumber="67" phylum="Annelida" rank="species" species="hartmanae">
<emphasis id="B956EAB23424FFEE1ADCFB1CFE7EFB60" box="[252,406,1243,1268]" italics="true" pageId="66" pageNumber="67">B. hartmanae</emphasis>
</taxonomicName>
the articles are medium-sized, becoming slightly longer distally, whereas in
<taxonomicName id="4C224D233424FFEE1F26FB1CFA98FB60" authorityName="Salazar-Vallejo" authorityYear="2017" box="[1286,1392,1243,1268]" class="Polychaeta" family="Flabelligeridae" genus="Bradabyssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="66" pageNumber="67" phylum="Annelida" rank="species" species="jirkovi">
<emphasis id="B956EAB23424FFEE1F26FB1CFA98FB60" box="[1286,1392,1243,1268]" italics="true" pageId="66" pageNumber="67">B. jirkovi</emphasis>
</taxonomicName>
the articles are much longer throughout the whole articulated region. Most other species have only a single superior notochaetae with long articles and all the rest of the notochaetae have shorter articles. Furthermore,
<taxonomicName id="4C224D233424FFEE1ECBFAE4FA6BFAA8" authorityName="Salazar-Vallejo" authorityYear="2017" box="[1259,1411,1315,1340]" class="Polychaeta" family="Flabelligeridae" genus="Bradabyssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="66" pageNumber="67" phylum="Annelida" rank="species" species="hartmanae">
<emphasis id="B956EAB23424FFEE1ECBFAE4FA6BFAA8" box="[1259,1411,1315,1340]" italics="true" pageId="66" pageNumber="67">B. hartmanae</emphasis>
</taxonomicName>
is unique in the genus by having only eight branchial filaments.
</paragraph>
<paragraph id="8B9D36A03424FFEE1AE7FAABFA6CF980" blockId="66.[151,1437,151,1844]" pageId="66" pageNumber="67">
On the other hand, as indicated above,
<taxonomicName id="4C224D233424FFEE18A1FAACFCF2FA10" authorityName="Salazar-Vallejo" authorityYear="2017" box="[641,794,1387,1412]" class="Polychaeta" family="Flabelligeridae" genus="Bradabyssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="66" pageNumber="67" phylum="Annelida" rank="species" species="hartmanae">
<emphasis id="B956EAB23424FFEE18A1FAACFCF2FA10" box="[641,794,1387,1412]" italics="true" pageId="66" pageNumber="67">B. hartmanae</emphasis>
</taxonomicName>
resembles what was regarded as
<emphasis id="B956EAB23424FFEE1EB4FAACFB45FA17" box="[1172,1197,1388,1411]" italics="true" pageId="66" pageNumber="67">B.</emphasis>
<emphasis id="B956EAB23424FFEE1EE1FAACFAC7FA17" box="[1217,1327,1388,1411]" italics="true" pageId="66" pageNumber="67">gangetica</emphasis>
” because both species have notochaetae with short to medium-sized basal articles, and 910 series of dorsal papillae per segment. The main difference is based on dorsal papillae density and gonopodial lobe pigmentation: in
<taxonomicName id="4C224D233424FFEE1FA4FA74FEF8FA64" authorityName="Salazar-Vallejo" authorityYear="2017" class="Polychaeta" family="Flabelligeridae" genus="Bradabyssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="66" pageNumber="67" phylum="Annelida" rank="species" species="hartmanae">
<emphasis id="B956EAB23424FFEE1FA4FA74FEF8FA64" italics="true" pageId="66" pageNumber="67">B. hartmanae</emphasis>
</taxonomicName>
papillae fuse laterally and the body wall is not visible, and gonopodial lobes are indistinct from the adjacent body wall, whereas in
<emphasis id="B956EAB23424FFEE1BD8FA3CFDE1F987" box="[504,521,1532,1555]" italics="true" pageId="66" pageNumber="67">B</emphasis>
. “
<emphasis id="B956EAB23424FFEE1803FA3CFD79F987" box="[547,657,1532,1555]" italics="true" pageId="66" pageNumber="67">gangetica</emphasis>
” papillae are distinct individually and gonopodial lobes are darker.
</paragraph>
<paragraph id="8B9D36A03424FFEE1AE7F9E0FBA9F930" blockId="66.[151,1437,151,1844]" pageId="66" pageNumber="67">
A posterior fragment collected in the Peru-Chile trench is regarded, with some hesitation, as conspecific because it shows similar dorsal, globose sediment balls as the
<typeStatus id="549988023424FFEE1968F984FC90F9C8" box="[840,888,1604,1628]" pageId="66" pageNumber="67">type</typeStatus>
material. The neurochaetae of this Pacific Ocean specimen are slightly longer than body width, which differs from the
<typeStatus id="549988023424FFEE1988F9A8FC30F914" box="[936,984,1640,1664]" pageId="66" pageNumber="67">type</typeStatus>
specimens of
<taxonomicName id="4C224D233424FFEE1E5CF9A8FAFDF914" authorityName="Salazar-Vallejo" authorityYear="2017" box="[1148,1301,1639,1664]" class="Polychaeta" family="Flabelligeridae" genus="Bradabyssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="66" pageNumber="67" phylum="Annelida" rank="species" species="hartmanae">
<emphasis id="B956EAB23424FFEE1E5CF9A8FAFDF914" box="[1148,1301,1639,1664]" italics="true" pageId="66" pageNumber="67">B. hartmanae</emphasis>
</taxonomicName>
; the lack of the anterior end and notochaetae means that it cannot be identified with confidence.
</paragraph>
</subSubSection>
<subSubSection id="C338652B3424FFEE1AE7F96FFEA5F89B" pageId="66" pageNumber="67" type="etymology">
<paragraph id="8B9D36A03424FFEE1AE7F96FFEA5F89B" blockId="66.[151,1437,151,1844]" pageId="66" pageNumber="67">
<emphasis id="B956EAB23424FFEE1AE7F96FFEAEF95C" bold="true" box="[199,326,1711,1736]" pageId="66" pageNumber="67">Etymology</emphasis>
. This species is named after the late Dr. Olga Hartman in recognition of her many publications, and because she originally proposed the genus
<taxonomicName id="4C224D233424FFEE185AF913FD16F978" authorityName="Hartman" authorityYear="1967" box="[634,766,1747,1772]" class="Polychaeta" family="Flabelligeridae" genus="Bradabyssa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="66" pageNumber="67" phylum="Annelida" rank="genus">
<emphasis id="B956EAB23424FFEE185AF913FD16F978" box="[634,766,1747,1772]" italics="true" pageId="66" pageNumber="67">Bradabyssa</emphasis>
</taxonomicName>
. Her series on Antarctic polychaetes have been very useful for my research.
</paragraph>
</subSubSection>
<subSubSection id="C338652B3424FFEE1AE7F8DBFC5CF8A0" box="[199,948,1819,1844]" pageId="66" pageNumber="67" type="distribution">
<paragraph id="8B9D36A03424FFEE1AE7F8DBFC5CF8A0" blockId="66.[151,1437,151,1844]" box="[199,948,1819,1844]" pageId="66" pageNumber="67">
<emphasis id="B956EAB23424FFEE1AE7F8DBFEB1F8A0" bold="true" box="[199,345,1819,1844]" pageId="66" pageNumber="67">Distribution</emphasis>
. Southwestern Atlantic Ocean, in over
<quantity id="4CDA9B453424FFEE1933F8DBFC80F8A7" box="[787,872,1819,1844]" metricMagnitude="3" metricUnit="m" metricValue="7.6" pageId="66" pageNumber="67" unit="m" value="7600.0">7600 m</quantity>
depth.
</paragraph>
</subSubSection>
</treatment>
</document>