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<document id="A66777B0B6CEE06BFE602CA9299DA3E5" ID-DOI="10.5852/ejt.2024.957.2651" ID-ISSN="2118-9773" ID-Zenodo-Dep="13773385" ID-ZooBank="9957F2DF-3F31-4B58-84BC-2F8E0F45C6E3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726582870537" checkinUser="plazi" docAuthor="Selnekovič, Dávid, Jäch, Manfred A. &amp; Kodada, Ján" docDate="2024" docId="038B5001DACEE786FDC9FB72FE5A23A9" docLanguage="en" docName="EJT.2024.957.1-229.pdf" docOrigin="European Journal of Taxonomy 957" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2651/12247" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Elmomorphus oblongus Selnekovič, Jäch &amp; Kodada, 2024, sp. nov." docType="treatment" docUuid="EBE16BA6-51C3-4D05-8231-D6D949DD8BEC" docUuidSource="ZooBank" docVersion="2" lastPageNumber="202" masterDocId="FFB22879DA08E74FFF9EFFBAFFE32440" masterDocTitle="Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae)" masterLastPageNumber="229" masterPageNumber="1" pageNumber="199" updateTime="1726584008522" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="674A5314283A957B7A8F8DC90CAACFC0">Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae)</mods:title>
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<mods:namePart id="DC754859F8E2E912C04D970747BC0FA0">Selnekovič, Dávid</mods:namePart>
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<mods:affiliation id="393B4C57FECB2819137040A630B09A0E">Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia.</mods:affiliation>
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<mods:namePart id="B42AC9BF3B607F99DD8D8A5A486807DE">Jäch, Manfred A.</mods:namePart>
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<mods:affiliation id="A26C8654BDB51DFF026B7A4CF81BB2C6">Naturhistorisches Museum Wien, Burgring 7, A- 1010 Wien, Austria.</mods:affiliation>
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<mods:namePart id="96A74B844AF9089B966FEC4E6ED10597">Kodada, Ján</mods:namePart>
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<mods:affiliation id="EC4DCA5F154AF4B2797FD665DD4BBB09">Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia.</mods:affiliation>
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<paragraph id="8B9DE117DACEE789FDC9FB72FC3F20A3" blockId="198.[356,1232,1224,1323]" box="[599,988,1224,1251]" pageId="198" pageNumber="199">
<heading id="D0D5567BDACEE789FDC9FB72FC3F20A3" box="[599,988,1224,1251]" centered="true" fontSize="11" level="2" pageId="198" pageNumber="199" reason="2">
<taxonomicName id="4C229A94DACEE789FDC9FB72FC9820A3" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[599,891,1224,1251]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="species" species="oblongus" status="sp. nov.">
<emphasis id="B9563D05DACEE789FDC9FB72FC9820A3" bold="true" box="[599,891,1224,1251]" italics="true" pageId="198" pageNumber="199">Elmomorphus oblongus</emphasis>
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<subSubSection id="C338B29CDACEE789FD14FAABFC4A216B" box="[650,937,1296,1323]" pageId="198" pageNumber="199" type="description">
<paragraph id="8B9DE117DACEE789FD14FAABFC4A216B" blockId="198.[356,1232,1224,1323]" box="[650,937,1296,1323]" pageId="198" pageNumber="199">
<figureCitation id="1319FD92DACEE789FD14FAABFCEC216B" box="[650,783,1297,1323]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="199.[189,232,1977,2003]" captionStartId-1="200.[189,232,1961,1987]" captionTargetBox-0="[189,1399,265,1928]" captionTargetBox-1="[349,1235,276,1923]" captionTargetId-0="figure-12@199.[189,1399,265,1928]" captionTargetId-1="figure-21@200.[327,1249,265,1931]" captionTargetPageId-0="199" captionTargetPageId-1="200" captionText-0="Fig. 98. Elmomorphus oblongus sp. nov., holotype, male (NMW), TL: 3.35 mm." captionText-1="Fig. 99. Elmomorphus oblongus sp. nov., holotype (NMW), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13773674" figureDoi-1="http://doi.org/10.5281/zenodo.13773678" httpUri-0="https://zenodo.org/record/13773674/files/figure.png" httpUri-1="https://zenodo.org/record/13773678/files/figure.png" pageId="198" pageNumber="199">Figs 98–99</figureCitation>
, 
<figureCitation id="1319FD92DACEE789FC85FAABFCBD216B" box="[795,862,1297,1323]" captionStart="Fig" captionStartId="214.[189,232,1893,1919]" captionTargetBox="[226,1364,267,1868]" captionTargetId="figure-68@214.[189,1399,259,1884]" captionTargetPageId="214" captionText="Fig. 106. Bursa copulatrix and vagina. A. Elmomorphus hamatus sp. nov., paratype from type locality (NMW). B. E. oblongus sp. nov., paratype from type locality (NMW). C. E. glabriclunis sp. nov., paratype from China, locality CWBS 258 (CKB). D. E. simplex sp. nov., paratype from type locality (CKB). E. E. comosiclunis sp. nov., paratype from China, locality CWBS 468. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773702" httpUri="https://zenodo.org/record/13773702/files/figure.png" pageId="198" pageNumber="199">106B</figureCitation>
, 
<figureCitation id="1319FD92DACEE789FCF4FAAAFC4A216B" box="[874,937,1296,1323]" captionStart="Fig" captionStartId="223.[189,232,1867,1893]" captionTargetBox="[254,1334,266,1829]" captionTargetId="figure-40@223.[189,1399,265,1830]" captionTargetPageId="223" captionText="Fig. 115. Distribution maps.A. Elmomorphus reticulatus sp. nov.B. E. superficialis sp. nov.C. E. hamatus sp. nov. D. E. oblongus sp. nov. E. E. glabriclunis sp. nov. F. E. simplex sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13773724" httpUri="https://zenodo.org/record/13773724/files/figure.png" pageId="198" pageNumber="199">115D</figureCitation>
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<paragraph id="8B9DE117DACEE789FF23FAE9FE2A212D" blockId="198.[189,457,1363,1389]" box="[189,457,1363,1389]" pageId="198" pageNumber="199">
<heading id="D0D5567BDACEE789FF23FAE9FE2A212D" bold="true" box="[189,457,1363,1389]" fontSize="11" level="3" pageId="198" pageNumber="199" reason="3">
<emphasis id="B9563D05DACEE789FF23FAE9FE2A212D" bold="true" box="[189,457,1363,1389]" pageId="198" pageNumber="199">Differential diagnosis</emphasis>
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<taxonomicName id="4C229A94DACEE789FF23FA38FE3421DC" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[189,471,1410,1436]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="species" species="oblongus" status="sp. nov.">
<emphasis id="B9563D05DACEE789FF23FA38FE3421DC" box="[189,471,1410,1436]" italics="true" pageId="198" pageNumber="199">Elmomorphus oblongus</emphasis>
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(
<figureCitation id="1319FD92DACEE789FDCEFA38FD4A21DC" box="[592,681,1410,1436]" captionStart="Fig" captionStartId="199.[189,232,1977,2003]" captionTargetBox="[189,1399,265,1928]" captionTargetId="figure-12@199.[189,1399,265,1928]" captionTargetPageId="199" captionText="Fig. 98. Elmomorphus oblongus sp. nov., holotype, male (NMW), TL: 3.35 mm." figureDoi="http://doi.org/10.5281/zenodo.13773674" httpUri="https://zenodo.org/record/13773674/files/figure.png" pageId="198" pageNumber="199">Fig. 98</figureCitation>
) is characterised by having elytra with scattered punctation lacking longitudinal rows of large punctures. The dorsal plastron is present only on the frontoclypeus reaching the level of the midlength of the eyes. Ventrite 1 is entirely covered with plastron; ventrite 5 is without plastron on the median area before the apex and the area without plastron extends laterad along the posterior margin. The species generally resembles 
<taxonomicName id="4C229A94DACEE789FCA6F9A9FC12226C" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[824,1009,1554,1580]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="species" species="comosiclunis" status="sp. nov.">
<emphasis id="B9563D05DACEE789FCA6F9A9FC12226C" box="[824,1009,1554,1580]" italics="true" pageId="198" pageNumber="199">E. comosiclunis</emphasis>
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<taxonomicNameLabel id="A265807EDACEE789FC66F9A9FBB6226D" box="[1016,1109,1555,1581]" pageId="198" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C229A94DACEE789FBFEF9A9FAEC226C" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[1120,1295,1554,1580]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="species" species="glabriclunis" status="sp. nov.">
<emphasis id="B9563D05DACEE789FBFEF9A9FAEC226C" box="[1120,1295,1554,1580]" italics="true" pageId="198" pageNumber="199">E. glabriclunis</emphasis>
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<taxonomicNameLabel id="A265807EDACEE789FA8BF9A9FA91226D" box="[1301,1394,1555,1581]" pageId="198" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
, and 
<taxonomicName id="4C229A94DACEE789FF6CF98DFE8C2211" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[242,367,1591,1617]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="species" species="simplex" status="sp. nov.">
<emphasis id="B9563D05DACEE789FF6CF98DFE8C2211" box="[242,367,1591,1617]" italics="true" pageId="198" pageNumber="199">E. simplex</emphasis>
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but differs from all of them in the presence of plastron on almost the entire frontoclypeus and in the oblong oval and less convex body (
<figureCitation id="1319FD92DACEE789FC1AF9E1FC3E2235" box="[900,989,1627,1653]" captionStart="Fig" captionStartId="199.[189,232,1977,2003]" captionTargetBox="[189,1399,265,1928]" captionTargetId="figure-12@199.[189,1399,265,1928]" captionTargetPageId="199" captionText="Fig. 98. Elmomorphus oblongus sp. nov., holotype, male (NMW), TL: 3.35 mm." figureDoi="http://doi.org/10.5281/zenodo.13773674" httpUri="https://zenodo.org/record/13773674/files/figure.png" pageId="198" pageNumber="199">Fig. 98</figureCitation>
). Moreover, from 
<taxonomicName id="4C229A94DACEE789FB2BF9E1FA8C2235" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[1205,1391,1627,1653]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="species" species="comosiclunis">
<emphasis id="B9563D05DACEE789FB2BF9E1FA8C2235" box="[1205,1391,1627,1653]" italics="true" pageId="198" pageNumber="199">E. comosiclunis</emphasis>
</taxonomicName>
, it differs in the absence of long setae on ventrite 5. From 
<taxonomicName id="4C229A94DACEE789FCC5F93AFBE922D9" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[859,1034,1663,1689]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="species" species="glabriclunis">
<emphasis id="B9563D05DACEE789FCC5F93AFBE922D9" box="[859,1034,1663,1689]" italics="true" pageId="198" pageNumber="199">E. glabriclunis</emphasis>
</taxonomicName>
, it differs in the evenly convex ventrite 5, without a large glabrous area.
</paragraph>
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<subSubSection id="C338B29CDACEE789FF23F95FFA94236E" box="[189,1399,1765,1839]" pageId="198" pageNumber="199" type="etymology">
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<heading id="D0D5567BDACEE789FF23F95FFEA722BF" bold="true" box="[189,324,1765,1791]" fontSize="11" level="3" pageId="198" pageNumber="199" reason="3">
<emphasis id="B9563D05DACEE789FF23F95FFEA722BF" bold="true" box="[189,324,1765,1791]" pageId="198" pageNumber="199">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DACEE789FF23F8AEFA94236E" blockId="198.[189,1399,1812,1839]" box="[189,1399,1812,1839]" pageId="198" pageNumber="199">The epithet ‘oblongus’ (‘oblong’) is a Latin adjective referring to the characteristic elongate body shape.</paragraph>
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<heading id="D0D5567BDACEE789FF23F8EDFE8F2331" bold="true" box="[189,364,1879,1905]" fontSize="11" level="3" pageId="198" pageNumber="199" reason="3">
<emphasis id="B9563D05DACEE789FF23F8EDFE8F2331" bold="true" box="[189,364,1879,1905]" pageId="198" pageNumber="199">Type material</emphasis>
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<paragraph id="8B9DE117DACEE789FF74F83CFEBF23E0" blockId="198.[189,1399,1926,2025]" box="[234,348,1926,1952]" pageId="198" pageNumber="199">
<emphasis id="B9563D05DACEE789FF74F83CFEBF23E0" bold="true" box="[234,348,1926,1952]" pageId="198" pageNumber="199">
<typeStatus id="54995FB5DACEE789FF74F83CFEBF23E0" box="[234,348,1926,1952]" pageId="198" pageNumber="199" type="holotype">Holotype</typeStatus>
</emphasis>
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<paragraph id="8B9DE117DACEE789FF23F810FC3E23A9" blockId="198.[189,1399,1926,2025]" pageId="198" pageNumber="199">
<materialsCitation id="3B4AEB4ADACEE789FF23F810FC3E23A9" collectionCode="NMW" country="Vietnam" county="Lang Bian" elevation="1665" latitude="12.05" location="Dembicky" longLatPrecision="1294" longitude="108.45" municipality="Pacholatko" pageId="198" pageNumber="199" specimenCount="1" specimenCount-male="1" stateProvince="Lam Dong" typeStatus="holotype">
<collectingCountry id="F335A187DACEE789FF23F810FEA72384" box="[189,324,1962,1988]" name="Vietnam" pageId="198" pageNumber="199">VIETNAM</collectingCountry>
– 
<emphasis id="B9563D05DACEE789FEF7F810FD892384" bold="true" box="[361,618,1962,1988]" pageId="198" pageNumber="199">
<collectingRegion id="49E62FF5DACEE789FEF7F810FD892384" box="[361,618,1962,1988]" country="Vietnam" name="Lam Dong" pageId="198" pageNumber="199">Lam Dong Province</collectingRegion>
</emphasis>
• 
<specimenCount id="9D242A9EDACEE789FD15F811FD472385" box="[651,676,1963,1989]" count="1" pageId="198" pageNumber="199" type="male">♂</specimenCount>
; “S-VIETNAM: 17.-21.4. 
<locationDeviation id="9A2FF835DACEE789FC77F810FB742384" box="[1001,1175,1962,1989]" location="Dalat" pageId="198" pageNumber="199">
<quantity id="4CDA4CF2DACEE789FC77F810FBCF2384" box="[1001,1068,1962,1988]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="198" pageNumber="199" unit="km" value="12.0">12km</quantity>
N Dalat
</locationDeviation>
1995 
<collectingCounty id="62FC999BDACEE789FB79F811FA862384" box="[1255,1381,1962,1989]" pageId="198" pageNumber="199">Lang Bian</collectingCounty>
ǀ 
<geoCoordinate id="EE1687D0DACEE789FF23F877FEFC23A9" box="[189,287,1997,2025]" degrees="12" direction="north" minutes="03" orientation="latitude" pageId="198" pageNumber="199" precision="925" value="12.05">12°03′N</geoCoordinate>
<geoCoordinate id="EE1687D0DACEE789FEB8F877FE7723A9" box="[294,404,1997,2025]" degrees="108" direction="east" minutes="27" orientation="longitude" pageId="198" pageNumber="199" precision="925" value="108.45">108°27′E</geoCoordinate>
<quantity id="4CDA4CF2DACEE789FE05F874FDA423A9" box="[411,583,1998,2025]" metricMagnitude="3" metricUnit="m" metricValue="1.6649999999999998" metricValueMax="1.75" metricValueMin="1.58" pageId="198" pageNumber="199" unit="m" value="1665.0" valueMax="1750.0" valueMin="1580.0">
<elevation id="000F0624DACEE789FE05F874FDA423A9" box="[411,583,1998,2025]" metricMagnitude="3" metricUnit="m" metricValue="1.6649999999999998" metricValueMax="1.75" metricValueMin="1.58" pageId="198" pageNumber="199" unit="m" value="1665.0" valueMax="1750.0" valueMin="1580.0">1580 - 1750 m</elevation>
</quantity>
<collectingMunicipality id="6BF97B6DDACEE789FDD0F874FD3323A8" box="[590,720,1998,2024]" pageId="198" pageNumber="199">Pacholatko</collectingMunicipality>
&amp; 
<location id="8EFDB7CCDACEE789FD68F874FC9423A8" LSID="urn:lsid:plazi:treatment:038B5001DACEE786FDC9FB72FE5A23A9:8EFDB7CCDACEE789FD68F874FC9423A8" box="[758,887,1998,2024]" country="Vietnam" county="Lang Bian" latitude="12.05" longLatPrecision="1294" longitude="108.45" municipality="Pacholatko" name="Dembicky" pageId="198" pageNumber="199" stateProvince="Lam Dong">Dembicky</location>
”; 
<collectionCode id="ED3379D2DACEE789FC13F875FC3A23A9" box="[909,985,1999,2025]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="198" pageNumber="199">NMW</collectionCode>
.
</materialsCitation>
</paragraph>
<caption id="DF5DB19FDACFE788FF23F803FB8D2393" ID-DOI="http://doi.org/10.5281/zenodo.13773674" ID-Zenodo-Dep="13773674" box="[189,1134,1977,2004]" httpUri="https://zenodo.org/record/13773674/files/figure.png" pageId="199" pageNumber="200" startId="199.[189,232,1977,2003]" targetBox="[189,1399,265,1928]" targetPageId="199">
<paragraph id="8B9DE117DACFE788FF23F803FB8D2393" blockId="199.[189,1134,1977,2004]" box="[189,1134,1977,2004]" pageId="199" pageNumber="200">
<emphasis id="B9563D05DACFE788FF23F803FEFA2393" bold="true" box="[189,281,1977,2003]" pageId="199" pageNumber="200">Fig. 98.</emphasis>
<taxonomicName id="4C229A94DACFE788FEBEF803FDDB2393" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[288,568,1977,2003]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="199" pageNumber="200" phylum="Arthropoda" rank="species" species="oblongus" status="sp. nov.">
<emphasis id="B9563D05DACFE788FEBEF803FDDB2393" box="[288,568,1977,2003]" italics="true" pageId="199" pageNumber="200">Elmomorphus oblongus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDACFE788FDA1F803FD7F2393" box="[575,668,1977,2003]" pageId="199" pageNumber="200" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, male (NMW), TL: 3.35 mm.
</paragraph>
</caption>
<caption id="DF5DB19FDAC0E787FF23F813FE1123A7" ID-DOI="http://doi.org/10.5281/zenodo.13773678" ID-Zenodo-Dep="13773678" httpUri="https://zenodo.org/record/13773678/files/figure.png" pageId="200" pageNumber="201" startId="200.[189,232,1961,1987]" targetBox="[349,1235,276,1923]" targetPageId="200">
<paragraph id="8B9DE117DAC0E787FF23F813FE1123A7" blockId="200.[189,1399,1961,2023]" pageId="200" pageNumber="201">
<emphasis id="B9563D05DAC0E787FF23F813FEFE2383" bold="true" box="[189,285,1961,1987]" pageId="200" pageNumber="201">Fig. 99.</emphasis>
<taxonomicName id="4C229A94DAC0E787FEB9F813FDA02383" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[295,579,1961,1987]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="200" pageNumber="201" phylum="Arthropoda" rank="species" species="oblongus" status="sp. nov.">
<emphasis id="B9563D05DAC0E787FEB9F813FDA02383" box="[295,579,1961,1987]" italics="true" pageId="200" pageNumber="201">Elmomorphus oblongus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDAC0E787FDD3F810FD4E2384" box="[589,685,1962,1988]" pageId="200" pageNumber="201" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (NMW), aedeagus. 
<emphasis id="B9563D05DAC0E787FB82F813FBD22383" bold="true" box="[1052,1073,1961,1987]" pageId="200" pageNumber="201">A</emphasis>
. Lateral aspect. 
<emphasis id="B9563D05DAC0E787FB63F813FAF22383" bold="true" box="[1277,1297,1961,1987]" pageId="200" pageNumber="201">B</emphasis>
. Ventral aspect. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8B9DE117DAC1E786FF74FEB3FE842563" blockId="201.[189,1324,265,325]" box="[234,359,265,291]" pageId="201" pageNumber="202">
<emphasis id="B9563D05DAC1E786FF74FEB3FE842563" bold="true" box="[234,359,265,291]" pageId="201" pageNumber="202">
<typeStatus id="54995FB5DAC1E786FF74FEB3FE842563" box="[234,359,265,291]" pageId="201" pageNumber="202" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B9DE117DAC1E786FF23FE90FACF2505" blockId="201.[189,1324,265,325]" box="[189,1324,298,325]" pageId="201" pageNumber="202">
<materialsCitation id="3B4AEB4ADAC1E786FF23FE90FACB2505" box="[189,1320,298,325]" collectionCode="CKB, NMW" country="Vietnam" county="Lang Bian" elevation="1665" latitude="12.05" location="Dembicky" longLatPrecision="1294" longitude="108.45" municipality="Pacholatko" pageId="201" pageNumber="202" specimenCount="139" stateProvince="Lam Dong" typeStatus="paratype">
<collectingCountry id="F335A187DAC1E786FF23FE90FEA72504" box="[189,324,298,324]" name="Vietnam" pageId="201" pageNumber="202">VIETNAM</collectingCountry>
– 
<emphasis id="B9563D05DAC1E786FEFCFE90FDBF2504" bold="true" box="[354,604,298,324]" pageId="201" pageNumber="202">
<collectingRegion id="49E62FF5DAC1E786FEFCFE90FDBF2504" box="[354,604,298,324]" country="Vietnam" name="Lam Dong" pageId="201" pageNumber="202">Lam Dong Province</collectingRegion>
</emphasis>
• 
<specimenCount id="9D242A9EDAC1E786FDEBFE90FD2D2505" box="[629,718,298,325]" count="139" pageId="201" pageNumber="202" type="generic">139 ex.</specimenCount>
; same collection data as for holotype; 
<collectionCode id="ED3379D2DAC1E786FB0FFE90FB332504" box="[1169,1232,298,324]" pageId="201" pageNumber="202">CKB</collectionCode>
, 
<collectionCode id="ED3379D2DAC1E786FB42FE91FACB2505" box="[1244,1320,299,325]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="201" pageNumber="202">NMW</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9DE117DAC1E786FF23FED0FEBE25C4" blockId="201.[189,349,362,388]" box="[189,349,362,388]" pageId="201" pageNumber="202">
<heading id="D0D5567BDAC1E786FF23FED0FEBE25C4" bold="true" box="[189,349,362,388]" fontSize="11" level="3" pageId="201" pageNumber="202" reason="3">
<emphasis id="B9563D05DAC1E786FF23FED0FEBE25C4" bold="true" box="[189,349,362,388]" pageId="201" pageNumber="202">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DAC1E786FF23FE2DFC7825F2" blockId="201.[189,923,406,434]" box="[189,923,406,434]" pageId="201" pageNumber="202">
<collectingCountry id="F335A187DAC1E786FF23FE2DFEC025F1" box="[189,291,407,433]" name="Vietnam" pageId="201" pageNumber="202">Vietnam</collectingCountry>
, 
<collectingRegion id="49E62FF5DAC1E786FEB1FE22FDC325F1" box="[303,544,407,434]" country="Vietnam" name="Lam Dong" pageId="201" pageNumber="202">Lam Dong Province</collectingRegion>
, Lang Bian, 
<geoCoordinate id="EE1687D0DAC1E786FD2BFE2CFCF025F2" box="[693,787,406,434]" degrees="12" direction="north" minutes="3" orientation="latitude" pageId="201" pageNumber="202" precision="925" value="12.05">12°3′ N</geoCoordinate>
, 
<geoCoordinate id="EE1687D0DAC1E786FC80FE2CFC7525F2" box="[798,918,406,434]" degrees="108" direction="east" minutes="27" orientation="longitude" pageId="201" pageNumber="202" precision="925" value="108.45">108°27′ E</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection id="C338B29CDAC1E786FF23FE6DFA902339" pageId="201" pageNumber="202" type="description">
<paragraph id="8B9DE117DAC1E786FF23FE6DFEAD25B1" blockId="201.[189,334,471,497]" box="[189,334,471,497]" pageId="201" pageNumber="202">
<heading id="D0D5567BDAC1E786FF23FE6DFEAD25B1" bold="true" box="[189,334,471,497]" fontSize="11" level="3" pageId="201" pageNumber="202" reason="3">
<emphasis id="B9563D05DAC1E786FF23FE6DFEAD25B1" bold="true" box="[189,334,471,497]" pageId="201" pageNumber="202">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DAC1E786FF23FDBFFC5726E5" blockId="201.[189,1399,516,678]" pageId="201" pageNumber="202">
Measurements (mm): TL: 
<specimenCount id="9D242A9EDAC1E786FE6CFDBFFDFD265F" box="[498,542,517,543]" count="1" pageId="201" pageNumber="202" type="male">♂♂</specimenCount>
3.21–3.50 (3.34± 0.09, n=8), 
<specimenCount id="9D242A9EDAC1E786FC1BFDBFFC52265F" box="[901,945,517,543]" count="1" pageId="201" pageNumber="202" type="female">♀♀</specimenCount>
3.42–3.58 (3.48± 0.05, n=8); PL: 
<specimenCount id="9D242A9EDAC1E786FAD4FDBFFA95265F" box="[1354,1398,517,543]" count="1" pageId="201" pageNumber="202" type="male">♂♂</specimenCount>
0.71–0.81 (0.76 ± 0.03, n= 8), 
<specimenCount id="9D242A9EDAC1E786FDAAFD9CFD832600" box="[564,608,550,576]" count="1" pageId="201" pageNumber="202" type="female">♀♀</specimenCount>
0.73–0.83 (0.79 ±0.03, n=8); PW: 
<specimenCount id="9D242A9EDAC1E786FBB6FD9CFBB72600" box="[1064,1108,550,576]" count="1" pageId="201" pageNumber="202" type="male">♂♂</specimenCount>
1.40–1.57 (1.51 ± 0.05, n= 8), 
<specimenCount id="9D242A9EDAC1E786FE8AFDF2FEA32622" box="[276,320,584,610]" count="1" pageId="201" pageNumber="202" type="female">♀♀</specimenCount>
1.50–1.60 (1.56± 0.03, n =8); EL: 
<specimenCount id="9D242A9EDAC1E786FC95FDF2FCD42622" box="[779,823,584,610]" count="1" pageId="201" pageNumber="202" type="male">♂♂</specimenCount>
2.54–2.75 (2.64± 0.06, n =8), 
<specimenCount id="9D242A9EDAC1E786FB5AFDF2FB132622" box="[1220,1264,584,610]" count="1" pageId="201" pageNumber="202" type="female">♀♀</specimenCount>
2.71–2.88 (2.76 ± 0.05, n =8); EW: 
<specimenCount id="9D242A9EDAC1E786FE73FDD0FDFA26C4" box="[493,537,618,644]" count="1" pageId="201" pageNumber="202" type="male">♂♂</specimenCount>
1.67–1.79 (1.74± 0.04, n= 8), 
<specimenCount id="9D242A9EDAC1E786FC0BFDD0FC2226C4" box="[917,961,618,644]" count="1" pageId="201" pageNumber="202" type="female">♀♀</specimenCount>
1.75–1.90 (1.83± 0.04, n= 8); PhL: 0.47–0.51 (0.49± 0.01, n =8); PrL: 0.39–0.42 (0.41 ±0.01, n =8).
</paragraph>
<paragraph id="8B9DE117DAC1E786FF23FD71FBE5272C" blockId="201.[189,1398,715,876]" pageId="201" pageNumber="202">
Body oblong oval, strongly convex, widest between first and second thirds of elytra (
<figureCitation id="1319FD92DAC1E786FB1CFD71FB3B26A5" box="[1154,1240,715,741]" captionStart="Fig" captionStartId="199.[189,232,1977,2003]" captionTargetBox="[189,1399,265,1928]" captionTargetId="figure-12@199.[189,1399,265,1928]" captionTargetPageId="199" captionText="Fig. 98. Elmomorphus oblongus sp. nov., holotype, male (NMW), TL: 3.35 mm." figureDoi="http://doi.org/10.5281/zenodo.13773674" httpUri="https://zenodo.org/record/13773674/files/figure.png" pageId="201" pageNumber="202">Fig. 98</figureCitation>
). Colouration black, antennae, mouthparts, and tarsi reddish brown, remaining parts of legs brown. Dorsal pubescence consisting of very short and thin yellowish setae. Plastron present in anterior portions of frontoclypeus, along lateral elytral edge in posterior half, and entire ventral surface except prosternal process, median part of metaventrite, and narrow area along posterior edge of ventrite 5.
</paragraph>
<paragraph id="8B9DE117DAC1E786FF23FC2BFD012014" blockId="201.[189,1399,913,1108]" pageId="201" pageNumber="202">
Dorsal cranial surface with coarse setiferous punctures, each approximately as wide as eye facet, separated by 0.5–1.0 × puncture diameter; plastron in anterior portions of frontoclypeus, reaching above antennal insertions. Labrum transverse, anterior margin weakly emarginate, exposed portion microreticulate, setae concentrated along anterior margin, equal in length in both sexes. Eyes oval, weakly protruding, ID: 
<specimenCount id="9D242A9EDAC1E786FE46FBA3FDE72073" box="[472,516,1049,1075]" count="1" pageId="201" pageNumber="202" type="male">♂♂</specimenCount>
<quantity id="4CDA4CF2DAC1E786FD90FBA2FD582073" box="[526,699,1048,1075]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" metricValueMax="5.699999999999999" metricValueMin="5.1" pageId="201" pageNumber="202" unit="mm" value="0.54" valueMax="0.57" valueMin="0.51">0.51–0.57 mm</quantity>
(0.55± 0.02, n =8), 
<specimenCount id="9D242A9EDAC1E786FC32FBA3FC3B2073" box="[940,984,1049,1075]" count="1" pageId="201" pageNumber="202" type="female">♀♀</specimenCount>
<quantity id="4CDA4CF2DAC1E786FC7CFBA2FB6D2073" box="[994,1166,1048,1075]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" metricValueMax="6.1" metricValueMin="5.5" pageId="201" pageNumber="202" unit="mm" value="0.58" valueMax="0.61" valueMin="0.55">0.55–0.61 mm</quantity>
(0.58± 0.01, n= 8). Antennae short, 10-segmented, densely setose.
</paragraph>
<paragraph id="8B9DE117DAC1E786FF23FBC0FEDA21A4" blockId="201.[189,1399,1145,1508]" pageId="201" pageNumber="202">
Pronotum transverse, moderately convex, widest at base, PW/PL: 
<specimenCount id="9D242A9EDAC1E786FC25FBC0FC0420D4" box="[955,999,1146,1172]" count="1" pageId="201" pageNumber="202" type="male">♂♂</specimenCount>
1.88–2.06 (1.97 ±0.05, n= 8), 
<specimenCount id="9D242A9EDAC1E786FAD4FBC0FA9520D4" box="[1354,1398,1146,1172]" count="1" pageId="201" pageNumber="202" type="female">♀♀</specimenCount>
1.92–2.09 (1.99 ±0.06, n=8); surface smooth, with round setiferous punctures; anterior angles protruding, deflexed; lateral sides convergent, weakly rounded. Prosternal process with lateral and posterior edges rounded; lateral portions weakly raised, without clusters of long setae; median keel arcuate. Scutellum slightly longer than wide, with several round punctures. Metaventral disc weakly convex, without clusters of long setae. Elytra oval, widest between first and second thirds, EL/EW: 
<specimenCount id="9D242A9EDAC1E786FAD4FA99FA95217D" box="[1354,1398,1315,1341]" count="1" pageId="201" pageNumber="202" type="male">♂♂</specimenCount>
1.48–1.55 (1.51 ±0.02, n=8), 
<specimenCount id="9D242A9EDAC1E786FDBBFAFEFDB2211E" box="[549,593,1348,1374]" count="1" pageId="201" pageNumber="202" type="female">♀♀</specimenCount>
1.46–1.57 (1.51 ± 0.04, n =8); surface very weakly microreticulate, with scattered round punctures; plastron present in posterior half in narrow bands along lateral edges. Tibiae weakly curved, protibia ca 1.5× as long as protarsus; PrTL/PL: 
<specimenCount id="9D242A9EDAC1E786FBB0FA32FBB921E2" box="[1070,1114,1416,1442]" count="1" pageId="201" pageNumber="202" type="male">♂♂</specimenCount>
0.99–1.10 (1.04± 0.04, n= 8), 
<specimenCount id="9D242A9EDAC1E786FE93FA13FEDA2183" box="[269,313,1449,1475]" count="1" pageId="201" pageNumber="202" type="female">♀♀</specimenCount>
0.82–1.12 (1.03± 0.09, n=8). Terminal protarsomere as long as three preceding segments combined.
</paragraph>
<paragraph id="8B9DE117DAC1E786FF23F9B0FDEF2228" blockId="201.[189,1399,1546,1640]" pageId="201" pageNumber="202">Ventrites covered with plastron, except on median area before apex of ventrite 5, which extends laterad along posterior edge. Ventrite 5 weakly emarginate at apex in males, rounded at apex, with short longitudinal keel in females.</paragraph>
<paragraph id="8B9DE117DAC1E786FF23F937FDC3234C" blockId="201.[189,1399,1677,1805]" pageId="201" pageNumber="202">
Aedeagus (
<figureCitation id="1319FD92DAC1E786FEA1F937FE7422E8" box="[319,407,1677,1704]" captionStart="Fig" captionStartId="200.[189,232,1961,1987]" captionTargetBox="[349,1235,276,1923]" captionTargetId="figure-21@200.[327,1249,265,1931]" captionTargetPageId="200" captionText="Fig. 99. Elmomorphus oblongus sp. nov., holotype (NMW), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773678" httpUri="https://zenodo.org/record/13773678/files/figure.png" pageId="201" pageNumber="202">Fig. 99</figureCitation>
): phallobase short, PhL/PrL: 1.16–1.27 (1.22 ±0.04, n =8); parameres relatively long, moderately bent ventrad in distal half, apices narrowly rounded (lateral aspect); penis narrowly rounded at apex, reaching two-thirds of parameral length; sclerotised fibula present. Bursa copulatrix (
<figureCitation id="1319FD92DAC1E786FB69F96BFA8D22AB" box="[1271,1390,1745,1771]" captionStart="Fig" captionStartId="214.[189,232,1893,1919]" captionTargetBox="[226,1364,267,1868]" captionTargetId="figure-68@214.[189,1399,259,1884]" captionTargetPageId="214" captionText="Fig. 106. Bursa copulatrix and vagina. A. Elmomorphus hamatus sp. nov., paratype from type locality (NMW). B. E. oblongus sp. nov., paratype from type locality (NMW). C. E. glabriclunis sp. nov., paratype from China, locality CWBS 258 (CKB). D. E. simplex sp. nov., paratype from type locality (CKB). E. E. comosiclunis sp. nov., paratype from China, locality CWBS 468. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773702" httpUri="https://zenodo.org/record/13773702/files/figure.png" pageId="201" pageNumber="202">Fig. 106B</figureCitation>
) with numerous microsclerites.
</paragraph>
<paragraph id="8B9DE117DAC1E786FF23F888FDD4230C" blockId="201.[189,567,1842,1869]" box="[189,567,1842,1869]" pageId="201" pageNumber="202">
<heading id="D0D5567BDAC1E786FF23F888FDD4230C" bold="true" box="[189,567,1842,1869]" fontSize="11" level="3" pageId="201" pageNumber="202" reason="3">
<emphasis id="B9563D05DAC1E786FF23F888FDD4230C" bold="true" box="[189,567,1842,1869]" pageId="201" pageNumber="202">Secondary sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DAC1E786FF23F8E5FA902339" blockId="201.[189,1395,1887,1914]" box="[189,1395,1887,1914]" pageId="201" pageNumber="202">
Ventrite 
<quantity id="4CDA4CF2DAC1E786FEBFF8E5FEAD2339" box="[289,334,1887,1914]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="201" pageNumber="202" unit="in" value="5.0">5 in</quantity>
males weakly emarginate at apex, in female rounded at apex, with short longitudinal keel.
</paragraph>
</subSubSection>
<subSubSection id="C338B29CDAC1E786FF23F825FE5A23A9" box="[189,441,1951,2025]" pageId="201" pageNumber="202" type="distribution">
<paragraph id="8B9DE117DAC1E786FF23F825FEB523F9" blockId="201.[189,342,1951,1977]" box="[189,342,1951,1977]" pageId="201" pageNumber="202">
<heading id="D0D5567BDAC1E786FF23F825FEB523F9" bold="true" box="[189,342,1951,1977]" fontSize="11" level="3" pageId="201" pageNumber="202" reason="3">
<emphasis id="B9563D05DAC1E786FF23F825FEB523F9" bold="true" box="[189,342,1951,1977]" pageId="201" pageNumber="202">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DAC1E786FF23F874FE5A23A9" blockId="201.[189,441,1998,2025]" box="[189,441,1998,2025]" pageId="201" pageNumber="202">
<collectingCountry id="F335A187DAC1E786FF23F874FEC323A8" box="[189,288,1998,2024]" name="Vietnam" pageId="201" pageNumber="202">Vietnam</collectingCountry>
(
<figureCitation id="1319FD92DAC1E786FEAEF874FE4F23A9" box="[304,428,1998,2025]" captionStart="Fig" captionStartId="223.[189,232,1867,1893]" captionTargetBox="[254,1334,266,1829]" captionTargetId="figure-40@223.[189,1399,265,1830]" captionTargetPageId="223" captionText="Fig. 115. Distribution maps.A. Elmomorphus reticulatus sp. nov.B. E. superficialis sp. nov.C. E. hamatus sp. nov. D. E. oblongus sp. nov. E. E. glabriclunis sp. nov. F. E. simplex sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13773724" httpUri="https://zenodo.org/record/13773724/files/figure.png" pageId="201" pageNumber="202">Fig. 115D</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>