149 lines
15 KiB
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149 lines
15 KiB
XML
<document id="DF04AC2019F5A020690C0A0EB7C5DDBA" ID-DOI="10.18590/euscorpius.2003.vol2003.iss11.1" ID-ISSN="1536-9307" ID-Zenodo-Dep="13237351" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="existingObjects" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1721353212353" checkinUser="felipe" docAuthor="Soleglad, Michael E. & Fet, Victor" docDate="2003" docId="038A87D5D73EF53BFF615AADFC6A5799" docLanguage="en" docName="Euscorpius.11.1-175.pdf" docOrigin="Euscorpius 2003 (11)" docSource="http://dx.doi.org/10.18590/euscorpius.2003.vol2003.iss11.1" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Hadrurinae Stahnke 1974" docType="treatment" docVersion="5" lastPageNumber="111" masterDocId="FFB3FFADD74CF549FFA15E77FFB0560F" masterDocTitle="High-level systematics and phylogeny of the extant scorpions (Scorpiones: Orthosterni)" masterLastPageNumber="175" masterPageNumber="1" pageNumber="111" updateTime="1722954143772" updateUser="ExternalLinkService">
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<mods:titleInfo id="0A384182E5CD66153C6639A7B588849B">
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<mods:title id="8D51177F1749DBE124F17BB6A6B1B778">High-level systematics and phylogeny of the extant scorpions (Scorpiones: Orthosterni)</mods:title>
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<mods:namePart id="20B3FA790517927F297C85DA9FBC5D9A">Soleglad, Michael E.</mods:namePart>
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<mods:namePart id="E3F2014614F71FD0C9C5902D588AE64D">Fet, Victor</mods:namePart>
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<mods:typeOfResource id="B339ADDD76638E46C1ED9C488F654801">text</mods:typeOfResource>
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<mods:title id="2FA679AB69A7131904DF23AFC24A85C7">Euscorpius</mods:title>
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<mods:part id="A0EA0B884D6B5C5698D2975036EB038C">
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<mods:date id="F8485B938E5CE4E4B459D9CC6B07922F">2003</mods:date>
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<mods:number id="ACC9779687108D62FDFDD835620B571F">2003-12-26</mods:number>
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<mods:number id="736343E85E57697BC30403F7976F207A">2003</mods:number>
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<mods:number id="0B0C22114DECF17785E345F42484D1E7">11</mods:number>
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<mods:url id="4D12F7C3FA2CCA449CA3D4C15DF772B5">http://dx.doi.org/10.18590/euscorpius.2003.vol2003.iss11.1</mods:url>
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<mods:classification id="E539FD3D6842931BF792C562556760F6">journal article</mods:classification>
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<mods:identifier id="A55DF9B858AE65005E122600C8A7CA2E" type="DOI">10.18590/euscorpius.2003.vol2003.iss11.1</mods:identifier>
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<mods:identifier id="082EF0C8DFC08DD008A4B40380458509" type="ISSN">1536-9307</mods:identifier>
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<treatment id="038A87D5D73EF53BFF615AADFC6A5799" ID-DOI="http://doi.org/10.5281/zenodo.12785267" ID-Zenodo-Dep="12785267" LSID="urn:lsid:plazi:treatment:038A87D5D73EF53BFF615AADFC6A5799" httpUri="http://treatment.plazi.org/id/038A87D5D73EF53BFF615AADFC6A5799" lastPageNumber="111" pageId="114" pageNumber="111">
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<subSubSection id="C3396548D73EF53BFF615AADFDD952FE" box="[192,617,1242,1265]" pageId="114" pageNumber="111" type="nomenclature">
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<paragraph id="8B9C36C3D73EF53BFF615AADFDD952FE" blockId="114.[192,802,1242,1910]" box="[192,617,1242,1265]" pageId="114" pageNumber="111">
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<heading id="D0D481AFD73EF53BFF615AADFDD952FE" box="[192,617,1242,1265]" fontSize="10" level="2" pageId="114" pageNumber="111" reason="5">
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<emphasis id="B957EAD1D73EF53BFF615AADFDD952FE" bold="true" box="[192,617,1242,1265]" pageId="114" pageNumber="111">
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Subfamily
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<taxonomicName id="4C234D40D73EF53BFE9D5AADFDD952FE" authority="Stahnke, 1974" authorityName="Stahnke" authorityYear="1974" box="[316,617,1242,1265]" class="Arachnida" family="Iuridae" kingdom="Animalia" order="Scorpiones" pageId="114" pageNumber="111" phylum="Arthropoda" rank="subFamily" subFamily="Hadrurinae">
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Hadrurinae
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<bibRefCitation id="EFB24B32D73EF53BFE685AADFDD952FE" author="Stahnke" box="[457,617,1242,1265]" firstAuthor="Stahnke" pageId="114" pageNumber="111" pagination="107 - 141" refId="ref97788" refString="STAHNKE, H. L. 1974. Revision and keys to the higher categories of Vejovidae. Journal of Arachnology, 1 (2): 107 - 141." type="journal article" year="1974">Stahnke, 1974</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3396548D73EF53BFF515A8FFE5A5342" pageId="114" pageNumber="111" type="type_taxon">
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<paragraph id="8B9C36C3D73EF53BFF515A8FFD375300" blockId="114.[192,802,1242,1910]" box="[240,647,1272,1295]" pageId="114" pageNumber="111">
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<typeStatus id="54988861D73EF53BFF515A8FFE955300" box="[240,293,1272,1295]" pageId="114" pageNumber="111">Type</typeStatus>
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Genus.
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<taxonomicName id="4C234D40D73EF53BFE215A8FFD345300" ID-CoL="62TPN" authority="Thorell, 1876" authorityName="Thorell" authorityYear="1876" box="[384,644,1272,1295]" class="Arachnida" family="Hadruridae" genus="Hadrurus" kingdom="Animalia" order="Scorpiones" pageId="114" pageNumber="111" phylum="Arthropoda" rank="genus">
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<emphasis id="B957EAD1D73EF53BFE215A8FFE585300" box="[384,488,1272,1295]" italics="true" pageId="114" pageNumber="111">Hadrurus</emphasis>
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<bibRefCitation id="EFB24B32D73EF53BFE4E5A8FFD345300" author="Thorell" box="[495,644,1272,1295]" firstAuthor="Thorell" pageId="114" pageNumber="111" pagination="1 - 15" refId="ref98099" refString="THORELL, T. 1876 a. On the classification of scorpions. Annals and Magazine of Natural History, 4 (17): 1 - 15." type="journal article" year="1876">Thorell, 1876</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8B9C36C3D73EF53BFF515B60FC925321" blockId="114.[192,802,1242,1910]" box="[240,802,1303,1326]" pageId="114" pageNumber="111">
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<emphasis id="B957EAD1D73EF53BFF515B60FE325321" bold="true" box="[240,386,1303,1326]" italics="true" pageId="114" pageNumber="111">Composition.</emphasis>
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This monotypic subfamily includes a
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</paragraph>
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<paragraph id="8B9C36C3D73EF53BFF515B41FE5A5342" blockId="114.[192,802,1242,1910]" box="[240,490,1334,1357]" pageId="114" pageNumber="111">
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single genus
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<taxonomicName id="4C234D40D73EF53BFEDA5B41FE535342" ID-CoL="62TPN" authorityName="Thorell" authorityYear="1876" box="[379,483,1334,1357]" class="Arachnida" family="Hadruridae" genus="Hadrurus" kingdom="Animalia" order="Scorpiones" pageId="114" pageNumber="111" phylum="Arthropoda" rank="genus">
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<emphasis id="B957EAD1D73EF53BFEDA5B41FE535342" box="[379,483,1334,1357]" italics="true" pageId="114" pageNumber="111">Hadrurus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3396548D73EF53BFF515B23FDF05091" pageId="114" pageNumber="111" type="distribution">
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<paragraph id="8B9C36C3D73EF53BFF515B23FD4D5364" blockId="114.[192,802,1242,1910]" box="[240,765,1364,1387]" pageId="114" pageNumber="111">
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<emphasis id="B957EAD1D73EF53BFF515B23FECC5364" bold="true" box="[240,380,1364,1387]" italics="true" pageId="114" pageNumber="111">Distribution.</emphasis>
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North America (
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<collectingCountry id="F3347653D73EF53BFD915B23FDD75364" box="[560,615,1364,1387]" name="United States of America" pageId="114" pageNumber="111">USA</collectingCountry>
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and
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<collectingCountry id="F3347653D73EF53BFD3A5B23FD415364" box="[667,753,1364,1387]" name="Mexico" pageId="114" pageNumber="111">Mexico</collectingCountry>
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).
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</paragraph>
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<paragraph id="8B9C36C3D73EF53BFF515B04FE1A53E9" blockId="114.[192,802,1242,1910]" pageId="114" pageNumber="111">
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<emphasis id="B957EAD1D73EF53BFF515B04FE755385" bold="true" box="[240,453,1395,1418]" italics="true" pageId="114" pageNumber="111">Taxonomic history</emphasis>
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. The content of
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<taxonomicName id="4C234D40D73EF53BFD2A5B04FCB35385" authorityName="Stahnke" authorityYear="1974" box="[651,771,1395,1418]" class="Arachnida" family="Iuridae" kingdom="Animalia" order="Scorpiones" pageId="114" pageNumber="111" phylum="Arthropoda" rank="subFamily" subFamily="Hadrurinae">Hadrurinae</taxonomicName>
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is changed here since we transfer the genus
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<taxonomicName id="4C234D40D73EF53BFDDF5BE5FCB653A6" box="[638,774,1426,1449]" class="Arachnida" family="Chactidae" genus="Anuroctonus" kingdom="Animalia" order="Scorpiones" pageId="114" pageNumber="111" phylum="Arthropoda" rank="genus">
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<emphasis id="B957EAD1D73EF53BFDDF5BE5FCB653A6" box="[638,774,1426,1449]" italics="true" pageId="114" pageNumber="111">Anuroctonus</emphasis>
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</taxonomicName>
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to
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<taxonomicName id="4C234D40D73EF53BFF615BC7FE9B53C8" baseAuthorityName="Stahnke" baseAuthorityYear="1940" box="[192,299,1456,1479]" class="Arachnida" family="Chactidae" kingdom="Animalia" order="Scorpiones" pageId="114" pageNumber="111" phylum="Arthropoda" rank="family">Chactidae</taxonomicName>
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(subfamily
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<taxonomicName id="4C234D40D73EF53BFE105BC7FD8953C8" authorityName="Mello-Leitao" authorityYear="1934" box="[433,569,1456,1479]" class="Arachnida" family="Vaejovidae" kingdom="Animalia" order="Scorpiones" pageId="114" pageNumber="111" phylum="Arthropoda" rank="subFamily" subFamily="Uroctoninae">Uroctoninae</taxonomicName>
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), so that the subfamily is now monotypic.
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</paragraph>
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<paragraph id="8B9C36C3D73EF53BFF515B99FDF05091" blockId="114.[192,802,1242,1910]" pageId="114" pageNumber="111">
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<emphasis id="B957EAD1D73EF53BFF515B99FE55500A" bold="true" box="[240,485,1518,1541]" italics="true" pageId="114" pageNumber="111">Biogeographic history</emphasis>
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. Species of
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<taxonomicName id="4C234D40D73EF53BFDCB5B99FD62500A" authorityName="Thorell" authorityYear="1876" box="[618,722,1518,1541]" class="Arachnida" family="Hadruridae" genus="Hadrurus" kingdom="Animalia" order="Scorpiones" pageId="114" pageNumber="111" phylum="Arthropoda" rank="genus">
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<emphasis id="B957EAD1D73EF53BFDCB5B99FD62500A" box="[618,722,1518,1541]" italics="true" pageId="114" pageNumber="111">Hadrurus</emphasis>
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</taxonomicName>
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are the largest desert scorpions in North America, and their evolution clearly is connected to aridization of this continent (Morafka et al., 1994). DNA phylogeny of this genus (
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<bibRefCitation id="EFB24B32D73EF53BFEAD581FFE005070" author="Fet" box="[268,432,1640,1663]" etAl="et al." firstAuthor="Fet" pageId="114" pageNumber="111" refId="ref96278" refString="SANTIAGO-BLAY, J. A., V. FET, M. E. SOLEGLAD, G. ROMERO & P. CRAIG. 2001. A Tertiary vaejovid scorpion from Mexico and another non-buthid cretaceous scorpion from Brazil. Geological Society of America. Annual Meeting and Exposition Abstracts, November 1 - 10, 2001, Boston, Massachusetts, p. 266." type="proceedings paper" year="2001">Fet et al., 2001</bibRefCitation>
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) agrees with its Mexican origin as suggested earlier (
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<bibRefCitation id="EFB24B32D73EF53BFE2358F0FD835091" author="Soleglad" box="[386,563,1671,1694]" firstAuthor="Soleglad" pageId="114" pageNumber="111" pagination="113 - 134" refId="ref97535" refString="SOLEGLAD, M. E. 1976 a. The taxonomy of the genus Hadrurus based on chela trichobothria (Scorpionida: Vejovidae). Journal of Arachnology, 3: 113 - 134." type="journal article" year="1976" yearSuffix="a">Soleglad, 1976a</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3396548D73EF53BFF5158D1FC665735" pageId="114" pageNumber="111" type="diagnosis">
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<paragraph id="8B9C36C3D73EF53BFF5158D1FC665735" blockId="114.[192,802,1242,1910]" lastBlockId="114.[830,1440,199,406]" pageId="114" pageNumber="111">
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<emphasis id="B957EAD1D73EF53BFF5158D1FEEE50B2" bold="true" box="[240,350,1702,1725]" italics="true" pageId="114" pageNumber="111">Diagnosis</emphasis>
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. Synapomorphies. Major neobothriotaxy present on ventral aspect of pedipalpal chela, and on ventral and external aspects of patella; ventral surface of leg tarsus with fused spinule cluster groups; leg tarsus unguicular spur well-developed; genital papillae of male absent. Important Symplesiomorphies. Chelal trichobothrium
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<emphasis id="B957EAD1D73EF53BFEB5592AFE855179" box="[276,309,1885,1910]" italics="true" pageId="114" pageNumber="111">
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Et
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<subScript id="17A73486D73EF53BFE8D591FFE855179" attach="left" box="[300,309,1896,1910]" fontSize="6" pageId="114" pageNumber="111">5</subScript>
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</emphasis>
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positioned on fixed finger; dorsal edge of cheliceral movable finger with
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<emphasis id="B957EAD1D73EF53BFB3F5EB0FB7456D1" box="[1182,1220,199,222]" italics="true" pageId="114" pageNumber="111">two</emphasis>
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subdistal denticles; leg coxae IV elongated; lateral carinae partially present on metasomal segment IV; chela with “10-carinae” configuration.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3396548D73EF53BFCCF5F35FC6A5799" pageId="114" pageNumber="111" type="discussion">
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<paragraph id="8B9C36C3D73EF53BFCCF5F35FC6A5799" blockId="114.[830,1440,199,406]" pageId="114" pageNumber="111">
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<emphasis id="B957EAD1D73EF53BFCCF5F35FC565756" bold="true" box="[878,998,322,345]" italics="true" pageId="114" pageNumber="111">Discussion</emphasis>
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. See discussion above under subfamily
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<taxonomicName id="4C234D40D73EF53BFC9F5F17FC725778" authorityName="Mello-Leitao" authorityYear="1934" box="[830,962,352,375]" class="Arachnida" family="Vaejovidae" kingdom="Animalia" order="Scorpiones" pageId="114" pageNumber="111" phylum="Arthropoda" rank="subFamily" subFamily="Uroctoninae">Uroctoninae</taxonomicName>
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for the reasons for removing
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<taxonomicName id="4C234D40D73EF53BFAB95F17FA105778" box="[1304,1440,352,375]" class="Arachnida" family="Chactidae" genus="Anuroctonus" kingdom="Animalia" order="Scorpiones" pageId="114" pageNumber="111" phylum="Arthropoda" rank="genus">
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<emphasis id="B957EAD1D73EF53BFAB95F17FA105778" box="[1304,1440,352,375]" italics="true" pageId="114" pageNumber="111">Anuroctonus</emphasis>
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</taxonomicName>
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from
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<taxonomicName id="4C234D40D73EF53BFCD85F08FC665799" authorityName="Thorell" authorityYear="1876" box="[889,982,383,406]" class="Arachnida" kingdom="Animalia" order="Scorpiones" pageId="114" pageNumber="184" phylum="Arthropoda" rank="superFamily" superFamily="Iuroidea">Iuroidea</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |