131 lines
13 KiB
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131 lines
13 KiB
XML
<document id="840F415708870362BFCBB077EF8ED85F" ID-DOI="10.18590/euscorpius.2003.vol2003.iss11.1" ID-ISSN="1536-9307" ID-Zenodo-Dep="13237351" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="existingObjects" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1721353212353" checkinUser="felipe" docAuthor="Soleglad, Michael E. & Fet, Victor" docDate="2003" docId="038A87D5D710F515FF615CC5FEA550D4" docLanguage="en" docName="Euscorpius.11.1-175.pdf" docOrigin="Euscorpius 2003 (11)" docSource="http://dx.doi.org/10.18590/euscorpius.2003.vol2003.iss11.1" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Pseudochactidae Gromov 1998" docType="treatment" docVersion="5" lastPageNumber="89" masterDocId="FFB3FFADD74CF549FFA15E77FFB0560F" masterDocTitle="High-level systematics and phylogeny of the extant scorpions (Scorpiones: Orthosterni)" masterLastPageNumber="175" masterPageNumber="1" pageNumber="89" updateTime="1722954143772" updateUser="ExternalLinkService">
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<mods:titleInfo id="4BACDEA71C4B8E385A4BA1087A730167">
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<mods:title id="4DE3ED41C35A1E65A63A387C5A2BF284">High-level systematics and phylogeny of the extant scorpions (Scorpiones: Orthosterni)</mods:title>
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<mods:namePart id="F3122435D35F2FD598B9FC829DA22815">Soleglad, Michael E.</mods:namePart>
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<mods:namePart id="172E35A6F77CABF81A31BB6806F409B6">Fet, Victor</mods:namePart>
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<mods:typeOfResource id="532C823C2682EE1534BD42F7A8326D5F">text</mods:typeOfResource>
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<mods:title id="A73692DE8B00D85D7EF99C367D2DBE76">Euscorpius</mods:title>
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</mods:titleInfo>
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<mods:part id="1E6B9A1BE3C8B54E3ACBD04930B3E2C1">
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<mods:date id="B6015296B03BD371EC5F67015CE24805">2003</mods:date>
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<mods:number id="560D87BEAAC2150938D3B558B89E295C">2003-12-26</mods:number>
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<mods:detail id="50051CE1A2B48522123547A23F4C8B78" type="volume">
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<mods:number id="F4CC35E8A5FA4B29599CE4ECAB08DE51">2003</mods:number>
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<mods:detail id="EE13E84A3A4E0816F1D9DA8A9B62024B" type="issue">
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<mods:number id="EF138EEBC6FE05AF809F3971D0DD2C3D">11</mods:number>
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<mods:url id="692E6223F175B03548ADB66A72926958">http://dx.doi.org/10.18590/euscorpius.2003.vol2003.iss11.1</mods:url>
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<mods:classification id="BABBADB73974D1345517E207C007FEC7">journal article</mods:classification>
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<mods:identifier id="DDB4B2472D457ACF8BB28ABC2BEF7B66" type="DOI">10.18590/euscorpius.2003.vol2003.iss11.1</mods:identifier>
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<mods:identifier id="CDFBAC8171365E824257A1B5C58CBF7C" type="ISSN">1536-9307</mods:identifier>
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<treatment id="038A87D5D710F515FF615CC5FEA550D4" ID-DOI="http://doi.org/10.5281/zenodo.12785173" ID-Zenodo-Dep="12785173" LSID="urn:lsid:plazi:treatment:038A87D5D710F515FF615CC5FEA550D4" httpUri="http://treatment.plazi.org/id/038A87D5D710F515FF615CC5FEA550D4" lastPageNumber="89" pageId="92" pageNumber="89">
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<subSubSection id="C3396548D710F515FF615CC5FD3354C6" box="[192,643,690,713]" pageId="92" pageNumber="89" type="nomenclature">
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<paragraph id="8B9C36C3D710F515FF615CC5FD3354C6" blockId="92.[192,802,690,1755]" box="[192,643,690,713]" pageId="92" pageNumber="89">
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<heading id="D0D481AFD710F515FF615CC5FD3354C6" box="[192,643,690,713]" fontSize="10" level="2" pageId="92" pageNumber="89" reason="5">
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<emphasis id="B957EAD1D710F515FF615CC5FDCC54C6" bold="true" box="[192,636,690,713]" pageId="92" pageNumber="89">
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Family
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<taxonomicName id="4C234D40D710F515FEB65CC5FDCC54C6" ID-CoL="B7XCK" authority="Gromov, 1998" authorityName="Gromov" authorityYear="1998" box="[279,636,690,713]" class="Arachnida" family="Pseudochactidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Scorpiones" pageId="92" pageNumber="89" phylum="Arthropoda" rank="family">Pseudochactidae Gromov, 1998</taxonomicName>
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</emphasis>
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.
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3396548D710F515FF515CA7FE8E554C" pageId="92" pageNumber="89" type="type_taxon">
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<paragraph id="8B9C36C3D710F515FF515CA7FD7A54E8" blockId="92.[192,802,690,1755]" box="[240,714,720,743]" pageId="92" pageNumber="89">
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<typeStatus id="54988861D710F515FF515CA7FE9554E8" box="[240,293,720,743]" pageId="92" pageNumber="89">Type</typeStatus>
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Genus.
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<taxonomicName id="4C234D40D710F515FE215CA7FD7654E8" ID-CoL="B7YV3" authority="Gromov, 1998" authorityName="Gromov" authorityYear="1998" box="[384,710,720,743]" class="Arachnida" family="Pseudochactidae" genus="Pseudochactas" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Scorpiones" pageId="92" pageNumber="89" phylum="Arthropoda" rank="genus">
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<emphasis id="B957EAD1D710F515FE215CA7FD9054E8" box="[384,544,720,743]" italics="true" pageId="92" pageNumber="89">Pseudochactas</emphasis>
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Gromov, 1998
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8B9C36C3D710F515FF515C98FE8E554C" blockId="92.[192,802,690,1755]" pageId="92" pageNumber="89">
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<emphasis id="B957EAD1D710F515FF515C98FE325509" bold="true" box="[240,386,751,774]" italics="true" pageId="92" pageNumber="89">Composition.</emphasis>
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This monotypic family includes a single monotypic genus
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<taxonomicName id="4C234D40D710F515FE6A5D79FDDB552A" ID-CoL="B7YV3" authorityName="Gromov" authorityYear="1998" box="[459,619,782,805]" class="Arachnida" family="Pseudochactidae" genus="Pseudochactas" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Scorpiones" pageId="92" pageNumber="89" phylum="Arthropoda" rank="genus">
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<emphasis id="B957EAD1D710F515FE6A5D79FDDB552A" box="[459,619,782,805]" italics="true" pageId="92" pageNumber="89">Pseudochactas</emphasis>
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</taxonomicName>
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(Gromov, 1998; Fet, 2000f).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3396548D710F515FF515D3CFE595364" pageId="92" pageNumber="89" type="distribution">
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<paragraph id="8B9C36C3D710F515FF515D3CFDB7558E" blockId="92.[192,802,690,1755]" pageId="92" pageNumber="89">
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<emphasis id="B957EAD1D710F515FF515D3CFEC5556D" bold="true" box="[240,373,843,866]" italics="true" pageId="92" pageNumber="89">Distribution</emphasis>
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. Central Asia (southeastern
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<collectingCountry id="F3347653D710F515FD6D5D3CFF5E558E" name="Uzbekistan" pageId="92" pageNumber="89">Uzbekistan</collectingCountry>
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, southwestern
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<collectingCountry id="F3347653D710F515FE2D5D1DFE4C558E" box="[396,508,874,897]" name="Tajikistan" pageId="92" pageNumber="89">Tajikistan</collectingCountry>
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).
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</paragraph>
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<paragraph id="8B9C36C3D710F515FF515DFFFE595364" blockId="92.[192,802,690,1755]" pageId="92" pageNumber="89">
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<emphasis id="B957EAD1D710F515FF515DFFFE595590" bold="true" box="[240,489,904,927]" italics="true" pageId="92" pageNumber="89">Biogeographic history</emphasis>
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. Unclear. The single genus
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<taxonomicName id="4C234D40D710F515FF615DD0FED055B1" authorityName="Gromov" authorityYear="1998" box="[192,352,935,958]" class="Arachnida" family="Pseudochactidae" genus="Pseudochactas" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Scorpiones" pageId="92" pageNumber="89" phylum="Arthropoda" rank="genus">
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<emphasis id="B957EAD1D710F515FF615DD0FED055B1" box="[192,352,935,958]" italics="true" pageId="92" pageNumber="89">Pseudochactas</emphasis>
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</taxonomicName>
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is found only in a restricted location in the mountains of Central Asia. The relict character of
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<taxonomicName id="4C234D40D710F515FF615D93FED055F4" authorityName="Gromov" authorityYear="1998" box="[192,352,996,1019]" class="Arachnida" family="Pseudochactidae" genus="Pseudochactas" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Scorpiones" pageId="92" pageNumber="89" phylum="Arthropoda" rank="genus">
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<emphasis id="B957EAD1D710F515FF615D93FED055F4" box="[192,352,996,1019]" italics="true" pageId="92" pageNumber="89">Pseudochactas</emphasis>
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</taxonomicName>
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could be due to its preservation in mildclimate, low-mountain depressions of Babatag and Gandzhino ranges (Gromov, 1998), with desert surroundings at lower altitudes. Ecologically, this is not a desert scorpion; it actively forages on wet mud along the temporary waterways, and likely spends most of dry season in hibernation (V. Fet & A. Gromov, pers. observation, 2002). It could represent one of the few faunal remnants of littoral fauna of the receded Tethys Ocean (Kryzhanovsky, 1965), elevated by Tertiary mountain uplift. Many such important floral elements of littoral origin are found in low mountains of
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<collectingCountry id="F3347653D710F515FDD75B41FD5E5342" box="[630,750,1334,1357]" name="Uzbekistan" pageId="92" pageNumber="89">Uzbekistan</collectingCountry>
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and
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<collectingCountry id="F3347653D710F515FF615B23FE9B5364" box="[192,299,1364,1387]" name="Tajikistan" pageId="92" pageNumber="89">Tajikistan</collectingCountry>
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(Kamelin, 1979).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3396548D710F515FF515B04FD775385" box="[240,711,1395,1418]" pageId="92" pageNumber="89" type="diagnosis">
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<paragraph id="8B9C36C3D710F515FF515B04FD775385" blockId="92.[192,802,690,1755]" box="[240,711,1395,1418]" pageId="92" pageNumber="89">
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<emphasis id="B957EAD1D710F515FF515B04FEEE5385" bold="true" box="[240,350,1395,1418]" italics="true" pageId="92" pageNumber="89">Diagnosis</emphasis>
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. As in parvorder
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<taxonomicName id="4C234D40D710F515FDB85B04FD735385" authorityName="Soleglad & Fet" authorityYear="2003" box="[537,707,1395,1418]" class="Arachnida" kingdom="Animalia" order="Scorpiones" pageId="92" pageNumber="89" parvOrder="Pseudochactida" phylum="Arthropoda" rank="parvOrder">Pseudochactida</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3396548D710F515FF515BE5FEA550D4" pageId="92" pageNumber="89" type="discussion">
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<paragraph id="8B9C36C3D710F515FF515BE5FEA550D4" blockId="92.[192,802,690,1755]" pageId="92" pageNumber="89">
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<emphasis id="B957EAD1D710F515FF515BE5FED853A6" bold="true" box="[240,360,1426,1449]" italics="true" pageId="92" pageNumber="89">Discussion</emphasis>
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. Gromov (1998: 1003) keenly observed that “It is possible that the representative of this new family (
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<taxonomicName id="4C234D40D710F515FEB85BB8FE7F53E9" authorityName="Gromov" authorityYear="1998" box="[281,463,1487,1510]" class="Arachnida" family="Pseudochactidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Scorpiones" pageId="92" pageNumber="89" phylum="Arthropoda" rank="family">Pseudochactidae</taxonomicName>
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) stands close to the common ancestor of all these families [the Recent scorpions]”. As
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<bibRefCitation id="EFB24B32D710F515FF61587BFE71502C" author="Soleglad & Fet" box="[192,449,1548,1571]" firstAuthor="Soleglad" pageId="92" pageNumber="89" pagination="1 - 38" refId="ref97606" refString="SOLEGLAD, M. E. & V. FET. 2001. Evolution of scorpion orthobothriotaxy: a cladistic approach. Euscorpius, 1: 1 - 38." type="journal article" year="2001">Soleglad & Fet (2001)</bibRefCitation>
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demonstrated based on trichobothrial patterns,
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<taxonomicName id="4C234D40D710F515FE2E585CFD9F504D" authorityName="Gromov" authorityYear="1998" box="[399,559,1579,1602]" class="Arachnida" family="Pseudochactidae" genus="Pseudochactas" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Scorpiones" pageId="92" pageNumber="89" phylum="Arthropoda" rank="genus">
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<emphasis id="B957EAD1D710F515FE2E585CFD9F504D" box="[399,559,1579,1602]" italics="true" pageId="92" pageNumber="89">Pseudochactas</emphasis>
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</taxonomicName>
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is certainly a basal member of Recent scorpions, and also shows significant affinity in some characters with the Carboniferous fossil scorpion family
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<taxonomicName id="4C234D40D710F515FE2058F0FDC25091" authorityName="Kjellesvig-Waering" authorityYear="1986" box="[385,626,1671,1694]" class="Arachnida" family="Palaeopisthacanthidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="92" pageNumber="89" phylum="Arthropoda" rank="family">Palaeopisthacanthidae</taxonomicName>
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. This veritable “platypus of a scorpion” deserves further study in all aspects.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |