176 lines
21 KiB
XML
176 lines
21 KiB
XML
<document id="FDAE84B0AFAD1893B8AD9487C6127C2F" ID-CLB-Dataset="27961" ID-DOI="10.11646/zootaxa.4521.2.5" ID-GBIF-Dataset="cd0f7302-9d9e-4e0e-aba7-be5ddc783a6f" ID-ISSN="1175-5326" ID-Zenodo-Dep="2609837" ID-ZooBank="F992EA3F-2AD0-49F8-9230-266B4D4C3ACE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553623770548" checkinUser="plazi" docAuthor="Kluge, Nikita J. & Bernal Vega, Juan A." docDate="2018" docId="4A1E8797FF79AC6CFF48822EFE4C9415" docLanguage="en" docName="zootaxa.4521.2.5.pdf" docOrigin="Zootaxa 4521 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Moribaetis salvini" docType="treatment" docVersion="9" lastPageNumber="256" masterDocId="B627FFEFFF61AC75FFDF8365FFF19701" masterDocTitle="Redescription of the Central American genus Moribaetis Waltz & McCafferty 1985 (Ephemeroptera, Baetidae)" masterLastPageNumber="257" masterPageNumber="231" pageNumber="255" updateTime="1698713718017" updateUser="ExternalLinkService">
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<mods:title id="D200820E74BB3BAFC143A380B4F55179">Redescription of the Central American genus Moribaetis Waltz & McCafferty 1985 (Ephemeroptera, Baetidae)</mods:title>
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<mods:namePart id="242A522FEB6D4EB1522378257858A5F8">Kluge, Nikita J.</mods:namePart>
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<mods:namePart id="6A056DE8F93D44189BF82C8CC9BE3B9B">Bernal Vega, Juan A.</mods:namePart>
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<mods:title id="B5DB3425AE1EB35B5B6830483185514F">Zootaxa</mods:title>
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<mods:date id="7B5E52E079D0D7D8489CD3944C4837FD">2018</mods:date>
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<mods:number id="82945CF94E4852BE97285C877BCD31B6">4521</mods:number>
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<treatment id="4A1E8797FF79AC6CFF48822EFE4C9415" ID-DOI="http://doi.org/10.5281/zenodo.5952432" ID-GBIF-Taxon="154498766" ID-Zenodo-Dep="5952432" LSID="urn:lsid:plazi:treatment:4A1E8797FF79AC6CFF48822EFE4C9415" httpUri="http://treatment.plazi.org/id/4A1E8797FF79AC6CFF48822EFE4C9415" lastPageId="25" lastPageNumber="256" pageId="24" pageNumber="255">
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<subSubSection id="8AAD650AFF79AC6DFF48822EFDE99667" box="[151,536,331,358]" pageId="24" pageNumber="255" type="nomenclature">
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<paragraph id="C2083681FF79AC6DFF48822EFDE99667" blockId="24.[151,536,331,391]" box="[151,536,331,358]" pageId="24" pageNumber="255">
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<heading id="994081EDFF79AC6DFF48822EFDE99667" bold="true" box="[151,536,331,358]" fontSize="11" level="1" pageId="24" pageNumber="255" reason="1">
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<taxonomicName id="05B74D02FF79AC6DFF48822EFDE99667" ID-CoL="44DBL" authority="(Eaton 1885)" baseAuthorityName="Eaton" baseAuthorityYear="1885" box="[151,536,331,358]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="24" pageNumber="255" phylum="Arthropoda" rank="species" species="salvini">
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<emphasis id="F0C3EA93FF79AC6DFF48822EFDE99667" bold="true" box="[151,536,331,358]" pageId="24" pageNumber="255">
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<emphasis id="F0C3EA93FF79AC6DFF48822EFE819664" bold="true" box="[151,368,331,357]" italics="true" pageId="24" pageNumber="255">Moribaetis salvini</emphasis>
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(Eaton 1885)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="8AAD650AFF79AC6DFF48820BFEB29686" box="[151,323,366,391]" pageId="24" pageNumber="255" type="description">
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<paragraph id="C2083681FF79AC6DFF48820BFEB29686" blockId="24.[151,536,331,391]" box="[151,323,366,391]" pageId="24" pageNumber="255">
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(
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<figureCitation id="5A8C2A04FF79AC6DFF40820BFECB9686" box="[159,314,366,391]" captionStart="FIGURES 102–105" captionStartId="24.[151,264,1973,1996]" captionTargetBox="[151,1436,574,1953]" captionTargetId="figure@24.[151,1436,574,1953]" captionTargetPageId="24" captionText="FIGURES 102–105. Moribaetis salvini (lectotype, photo by Ben Price). 102, lateral view; 103, fragment of cercus; 104, dorsal view and tube with genitals; 105, labels." httpUri="https://zenodo.org/record/2609869/files/figure.png" pageId="24" pageNumber="255">Figs 102–105</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="8AAD650AFF79AC6DFF4882DDFE41950B" pageId="24" pageNumber="255" type="reference_group">
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<paragraph id="C2083681FF79AC6DFF4882DDFE41950B" blockId="24.[151,1436,438,522]" pageId="24" pageNumber="255">
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<taxonomicName id="05B74D02FF79AC6DFF4882DDFE3E96CD" authority="Eaton 1885: 170" authorityName="Eaton" authorityPageNumber="170" authorityYear="1885" box="[151,463,438,461]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="24" pageNumber="255" phylum="Arthropoda" rank="species" species="salvini">
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<emphasis id="F0C3EA93FF79AC6DFF4882DDFED196CD" box="[151,288,438,461]" italics="true" pageId="24" pageNumber="255">Baetis salvini</emphasis>
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Eaton 1885: 170
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</taxonomicName>
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, Pl. XVI: figs 29a (male imago);
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<bibRefCitation id="A6264B70FF79AC6DFCFD82D3FC4B96CD" author="Kimmins, D. E." box="[802,954,438,460]" pageId="24" pageNumber="255" pagination="269 - 318" refId="ref11098" refString="Kimmins, D. E. (1960) The Ephemeroptera types of species described by A. E. Eaton, R. McLachlan and F. Walker, with particular reference to those in the British Museum (Natural History). Bulletin of the British Museum (Natural History), Entomology, 9 (4), 269 - 318." type="journal article" year="1960">Kimmins 1960</bibRefCitation>
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(
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<typeStatus id="1D0C8823FF79AC6DFC1982D3FBD596CD" box="[966,1060,438,460]" pageId="24" pageNumber="255" type="lectotype">lectotype</typeStatus>
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designation).
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<taxonomicName id="05B74D02FF79AC6DFF4882B0FD5596EA" ID-CoL="44DBL" authority=": Waltz & McCafferty 1985: 247" authorityName=": Waltz & McCafferty" authorityPageNumber="247" authorityYear="1985" box="[151,676,469,491]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="24" pageNumber="255" phylum="Arthropoda" rank="species" species="salvini">
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<emphasis id="F0C3EA93FF79AC6DFF4882B0FEA196EA" box="[151,336,469,491]" italics="true" pageId="24" pageNumber="255">Moribaetis salvini</emphasis>
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:
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<bibRefCitation id="A6264B70FF79AC6DFE8082B0FD8196EA" author="Waltz, R. D. & McCafferty, W. P." box="[351,624,469,491]" pageId="24" pageNumber="255" pagination="239 - 251" refId="ref11911" refString="Waltz, R. D. & McCafferty, W. P. (1985) Moribaetis: A new genus of Neotropical Baetidae (Ephemeroptera). Proceedings of the Entomological Society of Washington, 87 (1), 239 - 251." type="journal article" year="1985">Waltz & McCafferty 1985</bibRefCitation>
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: 247
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</taxonomicName>
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, figs 42–44 (male imago, not larva);
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<bibRefCitation id="A6264B70FF79AC6DFBFC82B0FA9A96EA" author="McCafferty, W. P. & Lugo-Ortiz, C. R." box="[1059,1387,469,491]" pageId="24" pageNumber="255" pagination="117 - 121" refId="ref11722" refString="McCafferty, W. P. & Lugo-Ortiz, C. R. (1998) The adult of Moribaetis macaferti (Ephemeroptera: Baetidae). Entomological News, 109 (2), 117 - 121." type="journal article" year="1998">McCafferty & Lugo-Ortiz 1998</bibRefCitation>
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: 120 (key to male imagoes).
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</paragraph>
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</subSubSection>
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<caption id="96C86609FF79AC6DFF4884D0FDDE90EC" httpUri="https://zenodo.org/record/2609869/files/figure.png" pageId="24" pageNumber="255" startId="24.[151,264,1973,1996]" targetBox="[151,1436,574,1953]" targetPageId="24">
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<paragraph id="C2083681FF79AC6DFF4884D0FDDE90EC" blockId="24.[151,1436,1973,2029]" pageId="24" pageNumber="255">
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<emphasis id="F0C3EA93FF79AC6DFF4884D0FE9D90CA" bold="true" box="[151,364,1973,1996]" pageId="24" pageNumber="255">FIGURES 102–105.</emphasis>
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<taxonomicName id="05B74D02FF79AC6DFEAD84D3FDDA90CD" authorityName=": Waltz & McCafferty" authorityYear="1985" box="[370,555,1974,1996]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="24" pageNumber="255" phylum="Arthropoda" rank="species" species="salvini">
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<emphasis id="F0C3EA93FF79AC6DFEAD84D3FDDA90CD" box="[370,555,1974,1996]" italics="true" pageId="24" pageNumber="255">Moribaetis salvini</emphasis>
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</taxonomicName>
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(lectotype, photo by Ben Price). 102, lateral view; 103, fragment of cercus; 104, dorsal view and tube with genitals; 105, labels.
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</paragraph>
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</caption>
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<subSubSection id="8AAD650AFF78AC6CFF4883F2FE4C9415" pageId="25" pageNumber="256" type="discussion">
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<paragraph id="C2083681FF78AC6CFF4883F2FEA9961C" blockId="25.[151,1436,151,789]" pageId="25" pageNumber="256">
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<emphasis id="F0C3EA93FF78AC6CFF4883F2FEEE97B1" bold="true" box="[151,287,151,176]" pageId="25" pageNumber="256">Comments.</emphasis>
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<taxonomicName id="05B74D02FF78AC6CFEF983FDFE4E97B1" authorityName="Eaton" authorityYear="1885" box="[294,447,152,176]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="25" pageNumber="256" phylum="Arthropoda" rank="species" species="salvini">
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<emphasis id="F0C3EA93FF78AC6CFEF983FDFE4E97B1" box="[294,447,152,176]" italics="true" pageId="25" pageNumber="256">Baetis salvini</emphasis>
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</taxonomicName>
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was originally described from
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<collectingCountry id="BAA07611FF78AC6CFCC083FDFC6897B1" box="[799,921,151,176]" name="Costa Rica" pageId="25" pageNumber="256">Costa Rica</collectingCountry>
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as one male imago and one female subimago (Eaton 1885). Fore wing, hind wing and genitals of the male imago are accurately figured in the original description (Eaton 1885: pl. XVI: fig. 29a). Subsequently, this male imago was designated as the
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<typeStatus id="1D0C8823FF78AC6CFAEA83BAFA6D97F9" box="[1333,1436,223,248]" pageId="25" pageNumber="256" type="lectotype">lectotype</typeStatus>
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(
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<bibRefCitation id="A6264B70FF78AC6CFF418261FEBA961C" author="Kimmins, D. E." box="[158,331,260,285]" pageId="25" pageNumber="256" pagination="269 - 318" refId="ref11098" refString="Kimmins, D. E. (1960) The Ephemeroptera types of species described by A. E. Eaton, R. McLachlan and F. Walker, with particular reference to those in the British Museum (Natural History). Bulletin of the British Museum (Natural History), Entomology, 9 (4), 269 - 318." type="journal article" year="1960">Kimmins 1960</bibRefCitation>
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).
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</paragraph>
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<paragraph id="C2083681FF78AC6CFF18824DFD7B96D1" blockId="25.[151,1436,151,789]" pageId="25" pageNumber="256">
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<bibRefCitation id="A6264B70FF78AC6CFF18824DFEA89641" author="Eaton, A. E." box="[199,345,296,320]" pageId="25" pageNumber="256" pagination="1 - 16" refId="ref10944" refString="Eaton, A. E. (1892) Ephemeridae. In: Biologia Centrali-Americana, Zoologia, Class Insecta, Order Neuroptera, pp. 1 - 16." type="book chapter" year="1892">Eaton (1892)</bibRefCitation>
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described one male imago, two female imagoes and one male subimago from various localities in
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<collectingCountry id="BAA07611FF78AC6CFF488229FEE59664" box="[151,276,332,357]" name="Guatemala" pageId="25" pageNumber="256">Guatemala</collectingCountry>
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; he determined them as belonging to
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<taxonomicName id="05B74D02FF78AC6CFD638228FCA79665" authorityName="Eaton" authorityYear="1885" box="[700,854,332,356]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="25" pageNumber="256" phylum="Arthropoda" rank="species" species="salvini">
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<emphasis id="F0C3EA93FF78AC6CFD638228FCA79665" box="[700,854,332,356]" italics="true" pageId="25" pageNumber="256">Baetis salvini</emphasis>
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</taxonomicName>
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, but noted their difference from the
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<typeStatus id="1D0C8823FF78AC6CFB2D8228FAD39664" box="[1266,1314,333,357]" pageId="25" pageNumber="256">type</typeStatus>
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specimen;
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<bibRefCitation id="A6264B70FF78AC6CFF48820AFEA29689" author="Kimmins, D. E." box="[151,339,367,392]" pageId="25" pageNumber="256" pagination="338 - 353" refId="ref11045" refString="Kimmins, D. E. (1934) Notes on the Ephemeroptera of the Godman and Salvin collection, with description of two new species. Annals and Magazine of Natural History, Series 1, 14, 338 - 353. https: // doi. org / 10.1080 / 00222933408654903" type="journal article" year="1934">Kimmins (1934)</bibRefCitation>
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described and figured genitals of this male imago. Judging by greatly elongated 1st segment of gonostylus (wrongly called «second joint») (
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<bibRefCitation id="A6264B70FF78AC6CFD4F82F1FC6796AD" author="Kimmins, D. E." box="[656,918,404,429]" pageId="25" pageNumber="256" pagination="338 - 353" refId="ref11045" refString="Kimmins, D. E. (1934) Notes on the Ephemeroptera of the Godman and Salvin collection, with description of two new species. Annals and Magazine of Natural History, Series 1, 14, 338 - 353. https: // doi. org / 10.1080 / 00222933408654903" type="journal article" year="1934">Kimmins 1934: fig. 13</bibRefCitation>
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), this specimen belongs to
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<taxonomicName id="05B74D02FF78AC6CFB1382F1FAB696AD" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[1228,1351,404,428]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="25" pageNumber="256" phylum="Arthropoda" rank="genus">
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<emphasis id="F0C3EA93FF78AC6CFB1382F1FAB696AD" box="[1228,1351,404,428]" italics="true" pageId="25" pageNumber="256">Moribaetis</emphasis>
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</taxonomicName>
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, but its exact systematic position should be verified.
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</paragraph>
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<paragraph id="C2083681FF78AC6CFF1882B9FEB995A6" blockId="25.[151,1436,151,789]" pageId="25" pageNumber="256">
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||
<bibRefCitation id="A6264B70FF78AC6CFF1882B9FDF496F4" author="Waltz, R. D. & McCafferty, W. P." box="[199,517,476,501]" pageId="25" pageNumber="256" pagination="239 - 251" refId="ref11911" refString="Waltz, R. D. & McCafferty, W. P. (1985) Moribaetis: A new genus of Neotropical Baetidae (Ephemeroptera). Proceedings of the Entomological Society of Washington, 87 (1), 239 - 251." type="journal article" year="1985">Waltz & McCafferty (1985)</bibRefCitation>
|
||
reported the
|
||
<typeStatus id="1D0C8823FF78AC6CFD7E82B9FCF996F4" box="[673,776,476,501]" pageId="25" pageNumber="256" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="05B74D02FF78AC6CFCEE82B9FC8D96F5" authorityName="Eaton" authorityYear="1885" box="[817,892,476,500]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="25" pageNumber="256" phylum="Arthropoda" rank="species" species="salvini">
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<emphasis id="F0C3EA93FF78AC6CFCEE82B9FC8D96F5" box="[817,892,476,500]" italics="true" pageId="25" pageNumber="256">salvini</emphasis>
|
||
</taxonomicName>
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||
[
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<taxonomicName id="05B74D02FF78AC6CFC5182B8FC2596F5" authorityName="Leach" authorityYear="1815" box="[910,980,477,500]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="25" pageNumber="256" phylum="Arthropoda" rank="genus">
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||
<emphasis id="F0C3EA93FF78AC6CFC5182B8FC2596F5" box="[910,980,477,500]" italics="true" pageId="25" pageNumber="256">Baetis</emphasis>
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</taxonomicName>
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] among material examined, but did not give additional description to this species; they only wrote that it was «Adequately characterized by Eaton (1885)», gave figures of wings redrawn from the original description and added a new drawing of its cercus (
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<bibRefCitation id="A6264B70FF78AC6CFAE68141FE489561" author="Waltz, R. D. & McCafferty, W. P." pageId="25" pageNumber="256" pagination="239 - 251" refId="ref11911" refString="Waltz, R. D. & McCafferty, W. P. (1985) Moribaetis: A new genus of Neotropical Baetidae (Ephemeroptera). Proceedings of the Entomological Society of Washington, 87 (1), 239 - 251." type="journal article" year="1985">Waltz & McCafferty 1985: fig. 42</bibRefCitation>
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). Photos of this specimen, kindly done by Ben Price, demonstrate that its coloration of fore wing and coloration of fore femur (
|
||
<figureCitation id="5A8C2A04FF78AC6CFDA78109FD2C9585" box="[632,733,620,645]" captionStart="FIGURES 102–105" captionStartId="24.[151,264,1973,1996]" captionTargetBox="[151,1436,574,1953]" captionTargetId="figure@24.[151,1436,574,1953]" captionTargetPageId="24" captionText="FIGURES 102–105. Moribaetis salvini (lectotype, photo by Ben Price). 102, lateral view; 103, fragment of cercus; 104, dorsal view and tube with genitals; 105, labels." httpUri="https://zenodo.org/record/2609869/files/figure.png" pageId="25" pageNumber="256">Fig. 102</figureCitation>
|
||
) are clearly different from that of
|
||
<taxonomicName id="05B74D02FF78AC6CFBA18108FACF9585" authorityName="Flowers" authorityYear="1979" box="[1150,1342,620,644]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="25" pageNumber="256" phylum="Arthropoda" rank="species" species="maculipennis">
|
||
<emphasis id="F0C3EA93FF78AC6CFBA18108FACF9585" box="[1150,1342,620,644]" italics="true" pageId="25" pageNumber="256">M. maculipennis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="05B74D02FF78AC6CFAA28108FEF795A9" authority="Kluge & Bernal Vega, 2018" authorityName="Kluge & Bernal Vega" authorityYear="2018" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="25" pageNumber="256" phylum="Arthropoda" rank="species" species="latipennis" status="sp. nov.">
|
||
<emphasis id="F0C3EA93FF78AC6CFAA28108FEF795A9" italics="true" pageId="25" pageNumber="256">M. latipennis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBF057E8FF78AC6CFED381F5FEB995A6" box="[268,328,656,679]" pageId="25" pageNumber="256" rank="species">sp. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="C2083681FF78AC6CFF1881D1FE4C9415" blockId="25.[151,1436,151,789]" pageId="25" pageNumber="256">
|
||
Judging by the figure in the original description (Eaton 1885: fig. 29a), gonostyli of
|
||
<taxonomicName id="05B74D02FF78AC6CFB5581D1FB2495CD" authorityName="Eaton" authorityYear="1885" box="[1162,1237,692,716]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="25" pageNumber="256" phylum="Arthropoda" rank="species" species="salvini">
|
||
<emphasis id="F0C3EA93FF78AC6CFB5581D1FB2495CD" box="[1162,1237,692,716]" italics="true" pageId="25" pageNumber="256">salvini</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<taxonomicName id="05B74D02FF78AC6CFB3981D0FADD95CD" authorityName="Leach" authorityYear="1815" box="[1254,1324,693,716]" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="25" pageNumber="256" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF78AC6CFB3981D0FADD95CD" box="[1254,1324,693,716]" italics="true" pageId="25" pageNumber="256">Baetis</emphasis>
|
||
</taxonomicName>
|
||
] have 1st segment longer than 2nd, and 3rd segment very short; this feature, as well as coloration of wings, allows to place this species to
|
||
<taxonomicName id="05B74D02FF78AC6CFEE48199FE479415" authorityName=": Lugo-Ortiz & McCafferty" authorityYear="1996" box="[315,438,764,788]" class="Insecta" family="Baetidae" genus="Moribaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="25" pageNumber="256" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F0C3EA93FF78AC6CFEE48199FE479415" box="[315,438,764,788]" italics="true" pageId="25" pageNumber="256">Moribaetis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |