300 lines
29 KiB
XML
300 lines
29 KiB
XML
<document id="A83349AE4402E964A01AEE6A866DE365" ID-DOI="10.11646/zootaxa.5374.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10148092" ID-ZooBank="D73FADAD-514B-4BE8-87B9-F4E66FD21406" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1700205240597" checkinUser="plazi" docAuthor="Tabell, Jukka, Kullberg, Jaakko, Mutanen, Marko, Tokár, Zdenko & Sihvonen, Pasi" docDate="2023" docId="038AB73454507824EAE2FE26FE0C58F2" docLanguage="en" docName="zootaxa.5374.2.1.pdf" docOrigin="Zootaxa 5374 (2)" docSource="https://mapress.com/zt/article/download/zootaxa.5374.2.1/52284" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coleophora carsteni Tabell & Kullberg & Mutanen & Tokár & Sihvonen 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="160" masterDocId="FFB3CF4C5459782DEA75FFA1FF925E41" masterDocTitle="New and little known Coleophora Hübner, 1822 species from Morocco. Part I (Lepidoptera, Coleophoridae)" masterLastPageNumber="195" masterPageNumber="151" pageNumber="160" updateTime="1700675694182" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title id="E50A49BB903C39B62869C06547615436">New and little known Coleophora Hübner, 1822 species from Morocco. Part I (Lepidoptera, Coleophoridae)</mods:title>
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<mods:namePart id="AF0B6443987CBCD8ACE682815E83AEBC">Tabell, Jukka</mods:namePart>
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<mods:nameIdentifier id="D9868462428B95883AB55B499F868B4E" type="email">jukka.tabell@phnet.fi</mods:nameIdentifier>
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<mods:nameIdentifier id="84105CA22924ED92D969BFAF86538351" type="email">jaakko.kullberg@gmail.com</mods:nameIdentifier>
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<mods:namePart id="7DC528A0F79DE831FF6DB1B65B5E38DC">Mutanen, Marko</mods:namePart>
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<mods:affiliation id="13B2F794A484BE1CDACC7A58CF605ABB">Ecology and Genetics Research Unit, Po Box 3000, FI- 90014 University of Oulu, Finland.</mods:affiliation>
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<mods:namePart id="709E7E65A243DF90C18312F64DD0F9E6">Tokár, Zdenko</mods:namePart>
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<mods:nameIdentifier id="710F7055A686872320D2B457CCC74EDA" type="email">zdeno.tokar@gmail.com</mods:nameIdentifier>
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<mods:namePart id="DF356B0FE4F4D4754F25419A09E1BED4">Sihvonen, Pasi</mods:namePart>
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<mods:affiliation id="900126C91237A6BC361C778CBF9DA912">Finnish Museum of Natural History, University of Helsinki, P. O. Box 17, FI- 00014, Finland.</mods:affiliation>
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<mods:nameIdentifier id="75640C7EBE798BE5B90760111AD26E29" type="email">pasi.sihvonen@helsinki.fi</mods:nameIdentifier>
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<mods:date id="16DD132C3B07210651ABCB19AAB6ABD2">2023</mods:date>
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<emphasis id="B957DA3054507824EAE2FE26FE1E5FE3" bold="true" box="[151,396,391,418]" italics="true" pageId="9" pageNumber="160">Coleophora carsteni</emphasis>
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Tabell
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,
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Barcode Index Number:
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<paragraph id="8B9C062254507824EAE2FE71FEA55FAB" blockId="9.[151,626,391,490]" box="[151,311,464,490]" pageId="9" pageNumber="160">Figs. 6, 21, 36</paragraph>
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♂ (GP
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J. Tabell):
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leg. http://id.luomus.fi/GBT.15 (coll.
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)
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(GP 5514 J. Tabell, DNA sample
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Lepid Phyl)
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,
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&
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leg.;
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|
||
<materialsCitation id="3B4B0C7F54507824EB98FD25FB7D5C83" collectingDate="2017-03-08" collectingDateMax="2017-03-10" collectingDateMin="2017-03-08" collectionCode="TAB" collectorName="C. Hviid & O. Karsholt & K. Larsen & D. Nilsson" country="Morocco" county="Lepid Phyl" elevation="1580" location="Tafraoute" municipality="Souss-Massa-Draa" pageId="9" pageNumber="160" specimenCode="27507" specimenCount="1" specimenCount-male="1" stateProvince="Anti-Atlas" typeStatus="paratype">
|
||
<specimenCount id="9D25CDAB54507824EB98FD25FD8A5CE1" box="[493,536,644,672]" count="1" pageId="9" pageNumber="160" type="male">1 ♂</specimenCount>
|
||
(DNA sample
|
||
<specimenCode id="DB85AE5954507824E8B7FD25FC9A5CDE" box="[706,776,644,671]" pageId="9" pageNumber="160">27507</specimenCode>
|
||
Lepid Phyl)
|
||
<collectingRegion id="49E7C8C054507824E9EFFD25FB875CDF" box="[922,1045,644,670]" pageId="9" pageNumber="160">Anti-Atlas</collectingRegion>
|
||
:
|
||
<collectingMunicipality id="6BF89C5854507824EE55FD25FB675CDF" box="[1056,1269,644,670]" pageId="9" pageNumber="160">Souss-Massa-Drâa</collectingMunicipality>
|
||
, Idikl
|
||
<locationDeviation id="9A2E1F0054507824EF49FD25FA0E5CDE" box="[1340,1436,644,671]" pageId="9" pageNumber="160">
|
||
<quantity id="4CDBABC754507824EF49FD25FA165CDF" box="[1340,1412,644,670]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="9" pageNumber="160" unit="km" value="16.0">16 km</quantity>
|
||
E
|
||
</locationDeviation>
|
||
<location id="8EFC50F954507824EAE2FD09FE975C83" LSID="urn:lsid:plazi:treatment:038AB73454507824EAE2FE26FE0C58F2:8EFC50F954507824EAE2FD09FE975C83" box="[151,261,680,706]" country="Morocco" county="Lepid Phyl" municipality="Souss-Massa-Draa" name="Tafraoute" pageId="9" pageNumber="160" stateProvince="Anti-Atlas">Tafraoute</location>
|
||
,
|
||
<quantity id="4CDBABC754507824EB65FD09FEF55C82" box="[272,359,680,707]" metricMagnitude="3" metricUnit="m" metricValue="1.58" pageId="9" pageNumber="160" unit="m" value="1580.0">
|
||
<elevation id="000EE11154507824EB65FD09FEF55C82" box="[272,359,680,707]" metricMagnitude="3" metricUnit="m" metricValue="1.58" pageId="9" pageNumber="160" unit="m" value="1580.0">1580 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FF9D20E254507824EB06FD09FD845C83" box="[371,534,680,706]" pageId="9" pageNumber="160" value="2017-03-08" valueMax="2017-03-10" valueMin="2017-03-08">
|
||
<collectingDate id="EFD9D90A54507824EB06FD09FD845C83" box="[371,534,680,706]" pageId="9" pageNumber="160" value="2017-03-08" valueMax="2017-03-10" valueMin="2017-03-08">8.–10.III.2017</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26D663F454507824E857FD09FD175C83" box="[546,645,680,706]" pageId="9" pageNumber="160">C. Hviid</collectorName>
|
||
,
|
||
<collectorName id="26D663F454507824E8E5FD09FC875C83" box="[656,789,680,706]" pageId="9" pageNumber="160">O. Karsholt</collectorName>
|
||
,
|
||
<collectorName id="26D663F454507824E954FD08FC005C82" box="[801,914,681,707]" pageId="9" pageNumber="160">K. Larsen</collectorName>
|
||
,
|
||
<collectorName id="26D663F454507824E9E8FD08FB845C83" box="[925,1046,680,707]" pageId="9" pageNumber="160">D. Nilsson</collectorName>
|
||
leg. (all coll.
|
||
<collectionCode id="ED329EE754507824EEC5FD09FB7A5C83" box="[1200,1256,680,706]" pageId="9" pageNumber="160">TAB</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33955A954507824EAB2FD6DFAC75D37" pageId="9" pageNumber="160" type="diagnosis">
|
||
<paragraph id="8B9C062254507824EAB2FD6DFAC75D37" blockId="9.[151,1437,536,1715]" pageId="9" pageNumber="160">
|
||
Diagnosis. Externally
|
||
<taxonomicName id="4C237DA154507824EBB5FD6DFDA85CA7" authorityName="Tabell & Kullberg & Mutanen & Tokár & Sihvonen" authorityYear="2023" box="[448,570,716,742]" class="Magnoliopsida" family="Thymelaeaceae" genus="Coleophora" kingdom="Plantae" order="Malvales" pageId="9" pageNumber="160" phylum="Tracheophyta" rank="species" species="carsteni">
|
||
<emphasis id="B957DA3054507824EBB5FD6DFDA85CA7" box="[448,570,716,742]" italics="true" pageId="9" pageNumber="160">C. carsteni</emphasis>
|
||
</taxonomicName>
|
||
is similar to several members of the
|
||
<taxonomicName id="4C237DA154507824E9A3FD6DFB005CA7" authorityName="Zeller" authorityYear="1849" box="[982,1170,716,742]" class="Magnoliopsida" family="Thymelaeaceae" genus="Coleophora" kingdom="Plantae" order="Malvales" pageId="9" pageNumber="160" phylum="Tracheophyta" rank="species" species="onobrychiella">
|
||
<emphasis id="B957DA3054507824E9A3FD6DFB005CA7" box="[982,1170,716,742]" italics="true" pageId="9" pageNumber="160">C. onobrychiella</emphasis>
|
||
</taxonomicName>
|
||
species group. By male genitalia, it resembles
|
||
<emphasis id="B957DA3054507824EBEFFD51FDA45D4B" box="[410,566,752,778]" italics="true" pageId="9" pageNumber="160">C. sisteronica</emphasis>
|
||
Toll, 1961 and
|
||
<emphasis id="B957DA3054507824E89BFD51FCEB5D4B" box="[750,889,752,778]" italics="true" pageId="9" pageNumber="160">C. hipponae</emphasis>
|
||
Baldizzone, 1993. In the male genitalia, longer sacculus and its shorter apex and unevenly curved ventral margin distinguish
|
||
<taxonomicName id="4C237DA154507824EE7DFCB5FB165D6F" authorityName="Tabell & Kullberg & Mutanen & Tokár & Sihvonen" authorityYear="2023" box="[1032,1156,788,814]" class="Magnoliopsida" family="Thymelaeaceae" genus="Coleophora" kingdom="Plantae" order="Malvales" pageId="9" pageNumber="160" phylum="Tracheophyta" rank="species" species="carsteni">
|
||
<emphasis id="B957DA3054507824EE7DFCB5FB165D6F" box="[1032,1156,788,814]" italics="true" pageId="9" pageNumber="160">C. carsteni</emphasis>
|
||
</taxonomicName>
|
||
from these species. Furthermore, the cucullus is shorter in
|
||
<emphasis id="B957DA3054507824E86BFC99FD345D13" box="[542,678,824,850]" italics="true" pageId="9" pageNumber="160">C. hipponae</emphasis>
|
||
and the abdominal structures different (transverse bar parallel-sided in
|
||
<emphasis id="B957DA3054507824EAC1FCFDFEAF5D37" box="[180,317,860,886]" italics="true" pageId="9" pageNumber="160">C. hipponae</emphasis>
|
||
, medially expanded in
|
||
<taxonomicName id="4C237DA154507824E836FCFDFD2C5D37" authorityName="Tabell & Kullberg & Mutanen & Tokár & Sihvonen" authorityYear="2023" box="[579,702,860,886]" class="Magnoliopsida" family="Thymelaeaceae" genus="Coleophora" kingdom="Plantae" order="Malvales" pageId="9" pageNumber="160" phylum="Tracheophyta" rank="species" species="carsteni">
|
||
<emphasis id="B957DA3054507824E836FCFDFD2C5D37" box="[579,702,860,886]" italics="true" pageId="9" pageNumber="160">C. carsteni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B957DA3054507824E881FCFDFC1D5D37" box="[756,911,860,886]" italics="true" pageId="9" pageNumber="160">C. sisteronica</emphasis>
|
||
). The female of
|
||
<taxonomicName id="4C237DA154507824EE3FFCFDFB565D37" authorityName="Tabell & Kullberg & Mutanen & Tokár & Sihvonen" authorityYear="2023" box="[1098,1220,860,886]" class="Magnoliopsida" family="Thymelaeaceae" genus="Coleophora" kingdom="Plantae" order="Malvales" pageId="9" pageNumber="160" phylum="Tracheophyta" rank="species" species="carsteni">
|
||
<emphasis id="B957DA3054507824EE3FFCFDFB565D37" box="[1098,1220,860,886]" italics="true" pageId="9" pageNumber="160">C. carsteni</emphasis>
|
||
</taxonomicName>
|
||
is unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33955A954507824EAB2FC21FD975863" pageId="9" pageNumber="160" type="description">
|
||
<paragraph id="8B9C062254507824EAB2FC21FDAD5DA3" blockId="9.[151,1437,536,1715]" pageId="9" pageNumber="160">
|
||
Molecular data.
|
||
<specimenCount id="9D25CDAB54507824EB0AFC20FDBC5DDB" box="[383,558,896,923]" count="2" pageId="9" pageNumber="160" type="generic">Two specimens</specimenCount>
|
||
of
|
||
<taxonomicName id="4C237DA154507824E820FC21FD425DDB" authorityName="Tabell & Kullberg & Mutanen & Tokár & Sihvonen" authorityYear="2023" box="[597,720,896,922]" class="Magnoliopsida" family="Thymelaeaceae" genus="Coleophora" kingdom="Plantae" order="Malvales" pageId="9" pageNumber="160" phylum="Tracheophyta" rank="species" species="carsteni">
|
||
<emphasis id="B957DA3054507824E820FC21FD425DDB" box="[597,720,896,922]" italics="true" pageId="9" pageNumber="160">C. carsteni</emphasis>
|
||
</taxonomicName>
|
||
were sequenced, resulting in 633 bp and 674 bp barcodes. The nearest neighbour to
|
||
<taxonomicName id="4C237DA154507824EBF0FC05FD935DFF" authorityName="Tabell & Kullberg & Mutanen & Tokár & Sihvonen" authorityYear="2023" box="[389,513,932,958]" class="Magnoliopsida" family="Thymelaeaceae" genus="Coleophora" kingdom="Plantae" order="Malvales" pageId="9" pageNumber="160" phylum="Tracheophyta" rank="species" species="carsteni">
|
||
<emphasis id="B957DA3054507824EBF0FC05FD935DFF" box="[389,513,932,958]" italics="true" pageId="9" pageNumber="160">C. carsteni</emphasis>
|
||
</taxonomicName>
|
||
is
|
||
<emphasis id="B957DA3054507824E856FC05FD2D5DFF" box="[547,703,932,958]" italics="true" pageId="9" pageNumber="160">C. sisteronica</emphasis>
|
||
, with a 7.6 % minimum divergence. The barcodes of
|
||
<taxonomicName id="4C237DA154507824EF54FC05FA0E5DFF" authorityName="Tabell & Kullberg & Mutanen & Tokár & Sihvonen" authorityYear="2023" box="[1313,1436,932,958]" class="Magnoliopsida" family="Thymelaeaceae" genus="Coleophora" kingdom="Plantae" order="Malvales" pageId="9" pageNumber="160" phylum="Tracheophyta" rank="species" species="carsteni">
|
||
<emphasis id="B957DA3054507824EF54FC05FA0E5DFF" box="[1313,1436,932,958]" italics="true" pageId="9" pageNumber="160">C. carsteni</emphasis>
|
||
</taxonomicName>
|
||
exhibit 0.46 % intraspecific variation.
|
||
</paragraph>
|
||
<paragraph id="8B9C062254507824EAB2FC4CFD745A46" blockId="9.[151,1437,536,1715]" box="[199,742,1004,1031]" pageId="9" pageNumber="160">
|
||
In BOLD there are no barcodes for
|
||
<emphasis id="B957DA3054507824E823FC4DFD4D5A47" box="[598,735,1004,1030]" italics="true" pageId="9" pageNumber="160">C. hipponae</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9C062254507824EAB2FBB1FE0F5A9F" blockId="9.[151,1437,536,1715]" pageId="9" pageNumber="160">
|
||
Description. Adult. Wingspan
|
||
<quantity id="4CDBABC754507824E86EFBB1FD3E5A6A" box="[539,684,1040,1067]" metricMagnitude="-2" metricUnit="m" metricValue="1.375" metricValueMax="1.4" metricValueMin="1.35" pageId="9" pageNumber="160" unit="mm" value="13.75" valueMax="14.0" valueMin="13.5">13.5–14 mm</quantity>
|
||
. Antenna white, indistinctly annulated with pale grey. Scape white, below ochre and tufted. Labial palp pale ochre mixed with white, second article 1.5x longer than third article. Head and thorax white with a pale ochre median line, tegula white mixed with pale ochre. Forewing ochreous brown, dorsal half slightly paler; costal stripe white, from base to ¾, slightly expanded apically; dorsal stripe distinct only at base. Costal fringe cilia ochreous brown, apically darker, dorsal fringe cilia ochreous brown. Hindwing pale grey, fringe cilia pale brown.
|
||
</paragraph>
|
||
<paragraph id="8B9C062254507824EAB2FB49FD485B0B" blockId="9.[151,1437,536,1715]" pageId="9" pageNumber="160">Abdominal structures. No tergal latero-posterior bar. Transverse bar broad, curved, proximal edge medially straight and sclerotized, distal edge strongly arched, sclerotized. Tergal sclerite about 4x as long as wide, weakly sclerotized, covered with 20 conical spines (on T3).</paragraph>
|
||
<paragraph id="8B9C062254507824EAB2FAF5FEAC5BBF" blockId="9.[151,1437,536,1715]" pageId="9" pageNumber="160">Male genitalia. Gnathos knob round. Tegumen constricted medially, pedunculum short, arched. Transtilla short, triangular. Valvula large, weakly delineated. Cucullus long, very narrow, club-shaped. Sacculus well sclerotized, elongated; ventral margin slightly sinuous, convex, apical half with long bristles, ending in sharp point; dorsal margin slightly concave. Phallotheca a conical tube. Vesica long, with several spiniform cornuti grouped into a tight curved bundle.</paragraph>
|
||
<paragraph id="8B9C062254507824EAB2F9A9FD975863" blockId="9.[151,1437,536,1715]" box="[199,517,1544,1570]" pageId="9" pageNumber="160">Female genitalia. Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33955A954507824EAB2F98DFDBF5807" box="[199,557,1580,1607]" pageId="9" pageNumber="160" type="biology_ecology">
|
||
<paragraph id="8B9C062254507824EAB2F98DFDBF5807" blockId="9.[151,1437,536,1715]" box="[199,557,1580,1607]" pageId="9" pageNumber="160">Biology. Early stages unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33955A954507824EAB2F9F1FB52582B" box="[199,1216,1616,1643]" pageId="9" pageNumber="160" type="distribution">
|
||
<paragraph id="8B9C062254507824EAB2F9F1FB52582B" blockId="9.[151,1437,536,1715]" box="[199,1216,1616,1643]" pageId="9" pageNumber="160">
|
||
Distribution.
|
||
<collectingCountry id="F33446B254507824EB28F9F0FE57582A" box="[349,453,1617,1643]" name="Morocco" pageId="9" pageNumber="160">Morocco</collectingCountry>
|
||
, known from a few localities in the High and Anti-Atlas Mountains.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33955A954507824EAB2F9D5FE0C58F2" pageId="9" pageNumber="160" type="etymology">
|
||
<paragraph id="8B9C062254507824EAB2F9D5FE0C58F2" blockId="9.[151,1437,536,1715]" pageId="9" pageNumber="160">
|
||
Etymology. The species is named in honour of Carsten Hviid, who was a participant of the Danish collecting expedition to
|
||
<collectingCountry id="F33446B254507824EB44F938FE0858F2" box="[305,410,1689,1715]" name="Morocco" pageId="9" pageNumber="160">Morocco</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |