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<document id="3FAAFB0371E27FDADC380871F188F89C" ID-CLB-Dataset="4362" ID-DOI="10.5852/ejt.2021.750.1369" ID-GBIF-Dataset="cd2883dd-811d-4617-a315-68c78d7e4490" ID-ISSN="2118-9773" ID-Zenodo-Dep="5451701" ID-ZooBank="FFDAD9F7-5E6A-40DA-9D2F-1713CE7AFF4A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630516139641" checkinUser="felipe" docAuthor="Delicado, Diana, Pešić, Vladimir &amp; Ramos, Marian A." docDate="2021" docId="038A87BDFF82677BFDCA82B0FC7DFDE1" docLanguage="en" docName="ejt-750_140-155.pdf" docOrigin="European Journal of Taxonomy 750 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.750.1369" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Docleiana Delicado &amp; Pesic 2021, gen. nov." docType="treatment" docUuid="04639DB3-91D2-46B7-BFAB-FFC1EA3C27BD" docUuidSource="ZooBank" docVersion="8" lastPageNumber="147" masterDocId="FFB3FFC5FF84677CFFC58662FFAFFFA9" masterDocTitle="Arganiella Giusti &amp; Pezzoli, 1980 (Caenogastropoda: Truncatelloidea: Hydrobiidae): a widespread genus or several narrow-range endemic genera?" masterLastPageNumber="155" masterPageNumber="140" pageNumber="146" updateTime="1699290064226" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8AB3D1B57996A77995FE5CC3EC761551">Arganiella Giusti &amp; Pezzoli, 1980 (Caenogastropoda: Truncatelloidea: Hydrobiidae): a widespread genus or several narrow-range endemic genera?</mods:title>
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<mods:namePart id="D55724D431EF0F47A3641C70D2175824">Delicado, Diana</mods:namePart>
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<mods:affiliation id="D99AD30C253705DACB72A7A6304E40C8">Justus Liebig University Giessen, Department of Animal Ecology &amp; Systematics, Heinrich-Buff-Ring 26 - 32 IFZD, 35392 Giessen, Germany.</mods:affiliation>
<mods:nameIdentifier id="F0D9743D0C43BF866B2AF33E95F4DF91" type="email">didelicado@gmail.com</mods:nameIdentifier>
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<mods:namePart id="4C3DE6CDF4CC04BF1C02E903EA8C8C8D">Pešić, Vladimir</mods:namePart>
<mods:nameIdentifier id="350A28BE75C8B7B3B4A6194C563A363F" type="LSID">2B46F869-1202-47A9-9359-0BE04D691F6C</mods:nameIdentifier>
<mods:affiliation id="AD42E2D09DA49837FBF6C59F1BF887B9">University of Montenegro, Faculty of Sciences, Department of Biology, Cetinjski put b. b., 81000 Podgorica, Montenegro.</mods:affiliation>
<mods:nameIdentifier id="8F91E96D3598E60FA586862080CAC546" type="email">vladopesic@gmail.com</mods:nameIdentifier>
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<mods:namePart id="EE9114BBC59CCDED4AE5B375FD804DAA">Ramos, Marian A.</mods:namePart>
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<mods:affiliation id="3EA6D74912E346D7E2522A9E9DAFC2E5">Museo Nacional de Ciencias Naturales (MNCN-CSIC), José Gutiérrez Abascal 2, 28006 Madrid, Spain.</mods:affiliation>
<mods:nameIdentifier id="D81446D0271E351499D0D51C969DF4FE" type="email">m.ramos@mncn.csic.es</mods:nameIdentifier>
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<mods:title id="EFDA077729DEA33CC271A31BE3E25595">European Journal of Taxonomy</mods:title>
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<mods:date id="91F309AAADA6B917FA2F57446D695755">2021</mods:date>
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<mods:number id="CC5F6840EB2D4C6F49CDB29ADDBA351C">2021-05-21</mods:number>
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<treatment id="038A87BDFF82677BFDCA82B0FC7DFDE1" ID-DOI="http://doi.org/10.5281/zenodo.5493136" ID-GBIF-Taxon="185454791" ID-Zenodo-Dep="5493136" LSID="urn:lsid:zoobank.org:act:04639DB3-91D2-46B7-BFAB-FFC1EA3C27BD" httpUri="http://treatment.plazi.org/id/038A87BDFF82677BFDCA82B0FC7DFDE1" lastPageId="7" lastPageNumber="147" pageId="6" pageNumber="146">
<subSubSection id="C3396520FF82677AFDCA82B0FB8BFB44" box="[527,1060,1233,1261]" pageId="6" pageNumber="146" type="nomenclature">
<paragraph id="8B9C36ABFF82677AFDCA82B0FB8BFB44" blockId="6.[356,1231,1233,1303]" box="[527,1060,1233,1261]" pageId="6" pageNumber="146">
<heading id="D0D481C7FF82677AFDCA82B0FB8BFB44" box="[527,1060,1233,1261]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="146" reason="2">
Genus
<taxonomicName id="4C234D28FF82677AFDA782B3FC1DFB45" ID-CoL="8H9TG" authority="Delicado &amp; Pesic" authorityName="Delicado &amp; Pesic" authorityYear="2021" box="[610,946,1233,1260]" class="Gastropoda" family="Hydrobiidae" genus="Docleiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="146" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B957EAB9FF82677AFDA782B3FD71FB45" bold="true" box="[610,734,1233,1260]" italics="true" pageId="6" pageNumber="146">Docleiana</emphasis>
Delicado &amp; Pešić
</taxonomicName>
<taxonomicNameLabel id="A26457C2FF82677AFC7F82B1FB8BFB44" box="[954,1060,1235,1261]" pageId="6" pageNumber="146" rank="genus">gen. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3396520FF82677AFEA1829FFB60FABE" box="[356,1231,1277,1303]" pageId="6" pageNumber="146" type="description">
<paragraph id="8B9C36ABFF82677AFEA1829FFB60FABE" blockId="6.[356,1231,1233,1303]" box="[356,1231,1277,1303]" pageId="6" pageNumber="146">
<uri id="FFB23AA9FF82677AFEA1829FFB60FABE" box="[356,1231,1277,1303]" pageId="6" pageNumber="146">
urn:lsid:zoobank.org:act:
<uuid id="FF850C7EFF82677AFD40829FFB60FABE" box="[645,1231,1277,1303]" pageId="6" pageNumber="146">04639DB3-91D2-46B7-BFAB-FFC1EA3C27BD</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3396520FF82677AFF78835CFC8AFA2E" pageId="6" pageNumber="146" type="type_taxon">
<paragraph id="8B9C36ABFF82677AFF78835CFEF7FAF1" blockId="6.[189,344,1342,1368]" box="[189,344,1342,1368]" pageId="6" pageNumber="146">
<heading id="D0D481C7FF82677AFF78835CFEF7FAF1" bold="true" box="[189,344,1342,1368]" fontSize="11" level="3" pageId="6" pageNumber="146" reason="3">
<emphasis id="B957EAB9FF82677AFF78835CFEF7FAF1" bold="true" box="[189,344,1342,1368]" pageId="6" pageNumber="146">
<typeStatus id="54988809FF82677AFF78835CFF55FAF1" box="[189,250,1342,1368]" pageId="6" pageNumber="146">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C36ABFF82677AFF78830FFC8AFA2E" blockId="6.[189,805,1389,1415]" box="[189,805,1389,1415]" pageId="6" pageNumber="146">
<taxonomicName id="4C234D28FF82677AFF78830FFC8EFA2E" ID-CoL="87LGR" authority="Boeters, Gloer &amp; Pesic, 2014" authorityName="Boeters, Gloer &amp; Pesic" authorityYear="2014" box="[189,801,1389,1415]" class="Gastropoda" family="Hydrobiidae" genus="Arganiella" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="146" phylum="Mollusca" rank="species" species="tabanensis">
<emphasis id="B957EAB9FF82677AFF78830FFE11FA2E" box="[189,446,1389,1415]" italics="true" pageId="6" pageNumber="146">Arganiella tabanensis</emphasis>
<bibRefCitation id="EFB24B5AFF82677AFE03830FFC8EFA2E" author="Boeters H. D. &amp; Gloer P. &amp; Pesic V." box="[454,801,1389,1415]" pageId="6" pageNumber="146" pagination="131 - 139" refId="ref7583" refString="Boeters H. D., Gloer P. &amp; Pesic V. 2014. Arganiella tabanensis n. sp. from Montenegro (Mollusca: Gastropoda: Hydrobiidae). Ecologica Montenegrina 1 (3): 131 - 139. https: // doi. org / 10.37828 / em. 2014.1.20" type="journal article" year="2014">Boeters, Glöer &amp; Pešić, 2014</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3396520FF82677AFF7883CCFE57F8B9" pageId="6" pageNumber="146" type="diagnosis">
<paragraph id="8B9C36ABFF82677AFF7883CCFE35FA61" blockId="6.[189,410,1454,1480]" box="[189,410,1454,1480]" pageId="6" pageNumber="146">
<heading id="D0D481C7FF82677AFF7883CCFE35FA61" bold="true" box="[189,410,1454,1480]" fontSize="11" level="3" pageId="6" pageNumber="146" reason="3">
<emphasis id="B957EAB9FF82677AFF7883CCFE35FA61" bold="true" box="[189,410,1454,1480]" pageId="6" pageNumber="146">Revised diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C36ABFF82677AFF7883BFFE57F8B9" blockId="6.[188,1399,1501,1808]" pageId="6" pageNumber="146">Shell valvatiform; whorls ca 3; aperture complete, from rounded to ellipsoidal; outer lip narrow, straight in lateral profile; umbilicus wide. Operculum corneous, yellowish, thin, pliable, rounded, paucispiral with a central nucleus, without peg. Two pairs of basal cusps on each central radular tooth. Ctenidium well developed, with 10–11 gill filaments. Animal unpigmented. Osphradium positioned opposite approximately to the middle of the ctenidium. Stomach without gastric caecum; rectum forms a gentle V-shape in the mantle cavity. Bursa copulatrix small, globular, pedunculated and positioned beyond the posterior edge of the albumen gland; bursal duct longer than bursal length; unpigmented renal oviduct; one pyriform seminal receptacle arising at the insertion point with the bursal duct loop. Penis small and simple, gradually tapering.</paragraph>
</subSubSection>
<subSubSection id="C3396520FF82677AFF78815AFBD0F80A" pageId="6" pageNumber="146" type="etymology">
<paragraph id="8B9C36ABFF82677AFF78815AFEEBF8FB" blockId="6.[189,324,1848,1874]" box="[189,324,1848,1874]" pageId="6" pageNumber="146">
<heading id="D0D481C7FF82677AFF78815AFEEBF8FB" bold="true" box="[189,324,1848,1874]" fontSize="11" level="3" pageId="6" pageNumber="146" reason="3">
<emphasis id="B957EAB9FF82677AFF78815AFEEBF8FB" bold="true" box="[189,324,1848,1874]" pageId="6" pageNumber="146">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C36ABFF82677AFF788104FBD0F80A" blockId="6.[189,1399,1894,1956]" pageId="6" pageNumber="146">The new genus is named after Doclea, the name of the Roman city located on the site of modern Podgorica on whose municipal territory a new genus was found; gender feminine.</paragraph>
</subSubSection>
<subSubSection id="C3396520FF83677BFF78876BFC7DFDE1" pageId="7" pageNumber="147" type="discussion">
<paragraph id="8B9C36ABFF83677BFF78876BFE80FE8A" blockId="7.[189,303,265,291]" box="[189,303,265,291]" pageId="7" pageNumber="147">
<heading id="D0D481C7FF83677BFF78876BFE80FE8A" bold="true" box="[189,303,265,291]" fontSize="11" level="3" pageId="7" pageNumber="147" reason="3">
<emphasis id="B957EAB9FF83677BFF78876BFE80FE8A" bold="true" box="[189,303,265,291]" pageId="7" pageNumber="147">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C36ABFF83677BFF788755FC7DFDE1" blockId="7.[189,1400,311,584]" pageId="7" pageNumber="147">
<taxonomicName id="4C234D28FF83677BFF788755FDBFFEF8" authority="Delicado &amp; Pesic" authorityName="Delicado &amp; Pesic" authorityYear="2021" box="[189,528,311,337]" class="Gastropoda" family="Hydrobiidae" genus="Docleiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="147" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B957EAB9FF83677BFF788755FE98FEF8" box="[189,311,311,337]" italics="true" pageId="7" pageNumber="147">Docleiana</emphasis>
Delicado &amp; Pešić
</taxonomicName>
<taxonomicNameLabel id="A26457C2FF83677BFDDD875AFD2AFEFB" box="[536,645,312,338]" pageId="7" pageNumber="147" rank="genus">gen. nov.</taxonomicNameLabel>
can be distinguished from
<taxonomicName id="4C234D28FF83677BFC0F8755FBE9FEF8" authorityName="Giusti &amp; Pezzoli" authorityYear="1980" box="[970,1094,311,337]" class="Gastropoda" family="Hydrobiidae" genus="Arganiella" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="147" phylum="Mollusca" rank="genus">
<emphasis id="B957EAB9FF83677BFC0F8755FBE9FEF8" box="[970,1094,311,337]" italics="true" pageId="7" pageNumber="147">Arganiella</emphasis>
</taxonomicName>
as the former has a more ellipsoidal shell aperture, a narrower base of the penis, a smaller bursa copulatrix that is positioned beyond the posterior edge of the albumen gland, a larger seminal receptacle and more cusps on the lateral radular teeth (
<figureCitation id="13182A2EFF83677BFE7187C3FE52FE12" box="[436,509,417,443]" captionStart="Fig" captionStartId="7.[189,232,1924,1950]" captionTargetBox="[189,1398,826,1886]" captionTargetId="figure-187@7.[189,1398,826,1886]" captionTargetPageId="7" captionText="Fig. 2. Shells, operculum and radulae of Docleiana tabanensis (Boeters, Glöer &amp; Pešić, 2014) (UGSB 18847). A–C. Shells. D–E. Operculum (D = inner side; E = outer side). F. Protoconch. G–I. Radulae. G. Portion of radula ribbon. H. Central radular teeth. I. Inner and outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.5451705" httpUri="https://zenodo.org/record/5451705/files/figure.png" pageId="7" pageNumber="147">Fig. 2</figureCitation>
; for comparison, see
<bibRefCitation id="EFB24B5AFF83677BFD3987C3FBA6FE12" author="Giusti F. &amp; Pezzoli E." box="[764,1033,417,443]" pageId="7" pageNumber="147" pagination="207 - 222" refId="ref8377" refString="Giusti F. &amp; Pezzoli E. 1980. Hydrobioidea nuove o poco conosciute dell Italia appenninica (Gastropoda: Prosobranchia). Archiv fur Molluskenkunde 111: 207 - 222." type="journal article" year="1980">Giusti &amp; Pezzoli 1980</bibRefCitation>
;
<bibRefCitation id="EFB24B5AFF83677BFBD387C3FB53FE12" author="Boeters H. D. &amp; Gloer P. &amp; Pesic V." box="[1046,1276,417,443]" pageId="7" pageNumber="147" pagination="131 - 139" refId="ref7583" refString="Boeters H. D., Gloer P. &amp; Pesic V. 2014. Arganiella tabanensis n. sp. from Montenegro (Mollusca: Gastropoda: Hydrobiidae). Ecologica Montenegrina 1 (3): 131 - 139. https: // doi. org / 10.37828 / em. 2014.1.20" type="journal article" year="2014">
Boeters
<emphasis id="B957EAB9FF83677BFBBC87C3FB1AFE12" box="[1145,1205,417,443]" italics="true" pageId="7" pageNumber="147">et al.</emphasis>
2014
</bibRefCitation>
). The new genus differs from the closely related genus
<taxonomicName id="4C234D28FF83677BFD1487A6FC8EFE77" authorityName="Radoman" authorityYear="1978" box="[721,801,452,478]" class="Gastropoda" family="Hydrobiidae" genus="Kerkia" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="147" phylum="Mollusca" rank="genus">
<emphasis id="B957EAB9FF83677BFD1487A6FC8EFE77" box="[721,801,452,478]" italics="true" pageId="7" pageNumber="147">Kerkia</emphasis>
</taxonomicName>
by its smaller shell, absence of lobes on the inner edge of the penis, a smaller bursa copulatrix, a more globular seminal receptacle and V-shaped rectum (see
<bibRefCitation id="EFB24B5AFF83677BFF328468FE00FD8D" author="Radoman P." box="[247,431,522,548]" pageId="7" pageNumber="147" pagination="27 - 44" refId="ref9055" refString="Radoman P. 1978. Neue Vertreter der Gruppe Hydrobioidea von der Balkanhalbinsel. Archiv fur Molluskenkunde 109 (1 - 3): 27 - 44." type="journal article" year="1978">Radoman 1978</bibRefCitation>
;
<bibRefCitation id="EFB24B5AFF83677BFE7B8468FD33FD8D" author="Bodon M. &amp; Manganelli G. &amp; Giusti F." box="[446,668,522,548]" pageId="7" pageNumber="147" pagination="103 - 215" refId="ref7501" refString="Bodon M., Manganelli G. &amp; Giusti F. 2001. A survey of the European valvatiform hydrobiid genera with special reference to Hauffenia Pollonera, 1898 (Gastropoda: Hydrobiidae). Malacologia 43: 103 - 215." type="journal article" year="2001">
Bodon
<emphasis id="B957EAB9FF83677BFDD38469FDFCFD8D" box="[534,595,522,548]" italics="true" pageId="7" pageNumber="147">et al.</emphasis>
2001
</bibRefCitation>
). Mean COI sequence divergence for
<taxonomicName id="4C234D28FF83677BFBAD8468FF54FDEE" authority="Delicado &amp; Pesic" authorityName="Delicado &amp; Pesic" authorityYear="2021" class="Gastropoda" family="Hydrobiidae" genus="Docleiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="147" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B957EAB9FF83677BFBAD8468FB4DFD8D" box="[1128,1250,522,548]" italics="true" pageId="7" pageNumber="147">Docleiana</emphasis>
Delicado &amp; Pešić
</taxonomicName>
<taxonomicNameLabel id="A26457C2FF83677BFEC7844CFEC2FDE1" box="[258,365,558,584]" pageId="7" pageNumber="147" rank="genus">gen. nov.</taxonomicNameLabel>
was 14.5% with
<taxonomicName id="4C234D28FF83677BFDFD844FFD1BFDEE" authorityName="Giusti &amp; Pezzoli" authorityYear="1980" box="[568,692,557,583]" class="Gastropoda" family="Hydrobiidae" genus="Arganiella" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="147" phylum="Mollusca" rank="genus">
<emphasis id="B957EAB9FF83677BFDFD844FFD1BFDEE" box="[568,692,557,583]" italics="true" pageId="7" pageNumber="147">Arganiella</emphasis>
</taxonomicName>
and 12.3% with
<taxonomicName id="4C234D28FF83677BFCB9844FFC63FDEE" authorityName="Radoman" authorityYear="1978" box="[892,972,557,583]" class="Gastropoda" family="Hydrobiidae" genus="Kerkia" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="147" phylum="Mollusca" rank="genus">
<emphasis id="B957EAB9FF83677BFCB9844FFC63FDEE" box="[892,972,557,583]" italics="true" pageId="7" pageNumber="147">Kerkia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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