455 lines
60 KiB
XML
455 lines
60 KiB
XML
<document id="1D45166EB665AF3BB1795925A81665D0" ID-CLB-Dataset="2631" ID-DOI="10.11646/zootaxa.5091.1.5" ID-GBIF-Dataset="b8e67802-3d14-4b63-85c6-6ee00ebc55d4" ID-ISSN="1175-5326" ID-Zenodo-Dep="5840635" ID-ZooBank="7E744DAE-E9E6-4FB5-9897-1CCF03D373BC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1641987074997" checkinUser="plazi" docAuthor="Xu, Xin, Yu, Li, Liu, Fengxiang & Li, Daiqin" docDate="2022" docId="038A8789FFA6FFC216FE5D2DC547FEC6" docLanguage="en" docName="zootaxa.5091.1.5.pdf" docOrigin="Zootaxa 5091 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Luthela yiyuan Xu & Yu & Liu & Li 2022, sp. nov." docType="treatment" docVersion="9" lastPageNumber="137" masterDocId="FFB3FFF1FFA2FFC416695C67C467FFA3" masterDocTitle="Delimitation of the segmented trapdoor spider genus Luthela gen. nov., with comments on the genus Sinothela from northern China (Araneae, Mesothelae, Liphistiidae)" masterLastPageNumber="154" masterPageNumber="131" pageNumber="135" updateTime="1699301488680" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="855E9B383A1EB371C53E81358FB26659">Delimitation of the segmented trapdoor spider genus Luthela gen. nov., with comments on the genus Sinothela from northern China (Araneae, Mesothelae, Liphistiidae)</mods:title>
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<mods:namePart id="F9F2FF2C3B4A1509B5201F808260B4F2">Xu, Xin</mods:namePart>
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<mods:affiliation id="1AFC64235D7D554D809FCC2505F5E650">College of Life Sciences, Hunan Normal University, Changsha, Hunan, China & School of Life and Environmental Sciences, University of Sydney, Sydney, NSW, Australia & State Key Laboratory of Biocatalysis and Enzyme Engineering & Centre for Behavioural Ecology and Evolution, School of Life Sciences, Hubei University, Wuhan, Hubei, China</mods:affiliation>
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<mods:roleTerm id="487AD366DD53632141FEE42D2E2D2191">Author</mods:roleTerm>
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<mods:namePart id="AC1B549BB2CA82E77E79470EDC933642">Yu, Li</mods:namePart>
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<mods:affiliation id="408BE78DED76D5DCF646C9BD17874563">College of Life Sciences, Hunan Normal University, Changsha, Hunan, China</mods:affiliation>
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<mods:namePart id="BB70F9BC15D15AA697B3445D19857C0C">Liu, Fengxiang</mods:namePart>
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<mods:affiliation id="DA367CAE76815292883247AE5ACBF588">State Key Laboratory of Biocatalysis and Enzyme Engineering & Centre for Behavioural Ecology and Evolution, School of Life Sciences, Hubei University, Wuhan, Hubei, China</mods:affiliation>
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<mods:namePart id="A8536D633E630565EEE1D791097304E0">Li, Daiqin</mods:namePart>
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<mods:affiliation id="43DA3E5E1EAB04C1D22307D15CCEFBD0">Department of Biological Sciences, National University of Singapore, Singapore</mods:affiliation>
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<mods:title id="83D113DA51CAAEC658B143A5B6C901D4">Zootaxa</mods:title>
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<treatment id="038A8789FFA6FFC216FE5D2DC547FEC6" ID-DOI="http://doi.org/10.5281/zenodo.5843599" ID-GBIF-Taxon="192044611" ID-Zenodo-Dep="5843599" LSID="urn:lsid:plazi:treatment:038A8789FFA6FFC216FE5D2DC547FEC6" httpUri="http://treatment.plazi.org/id/038A8789FFA6FFC216FE5D2DC547FEC6" lastPageId="6" lastPageNumber="137" pageId="4" pageNumber="135">
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<taxonomicName id="4C234D1CFFA6FFC016FE5D2DC529FEC6" authority="Xu & Yu & Liu & Li, 2022" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" box="[151,334,330,357]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="135" phylum="Arthropoda" rank="species" species="yiyuan" status="sp. nov.">
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<emphasis id="B957EA8DFFA6FFC016FE5D2DC529FEC6" bold="true" box="[151,334,330,357]" italics="true" pageId="4" pageNumber="135">Luthela yiyuan</emphasis>
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<taxonomicNameLabel id="A26457F6FFA6FFC0173F5D2CC5D4FEC6" box="[342,435,331,357]" pageId="4" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C3396514FFA6FFC016FE5D08C512FE2A" box="[151,373,367,393]" pageId="4" pageNumber="135" type="description">
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<paragraph id="8B9C369FFFA6FFC016FE5D08C512FE2A" blockId="4.[151,435,330,393]" box="[151,373,367,393]" pageId="4" pageNumber="135">
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(
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<figureCitation id="13182A1AFFA6FFC016F65D08C578FE2A" box="[159,287,367,393]" captionStart="FIGURE 3" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[170,1417,237,1695]" captionTargetId="figure-43@5.[170,1417,237,1695]" captionTargetPageId="5" captionText="FIGURE 3. Male genital anatomy of Luthela yiyuan sp. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view. A–G, XUX-2012-051 (holotype). H–K, XUX-2012-045. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840647" httpUri="https://zenodo.org/record/5840647/files/figure.png" pageId="4" pageNumber="135">Figs 3A–K</figureCitation>
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,
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<figureCitation id="13182A1AFFA6FFC017435D08C50AFE2A" box="[298,365,367,393]" captionStart="FIGURE 4" captionStartId="6.[151,250,1251,1275]" captionTargetBox="[164,1423,381,1228]" captionTargetId="figure-115@6.[164,1423,381,1228]" captionTargetPageId="6" captionText="FIGURE 4. Female genital anatomy of Luthela yiyuan sp. nov. A–E, K–O, vulva dorsal view. F–J, P–T, vulva ventral view. A, F, XUX-2012-034. B, G, XUX-2012-035. C, H, XUX-2012-049. D, I, XUX-2016-101. E, J, XUX-2016-107. K, P, XUX-2012- 042. L, Q, XUX-2012-040. M, R, XUX-2012-046. N, S, XUX-2016-103. O, T, XUX-2016-108. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840651" httpUri="https://zenodo.org/record/5840651/files/figure.png" pageId="4" pageNumber="135">4O–T</figureCitation>
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)
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</paragraph>
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<taxonomicName id="4C234D1CFFA6FFC016FE5DD4C653FE68" authority=": Xu et al., 2015 a: 139" authorityName="Xu" authorityPageNumber="139" authorityYear="2015" box="[151,564,435,459]" class="Arachnida" family="Liphistiidae" genus="Sinothela" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="135" phylum="Arthropoda" rank="species" species="sinensis">
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<emphasis id="B957EA8DFFA6FFC016FE5DD4C52AFE68" box="[151,333,435,459]" italics="true" pageId="4" pageNumber="135">Sinothela sinensis</emphasis>
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:
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<bibRefCitation id="EFB24B6EFFA6FFC017325DD3C653FE68" author="Xu, X. & Liu, F. & Chen, J. & Ono, H. & Li, D. & Kuntner, M." box="[347,564,435,459]" pageId="4" pageNumber="135" pagination="121 - 151" refId="ref14523" refString="Xu, X., Liu, F., Chen, J., Ono, H., Li, D. & Kuntner, M. (2015 a) A genus-level taxonomic review of primitively segmented spiders (Mesothelae, Liphistiidae). Zookeys, 488, 121 - 151. https: // doi. org / 10.3897 / zookeys. 488.8726" type="journal article" year="2015">
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Xu
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<emphasis id="B957EA8DFFA6FFC017E95DD3C5CAFE68" box="[384,429,435,459]" italics="true" pageId="4" pageNumber="135">et al</emphasis>
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., 2015a: 139
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</taxonomicName>
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; misidentification.
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<typeStatus id="5498883DFFA6FFC016FE5D9EC561FDB0" box="[151,262,505,531]" pageId="4" pageNumber="135" type="holotype">Holotype</typeStatus>
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.
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<location id="8EFC6044FFA6FFC0177D5D9EC529FDB0" LSID="urn:lsid:plazi:treatment:038A8789FFA6FFC216FE5D2DC547FEC6:8EFC6044FFA6FFC0177D5D9EC529FDB0" box="[276,334,505,531]" country="China" county="Yiyuan County" latitude="36.152" longLatPrecision="70" longitude="118.334" municipality="Zibo City" name="Male" pageId="4" pageNumber="135" stateProvince="Shandong">Male</location>
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(XUX-2012-051, matured on
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at CBEE),
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,
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,
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,
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,
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<collectingCountry id="F334760FFFA6FFC017E85E7AC5A0FD94" box="[385,455,541,567]" name="China" pageId="4" pageNumber="135">China</collectingCountry>
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,
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,
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,
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, collected on
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, by
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,
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<collectorName id="26D65349FFA6FFC0128A5E79C122FD94" box="[1251,1349,541,568]" pageId="4" pageNumber="135">Z.L. Liu</collectorName>
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, and
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<collectorName id="26D65349FFA6FFC013E85E79C4DBFDFF" pageId="4" pageNumber="135">X. Xu</collectorName>
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, CBEE.
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<typeStatus id="5498883DFFA6FFC016AE5E02C526FDDC" box="[199,321,613,639]" pageId="4" pageNumber="135" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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<specimenCount id="9D25FD16FFA6FFC017385E02C5A6FDDC" box="[337,449,613,639]" pageId="4" pageNumber="135" type="female">3 females</specimenCount>
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(XUX-2012-046/049/050) and
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<materialsCitation id="3B4B3CC2FFA6FFC015585E02C656FD00" ID-GBIF-Occurrence="3435942303" collectingDate="2012-07-29" collectingDateMax="2012-07-29" collectingDateMin="2012-05-01" collectorName="F. X. Liu & Z. L. Liu & X. Xu" country="China" county="Yiyuan County" elevation="275" latitude="36.152" location="Male" longLatPrecision="70" longitude="118.334" municipality="Zibo City" pageId="4" pageNumber="135" specimenCount="1" specimenCount-male="1" stateProvince="Shandong" typeStatus="paratype">
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(XUX-2012-045, matured on
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||
</date>
|
||
at CBEE), same data as the holotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B4B3CC2FFA6FFC014285EEEC5A6FD48" ID-GBIF-Occurrence="3435942305" collectingDate="2012-04-30" collectingDateMax="2012-07-10" collectingDateMin="2012-04-30" collectorName="Zhangqiu City" country="China" elevation="100" latitude="36.728" location="Puji Town" longLatPrecision="70" longitude="117.611" pageId="4" pageNumber="135" specimenCount="6" specimenCount-female="6" stateProvince="Shandong" typeStatus="paratype">
|
||
<specimenCount id="9D25FD16FFA6FFC014285EEEC6D6FD00" box="[577,689,649,675]" pageId="4" pageNumber="135" type="female">6 females</specimenCount>
|
||
(XUX-2012-034/035/036/037/040/042, XUX-2012-036 matured on
|
||
<date id="FF9D105FFFA6FFC016D55ECAC537FD64" box="[188,336,685,711]" pageId="4" pageNumber="135" value="2012-07-10">
|
||
<collectingDate id="EFD9E9B7FFA6FFC016D55ECAC537FD64" box="[188,336,685,711]" pageId="4" pageNumber="135" value="2012-07-10">10 July 2012</collectingDate>
|
||
</date>
|
||
at CBEE),
|
||
<location id="8EFC6044FFA6FFC017BF5ECAC629FD6B" LSID="urn:lsid:plazi:treatment:038A8789FFA6FFC216FE5D2DC547FEC6:8EFC6044FFA6FFC017BF5ECAC629FD6B" box="[470,590,685,712]" country="China" latitude="36.728" longLatPrecision="70" longitude="117.611" name="Puji Town" pageId="4" pageNumber="135" stateProvince="Shandong">Puji Town</location>
|
||
,
|
||
<collectorName id="26D65349FFA6FFC014325ECAC767FD64" box="[603,768,685,711]" pageId="4" pageNumber="135">Zhangqiu City</collectorName>
|
||
,
|
||
<collectingRegion id="49E7F87DFFA6FFC015645ECAC78BFD64" box="[781,1004,685,711]" country="China" name="Shandong" pageId="4" pageNumber="135">Shandong Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F334760FFFA6FFC015905ECAC058FD64" box="[1017,1087,685,711]" name="China" pageId="4" pageNumber="135">China</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE175058FFA6FFC012255ECAC0DCFD64" box="[1100,1211,685,711]" degrees="36.728" direction="north" orientation="latitude" pageId="4" pageNumber="135" precision="55" value="36.728">36.728°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE175058FFA6FFC012A15ECAC127FD64" box="[1224,1344,685,711]" degrees="117.611" direction="east" orientation="longitude" pageId="4" pageNumber="135" precision="55" value="117.611">117.611°E</geoCoordinate>
|
||
,
|
||
<quantity id="4CDB9B7AFFA6FFC013245ECAC1F0FD6B" box="[1357,1431,685,712]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="135" unit="m" value="100.0">
|
||
<elevation id="000ED1ACFFA6FFC013245ECAC1F0FD6B" box="[1357,1431,685,712]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="135" unit="m" value="100.0">100 m</elevation>
|
||
</quantity>
|
||
, collected on
|
||
<date id="FF9D105FFFA6FFC0174D5EB6C5A6FD48" box="[292,449,721,747]" pageId="4" pageNumber="135" value="2012-04-30">
|
||
<collectingDate id="EFD9E9B7FFA6FFC0174D5EB6C5A6FD48" box="[292,449,721,747]" pageId="4" pageNumber="135" value="2012-04-30">30 April 2012</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B4B3CC2FFA6FFC017A35EB6C7F7FCAC" ID-GBIF-Occurrence="3435942317" collectingDate="2016-08-05" country="China" elevation="184" latitude="36.674" location="Hushan" longLatPrecision="70" longitude="117.553" municipality="Zhangqiu City" pageId="4" pageNumber="135" specimenCount="1" specimenCount-female="1" stateProvince="Shandong" typeStatus="paratype">
|
||
<specimenCount id="9D25FD16FFA6FFC017A35EB6C64DFD48" box="[458,554,721,747]" pageId="4" pageNumber="135" type="female">1 female</specimenCount>
|
||
(XUX-2016-096),
|
||
<location id="8EFC6044FFA6FFC015695EB6C731FD48" LSID="urn:lsid:plazi:treatment:038A8789FFA6FFC216FE5D2DC547FEC6:8EFC6044FFA6FFC015695EB6C731FD48" box="[768,854,721,747]" country="China" latitude="36.674" longLatPrecision="70" longitude="117.553" municipality="Zhangqiu City" name="Hushan" pageId="4" pageNumber="135" stateProvince="Shandong">Hushan</location>
|
||
forestry station,
|
||
<collectingMunicipality id="6BF8ACE5FFA6FFC012795EB6C0D5FD48" box="[1040,1202,721,747]" pageId="4" pageNumber="135">Zhangqiu City</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49E7F87DFFA6FFC012D55EB6C1FFFD48" box="[1212,1432,721,747]" country="China" name="Shandong" pageId="4" pageNumber="135">Shandong Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F334760FFFA6FFC016FE5E92C4BAFCAC" box="[151,221,757,783]" name="China" pageId="4" pageNumber="135">China</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE175058FFA6FFC016825E92C53DFCAC" box="[235,346,757,783]" degrees="36.674" direction="north" orientation="latitude" pageId="4" pageNumber="135" precision="55" value="36.674">36.674°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE175058FFA6FFC017005E92C585FCB3" box="[361,482,757,784]" degrees="117.553" direction="east" orientation="longitude" pageId="4" pageNumber="135" precision="55" value="117.553">117.553°E</geoCoordinate>
|
||
,
|
||
<quantity id="4CDB9B7AFFA6FFC017985E92C65AFCB3" box="[497,573,757,784]" metricMagnitude="2" metricUnit="m" metricValue="1.8399999999999999" pageId="4" pageNumber="135" unit="m" value="184.0">
|
||
<elevation id="000ED1ACFFA6FFC017985E92C65AFCB3" box="[497,573,757,784]" metricMagnitude="2" metricUnit="m" metricValue="1.8399999999999999" pageId="4" pageNumber="135" unit="m" value="184.0">184 m</elevation>
|
||
</quantity>
|
||
, collected on
|
||
<date id="FF9D105FFFA6FFC0148B5E92C7F7FCAC" box="[738,912,757,784]" pageId="4" pageNumber="135" value="2016-08-05">
|
||
<collectingDate id="EFD9E9B7FFA6FFC0148B5E92C7F7FCAC" box="[738,912,757,784]" pageId="4" pageNumber="135" value="2016-08-05">5 August 2016</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B4B3CC2FFA6FFC015F75E92C171FC90" ID-GBIF-Occurrence="3435942301" collectingDate="2016-08-07" country="China" elevation="106" latitude="36.728" location="Puji Town" longLatPrecision="70" longitude="117.613" municipality="Zhangqiu City" pageId="4" pageNumber="135" specimenCount="2" specimenCount-female="2" stateProvince="Shandong" typeStatus="paratype">
|
||
<specimenCount id="9D25FD16FFA6FFC015F75E92C06AFCAC" box="[926,1037,757,783]" pageId="4" pageNumber="135" type="female">2 females</specimenCount>
|
||
(XUX-2016-097/099),
|
||
<location id="8EFC6044FFA6FFC013765E92C1FFFCB3" LSID="urn:lsid:plazi:treatment:038A8789FFA6FFC216FE5D2DC547FEC6:8EFC6044FFA6FFC013765E92C1FFFCB3" box="[1311,1432,757,784]" country="China" latitude="36.728" longLatPrecision="70" longitude="117.613" municipality="Zhangqiu City" name="Puji Town" pageId="4" pageNumber="135" stateProvince="Shandong">Puji Town</location>
|
||
,
|
||
<collectingMunicipality id="6BF8ACE5FFA6FFC016FE5F7EC55CFC90" box="[151,315,793,819]" pageId="4" pageNumber="135">Zhangqiu City</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49E7F87DFFA6FFC017215F7EC641FC90" box="[328,550,793,819]" country="China" name="Shandong" pageId="4" pageNumber="135">Shandong Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F334760FFFA6FFC0145B5F7EC61FFC90" box="[562,632,793,819]" name="China" pageId="4" pageNumber="135">China</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE175058FFA6FFC014ED5F7EC694FC90" box="[644,755,793,819]" degrees="36.728" direction="north" orientation="latitude" pageId="4" pageNumber="135" precision="55" value="36.728">36.728°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE175058FFA6FFC014965F7EC71FFC90" box="[767,888,793,819]" degrees="117.613" direction="east" orientation="longitude" pageId="4" pageNumber="135" precision="55" value="117.613">117.613°E</geoCoordinate>
|
||
,
|
||
<quantity id="4CDB9B7AFFA6FFC015EC5F7EC7A8FC97" box="[901,975,793,820]" metricMagnitude="2" metricUnit="m" metricValue="1.06" pageId="4" pageNumber="135" unit="m" value="106.0">
|
||
<elevation id="000ED1ACFFA6FFC015EC5F7EC7A8FC97" box="[901,975,793,820]" metricMagnitude="2" metricUnit="m" metricValue="1.06" pageId="4" pageNumber="135" unit="m" value="106.0">106 m</elevation>
|
||
</quantity>
|
||
, collected on
|
||
<date id="FF9D105FFFA6FFC012055F7DC171FC90" box="[1132,1302,793,820]" pageId="4" pageNumber="135" value="2016-08-07">
|
||
<collectingDate id="EFD9E9B7FFA6FFC012055F7DC171FC90" box="[1132,1302,793,820]" pageId="4" pageNumber="135" value="2016-08-07">7 August 2016</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B4B3CC2FFA6FFC013485F7EC6C1FCD8" ID-GBIF-Occurrence="3435942315" collectingDate="2016-08-08" country="China" elevation="103" latitude="36.669" location="Zhangqiu City" longLatPrecision="70" longitude="117.428" pageId="4" pageNumber="135" specimenCount="10" specimenCount-female="10" stateProvince="Shandong" typeStatus="paratype">
|
||
<specimenCount id="9D25FD16FFA6FFC013485F7EC1FBFC90" box="[1313,1436,793,819]" pageId="4" pageNumber="135" type="female">10 females</specimenCount>
|
||
(XUX-2016-100/101/102/103/104/105/106/ 107/108/109),
|
||
<location id="8EFC6044FFA6FFC0155C5F5AC7BCFCF4" LSID="urn:lsid:plazi:treatment:038A8789FFA6FFC216FE5D2DC547FEC6:8EFC6044FFA6FFC0155C5F5AC7BCFCF4" box="[821,987,829,855]" country="China" latitude="36.669" longLatPrecision="70" longitude="117.428" name="Zhangqiu City" pageId="4" pageNumber="135" stateProvince="Shandong">Zhangqiu City</location>
|
||
,
|
||
<collectingRegion id="49E7F87DFFA6FFC015815F5AC0AFFCF4" box="[1000,1224,829,855]" country="China" name="Shandong" pageId="4" pageNumber="135">Shandong Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F334760FFFA6FFC012BF5F5AC17BFCF4" box="[1238,1308,829,855]" name="China" pageId="4" pageNumber="135">China</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE175058FFA6FFC013405F5AC1FFFCF4" box="[1321,1432,829,855]" degrees="36.669" direction="north" orientation="latitude" pageId="4" pageNumber="135" precision="55" value="36.669">36.669°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE175058FFA6FFC016FE5F06C577FCD8" box="[151,272,865,891]" degrees="117.428" direction="east" orientation="longitude" pageId="4" pageNumber="135" precision="55" value="117.428">117.428°E</geoCoordinate>
|
||
,
|
||
<quantity id="4CDB9B7AFFA6FFC017725F06C503FCDF" box="[283,356,865,892]" metricMagnitude="2" metricUnit="m" metricValue="1.03" pageId="4" pageNumber="135" unit="m" value="103.0">
|
||
<elevation id="000ED1ACFFA6FFC017725F06C503FCDF" box="[283,356,865,892]" metricMagnitude="2" metricUnit="m" metricValue="1.03" pageId="4" pageNumber="135" unit="m" value="103.0">103 m</elevation>
|
||
</quantity>
|
||
, collected on
|
||
<date id="FF9D105FFFA6FFC017965F06C6C1FCD8" box="[511,678,865,892]" pageId="4" pageNumber="135" value="2016-08-08">
|
||
<collectingDate id="EFD9E9B7FFA6FFC017965F06C6C1FCD8" box="[511,678,865,892]" pageId="4" pageNumber="135" value="2016-08-08">8 August 2016</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B4B3CC2FFA6FFC014D85F06C144FCD8" ID-GBIF-Occurrence="3435942316" box="[689,1315,865,892]" collectionCode="T" collectorName="F. Li & F. X. Liu" country="China" location="Liu" municipality="Zhang. All" pageId="4" pageNumber="135" specimenCount="1" stateProvince="Shandong" typeStatus="paratype">
|
||
by
|
||
<collectorName id="26D65349FFA6FFC014BD5F05C76DFCD8" box="[724,778,865,892]" pageId="4" pageNumber="135">F. Li</collectorName>
|
||
,
|
||
<collectorName id="26D65349FFA6FFC0157F5F05C711FCD8" box="[790,886,865,892]" pageId="4" pageNumber="135">F.X. Liu</collectorName>
|
||
, H.
|
||
<location id="8EFC6044FFA6FFC015CA5F06C7AAFCD8" LSID="urn:lsid:plazi:treatment:038A8789FFA6FFC216FE5D2DC547FEC6:8EFC6044FFA6FFC015CA5F06C7AAFCD8" box="[931,973,865,891]" country="China" municipality="Zhang. All" name="Liu" pageId="4" pageNumber="135" stateProvince="Shandong">Liu</location>
|
||
, and Z.
|
||
<collectionCode id="ED32AE5AFFA6FFC012765F05C055FCDF" box="[1055,1074,866,892]" name="Tavera, Department of Geology and Geophysics" pageId="4" pageNumber="135">T</collectionCode>
|
||
.
|
||
<collectingMunicipality id="6BF8ACE5FFA6FFC012545F06C0D2FCD8" box="[1085,1205,865,891]" pageId="4" pageNumber="135">Zhang. All</collectingMunicipality>
|
||
in CBEE
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3396514FFA6FFC016AE5FE2C4D2FB1C" pageId="4" pageNumber="135" type="diagnosis">
|
||
<paragraph id="8B9C369FFFA6FFC016AE5FE2C4D2FB1C" blockId="4.[151,1437,505,2008]" pageId="4" pageNumber="135">
|
||
<emphasis id="B957EA8DFFA6FFC016AE5FE2C525FC3C" bold="true" box="[199,322,901,927]" pageId="4" pageNumber="135">Diagnosis.</emphasis>
|
||
Males of
|
||
<taxonomicName id="4C234D1CFFA6FFC017D05FE1C644FC3C" authority="Xu & Yu & Liu & Li, 2022" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" box="[441,547,902,927]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="135" phylum="Arthropoda" rank="species" species="yiyuan" status="sp. nov.">
|
||
<emphasis id="B957EA8DFFA6FFC017D05FE1C644FC3C" box="[441,547,902,927]" italics="true" pageId="4" pageNumber="135">L. yiyuan</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B957EA8DFFA6FFC014455FE2C6EFFC3C" bold="true" box="[556,648,901,927]" pageId="4" pageNumber="135">
|
||
<taxonomicNameLabel id="A26457F6FFA6FFC014455FE2C6EFFC3C" box="[556,648,901,927]" pageId="4" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
resemble those of
|
||
<taxonomicName id="4C234D1CFFA6FFC0150D5FE1C06AFC3C" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" baseAuthorityName="Schenkel" baseAuthorityYear="1953" box="[868,1037,901,927]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="135" phylum="Arthropoda" rank="species" species="schensiensis">
|
||
<emphasis id="B957EA8DFFA6FFC0150D5FE1C06AFC3C" box="[868,1037,901,927]" italics="true" pageId="4" pageNumber="135">L. schensiensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C234D1CFFA6FFC012205FE1C0B4FC3C" authority="Xu & Yu & Liu & Li, 2022" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" box="[1097,1235,901,927]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="135" phylum="Arthropoda" rank="species" species="yuncheng" status="sp. nov.">
|
||
<emphasis id="B957EA8DFFA6FFC012205FE1C0B4FC3C" box="[1097,1235,901,927]" italics="true" pageId="4" pageNumber="135">L. yuncheng</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B957EA8DFFA6FFC012B45FE2C15FFC3C" bold="true" box="[1245,1336,901,927]" pageId="4" pageNumber="135">
|
||
<taxonomicNameLabel id="A26457F6FFA6FFC012B45FE2C15FFC3C" box="[1245,1336,901,927]" pageId="4" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, but can be distinguished from those of
|
||
<taxonomicName id="4C234D1CFFA6FFC0179E5FCDC6F8FC60" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" baseAuthorityName="Schenkel" baseAuthorityYear="1953" box="[503,671,937,963]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="135" phylum="Arthropoda" rank="species" species="schensiensis">
|
||
<emphasis id="B957EA8DFFA6FFC0179E5FCDC6F8FC60" box="[503,671,937,963]" italics="true" pageId="4" pageNumber="135">L. schensiensis</emphasis>
|
||
</taxonomicName>
|
||
by the tegulum having a slightly larger terminal apophysis, and the embolus with less longitudinal ribs retrolaterally (
|
||
<figureCitation id="13182A1AFFA6FFC014AE5FAAC72FFC44" box="[711,840,973,999]" captionStart="FIGURE 3" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[170,1417,237,1695]" captionTargetId="figure-43@5.[170,1417,237,1695]" captionTargetPageId="5" captionText="FIGURE 3. Male genital anatomy of Luthela yiyuan sp. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view. A–G, XUX-2012-051 (holotype). H–K, XUX-2012-045. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840647" httpUri="https://zenodo.org/record/5840647/files/figure.png" pageId="4" pageNumber="135">Fig. 3J, 3K</figureCitation>
|
||
); from those of
|
||
<taxonomicName id="4C234D1CFFA6FFC015955FA9C0E3FC44" authority="Xu & Yu & Liu & Li, 2022" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" box="[1020,1156,973,999]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="135" phylum="Arthropoda" rank="species" species="yuncheng" status="sp. nov.">
|
||
<emphasis id="B957EA8DFFA6FFC015955FA9C0E3FC44" box="[1020,1156,973,999]" italics="true" pageId="4" pageNumber="135">L. yuncheng</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B957EA8DFFA6FFC012E55FAAC081FC44" bold="true" box="[1164,1254,973,999]" pageId="4" pageNumber="135">
|
||
<taxonomicNameLabel id="A26457F6FFA6FFC012E55FAAC081FC44" box="[1164,1254,973,999]" pageId="4" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the marginal tegular apophysis and the dorsal extension of the terminal apophysis of the tegulum having irregular teeth (
|
||
<figureCitation id="13182A1AFFA6FFC013565F96C4DAFB8C" captionStart="FIGURE 3" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[170,1417,237,1695]" captionTargetId="figure-43@5.[170,1417,237,1695]" captionTargetPageId="5" captionText="FIGURE 3. Male genital anatomy of Luthela yiyuan sp. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view. A–G, XUX-2012-051 (holotype). H–K, XUX-2012-045. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840647" httpUri="https://zenodo.org/record/5840647/files/figure.png" pageId="4" pageNumber="135">Fig. 3G, 3K</figureCitation>
|
||
); from those of
|
||
<taxonomicName id="4C234D1CFFA6FFC017195871C593FB8C" authority="Xu & Yu & Liu & Li, 2022" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" box="[368,500,1045,1071]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="135" phylum="Arthropoda" rank="species" species="dengfeng" status="sp. nov.">
|
||
<emphasis id="B957EA8DFFA6FFC017195871C593FB8C" box="[368,500,1045,1071]" italics="true" pageId="4" pageNumber="135">L. dengfeng</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B957EA8DFFA6FFC017925872C632FB8C" bold="true" box="[507,597,1045,1071]" pageId="4" pageNumber="135">
|
||
<taxonomicNameLabel id="A26457F6FFA6FFC017925872C632FB8C" box="[507,597,1045,1071]" pageId="4" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the conductor having a longer lower spine (
|
||
<figureCitation id="13182A1AFFA6FFC0120F5872C097FB8C" box="[1126,1264,1045,1071]" captionStart="FIGURE 3" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[170,1417,237,1695]" captionTargetId="figure-43@5.[170,1417,237,1695]" captionTargetPageId="5" captionText="FIGURE 3. Male genital anatomy of Luthela yiyuan sp. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view. A–G, XUX-2012-051 (holotype). H–K, XUX-2012-045. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840647" httpUri="https://zenodo.org/record/5840647/files/figure.png" pageId="4" pageNumber="135">Fig. 3G, 3K</figureCitation>
|
||
); from those of
|
||
<taxonomicName id="4C234D1CFFA6FFC016FE585DC56EFBF0" authority="Xu & Yu & Liu & Li, 2022" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" box="[151,265,1081,1107]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="135" phylum="Arthropoda" rank="species" species="handan" status="sp. nov.">
|
||
<emphasis id="B957EA8DFFA6FFC016FE585DC56EFBF0" box="[151,265,1081,1107]" italics="true" pageId="4" pageNumber="135">L. handan</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B957EA8DFFA6FFC01766585EC50EFBF0" bold="true" box="[271,361,1081,1107]" pageId="4" pageNumber="135">
|
||
<taxonomicNameLabel id="A26457F6FFA6FFC01766585EC50EFBF0" box="[271,361,1081,1107]" pageId="4" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by contrategulum margin having slightly small teeth (
|
||
<figureCitation id="13182A1AFFA6FFC015AE585EC036FBF0" box="[967,1105,1081,1107]" captionStart="FIGURE 3" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[170,1417,237,1695]" captionTargetId="figure-43@5.[170,1417,237,1695]" captionTargetPageId="5" captionText="FIGURE 3. Male genital anatomy of Luthela yiyuan sp. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view. A–G, XUX-2012-051 (holotype). H–K, XUX-2012-045. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840647" httpUri="https://zenodo.org/record/5840647/files/figure.png" pageId="4" pageNumber="135">Fig. 3D, 3K</figureCitation>
|
||
); from those of
|
||
<taxonomicName id="4C234D1CFFA6FFC0136A585DC113FBF0" authority="Xu & Yu & Liu & Li, 2022" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" box="[1283,1396,1081,1107]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="135" phylum="Arthropoda" rank="species" species="badong" status="sp. nov.">
|
||
<emphasis id="B957EA8DFFA6FFC0136A585DC113FBF0" box="[1283,1396,1081,1107]" italics="true" pageId="4" pageNumber="135">L. badong</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B957EA8DFFA6FFC01312585EC4AFFBD4" bold="true" pageId="4" pageNumber="135">
|
||
<taxonomicNameLabel id="A26457F6FFA6FFC01312585EC4AFFBD4" pageId="4" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C234D1CFFA6FFC017695839C53EFBD4" authority="Xu & Yu & Liu & Li, 2022" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" box="[256,345,1118,1143]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="135" phylum="Arthropoda" rank="species" species="taian" status="sp. nov.">
|
||
<emphasis id="B957EA8DFFA6FFC017695839C53EFBD4" box="[256,345,1118,1143]" italics="true" pageId="4" pageNumber="135">L. taian</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B957EA8DFFA6FFC01709583AC5DDFBD4" bold="true" box="[352,442,1117,1143]" pageId="4" pageNumber="135">
|
||
<taxonomicNameLabel id="A26457F6FFA6FFC01709583AC5DDFBD4" box="[352,442,1117,1143]" pageId="4" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the conductor with a small middle spine (
|
||
<figureCitation id="13182A1AFFA6FFC015D2583AC05EFBD4" box="[955,1081,1117,1143]" captionStart="FIGURE 3" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[170,1417,237,1695]" captionTargetId="figure-43@5.[170,1417,237,1695]" captionTargetPageId="5" captionText="FIGURE 3. Male genital anatomy of Luthela yiyuan sp. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view. A–G, XUX-2012-051 (holotype). H–K, XUX-2012-045. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840647" httpUri="https://zenodo.org/record/5840647/files/figure.png" pageId="4" pageNumber="135">Fig. 3A–G</figureCitation>
|
||
). Females of
|
||
<taxonomicName id="4C234D1CFFA6FFC012B85839C15DFBD4" authority="Xu & Yu & Liu & Li, 2022" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" box="[1233,1338,1118,1143]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="135" phylum="Arthropoda" rank="species" species="yiyuan" status="sp. nov.">
|
||
<emphasis id="B957EA8DFFA6FFC012B85839C15DFBD4" box="[1233,1338,1118,1143]" italics="true" pageId="4" pageNumber="135">L. yiyuan</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B957EA8DFFA6FFC0132B583AC1FBFBD4" bold="true" box="[1346,1436,1117,1143]" pageId="4" pageNumber="135">
|
||
<taxonomicNameLabel id="A26457F6FFA6FFC0132B583AC1FBFBD4" box="[1346,1436,1117,1143]" pageId="4" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differ from those of
|
||
<taxonomicName id="4C234D1CFFA6FFC0171E58E5C59CFB38" authority="Xu & Yu & Liu & Li, 2022" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" box="[375,507,1153,1179]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="135" phylum="Arthropoda" rank="species" species="dengfeng" status="sp. nov.">
|
||
<emphasis id="B957EA8DFFA6FFC0171E58E5C59CFB38" box="[375,507,1153,1179]" italics="true" pageId="4" pageNumber="135">L. dengfeng</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B957EA8DFFA6FFC0146858E6C63DFB38" bold="true" box="[513,602,1153,1179]" pageId="4" pageNumber="135">
|
||
<taxonomicNameLabel id="A26457F6FFA6FFC0146858E6C63DFB38" box="[513,602,1153,1179]" pageId="4" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the bases of middle receptacular clusters being close to each other (
|
||
<figureCitation id="13182A1AFFA6FFC0130658E6C4CFFB1C" captionStart="FIGURE 4" captionStartId="6.[151,250,1251,1275]" captionTargetBox="[164,1423,381,1228]" captionTargetId="figure-115@6.[164,1423,381,1228]" captionTargetPageId="6" captionText="FIGURE 4. Female genital anatomy of Luthela yiyuan sp. nov. A–E, K–O, vulva dorsal view. F–J, P–T, vulva ventral view. A, F, XUX-2012-034. B, G, XUX-2012-035. C, H, XUX-2012-049. D, I, XUX-2016-101. E, J, XUX-2016-107. K, P, XUX-2012- 042. L, Q, XUX-2012-040. M, R, XUX-2012-046. N, S, XUX-2016-103. O, T, XUX-2016-108. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840651" httpUri="https://zenodo.org/record/5840651/files/figure.png" pageId="4" pageNumber="135">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3396514FFA6FFC116AE58AEC6F6FF76" lastPageId="5" lastPageNumber="136" pageId="4" pageNumber="135" type="description">
|
||
<paragraph id="8B9C369FFFA6FFC016AE58AEC558FA1F" blockId="4.[151,1437,505,2008]" pageId="4" pageNumber="135">
|
||
<emphasis id="B957EA8DFFA6FFC016AE58AEC53EFB40" bold="true" box="[199,345,1225,1251]" pageId="4" pageNumber="135">Description.</emphasis>
|
||
Male (
|
||
<typeStatus id="5498883DFFA6FFC017C358AEC67EFB47" box="[426,537,1225,1252]" pageId="4" pageNumber="135" type="holotype">Holotype</typeStatus>
|
||
). Carapace brown; opisthosoma light brown, with dark brown tergites; sternum narrow, much longer than wide; few long pointed hairs over ocular mound in longitudinal row; chelicerae robust, promargin of groove with 11 denticles of variable size; legs with strong hairs and spines; opisthosoma with 12 tergites, the second to fifth larger than others and the others small; 7 spinnerets. Measurements: BL 16.40, CL 6.65, CW 5.50, OL 8.80, OW 6.20; ALE> PLE> PME> AME; leg I 18.40 (5.45 + 2.30+ 3.65 + 4.70 + 2.30), leg II 18.10 (5.00 + 2.35 + 3.45 + 4.70 + 2.60), leg III 19.55 (4.95 + 2.35+ 3.50 + 5.60 + 3.15), leg IV 24.60 (6.25 + 2.50 + 4.80 + 7.55 + 3.50).
|
||
</paragraph>
|
||
<paragraph id="8B9C369FFFA6FFC016AE59A2C154F914" blockId="4.[151,1437,505,2008]" pageId="4" pageNumber="135">
|
||
Palp: Prolateral paracymbium unpigmented and unsclerotized, numerous setae and spines at tip of paracymbium (
|
||
<figureCitation id="13182A1AFFA6FFC016C9598EC57BF9A0" box="[160,284,1513,1539]" captionStart="FIGURE 3" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[170,1417,237,1695]" captionTargetId="figure-43@5.[170,1417,237,1695]" captionTargetPageId="5" captionText="FIGURE 3. Male genital anatomy of Luthela yiyuan sp. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view. A–G, XUX-2012-051 (holotype). H–K, XUX-2012-045. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840647" httpUri="https://zenodo.org/record/5840647/files/figure.png" pageId="4" pageNumber="135">Fig. 3A–C</figureCitation>
|
||
). Contrategulum with 7 or 8 teeth along margin, distal tooth with 3 serrations (
|
||
<figureCitation id="13182A1AFFA6FFC012F0598EC142F9A0" box="[1177,1317,1513,1539]" captionStart="FIGURE 3" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[170,1417,237,1695]" captionTargetId="figure-43@5.[170,1417,237,1695]" captionTargetPageId="5" captionText="FIGURE 3. Male genital anatomy of Luthela yiyuan sp. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view. A–G, XUX-2012-051 (holotype). H–K, XUX-2012-045. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840647" httpUri="https://zenodo.org/record/5840647/files/figure.png" pageId="4" pageNumber="135">Fig. 3D, 3H</figureCitation>
|
||
). Tegulum with irregular dentate marginal apophysis, irregular dorsal extension of terminal apophysis, and smooth terminal apophysis (
|
||
<figureCitation id="13182A1AFFA6FFC0177F5A56C5C7F9E8" box="[278,416,1585,1611]" captionStart="FIGURE 3" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[170,1417,237,1695]" captionTargetId="figure-43@5.[170,1417,237,1695]" captionTargetPageId="5" captionText="FIGURE 3. Male genital anatomy of Luthela yiyuan sp. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view. A–G, XUX-2012-051 (holotype). H–K, XUX-2012-045. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840647" httpUri="https://zenodo.org/record/5840647/files/figure.png" pageId="4" pageNumber="135">Fig. 3G, 3K</figureCitation>
|
||
). Conductor situated ventro-proximally on embolus, with two long spines, upper one fits to furrow of embolus, lower one pointed to contrategulum, and one small tooth situated at between two long spines, and close to upper one (
|
||
<figureCitation id="13182A1AFFA6FFC017F65A1EC67BF930" box="[415,540,1657,1683]" captionStart="FIGURE 3" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[170,1417,237,1695]" captionTargetId="figure-43@5.[170,1417,237,1695]" captionTargetPageId="5" captionText="FIGURE 3. Male genital anatomy of Luthela yiyuan sp. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view. A–G, XUX-2012-051 (holotype). H–K, XUX-2012-045. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840647" httpUri="https://zenodo.org/record/5840647/files/figure.png" pageId="4" pageNumber="135">Fig. 3A–G</figureCitation>
|
||
), or an inconspicuous tooth located at the middle of two long spines (
|
||
<figureCitation id="13182A1AFFA6FFC013745A1EC1F7F930" box="[1309,1424,1657,1683]" captionStart="FIGURE 3" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[170,1417,237,1695]" captionTargetId="figure-43@5.[170,1417,237,1695]" captionTargetPageId="5" captionText="FIGURE 3. Male genital anatomy of Luthela yiyuan sp. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view. A–G, XUX-2012-051 (holotype). H–K, XUX-2012-045. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840647" httpUri="https://zenodo.org/record/5840647/files/figure.png" pageId="4" pageNumber="135">Fig. 3H–J</figureCitation>
|
||
). Embolus largely sclerotized, with a flat opening and several longitudinal ribs retrolaterally (
|
||
<figureCitation id="13182A1AFFA6FFC012F55AFAC141F914" box="[1180,1318,1693,1719]" captionStart="FIGURE 3" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[170,1417,237,1695]" captionTargetId="figure-43@5.[170,1417,237,1695]" captionTargetPageId="5" captionText="FIGURE 3. Male genital anatomy of Luthela yiyuan sp. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view. A–G, XUX-2012-051 (holotype). H–K, XUX-2012-045. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840647" httpUri="https://zenodo.org/record/5840647/files/figure.png" pageId="4" pageNumber="135">Fig. 3D, 3K</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9C369FFFA6FFC016AE5AA6C604F833" blockId="4.[151,1437,505,2008]" pageId="4" pageNumber="135">Female (XUX-2012-035). Carapace reddish brown; opisthosoma light brown, with brown tergites; chelicerae robust with promargin of groove with 10–11 strong denticles of variable size; legs with strong hairs and spines; opisthosoma with 12 tergites, similar to male; 7 spinnerets. Measurements: BL 21.70, CL 8.40, CW 8.10, OL 12.90, OW 12.20; ALE> PLE> PME> AME; palp 16.40 (5.20 + 2.90 + 3.90 + 4.40), leg I 17.90 (5.60 + 3.30 + 3.40 + 3.70 + 1.90), leg II 16.40 (4.50 + 3.20 + 3.10 + 3.50 + 2.10), leg III 18.40 (5.40 + 3.20 + 3.00 + 4.40 + 2.40), leg IV 27.60 (7.90 + 3.90 + 4.80 + 7.40 + 3.60).</paragraph>
|
||
<paragraph id="8B9C369FFFA6FFC116AE5BFEC6F6FF76" blockId="4.[151,1437,505,2008]" lastBlockId="5.[151,1437,151,213]" lastPageId="5" lastPageNumber="136" pageId="4" pageNumber="135">
|
||
Genitalia: Posterior margin of genital sternite slightly incurved, two pairs of receptacular clusters similar-sized, middle ones with stalks, close to each other (
|
||
<figureCitation id="13182A1AFFA6FFC014FC5BDAC77FF874" box="[661,792,1981,2007]" captionStart="FIGURE 4" captionStartId="6.[151,250,1251,1275]" captionTargetBox="[164,1423,381,1228]" captionTargetId="figure-115@6.[164,1423,381,1228]" captionTargetPageId="6" captionText="FIGURE 4. Female genital anatomy of Luthela yiyuan sp. nov. A–E, K–O, vulva dorsal view. F–J, P–T, vulva ventral view. A, F, XUX-2012-034. B, G, XUX-2012-035. C, H, XUX-2012-049. D, I, XUX-2016-101. E, J, XUX-2016-107. K, P, XUX-2012- 042. L, Q, XUX-2012-040. M, R, XUX-2012-046. N, S, XUX-2016-103. O, T, XUX-2016-108. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840651" httpUri="https://zenodo.org/record/5840651/files/figure.png" pageId="4" pageNumber="135">Fig. 4A–M</figureCitation>
|
||
), or fused together as one large receptacular cluster (
|
||
<figureCitation id="13182A1AFFA6FFC113065BDAC48BFF12" captionStart="FIGURE 4" captionStartId="6.[151,250,1251,1275]" captionTargetBox="[164,1423,381,1228]" captionTargetId="figure-115@6.[164,1423,381,1228]" captionTargetPageId="6" captionText="FIGURE 4. Female genital anatomy of Luthela yiyuan sp. nov. A–E, K–O, vulva dorsal view. F–J, P–T, vulva ventral view. A, F, XUX-2012-034. B, G, XUX-2012-035. C, H, XUX-2012-049. D, I, XUX-2016-101. E, J, XUX-2016-107. K, P, XUX-2012- 042. L, Q, XUX-2012-040. M, R, XUX-2012-046. N, S, XUX-2016-103. O, T, XUX-2016-108. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840651" httpUri="https://zenodo.org/record/5840651/files/figure.png" lastPageId="5" lastPageNumber="136" pageId="4" pageNumber="135">Fig. 4N, 4O</figureCitation>
|
||
), situated at anterior margin of bursa copulatrix, lateral ones situated at slightly dorsolateral position of bursa copulatrix with inconspicuous stalks (
|
||
<figureCitation id="13182A1AFFA7FFC114565CDCC6E3FF76" box="[575,644,187,213]" captionStart="FIGURE 4" captionStartId="6.[151,250,1251,1275]" captionTargetBox="[164,1423,381,1228]" captionTargetId="figure-115@6.[164,1423,381,1228]" captionTargetPageId="6" captionText="FIGURE 4. Female genital anatomy of Luthela yiyuan sp. nov. A–E, K–O, vulva dorsal view. F–J, P–T, vulva ventral view. A, F, XUX-2012-034. B, G, XUX-2012-035. C, H, XUX-2012-049. D, I, XUX-2016-101. E, J, XUX-2016-107. K, P, XUX-2012- 042. L, Q, XUX-2012-040. M, R, XUX-2012-046. N, S, XUX-2016-103. O, T, XUX-2016-108. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840651" httpUri="https://zenodo.org/record/5840651/files/figure.png" pageId="5" pageNumber="136">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5C6617FFA7FFC116FE5AD1C115F951" ID-DOI="http://doi.org/10.5281/zenodo.5840647" ID-Zenodo-Dep="5840647" httpUri="https://zenodo.org/record/5840647/files/figure.png" pageId="5" pageNumber="136" startId="5.[151,250,1718,1742]" targetBox="[170,1417,237,1695]" targetPageId="5">
|
||
<paragraph id="8B9C369FFFA7FFC116FE5AD1C115F951" blockId="5.[151,1437,1718,1778]" pageId="5" pageNumber="136">
|
||
<emphasis id="B957EA8DFFA7FFC116FE5AD1C574F96D" bold="true" box="[151,275,1718,1742]" pageId="5" pageNumber="136">FIGURE 3.</emphasis>
|
||
Male genital anatomy of
|
||
<taxonomicName id="4C234D1CFFA7FFC114755AD1C6D3F96D" authority="Xu & Yu & Liu & Li, 2022" authorityName="Xu & Yu & Liu & Li" authorityYear="2022" box="[540,692,1718,1742]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="136" phylum="Arthropoda" rank="species" species="yiyuan" status="sp. nov.">
|
||
<emphasis id="B957EA8DFFA7FFC114755AD1C6D3F96D" box="[540,692,1718,1742]" italics="true" pageId="5" pageNumber="136">Luthela yiyuan</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B957EA8DFFA7FFC114D35AD1C76CF96D" bold="true" box="[698,779,1718,1742]" pageId="5" pageNumber="136">
|
||
<taxonomicNameLabel id="A26457F6FFA7FFC114D35AD1C76CF96D" box="[698,779,1718,1742]" pageId="5" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view. A–G, XUX-2012-051 (holotype). H–K, XUX-2012-045. Scale bars: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3396514FFA7FFC216AE5B46C547FEC6" lastPageId="6" lastPageNumber="137" pageId="5" pageNumber="136" type="etymology">
|
||
<paragraph id="8B9C369FFFA7FFC116AE5B46C02FF899" blockId="5.[151,1437,1824,2031]" box="[199,1096,1824,1851]" pageId="5" pageNumber="136">
|
||
<emphasis id="B957EA8DFFA7FFC116AE5B46C529F898" bold="true" box="[199,334,1825,1851]" pageId="5" pageNumber="136">Etymology.</emphasis>
|
||
The species epithet, a noun in apposition, refers to the
|
||
<typeStatus id="5498883DFFA7FFC115DE5B46C780F898" box="[951,999,1825,1851]" pageId="5" pageNumber="136">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
<paragraph id="8B9C369FFFA7FFC116AE5B23C76DF8FC" blockId="5.[151,1437,1824,2031]" box="[199,778,1860,1887]" pageId="5" pageNumber="136">
|
||
<emphasis id="B957EA8DFFA7FFC116AE5B23C506F8FD" bold="true" box="[199,353,1860,1886]" pageId="5" pageNumber="136">Distribution.</emphasis>
|
||
<collectingRegion id="49E7F87DFFA7FFC1170E5B23C625F8FD" box="[359,578,1860,1886]" country="China" name="Shandong" pageId="5" pageNumber="136">Shandong Province</collectingRegion>
|
||
(Zhangqiu, Zibo)
|
||
</paragraph>
|
||
<paragraph id="8B9C369FFFA7FFC216AE5B0EC7ECFF5A" blockId="5.[151,1437,1824,2031]" lastBlockId="6.[151,1437,151,357]" lastPageId="6" lastPageNumber="137" pageId="5" pageNumber="136">
|
||
<emphasis id="B957EA8DFFA7FFC116AE5B0EC55CF820" bold="true" box="[199,315,1897,1923]" pageId="5" pageNumber="136">Remarks.</emphasis>
|
||
Males and females vary in body size: males (
|
||
<emphasis id="B957EA8DFFA7FFC1155C5B0EC72FF821" box="[821,840,1897,1922]" italics="true" pageId="5" pageNumber="136">N</emphasis>
|
||
= 2): BL 14.75–16.40, CL 6.35–6.65, CW 5.50–5.85, OL 7.40–8.80, OW 5.00–6.20; females (
|
||
<emphasis id="B957EA8DFFA7FFC114355BEAC608F805" box="[604,623,1933,1958]" italics="true" pageId="5" pageNumber="136">N</emphasis>
|
||
= 22): BL 16.70–24.90, CL 8.00–10.75, CW 7.10–8.90, OL 8.80–13.40, OW 6.90–13.20. There was intraspecific variation in male and female genital morphology and molecular markers. For male genitalia, the conductor has a small middle spine close to the upper spine (
|
||
<figureCitation id="13182A1AFFA7FFC112585BB3C0CAF84C" box="[1073,1197,2004,2031]" captionStart="FIGURE 3" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[170,1417,237,1695]" captionTargetId="figure-43@5.[170,1417,237,1695]" captionTargetPageId="5" captionText="FIGURE 3. Male genital anatomy of Luthela yiyuan sp. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view. A–G, XUX-2012-051 (holotype). H–K, XUX-2012-045. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840647" httpUri="https://zenodo.org/record/5840647/files/figure.png" pageId="5" pageNumber="136">Fig. 3A–G</figureCitation>
|
||
), or an inconspicuous middle spine close to the lower spine (
|
||
<figureCitation id="13182A1AFFA4FFC214395CF0C6CCFF12" box="[592,683,151,177]" captionStart="FIGURE 3" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[170,1417,237,1695]" captionTargetId="figure-43@5.[170,1417,237,1695]" captionTargetPageId="5" captionText="FIGURE 3. Male genital anatomy of Luthela yiyuan sp. nov. A, D, H, palp prolateral view. B, E, I, palp ventral view. C, F, J, palp retrolateral view. G, K, palp distal view. A–G, XUX-2012-051 (holotype). H–K, XUX-2012-045. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840647" httpUri="https://zenodo.org/record/5840647/files/figure.png" pageId="6" pageNumber="137">Fig. 3H</figureCitation>
|
||
). For female genitalia, the posterior margin of genital sternite was slightly incurved (
|
||
<figureCitation id="13182A1AFFA4FFC217005CDCC6F8FF76" box="[361,671,187,213]" captionStart="FIGURE 4" captionStartId="6.[151,250,1251,1275]" captionTargetBox="[164,1423,381,1228]" captionTargetId="figure-115@6.[164,1423,381,1228]" captionTargetPageId="6" captionText="FIGURE 4. Female genital anatomy of Luthela yiyuan sp. nov. A–E, K–O, vulva dorsal view. F–J, P–T, vulva ventral view. A, F, XUX-2012-034. B, G, XUX-2012-035. C, H, XUX-2012-049. D, I, XUX-2016-101. E, J, XUX-2016-107. K, P, XUX-2012- 042. L, Q, XUX-2012-040. M, R, XUX-2012-046. N, S, XUX-2016-103. O, T, XUX-2016-108. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840651" httpUri="https://zenodo.org/record/5840651/files/figure.png" pageId="6" pageNumber="137">Fig. 4F, 4G, 4I, 4J, 4P, 4Q</figureCitation>
|
||
) or straight (
|
||
<figureCitation id="13182A1AFFA4FFC215515CDCC781FF76" box="[824,998,187,213]" captionStart="FIGURE 4" captionStartId="6.[151,250,1251,1275]" captionTargetBox="[164,1423,381,1228]" captionTargetId="figure-115@6.[164,1423,381,1228]" captionTargetPageId="6" captionText="FIGURE 4. Female genital anatomy of Luthela yiyuan sp. nov. A–E, K–O, vulva dorsal view. F–J, P–T, vulva ventral view. A, F, XUX-2012-034. B, G, XUX-2012-035. C, H, XUX-2012-049. D, I, XUX-2016-101. E, J, XUX-2016-107. K, P, XUX-2012- 042. L, Q, XUX-2012-040. M, R, XUX-2012-046. N, S, XUX-2016-103. O, T, XUX-2016-108. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840651" httpUri="https://zenodo.org/record/5840651/files/figure.png" pageId="6" pageNumber="137">Fig. 4H, 4R–T</figureCitation>
|
||
); there were two pairs of receptacular clusters or the middle ones were fused as a large one (
|
||
<figureCitation id="13182A1AFFA4FFC2149A5CB8C719FF5A" box="[755,894,223,249]" captionStart="FIGURE 4" captionStartId="6.[151,250,1251,1275]" captionTargetBox="[164,1423,381,1228]" captionTargetId="figure-115@6.[164,1423,381,1228]" captionTargetPageId="6" captionText="FIGURE 4. Female genital anatomy of Luthela yiyuan sp. nov. A–E, K–O, vulva dorsal view. F–J, P–T, vulva ventral view. A, F, XUX-2012-034. B, G, XUX-2012-035. C, H, XUX-2012-049. D, I, XUX-2016-101. E, J, XUX-2016-107. K, P, XUX-2012- 042. L, Q, XUX-2012-040. M, R, XUX-2012-046. N, S, XUX-2016-103. O, T, XUX-2016-108. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840651" httpUri="https://zenodo.org/record/5840651/files/figure.png" pageId="6" pageNumber="137">Fig. 4N, 4O</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9C369FFFA4FFC216AE5D64C547FEC6" blockId="6.[151,1437,151,357]" pageId="6" pageNumber="137">
|
||
Intraspecific genetic distance is between 0% and 3.4% based on the K2P substitution model and
|
||
<emphasis id="B957EA8DFFA4FFC212985D64C146FEBE" box="[1265,1313,259,285]" italics="true" pageId="6" pageNumber="137">COI</emphasis>
|
||
nucleotide sequences (
|
||
<emphasis id="B957EA8DFFA4FFC217495D40C554FEE3" box="[288,307,295,320]" italics="true" pageId="6" pageNumber="137">N</emphasis>
|
||
= 21, including juveniles). The GenBank accession code of the
|
||
<typeStatus id="5498883DFFA4FFC212215D40C0CDFEE2" box="[1096,1194,295,321]" pageId="6" pageNumber="137" type="holotype">holotype</typeStatus>
|
||
(XUX-2012-051) is
|
||
<accessionNumber id="9470AB7CFFA4FFC216FE5D2CC57BFEC6" box="[151,284,331,357]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH172727" pageId="6" pageNumber="137">MH172727</accessionNumber>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |