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<document id="C4DBAAA089BC048019935450968990EA" ID-DOI="10.11646/zootaxa.5315.4.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="8142274" ID-ZooBank="29F4CB0A-16C0-4BAE-88F1-958762C99938" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1689235878063" checkinUser="plazi" docAuthor="Provenzano-Rizzi, Francisco" docDate="2023" docId="094C87FDF6589005FDB94078161DFC8C" docLanguage="en" docName="zootaxa.5315.4.2.pdf" docOrigin="Zootaxa 5315 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5315.4.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cheilonimata minuta Provenzano-Rizzi 2023, new species" docType="treatment" docVersion="3" lastPageNumber="322" masterDocId="F575FF85F65B9002FD2E41321709FFD1" masterDocTitle="A new genus and species of whiptail armored catfish (Siluriformes: Loricariinae) from southern Venezuela" masterLastPageNumber="328" masterPageNumber="315" pageNumber="318" updateTime="1689278290879" updateUser="julia" zenodo-license-document="CLOSED">
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<mods:title id="C4B3CAF54DC86F8CAECE45B1631EE399">A new genus and species of whiptail armored catfish (Siluriformes: Loricariinae) from southern Venezuela</mods:title>
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<mods:date id="FEE3FA5C7972B0FA210DDD5E8E7AC22C">2023</mods:date>
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<taxonomicName id="46E54D68F6589001FDB940781690FEB4" authority="Provenzano-Rizzi, 2023" authorityName="Provenzano-Rizzi" authorityYear="2023" box="[151,409,330,357]" family="Loricariidae" genus="Cheilonimata" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="318" phylum="Chordata" rank="species" species="minuta" status="new species">
<emphasis id="B391EAF9F6589001FDB940781690FEB4" bold="true" box="[151,409,330,357]" italics="true" pageId="3" pageNumber="318">Cheilonimata minuta</emphasis>
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<figureCitation id="19DE2A6EF6589001FDB9405D160BFE58" box="[151,258,367,393]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1897,1921]" captionStartId-1="5.[151,250,1944,1968]" captionStartId-2="6.[151,250,1795,1819]" captionTargetBox-0="[183,1405,698,1872]" captionTargetBox-1="[263,1324,180,1919]" captionTargetBox-2="[176,1411,884,1771]" captionTargetId-0="figure-114@3.[183,1405,698,1872]" captionTargetId-1="figure-15@5.[263,1324,180,1919]" captionTargetId-2="figure-335@6.[176,1411,884,1771]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 1. Cheilonimata minuta, MBUCV-V-32954, 54.3 mm SL, holotype. Lateral, dorsal and ventral views. Scale bar 2 mm. Photos by C. Do Nascimiento." captionText-1="FIGURE 2. Cheilonimata minuta, MBUCV-V-32954, 54.3 mm SL, holotype. a) Detail of ventral view to show: head contour, mouth and abdomen cover. b) Close-up of mouth, detail of lower lip cover and translucent membrane on upper lip margin (red arrows). Scale bar 2 mm. c) Photos by C. Do Nascimiento." captionText-2="FIGURE 3. Cheilonimata minuta, ANSP 162156, 39.5 mm SL, paratype, detail of buccal ornamentation on mouth roof and behind premaxillaries (red arrow). Scale bar 1 mm. Photo by F. Provenzano." figureDoi-0="http://doi.org/10.5281/zenodo.8142276" figureDoi-1="http://doi.org/10.5281/zenodo.8142278" figureDoi-2="http://doi.org/10.5281/zenodo.8142280" httpUri-0="https://zenodo.org/record/8142276/files/figure.png" httpUri-1="https://zenodo.org/record/8142278/files/figure.png" httpUri-2="https://zenodo.org/record/8142280/files/figure.png" pageId="3" pageNumber="318">Figs. 13</figureCitation>
,
<tableCitation id="CC670350F6589001FC23405D1669FE58" box="[269,352,367,393]" captionStart="TABLE 1" captionStartId="4.[151,239,974,998]" captionTargetPageId="4" captionText="TABLE 1. Morphometric data of Cheilonimata minuta expressed as percentage of standard length (n=12)." pageId="3" pageNumber="318">Table 1</tableCitation>
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New genus and species “L3”.
<bibRefCitation id="E5744B1AF6589001FCE640EA15CEFE3E" author="Provenzano, F." box="[456,711,471,495]" pageId="3" pageNumber="318" refId="ref9705" refString="Provenzano, F. (2011) Estudio sobre las relaciones filogeneticas de las especies incluidas en la subfamilia Loricariinae (Siluriformes, Loricariidae). Unpublished Doctor in Science Dissertation, Universidad Central de Venezuela, Caracas, 266 pp. [http: // saber. ucv. ve / handle / 123456789 / 1552]" type="book" year="2011">Provenzano, 2011:7275</bibRefCitation>
, Figs 4142.
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<materialsCitation id="318D3CB6F6589001FDB9432F15DBFDAE" collectingDate="1987-03-08" collectionCode="V" collectorName="B. Chernoff &amp; W. G. Saul &amp; H. Lopez-Rojas &amp; J. Fernandez &amp; O. Castillo &amp; M. E. Antonio" latitude="3.9083335" location="Orinoco River" longLatPrecision="21" longitude="-66.99166" pageId="3" pageNumber="318" specimenCode="V87-3" specimenCount="1" typeStatus="holotype">
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<typeStatus id="5E5E8849F6589001FDB9432F160DFDE6" box="[151,260,541,567]" pageId="3" pageNumber="318">Holotype</typeStatus>
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MBUCV-V-32954,
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SL,
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at sand island ca. 1/
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<quantity id="461D9B0EF6589001F932432F1350FDE6" box="[1052,1113,541,567]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="318" unit="km" value="2.0">2 km</quantity>
upstream from Guachapana
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(Guachipana), approx.
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,
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,
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,
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,
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,
<collectorName id="2C10533DF6589001F9ED43701272FD8A" box="[1219,1403,577,604]" pageId="3" pageNumber="318">H. López-Rojas</collectorName>
,
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,
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and
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, (
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).
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(
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from
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,
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)
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.
</paragraph>
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<emphasis id="B391EAF9F6589001FDB9465B161BF850" bold="true" box="[151,274,1897,1921]" pageId="3" pageNumber="318">FIGURE 1.</emphasis>
<taxonomicName id="46E54D68F6589001FC39465B16E5F850" authorityName="Provenzano-Rizzi" authorityYear="2023" box="[279,492,1897,1921]" family="Loricariidae" genus="Cheilonimata" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="318" phylum="Chordata" rank="species" species="minuta">
<emphasis id="B391EAF9F6589001FC39465B16E5F850" box="[279,492,1897,1921]" italics="true" pageId="3" pageNumber="318">Cheilonimata minuta</emphasis>
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, MBUCV-V-32954, 54.3 mm SL, holotype. Lateral, dorsal and ventral views. Scale bar 2 mm. Photos by C. Do Nascimiento.
</paragraph>
</caption>
<paragraph id="815A36EBF65F9006FDE941A4123EFE7C" blockId="4.[151,1437,150,933]" pageId="4" pageNumber="319">
<materialsCitation id="318D3CB6F65F9006FDE941A41445FF60" box="[199,844,150,177]" collectingDate="1987-03-08" collectionCode="ANSP" collectorName="B. Chernoff &amp; W. G. Saul &amp; H. Lopez-Rojas &amp; J. Fernandez &amp; O. Castillo &amp; M. E. Antonio" latitude="3.9083335" location="Orinoco River" longLatPrecision="21" longitude="-66.99166" pageId="4" pageNumber="319" specimenCode="ANSP 162156, 2" specimenCount="1">
MBUCV-V-29154,
<quantity id="461D9B0EF65F9006FC8D41A41500FF60" box="[419,521,150,177]" metricMagnitude="-2" metricUnit="m" metricValue="3.56" pageId="4" pageNumber="319" unit="mm" value="35.6">35.6 mm</quantity>
SL, same data as holotype.
</materialsCitation>
<materialsCitation id="318D3CB6F65F9006FE7B41A517F6FF04" collectingDate="1987-03-08" collectingDateMax="1987-03-20" collectingDateMin="1987-03-10" collectionCode="ANSP" collectorName="B. Chernoff &amp; W. G. Saul &amp; H. Lopez-Rojas &amp; J. Fernandez &amp; O. Castillo &amp; M. E. Antonio" county="Pamoni River" latitude="3.9083335" location="Orinoco River" longLatPrecision="21" longitude="-66.99166" municipality="Casiquiare River" pageId="4" pageNumber="319" specimenCode="ANSP 162156, 2" specimenCount="1">
<specimenCode id="D1439E90F65F9006FE7B41A514F3FF60" box="[853,1018,150,177]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="4" pageNumber="319">ANSP 162156</specimenCode>
,
<specimenCode id="D1439E90F65F9006F92741A51310FF60" box="[1033,1049,151,177]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="4" pageNumber="319">2</specimenCode>
,
<quantity id="461D9B0EF65F9006F90741A413C4FF60" box="[1065,1229,150,177]" metricMagnitude="-2" metricUnit="m" metricValue="4.285" metricValueMax="4.62" metricValueMin="3.95" pageId="4" pageNumber="319" unit="mm" value="42.85" valueMax="46.2" valueMin="39.5">39.546.2 mm</quantity>
SL, same data as holotype.
</materialsCitation>
<materialsCitation id="318D3CB6F65F9006FC2A418815DDFECC" collectingDate="1987-03-08" collectingDateMax="1987-03-20" collectingDateMin="1987-03-10" collectionCode="ANSP" collectorName="B. Chernoff &amp; W. G. Saul &amp; H. Lopez-Rojas &amp; J. Fernandez &amp; O. Castillo &amp; M. E. Antonio" county="Pamoni River" latitude="3.9083335" location="Orinoco River" longLatPrecision="21" longitude="-66.99166" municipality="Casiquiare River" pageId="4" pageNumber="319" specimenCode="ANSP 162156, 2" specimenCount="1">
MBUCV-V-17551, 2 alc, 1 c&amp;s,
<quantity id="461D9B0EF65F9006FF4741881401FF04" box="[617,776,186,213]" metricMagnitude="-2" metricUnit="m" metricValue="5.47" metricValueMax="5.4799999999999995" metricValueMin="5.46" pageId="4" pageNumber="319" unit="mm" value="54.7" valueMax="54.8" valueMin="54.6">54.654.8 mm</quantity>
SL,
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, at beach 1/2 h upstream from
<location id="843A6030F65F9006F808418917D4FF28" LSID="urn:lsid:plazi:treatment:094C87FDF6589005FDB94078161DFC8C:843A6030F65F9006F808418917D4FF28" county="Pamoni River" latitude="3.0969443" longLatPrecision="21" longitude="-65.89611" municipality="Casiquiare River" name="Temblador Island" pageId="4" pageNumber="319">Temblador Island</location>
, approx.
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,
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,
<date id="F55B102BF65F9006FF5241ED1424FF28" box="[636,813,223,249]" pageId="4" pageNumber="319" value="1987-03-10">
<collectingDate id="E51FE9C3F65F9006FF5241ED1424FF28" box="[636,813,223,249]" pageId="4" pageNumber="319" value="1987-03-10">10 March 1987</collectingDate>
</date>
,
<collectorName id="2C10533DF65F9006FE1241ED14CFFF28" box="[828,966,223,249]" pageId="4" pageNumber="319">B. Chernoff</collectorName>
,
<collectorName id="2C10533DF65F9006FEFD41ED1381FF28" box="[979,1160,223,249]" pageId="4" pageNumber="319">H. López-Rojas</collectorName>
,
<collectorName id="2C10533DF65F9006F9B841ED1205FF28" box="[1174,1292,223,249]" pageId="4" pageNumber="319">W.G. Saul</collectorName>
,
<collectorName id="2C10533DF65F9006F83541ED1291FF28" box="[1307,1432,223,249]" pageId="4" pageNumber="319">O. Castillo</collectorName>
,
<collectorName id="2C10533DF65F9006FDB940311639FECC" box="[151,304,259,285]" pageId="4" pageNumber="319">M.E. Antonio</collectorName>
,
<collectorName id="2C10533DF65F9006FC14403116CDFECC" box="[314,452,259,285]" pageId="4" pageNumber="319">J. Fernández</collectorName>
and
<collectorName id="2C10533DF65F9006FCD740311562FECC" box="[505,619,259,285]" pageId="4" pageNumber="319">J. Moreno</collectorName>
, (
<specimenCode id="D1439E90F65F9006FF53403115C1FECC" box="[637,712,259,285]" collectionCode="V" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="319" type="Museum">V87-6</specimenCode>
).
</materialsCitation>
<materialsCitation id="318D3CB6F65F9006FFF44030137EFEB4" collectingDate="1987-03-08" collectingDateMax="1987-03-20" collectingDateMin="1987-03-10" collectionCode="ANSP" collectorName="B. Chernoff &amp; W. G. Saul &amp; H. Lopez-Rojas &amp; J. Fernandez &amp; O. Castillo &amp; M. E. Antonio" county="Pamoni River" latitude="3.9083335" location="Orinoco River" longLatPrecision="21" longitude="-66.99166" municipality="Casiquiare River" pageId="4" pageNumber="319" specimenCode="ANSP 162156, 2" specimenCount="1">
MBUCV-V-17580,
<quantity id="461D9B0EF65F9006FE9C4031131AFECC" box="[946,1043,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="4.11" pageId="4" pageNumber="319" unit="mm" value="41.1">41.1 mm</quantity>
SL,
<collectingMunicipality id="613EAC91F65F9006F96940311208FECC" box="[1095,1281,259,285]" pageId="4" pageNumber="319">Casiquiare River</collectingMunicipality>
, approx.
<quantity id="461D9B0EF65F9006F84B40311295FECC" box="[1381,1436,259,285]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="319" unit="km" value="7.0">7 km</quantity>
downstream from the mouth of the
<collectingCounty id="683B4E67F65F9006FF33401515BEFE90" box="[541,695,295,321]" pageId="4" pageNumber="319">Pamoni River</collectingCounty>
, small caño at E side, approx.
<geoCoordinate id="E4D1502CF65F9006F92840141386FE90" box="[1030,1167,294,321]" degrees="03" direction="north" minutes="05" orientation="latitude" pageId="4" pageNumber="319" precision="15" seconds="49" value="3.0969443">03°0549”N</geoCoordinate>
,
<geoCoordinate id="E4D1502CF65F9006F9B44014122FFE90" box="[1178,1318,294,321]" degrees="65" direction="west" minutes="53" orientation="longitude" pageId="4" pageNumber="319" precision="15" seconds="46" value="-65.89611">65°5346”W</geoCoordinate>
,
<date id="F55B102BF65F9006F81C401517D9FEB4" pageId="4" pageNumber="319" value="1987-03-20">
<collectingDate id="E51FE9C3F65F9006F81C401517D9FEB4" pageId="4" pageNumber="319" value="1987-03-20">20 March 1987</collectingDate>
</date>
,
<collectorName id="2C10533DF65F9006FDF540791668FEB4" box="[219,353,331,357]" pageId="4" pageNumber="319">B. Chernoff</collectorName>
,
<collectorName id="2C10533DF65F9006FC4440791513FEB4" box="[362,538,331,357]" pageId="4" pageNumber="319">H. López-Rojas</collectorName>
,
<collectorName id="2C10533DF65F9006FF0B407915BBFEB4" box="[549,690,331,357]" pageId="4" pageNumber="319">J. Fernández</collectorName>
,
<collectorName id="2C10533DF65F9006FF924079143BFEB4" box="[700,818,331,357]" pageId="4" pageNumber="319">O. Castillo</collectorName>
and
<collectorName id="2C10533DF65F9006FE4940791309FEB4" box="[871,1024,331,357]" pageId="4" pageNumber="319">M.E. Antonio</collectorName>
, (
<specimenCode id="D1439E90F65F9006F93C40791363FEB4" box="[1042,1130,331,357]" collectionCode="V" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="319" type="Museum">V87-38</specimenCode>
).
</materialsCitation>
<materialsCitation id="318D3CB6F65F9006F9554079123AFE7C" collectingDate="1987-03-08" collectingDateMax="1987-03-20" collectingDateMin="1987-03-10" collectionCode="ANSP" collectorName="B. Chernoff &amp; W. G. Saul &amp; H. Lopez-Rojas &amp; J. Fernandez &amp; O. Castillo &amp; M. E. Antonio" county="Pamoni River" latitude="3.9083335" location="Orinoco River" longLatPrecision="21" longitude="-66.99166" municipality="Casiquiare River" pageId="4" pageNumber="319" specimenCode="ANSP 162156, 2" specimenCount="1">
<specimenCode id="D1439E90F65F9006F95540791200FEB4" box="[1147,1289,330,357]" collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="4" pageNumber="319" type="University or college">AUM 56760</specimenCode>
,
<specimenCode id="D1439E90F65F9006F83A4079122DFEB4" box="[1300,1316,331,357]" collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="4" pageNumber="319" type="University or college">4</specimenCode>
,
<quantity id="461D9B0EF65F9006F800407817CBFE58" metricMagnitude="-2" metricUnit="m" metricValue="6.984999999999999" metricValueMax="7.43" metricValueMin="6.54" pageId="4" pageNumber="319" unit="mm" value="69.85" valueMax="74.3" valueMin="65.4">65.474.3 mm</quantity>
SL,
<location id="843A6030F65F9006FDD8405D169CFE58" LSID="urn:lsid:plazi:treatment:094C87FDF6589005FDB94078161DFC8C:843A6030F65F9006FDD8405D169CFE58" box="[246,405,367,393]" latitude="3.7025" longLatPrecision="21" longitude="-66.954994" municipality="Ventuari River" name="Orinoco River" pageId="4" pageNumber="319">Orinoco River</location>
, beach in the middle of the river, upstream mouth of
<collectingMunicipality id="613EAC91F65F9006FEF8405D1371FE58" box="[982,1144,367,393]" pageId="4" pageNumber="319">Ventuari River</collectingMunicipality>
,
<locationDeviation id="90E82FC9F65F9006F9AC405D161FFE7C" location="San Fernando de Atabapo" pageId="4" pageNumber="319">
<quantity id="461D9B0EF65F9006F9AC405D13CEFE58" box="[1154,1223,367,393]" metricMagnitude="4" metricUnit="m" metricValue="9.1" pageId="4" pageNumber="319" unit="km" value="91.0">91 km</quantity>
ESE San Fernando de Atabapo
</locationDeviation>
,
<geoCoordinate id="E4D1502CF65F9006FC0F40A116A6FE7C" box="[289,431,403,429]" degrees="03" direction="north" minutes="42" orientation="latitude" pageId="4" pageNumber="319" precision="15" seconds="09" value="3.7025">03°4209” N</geoCoordinate>
<geoCoordinate id="E4D1502CF65F9006FC9B40A01541FE7C" box="[437,584,402,429]" degrees="66" direction="west" minutes="57" orientation="longitude" pageId="4" pageNumber="319" precision="15" seconds="18" value="-66.954994">66°5718” W</geoCoordinate>
,
<date id="F55B102BF65F9006FF7D40A115F7FE7C" box="[595,766,402,429]" pageId="4" pageNumber="319" value="2005-03-03">
<collectingDate id="E51FE9C3F65F9006FF7D40A115F7FE7C" box="[595,766,402,429]" pageId="4" pageNumber="319" value="2005-03-03">03 March 2005</collectingDate>
</date>
,
<collectorName id="2C10533DF65F9006FE2740A11481FE7C" box="[777,904,403,429]" pageId="4" pageNumber="319">N.K. Lujan</collectorName>
,
<collectorName id="2C10533DF65F9006FEBD40A11311FE7C" box="[915,1048,403,429]" pageId="4" pageNumber="319">D. Werneke</collectorName>
,
<collectorName id="2C10533DF65F9006F90C40A113A9FE7C" box="[1058,1184,403,429]" pageId="4" pageNumber="319">M.H. Sabaj</collectorName>
and
<collectorName id="2C10533DF65F9006F9FB40A1123AFE7C" box="[1237,1331,403,429]" pageId="4" pageNumber="319">M. Arce</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C9FF6560F65F9006FDE940841247FDB0" pageId="4" pageNumber="319" type="diagnosis">
<paragraph id="815A36EBF65F9006FDE940841247FDB0" blockId="4.[151,1437,150,933]" pageId="4" pageNumber="319">
<emphasis id="B391EAF9F65F9006FDE940841636FE01" bold="true" box="[199,319,438,464]" pageId="4" pageNumber="319">Diagnosis.</emphasis>
As given for genus. In addition,
<taxonomicName id="46E54D68F65F9006FF894085141AFE01" authorityName="Provenzano-Rizzi" authorityYear="2023" box="[679,787,439,465]" family="Loricariidae" genus="Cheilonimata" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="319" phylum="Chordata" rank="species" species="minuta">
<emphasis id="B391EAF9F65F9006FF894085141AFE01" box="[679,787,439,465]" italics="true" pageId="4" pageNumber="319">C. minuta</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="46E54D68F65F9006FEF740851356FE00" authorityName="Provenzano-Rizzi" authorityYear="2023" baseAuthorityName="Crispim-Rodrigues, Bernt, Waltz, Silva, Benine, Oliveira, Covain, and Roxo" baseAuthorityYear="2023" box="[985,1119,439,465]" family="Loricariidae" genus="Cheilonimata" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="319" phylum="Chordata" rank="species" species="papillosa">
<emphasis id="B391EAF9F65F9006FEF740851356FE00" box="[985,1119,439,465]" italics="true" pageId="4" pageNumber="319">C. papillosa</emphasis>
</taxonomicName>
, and can be distinguished by maxillary barbel reaches gill opening
<emphasis id="B391EAF9F65F9006FF1B40E9155AFE25" box="[565,595,475,500]" italics="true" pageId="4" pageNumber="319">vs.</emphasis>
maxillary barbel reaches middle of pectoral-fin in
<taxonomicName id="46E54D68F65F9006F95140E9120DFE24" authorityName="Provenzano-Rizzi" authorityYear="2023" baseAuthorityName="Crispim-Rodrigues, Bernt, Waltz, Silva, Benine, Oliveira, Covain, and Roxo" baseAuthorityYear="2023" box="[1151,1284,475,501]" family="Loricariidae" genus="Cheilonimata" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="319" phylum="Chordata" rank="species" species="papillosa">
<emphasis id="B391EAF9F65F9006F95140E9120DFE24" box="[1151,1284,475,501]" italics="true" pageId="4" pageNumber="319">C. papillosa</emphasis>
</taxonomicName>
; lateral series plates 2932 (mode 32)
<emphasis id="B391EAF9F65F9006FC8640CD16CFFDC9" box="[424,454,511,536]" italics="true" pageId="4" pageNumber="319">vs.</emphasis>
lateral series plates 2729 and pectoral-fin with two or three transversal dark stripes on unbranched and branched rays
<emphasis id="B391EAF9F65F9006FCCB4311150AFDED" box="[485,515,547,572]" italics="true" pageId="4" pageNumber="319">vs.</emphasis>
pectoral-fin with one wide stripe on unbranched and branched rays. Other differences related to the number teeth in upper jaw and body color pattern need to be confirmed with direct examination.
</paragraph>
</subSubSection>
<subSubSection id="C9FF6560F65F9005FDE94358157CFCC4" lastPageId="7" lastPageNumber="322" pageId="4" pageNumber="319" type="description">
<paragraph id="815A36EBF65F9006FDE943581678FC74" blockId="4.[151,1437,150,933]" pageId="4" pageNumber="319">
<emphasis id="B391EAF9F65F9006FDE943581650FD55" bold="true" box="[199,345,618,644]" pageId="4" pageNumber="319">Description.</emphasis>
Morphometric data is presented in
<tableCitation id="CC670350F65F9006FFC743591437FD54" box="[745,830,619,645]" captionStart="TABLE 1" captionStartId="4.[151,239,974,998]" captionTargetPageId="4" captionText="TABLE 1. Morphometric data of Cheilonimata minuta expressed as percentage of standard length (n=12)." pageId="4" pageNumber="319">Table 1</tableCitation>
. Specimens small with fragile or delicate appearance. In lateral view, head and body very depressed, caudal peduncle without adipose-fin; body deeper at eye level and gradually tapering towards caudal-fin origin (
<figureCitation id="19DE2A6EF65F9006FFB9438115D1FD1C" box="[663,728,691,717]" captionStart="FIGURE 1" captionStartId="3.[151,250,1897,1921]" captionTargetBox="[183,1405,698,1872]" captionTargetId="figure-114@3.[183,1405,698,1872]" captionTargetPageId="3" captionText="FIGURE 1. Cheilonimata minuta, MBUCV-V-32954, 54.3 mm SL, holotype. Lateral, dorsal and ventral views. Scale bar 2 mm. Photos by C. Do Nascimiento." figureDoi="http://doi.org/10.5281/zenodo.8142276" httpUri="https://zenodo.org/record/8142276/files/figure.png" pageId="4" pageNumber="319">Fig. 1</figureCitation>
). Dorsal profile of body from tip of snout through anterior eye border straight and ascending with 45° slope approximately, from this point to caudal-fin origin descending straight. Ventral profile of head and body flat and straight (
<figureCitation id="19DE2A6EF65F9006FFE143C91418FCC4" box="[719,785,763,789]" captionStart="FIGURE 1" captionStartId="3.[151,250,1897,1921]" captionTargetBox="[183,1405,698,1872]" captionTargetId="figure-114@3.[183,1405,698,1872]" captionTargetPageId="3" captionText="FIGURE 1. Cheilonimata minuta, MBUCV-V-32954, 54.3 mm SL, holotype. Lateral, dorsal and ventral views. Scale bar 2 mm. Photos by C. Do Nascimiento." figureDoi="http://doi.org/10.5281/zenodo.8142276" httpUri="https://zenodo.org/record/8142276/files/figure.png" pageId="4" pageNumber="319">Fig. 1</figureCitation>
). In dorsal view, head contour elliptical with sides gently convex; snout not projected, tip rounded; body wider at branchial opening or pectoral-fin insertion and tapering gradually and continuously to caudal-fin origin (
<figureCitation id="19DE2A6EF65F9006FF96427115F3FC8C" box="[696,762,835,861]" captionStart="FIGURE 1" captionStartId="3.[151,250,1897,1921]" captionTargetBox="[183,1405,698,1872]" captionTargetId="figure-114@3.[183,1405,698,1872]" captionTargetPageId="3" captionText="FIGURE 1. Cheilonimata minuta, MBUCV-V-32954, 54.3 mm SL, holotype. Lateral, dorsal and ventral views. Scale bar 2 mm. Photos by C. Do Nascimiento." figureDoi="http://doi.org/10.5281/zenodo.8142276" httpUri="https://zenodo.org/record/8142276/files/figure.png" pageId="4" pageNumber="319">Fig. 1</figureCitation>
). Pectoral-fin insertion slightly posterior to vertical through posterior margin of orbit. Dorsal-fin origin opposite to pelvic-fin insertion. Anal-fin origin at lateral plate number nine or ten (
<figureCitation id="19DE2A6EF65F9006FC3042B91669FC74" box="[286,352,907,933]" captionStart="FIGURE 1" captionStartId="3.[151,250,1897,1921]" captionTargetBox="[183,1405,698,1872]" captionTargetId="figure-114@3.[183,1405,698,1872]" captionTargetPageId="3" captionText="FIGURE 1. Cheilonimata minuta, MBUCV-V-32954, 54.3 mm SL, holotype. Lateral, dorsal and ventral views. Scale bar 2 mm. Photos by C. Do Nascimiento." figureDoi="http://doi.org/10.5281/zenodo.8142276" httpUri="https://zenodo.org/record/8142276/files/figure.png" pageId="4" pageNumber="319">Fig. 1</figureCitation>
).
</paragraph>
<caption id="D59A6663F65F9006FDB942FC1201FC37" box="[151,1288,974,998]" pageId="4" pageNumber="319" startId="4.[151,239,974,998]" targetBox="[163,1373,1013,1995]" targetIsTable="true" targetPageId="4" targetType="table">
<paragraph id="815A36EBF65F9006FDB942FC1201FC37" blockId="4.[151,1288,974,998]" box="[151,1288,974,998]" pageId="4" pageNumber="319">
<emphasis id="B391EAF9F65F9006FDB942FC1603FC37" bold="true" box="[151,266,974,998]" pageId="4" pageNumber="319">TABLE 1.</emphasis>
Morphometric data of
<taxonomicName id="46E54D68F65F9006FF2C42FC15EAFC34" authorityName="Provenzano-Rizzi" authorityYear="2023" box="[514,739,974,998]" family="Loricariidae" genus="Cheilonimata" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="319" phylum="Chordata" rank="species" species="minuta">
<emphasis id="B391EAF9F65F9006FF2C42FC15EAFC34" box="[514,739,974,998]" italics="true" pageId="4" pageNumber="319">Cheilonimata minuta</emphasis>
</taxonomicName>
expressed as percentage of standard length (n=12).
</paragraph>
</caption>
<paragraph id="815A36EBF65F9006FD8D42C7125EF81A" pageId="4" pageNumber="319">
<table id="F3E5C44BF65F6FFDFD8D42C71254F81A" box="[163,1373,1013,1995]" gridcols="6" gridrows="24" pageId="4" pageNumber="319">
<tr id="3FD534A9F65F6FFDFD8D42C71254FBDF" box="[163,1373,1013,1038]" gridrow="0" pageId="4" pageNumber="319">
<th id="7C045DD5F65F6FFDFD8D42C716A4FBDF" box="[163,429,1013,1038]" gridcol="0" gridrow="0" pageId="4" pageNumber="319">
<emphasis id="B391EAF9F65F9006FD8D42C71615FBDC" bold="true" box="[163,284,1013,1037]" pageId="4" pageNumber="319">Characters</emphasis>
</th>
<th id="7C045DD5F65F6FFDFF7242C715B6FBDF" box="[604,703,1013,1038]" gridcol="1" gridrow="0" pageId="4" pageNumber="319">
<emphasis id="B391EAF9F65F9006FF7242C415B6FBDF" bold="true" box="[604,703,1014,1038]" pageId="4" pageNumber="319">Holotype</emphasis>
</th>
<th id="7C045DD5F65F6FFDFE0042C71465FBDF" box="[814,876,1013,1038]" gridcol="2" gridrow="0" pageId="4" pageNumber="319">
<emphasis id="B391EAF9F65F9006FE0042C41465FBDF" bold="true" box="[814,876,1014,1038]" pageId="4" pageNumber="319">Mean</emphasis>
</th>
<th id="7C045DD5F65F6FFDFEF242C71304FBDF" box="[988,1037,1013,1038]" gridcol="3" gridrow="0" pageId="4" pageNumber="319">
<emphasis id="B391EAF9F65F9006FEF242C71304FBDC" bold="true" box="[988,1037,1013,1037]" pageId="4" pageNumber="319">STD</emphasis>
</th>
<th id="7C045DD5F65F6FFDF95042C713B8FBDF" box="[1150,1201,1013,1038]" gridcol="4" gridrow="0" pageId="4" pageNumber="319">
<emphasis id="B391EAF9F65F9006F95042C413A2FBDF" bold="true" box="[1150,1195,1014,1038]" pageId="4" pageNumber="319">Min</emphasis>
.
</th>
<th id="7C045DD5F65F6FFDF80842C71254FBDF" box="[1318,1373,1013,1038]" gridcol="5" gridrow="0" pageId="4" pageNumber="319">
<emphasis id="B391EAF9F65F9006F80842C4125EFBDF" bold="true" box="[1318,1367,1014,1038]" pageId="4" pageNumber="319">Max</emphasis>
.
</th>
</tr>
<tr id="3FD534A9F65F6FFDFD8D452D1254FBE9" box="[163,1373,1055,1080]" gridrow="1" pageId="4" pageNumber="319" rowspan-2="1" rowspan-3="1">
<th id="7C045DD5F65F6FFDFD8D452D16A4FBE9" box="[163,429,1055,1080]" gridcol="0" gridrow="1" pageId="4" pageNumber="319">SL (mm)</th>
<td id="7C045DD5F65F6FFDFF72452D15B6FBE9" box="[604,703,1055,1080]" gridcol="1" gridrow="1" pageId="4" pageNumber="319">54.3</td>
<td id="7C045DD5F65F6FFDF950452D13B8FBE9" box="[1150,1201,1055,1080]" gridcol="4" gridrow="1" pageId="4" pageNumber="319">35.6</td>
<td id="7C045DD5F65F6FFDF808452D1254FBE9" box="[1318,1373,1055,1080]" gridcol="5" gridrow="1" pageId="4" pageNumber="319">74.3</td>
</tr>
<tr id="3FD534A9F65F6FFDFD8D457B1254FBB0" box="[163,1373,1097,1121]" gridrow="2" pageId="4" pageNumber="319">
<th id="7C045DD5F65F6FFDFD8D457B16A4FBB0" box="[163,429,1097,1121]" gridcol="0" gridrow="2" pageId="4" pageNumber="319">Head length</th>
<td id="7C045DD5F65F6FFDFF72457B15B6FBB0" box="[604,703,1097,1121]" gridcol="1" gridrow="2" pageId="4" pageNumber="319">25.2</td>
<td id="7C045DD5F65F6FFDFE00457B1465FBB0" box="[814,876,1097,1121]" gridcol="2" gridrow="2" pageId="4" pageNumber="319">23.9</td>
<td id="7C045DD5F65F6FFDFEF2457B1304FBB0" box="[988,1037,1097,1121]" gridcol="3" gridrow="2" pageId="4" pageNumber="319">2.2</td>
<td id="7C045DD5F65F6FFDF950457B13B8FBB0" box="[1150,1201,1097,1121]" gridcol="4" gridrow="2" pageId="4" pageNumber="319">21.1</td>
<td id="7C045DD5F65F6FFDF808457B1254FBB0" box="[1318,1373,1097,1121]" gridcol="5" gridrow="2" pageId="4" pageNumber="319">26.5</td>
</tr>
<tr id="3FD534A9F65F6FFDFD8D45411254FB5A" box="[163,1373,1139,1163]" gridrow="3" pageId="4" pageNumber="319">
<th id="7C045DD5F65F6FFDFD8D454116A4FB5A" box="[163,429,1139,1163]" gridcol="0" gridrow="3" pageId="4" pageNumber="319">Predorsal length</th>
<td id="7C045DD5F65F6FFDFF72454115B6FB5A" box="[604,703,1139,1163]" gridcol="1" gridrow="3" pageId="4" pageNumber="319">33.2</td>
<td id="7C045DD5F65F6FFDFE0045411465FB5A" box="[814,876,1139,1163]" gridcol="2" gridrow="3" pageId="4" pageNumber="319">33.4</td>
<td id="7C045DD5F65F6FFDFEF245411304FB5A" box="[988,1037,1139,1163]" gridcol="3" gridrow="3" pageId="4" pageNumber="319">0.8</td>
<td id="7C045DD5F65F6FFDF950454113B8FB5A" box="[1150,1201,1139,1163]" gridcol="4" gridrow="3" pageId="4" pageNumber="319">32.3</td>
<td id="7C045DD5F65F6FFDF80845411254FB5A" box="[1318,1373,1139,1163]" gridcol="5" gridrow="3" pageId="4" pageNumber="319">35.4</td>
</tr>
<tr id="3FD534A9F65F6FFDFD8D45AE1254FB64" box="[163,1373,1180,1205]" gridrow="4" pageId="4" pageNumber="319">
<th id="7C045DD5F65F6FFDFD8D45AE16A4FB64" box="[163,429,1180,1205]" gridcol="0" gridrow="4" pageId="4" pageNumber="319">Postdorsal length</th>
<td id="7C045DD5F65F6FFDFF7245AE15B6FB64" box="[604,703,1180,1205]" gridcol="1" gridrow="4" pageId="4" pageNumber="319">59.7</td>
<td id="7C045DD5F65F6FFDFE0045AE1465FB64" box="[814,876,1180,1205]" gridcol="2" gridrow="4" pageId="4" pageNumber="319">58.3</td>
<td id="7C045DD5F65F6FFDFEF245AE1304FB64" box="[988,1037,1180,1205]" gridcol="3" gridrow="4" pageId="4" pageNumber="319">1.7</td>
<td id="7C045DD5F65F6FFDF95045AE13B8FB64" box="[1150,1201,1180,1205]" gridcol="4" gridrow="4" pageId="4" pageNumber="319">55.8</td>
<td id="7C045DD5F65F6FFDF80845AE1254FB64" box="[1318,1373,1180,1205]" gridcol="5" gridrow="4" pageId="4" pageNumber="319">60.4</td>
</tr>
<tr id="3FD534A9F65F6FFDFD8D45F41254FB0F" box="[163,1373,1222,1246]" gridrow="5" pageId="4" pageNumber="319">
<th id="7C045DD5F65F6FFDFD8D45F416A4FB0F" box="[163,429,1222,1246]" gridcol="0" gridrow="5" pageId="4" pageNumber="319">Snout-anus length</th>
<td id="7C045DD5F65F6FFDFF7245F415B6FB0F" box="[604,703,1222,1246]" gridcol="1" gridrow="5" pageId="4" pageNumber="319">35.2</td>
<td id="7C045DD5F65F6FFDFE0045F41465FB0F" box="[814,876,1222,1246]" gridcol="2" gridrow="5" pageId="4" pageNumber="319">36.2</td>
<td id="7C045DD5F65F6FFDFEF245F41304FB0F" box="[988,1037,1222,1246]" gridcol="3" gridrow="5" pageId="4" pageNumber="319">1.2</td>
<td id="7C045DD5F65F6FFDF95045F413B8FB0F" box="[1150,1201,1222,1246]" gridcol="4" gridrow="5" pageId="4" pageNumber="319">34.4</td>
<td id="7C045DD5F65F6FFDF80845F41254FB0F" box="[1318,1373,1222,1246]" gridcol="5" gridrow="5" pageId="4" pageNumber="319">37.7</td>
</tr>
<tr id="3FD534A9F65F6FFDFD8D45DD1254FAD9" box="[163,1373,1263,1288]" gridrow="6" pageId="4" pageNumber="319">
<th id="7C045DD5F65F6FFDFD8D45DD16A4FAD9" box="[163,429,1263,1288]" gridcol="0" gridrow="6" pageId="4" pageNumber="319">Preanal length</th>
<td id="7C045DD5F65F6FFDFF7245DD15B6FAD9" box="[604,703,1263,1288]" gridcol="1" gridrow="6" pageId="4" pageNumber="319">45.7</td>
<td id="7C045DD5F65F6FFDFE0045DD1465FAD9" box="[814,876,1263,1288]" gridcol="2" gridrow="6" pageId="4" pageNumber="319">47.2</td>
<td id="7C045DD5F65F6FFDFEF245DD1304FAD9" box="[988,1037,1263,1288]" gridcol="3" gridrow="6" pageId="4" pageNumber="319">1.3</td>
<td id="7C045DD5F65F6FFDF95045DD13B8FAD9" box="[1150,1201,1263,1288]" gridcol="4" gridrow="6" pageId="4" pageNumber="319">44.8</td>
<td id="7C045DD5F65F6FFDF80845DD1254FAD9" box="[1318,1373,1263,1288]" gridcol="5" gridrow="6" pageId="4" pageNumber="319">48.7</td>
</tr>
<tr id="3FD534A9F65F6FFDFD8D442B1254FAE0" box="[163,1373,1305,1329]" gridrow="7" pageId="4" pageNumber="319">
<th id="7C045DD5F65F6FFDFD8D442B16A4FAE0" box="[163,429,1305,1329]" gridcol="0" gridrow="7" pageId="4" pageNumber="319">Postanal length</th>
<td id="7C045DD5F65F6FFDFF72442B15B6FAE0" box="[604,703,1305,1329]" gridcol="1" gridrow="7" pageId="4" pageNumber="319">48.6</td>
<td id="7C045DD5F65F6FFDFE00442B1465FAE0" box="[814,876,1305,1329]" gridcol="2" gridrow="7" pageId="4" pageNumber="319">49.0</td>
<td id="7C045DD5F65F6FFDFEF2442B1304FAE0" box="[988,1037,1305,1329]" gridcol="3" gridrow="7" pageId="4" pageNumber="319">1.7</td>
<td id="7C045DD5F65F6FFDF950442B13B8FAE0" box="[1150,1201,1305,1329]" gridcol="4" gridrow="7" pageId="4" pageNumber="319">46.4</td>
<td id="7C045DD5F65F6FFDF808442B1254FAE0" box="[1318,1373,1305,1329]" gridcol="5" gridrow="7" pageId="4" pageNumber="319">52.2</td>
</tr>
<tr id="3FD534A9F65F6FFDFD8D44711254FA8A" box="[163,1373,1347,1371]" gridrow="8" pageId="4" pageNumber="319">
<th id="7C045DD5F65F6FFDFD8D447116A4FA8A" box="[163,429,1347,1371]" gridcol="0" gridrow="8" pageId="4" pageNumber="319">Prepectoral length</th>
<td id="7C045DD5F65F6FFDFF72447115B6FA8A" box="[604,703,1347,1371]" gridcol="1" gridrow="8" pageId="4" pageNumber="319">16.7</td>
<td id="7C045DD5F65F6FFDFE0044711465FA8A" box="[814,876,1347,1371]" gridcol="2" gridrow="8" pageId="4" pageNumber="319">18.1</td>
<td id="7C045DD5F65F6FFDFEF244711304FA8A" box="[988,1037,1347,1371]" gridcol="3" gridrow="8" pageId="4" pageNumber="319">1.0</td>
<td id="7C045DD5F65F6FFDF950447113B8FA8A" box="[1150,1201,1347,1371]" gridcol="4" gridrow="8" pageId="4" pageNumber="319">16.7</td>
<td id="7C045DD5F65F6FFDF80844711254FA8A" box="[1318,1373,1347,1371]" gridcol="5" gridrow="8" pageId="4" pageNumber="319">19.2</td>
</tr>
<tr id="3FD534A9F65F6FFDFD8D445E1254FA54" box="[163,1373,1388,1413]" gridrow="9" pageId="4" pageNumber="319">
<th id="7C045DD5F65F6FFDFD8D445E16A4FA54" box="[163,429,1388,1413]" gridcol="0" gridrow="9" pageId="4" pageNumber="319">Prepelvic length</th>
<td id="7C045DD5F65F6FFDFF72445E15B6FA54" box="[604,703,1388,1413]" gridcol="1" gridrow="9" pageId="4" pageNumber="319">30.3</td>
<td id="7C045DD5F65F6FFDFE00445E1465FA54" box="[814,876,1388,1413]" gridcol="2" gridrow="9" pageId="4" pageNumber="319">31.0</td>
<td id="7C045DD5F65F6FFDFEF2445E1304FA54" box="[988,1037,1388,1413]" gridcol="3" gridrow="9" pageId="4" pageNumber="319">0.8</td>
<td id="7C045DD5F65F6FFDF950445E13B8FA54" box="[1150,1201,1388,1413]" gridcol="4" gridrow="9" pageId="4" pageNumber="319">29.8</td>
<td id="7C045DD5F65F6FFDF808445E1254FA54" box="[1318,1373,1388,1413]" gridcol="5" gridrow="9" pageId="4" pageNumber="319">32.0</td>
</tr>
<tr id="3FD534A9F65F6FFDFD8D44A41254FA7F" box="[163,1373,1430,1454]" gridrow="10" pageId="4" pageNumber="319">
<th id="7C045DD5F65F6FFDFD8D44A416A4FA7F" box="[163,429,1430,1454]" gridcol="0" gridrow="10" pageId="4" pageNumber="319">Thoracic length</th>
<td id="7C045DD5F65F6FFDFF7244A415B6FA7F" box="[604,703,1430,1454]" gridcol="1" gridrow="10" pageId="4" pageNumber="319">14.7</td>
<td id="7C045DD5F65F6FFDFE0044A41465FA7F" box="[814,876,1430,1454]" gridcol="2" gridrow="10" pageId="4" pageNumber="319">15.9</td>
<td id="7C045DD5F65F6FFDFEF244A41304FA7F" box="[988,1037,1430,1454]" gridcol="3" gridrow="10" pageId="4" pageNumber="319">1.0</td>
<td id="7C045DD5F65F6FFDF95044A413B8FA7F" box="[1150,1201,1430,1454]" gridcol="4" gridrow="10" pageId="4" pageNumber="319">14.4</td>
<td id="7C045DD5F65F6FFDF80844A41254FA7F" box="[1318,1373,1430,1454]" gridcol="5" gridrow="10" pageId="4" pageNumber="319">17.7</td>
</tr>
<tr id="3FD534A9F65F6FFDFD8D448D1254FA09" box="[163,1373,1471,1496]" gridrow="11" pageId="4" pageNumber="319">
<th id="7C045DD5F65F6FFDFD8D448D16A4FA09" box="[163,429,1471,1496]" gridcol="0" gridrow="11" pageId="4" pageNumber="319">Abdominal length</th>
<td id="7C045DD5F65F6FFDFF72448D15B6FA09" box="[604,703,1471,1496]" gridcol="1" gridrow="11" pageId="4" pageNumber="319">17.5</td>
<td id="7C045DD5F65F6FFDFE00448D1465FA09" box="[814,876,1471,1496]" gridcol="2" gridrow="11" pageId="4" pageNumber="319">17.2</td>
<td id="7C045DD5F65F6FFDFEF2448D1304FA09" box="[988,1037,1471,1496]" gridcol="3" gridrow="11" pageId="4" pageNumber="319">0.7</td>
<td id="7C045DD5F65F6FFDF950448D13B8FA09" box="[1150,1201,1471,1496]" gridcol="4" gridrow="11" pageId="4" pageNumber="319">16.0</td>
<td id="7C045DD5F65F6FFDF808448D1254FA09" box="[1318,1373,1471,1496]" gridcol="5" gridrow="11" pageId="4" pageNumber="319">18.3</td>
</tr>
<tr id="3FD534A9F65F6FFDFD8D44DB1254F9D0" box="[163,1373,1513,1537]" gridrow="12" pageId="4" pageNumber="319">
<th id="7C045DD5F65F6FFDFD8D44DB16A4F9D0" box="[163,429,1513,1537]" gridcol="0" gridrow="12" pageId="4" pageNumber="319">Cleithral width</th>
<td id="7C045DD5F65F6FFDFF7244DB15B6F9D0" box="[604,703,1513,1537]" gridcol="1" gridrow="12" pageId="4" pageNumber="319">19.1</td>
<td id="7C045DD5F65F6FFDFE0044DB1465F9D0" box="[814,876,1513,1537]" gridcol="2" gridrow="12" pageId="4" pageNumber="319">19.0</td>
<td id="7C045DD5F65F6FFDFEF244DB1304F9D0" box="[988,1037,1513,1537]" gridcol="3" gridrow="12" pageId="4" pageNumber="319">1.6</td>
<td id="7C045DD5F65F6FFDF95044DB13B8F9D0" box="[1150,1201,1513,1537]" gridcol="4" gridrow="12" pageId="4" pageNumber="319">15.9</td>
<td id="7C045DD5F65F6FFDF80844DB1254F9D0" box="[1318,1373,1513,1537]" gridcol="5" gridrow="12" pageId="4" pageNumber="319">21.1</td>
</tr>
<tr id="3FD534A9F65F6FFDFD8D47211254F9FA" box="[163,1373,1555,1579]" gridrow="13" pageId="4" pageNumber="319">
<th id="7C045DD5F65F6FFDFD8D472116A4F9FA" box="[163,429,1555,1579]" gridcol="0" gridrow="13" pageId="4" pageNumber="319">Body width at pelvic fin</th>
<td id="7C045DD5F65F6FFDFF72472115B6F9FA" box="[604,703,1555,1579]" gridcol="1" gridrow="13" pageId="4" pageNumber="319">12.9</td>
<td id="7C045DD5F65F6FFDFE0047211465F9FA" box="[814,876,1555,1579]" gridcol="2" gridrow="13" pageId="4" pageNumber="319">13.5</td>
<td id="7C045DD5F65F6FFDFEF247211304F9FA" box="[988,1037,1555,1579]" gridcol="3" gridrow="13" pageId="4" pageNumber="319">0.5</td>
<td id="7C045DD5F65F6FFDF950472113B8F9FA" box="[1150,1201,1555,1579]" gridcol="4" gridrow="13" pageId="4" pageNumber="319">12.9</td>
<td id="7C045DD5F65F6FFDF80847211254F9FA" box="[1318,1373,1555,1579]" gridcol="5" gridrow="13" pageId="4" pageNumber="319">14.2</td>
</tr>
<tr id="3FD534A9F65F6FFDFD8D470E1254F984" box="[163,1373,1596,1621]" gridrow="14" pageId="4" pageNumber="319">
<th id="7C045DD5F65F6FFDFD8D470E16A4F984" box="[163,429,1596,1621]" gridcol="0" gridrow="14" pageId="4" pageNumber="319">Body width at anal fin</th>
<td id="7C045DD5F65F6FFDFF72470E15B6F984" box="[604,703,1596,1621]" gridcol="1" gridrow="14" pageId="4" pageNumber="319">11.3</td>
<td id="7C045DD5F65F6FFDFE00470E1465F984" box="[814,876,1596,1621]" gridcol="2" gridrow="14" pageId="4" pageNumber="319">11.5</td>
<td id="7C045DD5F65F6FFDFEF2470E1304F984" box="[988,1037,1596,1621]" gridcol="3" gridrow="14" pageId="4" pageNumber="319">0.5</td>
<td id="7C045DD5F65F6FFDF950470E13B8F984" box="[1150,1201,1596,1621]" gridcol="4" gridrow="14" pageId="4" pageNumber="319">10.7</td>
<td id="7C045DD5F65F6FFDF808470E1254F984" box="[1318,1373,1596,1621]" gridcol="5" gridrow="14" pageId="4" pageNumber="319">12.0</td>
</tr>
<tr id="3FD534A9F65F6FFDFD8D47541254F9AF" box="[163,1373,1638,1662]" gridrow="15" pageId="4" pageNumber="319">
<th id="7C045DD5F65F6FFDFD8D475416A4F9AF" box="[163,429,1638,1662]" gridcol="0" gridrow="15" pageId="4" pageNumber="319">Head depth</th>
<td id="7C045DD5F65F6FFDFF72475415B6F9AF" box="[604,703,1638,1662]" gridcol="1" gridrow="15" pageId="4" pageNumber="319">7.4</td>
<td id="7C045DD5F65F6FFDFE0047541465F9AF" box="[814,876,1638,1662]" gridcol="2" gridrow="15" pageId="4" pageNumber="319">6.8</td>
<td id="7C045DD5F65F6FFDFEF247541304F9AF" box="[988,1037,1638,1662]" gridcol="3" gridrow="15" pageId="4" pageNumber="319">0.9</td>
<td id="7C045DD5F65F6FFDF950475413B8F9AF" box="[1150,1201,1638,1662]" gridcol="4" gridrow="15" pageId="4" pageNumber="319">5.4</td>
<td id="7C045DD5F65F6FFDF80847541254F9AF" box="[1318,1373,1638,1662]" gridcol="5" gridrow="15" pageId="4" pageNumber="319">8.2</td>
</tr>
<tr id="3FD534A9F65F6FFDFD8D47A21254F979" box="[163,1373,1680,1704]" gridrow="16" pageId="4" pageNumber="319">
<th id="7C045DD5F65F6FFDFD8D47A216A4F979" box="[163,429,1680,1704]" gridcol="0" gridrow="16" pageId="4" pageNumber="319">Snout length</th>
<td id="7C045DD5F65F6FFDFF7247A215B6F979" box="[604,703,1680,1704]" gridcol="1" gridrow="16" pageId="4" pageNumber="319">11.2</td>
<td id="7C045DD5F65F6FFDFE0047A21465F979" box="[814,876,1680,1704]" gridcol="2" gridrow="16" pageId="4" pageNumber="319">11.8</td>
<td id="7C045DD5F65F6FFDFEF247A21304F979" box="[988,1037,1680,1704]" gridcol="3" gridrow="16" pageId="4" pageNumber="319">0.7</td>
<td id="7C045DD5F65F6FFDF95047A213B8F979" box="[1150,1201,1680,1704]" gridcol="4" gridrow="16" pageId="4" pageNumber="319">11.0</td>
<td id="7C045DD5F65F6FFDF80847A21254F979" box="[1318,1373,1680,1704]" gridcol="5" gridrow="16" pageId="4" pageNumber="319">13.4</td>
</tr>
<tr id="3FD534A9F65F6FFDFD8D478B1254F900" box="[163,1373,1721,1745]" gridrow="17" pageId="4" pageNumber="319">
<th id="7C045DD5F65F6FFDFD8D478B16A4F900" box="[163,429,1721,1745]" gridcol="0" gridrow="17" pageId="4" pageNumber="319">Interorbital width</th>
<td id="7C045DD5F65F6FFDFF72478B15B6F900" box="[604,703,1721,1745]" gridcol="1" gridrow="17" pageId="4" pageNumber="319">3.5</td>
<td id="7C045DD5F65F6FFDFE00478B1465F900" box="[814,876,1721,1745]" gridcol="2" gridrow="17" pageId="4" pageNumber="319">4.0</td>
<td id="7C045DD5F65F6FFDFEF2478B1304F900" box="[988,1037,1721,1745]" gridcol="3" gridrow="17" pageId="4" pageNumber="319">1.0</td>
<td id="7C045DD5F65F6FFDF950478B13B8F900" box="[1150,1201,1721,1745]" gridcol="4" gridrow="17" pageId="4" pageNumber="319">3.2</td>
<td id="7C045DD5F65F6FFDF808478B1254F900" box="[1318,1373,1721,1745]" gridcol="5" gridrow="17" pageId="4" pageNumber="319">6.8</td>
</tr>
<tr id="3FD534A9F65F6FFDFD8D47D11254F92A" box="[163,1373,1763,1787]" gridrow="18" pageId="4" pageNumber="319">
<th id="7C045DD5F65F6FFDFD8D47D116A4F92A" box="[163,429,1763,1787]" gridcol="0" gridrow="18" pageId="4" pageNumber="319">Maximun orbital diameter</th>
<td id="7C045DD5F65F6FFDFF7247D115B6F92A" box="[604,703,1763,1787]" gridcol="1" gridrow="18" pageId="4" pageNumber="319">4.7</td>
<td id="7C045DD5F65F6FFDFE0047D11465F92A" box="[814,876,1763,1787]" gridcol="2" gridrow="18" pageId="4" pageNumber="319">5.3</td>
<td id="7C045DD5F65F6FFDFEF247D11304F92A" box="[988,1037,1763,1787]" gridcol="3" gridrow="18" pageId="4" pageNumber="319">0.9</td>
<td id="7C045DD5F65F6FFDF95047D113B8F92A" box="[1150,1201,1763,1787]" gridcol="4" gridrow="18" pageId="4" pageNumber="319">4.4</td>
<td id="7C045DD5F65F6FFDF80847D11254F92A" box="[1318,1373,1763,1787]" gridcol="5" gridrow="18" pageId="4" pageNumber="319">7.7</td>
</tr>
<tr id="3FD534A9F65F6FFDFD8D463E1254F8F4" box="[163,1373,1804,1829]" gridrow="19" pageId="4" pageNumber="319">
<th id="7C045DD5F65F6FFDFD8D463E16A4F8F4" box="[163,429,1804,1829]" gridcol="0" gridrow="19" pageId="4" pageNumber="319">Dorsal fin base</th>
<td id="7C045DD5F65F6FFDFF72463E15B6F8F4" box="[604,703,1804,1829]" gridcol="1" gridrow="19" pageId="4" pageNumber="319">6.6</td>
<td id="7C045DD5F65F6FFDFE00463E1465F8F4" box="[814,876,1804,1829]" gridcol="2" gridrow="19" pageId="4" pageNumber="319">7.7</td>
<td id="7C045DD5F65F6FFDFEF2463E1304F8F4" box="[988,1037,1804,1829]" gridcol="3" gridrow="19" pageId="4" pageNumber="319">1.1</td>
<td id="7C045DD5F65F6FFDF950463E13B8F8F4" box="[1150,1201,1804,1829]" gridcol="4" gridrow="19" pageId="4" pageNumber="319">6.1</td>
<td id="7C045DD5F65F6FFDF808463E1254F8F4" box="[1318,1373,1804,1829]" gridcol="5" gridrow="19" pageId="4" pageNumber="319">9.8</td>
</tr>
<tr id="3FD534A9F65F6FFDFD8D46041254F89F" box="[163,1373,1846,1870]" gridrow="20" pageId="4" pageNumber="319">
<th id="7C045DD5F65F6FFDFD8D460416A4F89F" box="[163,429,1846,1870]" gridcol="0" gridrow="20" pageId="4" pageNumber="319">Dorsal spine length</th>
<td id="7C045DD5F65F6FFDFF72460415B6F89F" box="[604,703,1846,1870]" gridcol="1" gridrow="20" pageId="4" pageNumber="319">18.4</td>
<td id="7C045DD5F65F6FFDFE0046041465F89F" box="[814,876,1846,1870]" gridcol="2" gridrow="20" pageId="4" pageNumber="319">18.6</td>
<td id="7C045DD5F65F6FFDFEF246041304F89F" box="[988,1037,1846,1870]" gridcol="3" gridrow="20" pageId="4" pageNumber="319">0.9</td>
<td id="7C045DD5F65F6FFDF950460413B8F89F" box="[1150,1201,1846,1870]" gridcol="4" gridrow="20" pageId="4" pageNumber="319">16.7</td>
<td id="7C045DD5F65F6FFDF80846041254F89F" box="[1318,1373,1846,1870]" gridcol="5" gridrow="20" pageId="4" pageNumber="319">20.7</td>
</tr>
<tr id="3FD534A9F65F6FFDFD8D46521254F8A9" box="[163,1373,1888,1912]" gridrow="21" pageId="4" pageNumber="319">
<th id="7C045DD5F65F6FFDFD8D465216A4F8A9" box="[163,429,1888,1912]" gridcol="0" gridrow="21" pageId="4" pageNumber="319">Pectoral spine length</th>
<td id="7C045DD5F65F6FFDFF72465215B6F8A9" box="[604,703,1888,1912]" gridcol="1" gridrow="21" pageId="4" pageNumber="319">21.8</td>
<td id="7C045DD5F65F6FFDFE0046521465F8A9" box="[814,876,1888,1912]" gridcol="2" gridrow="21" pageId="4" pageNumber="319">21.8</td>
<td id="7C045DD5F65F6FFDFEF246521304F8A9" box="[988,1037,1888,1912]" gridcol="3" gridrow="21" pageId="4" pageNumber="319">1.4</td>
<td id="7C045DD5F65F6FFDF950465213B8F8A9" box="[1150,1201,1888,1912]" gridcol="4" gridrow="21" pageId="4" pageNumber="319">19.8</td>
<td id="7C045DD5F65F6FFDF80846521254F8A9" box="[1318,1373,1888,1912]" gridcol="5" gridrow="21" pageId="4" pageNumber="319">24.3</td>
</tr>
<tr id="3FD534A9F65F6FFDFD8D46BB1254F870" box="[163,1373,1929,1953]" gridrow="22" pageId="4" pageNumber="319">
<th id="7C045DD5F65F6FFDFD8D46BB16A4F870" box="[163,429,1929,1953]" gridcol="0" gridrow="22" pageId="4" pageNumber="319">Pelvic spine length</th>
<td id="7C045DD5F65F6FFDFF7246BB15B6F870" box="[604,703,1929,1953]" gridcol="1" gridrow="22" pageId="4" pageNumber="319">14.9</td>
<td id="7C045DD5F65F6FFDFE0046BB1465F870" box="[814,876,1929,1953]" gridcol="2" gridrow="22" pageId="4" pageNumber="319">14.0</td>
<td id="7C045DD5F65F6FFDFEF246BB1304F870" box="[988,1037,1929,1953]" gridcol="3" gridrow="22" pageId="4" pageNumber="319">1.8</td>
<td id="7C045DD5F65F6FFDF95046BB13B8F870" box="[1150,1201,1929,1953]" gridcol="4" gridrow="22" pageId="4" pageNumber="319">11.4</td>
<td id="7C045DD5F65F6FFDF80846BB1254F870" box="[1318,1373,1929,1953]" gridcol="5" gridrow="22" pageId="4" pageNumber="319">17.8</td>
</tr>
<tr id="3FD534A9F65F6FFDFD8D46811254F81A" box="[163,1373,1971,1995]" gridrow="23" pageId="4" pageNumber="319">
<th id="7C045DD5F65F6FFDFD8D468116A4F81A" box="[163,429,1971,1995]" gridcol="0" gridrow="23" pageId="4" pageNumber="319">Anal spine length</th>
<td id="7C045DD5F65F6FFDFF72468115B6F81A" box="[604,703,1971,1995]" gridcol="1" gridrow="23" pageId="4" pageNumber="319">17.9</td>
<td id="7C045DD5F65F6FFDFE0046811465F81A" box="[814,876,1971,1995]" gridcol="2" gridrow="23" pageId="4" pageNumber="319">14.9</td>
<td id="7C045DD5F65F6FFDFEF246811304F81A" box="[988,1037,1971,1995]" gridcol="3" gridrow="23" pageId="4" pageNumber="319">3.1</td>
<td id="7C045DD5F65F6FFDF950468113B8F81A" box="[1150,1201,1971,1995]" gridcol="4" gridrow="23" pageId="4" pageNumber="319">10.4</td>
<td id="7C045DD5F65F6FFDF80846811254F81A" box="[1318,1373,1971,1995]" gridcol="5" gridrow="23" pageId="4" pageNumber="319">17.9</td>
</tr>
</table>
</paragraph>
<caption id="D59A6663F65E9007FDB946AA15E5F829" ID-DOI="http://doi.org/10.5281/zenodo.8142278" ID-Zenodo-Dep="8142278" httpUri="https://zenodo.org/record/8142278/files/figure.png" pageId="5" pageNumber="320" startId="5.[151,250,1944,1968]" targetBox="[263,1324,180,1919]" targetPageId="5" targetType="figure">
<paragraph id="815A36EBF65E9007FDB946AA15E5F829" blockId="5.[151,1436,1944,2040]" pageId="5" pageNumber="320">
<emphasis id="B391EAF9F65E9007FDB946AA161AF861" bold="true" box="[151,275,1944,1968]" pageId="5" pageNumber="320">FIGURE 2.</emphasis>
<taxonomicName id="46E54D68F65E9007FC3446AA16F9F861" authorityName="Provenzano-Rizzi" authorityYear="2023" box="[282,496,1944,1968]" family="Loricariidae" genus="Cheilonimata" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="320" phylum="Chordata" rank="species" species="minuta">
<emphasis id="B391EAF9F65E9007FC3446AA16F9F861" box="[282,496,1944,1968]" italics="true" pageId="5" pageNumber="320">Cheilonimata minuta</emphasis>
</taxonomicName>
, MBUCV-V-32954, 54.3 mm SL, holotype. a) Detail of ventral view to show: head contour, mouth and abdomen cover. b) Close-up of mouth, detail of lower lip cover and translucent membrane on upper lip margin (red arrows). Scale bar 2 mm. c) Photos by C. Do Nascimiento.
</paragraph>
</caption>
<paragraph id="815A36EBF65D9004FDE941A51558FE90" blockId="6.[151,1437,151,861]" pageId="6" pageNumber="321">
Eyes located dorsally, relatively large, orbit with shallow posterior notch, and with ventral extension; eyeball appears internally withdrawn from the ventral edge of the bone orbit.From nostrils until anterior edge of supraoccipital two parallel low keels pass between eyes. Then over supraoccipital, keels converge posteriorly becoming single keel, until ending at supraoccipital tip. Predorsal region with three unpaired plates, first and second plates double keeled, third with a single keel (
<figureCitation id="19DE2A6EF65D9004FCD040151549FE90" box="[510,576,295,321]" captionStart="FIGURE 1" captionStartId="3.[151,250,1897,1921]" captionTargetBox="[183,1405,698,1872]" captionTargetId="figure-114@3.[183,1405,698,1872]" captionTargetPageId="3" captionText="FIGURE 1. Cheilonimata minuta, MBUCV-V-32954, 54.3 mm SL, holotype. Lateral, dorsal and ventral views. Scale bar 2 mm. Photos by C. Do Nascimiento." figureDoi="http://doi.org/10.5281/zenodo.8142276" httpUri="https://zenodo.org/record/8142276/files/figure.png" pageId="6" pageNumber="321">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="815A36EBF65D9004FDE9407913A9FDC8" blockId="6.[151,1437,151,861]" pageId="6" pageNumber="321">
Ventral surface of head without plates.Abdomen, in larger specimens, nearly naked with only three longitudinal and discontinuous lines or rows of very minute, not clumped, irregularly shaped plates; plate rows include one midventral almost reaching anus, and two lateral rows between pectoral and pelvic fins. In small specimens, abdomen naked with only few and scattered very minute plates, difficult to observe with naked eye. Thoracic plates only observed in specimens more than 65.0 mm SL (
<figureCitation id="19DE2A6EF65D9004FF8B40E915FFFE24" box="[677,758,475,501]" captionStart="FIGURE 2" captionStartId="5.[151,250,1944,1968]" captionTargetBox="[263,1324,180,1919]" captionTargetId="figure-15@5.[263,1324,180,1919]" captionTargetPageId="5" captionText="FIGURE 2. Cheilonimata minuta, MBUCV-V-32954, 54.3 mm SL, holotype. a) Detail of ventral view to show: head contour, mouth and abdomen cover. b) Close-up of mouth, detail of lower lip cover and translucent membrane on upper lip margin (red arrows). Scale bar 2 mm. c) Photos by C. Do Nascimiento." figureDoi="http://doi.org/10.5281/zenodo.8142278" httpUri="https://zenodo.org/record/8142278/files/figure.png" pageId="6" pageNumber="321">Fig. 2a</figureCitation>
). Anus projected as very small tube, delimited by naked area. Urogenital papilla not visible, apparently attached to posterior surface of anal tube (
<figureCitation id="19DE2A6EF65D9004F96F40CD139AFDC8" box="[1089,1171,511,537]" captionStart="FIGURE 2" captionStartId="5.[151,250,1944,1968]" captionTargetBox="[263,1324,180,1919]" captionTargetId="figure-15@5.[263,1324,180,1919]" captionTargetPageId="5" captionText="FIGURE 2. Cheilonimata minuta, MBUCV-V-32954, 54.3 mm SL, holotype. a) Detail of ventral view to show: head contour, mouth and abdomen cover. b) Close-up of mouth, detail of lower lip cover and translucent membrane on upper lip margin (red arrows). Scale bar 2 mm. c) Photos by C. Do Nascimiento." figureDoi="http://doi.org/10.5281/zenodo.8142278" httpUri="https://zenodo.org/record/8142278/files/figure.png" pageId="6" pageNumber="321">Fig. 2a</figureCitation>
).
</paragraph>
<paragraph id="815A36EBF65D9004FDE9431113E2FC8C" blockId="6.[151,1437,151,861]" pageId="6" pageNumber="321">
Mouth ventral; maxillary barbel reaches pectoral-fin spine origin. Upper lip border and maxillary barbel with elongated, cylindrical, unbranched, fleshy barblets embedded and interconnected by translucent membrane, leaving only barblet tips free (
<figureCitation id="19DE2A6EF65D9004FCBB435916E3FD54" box="[405,490,619,645]" captionStart="FIGURE 2" captionStartId="5.[151,250,1944,1968]" captionTargetBox="[263,1324,180,1919]" captionTargetId="figure-15@5.[263,1324,180,1919]" captionTargetPageId="5" captionText="FIGURE 2. Cheilonimata minuta, MBUCV-V-32954, 54.3 mm SL, holotype. a) Detail of ventral view to show: head contour, mouth and abdomen cover. b) Close-up of mouth, detail of lower lip cover and translucent membrane on upper lip margin (red arrows). Scale bar 2 mm. c) Photos by C. Do Nascimiento." figureDoi="http://doi.org/10.5281/zenodo.8142278" httpUri="https://zenodo.org/record/8142278/files/figure.png" pageId="6" pageNumber="321">Fig. 2b</figureCitation>
). Translucent membrane occurs along upper lip margin and on proximal two thirds of maxillary barbel. Also, translucent membrane connects maxillary barbel with lower lip border, laterally and internally. Upper lip surface very narrow and continuous with maxillary barbel and has few elongated, cylindrical, unbranched fleshy barblets (
<figureCitation id="19DE2A6EF65D9004FCF943E51522FD20" box="[471,555,727,753]" captionStart="FIGURE 2" captionStartId="5.[151,250,1944,1968]" captionTargetBox="[263,1324,180,1919]" captionTargetId="figure-15@5.[263,1324,180,1919]" captionTargetPageId="5" captionText="FIGURE 2. Cheilonimata minuta, MBUCV-V-32954, 54.3 mm SL, holotype. a) Detail of ventral view to show: head contour, mouth and abdomen cover. b) Close-up of mouth, detail of lower lip cover and translucent membrane on upper lip margin (red arrows). Scale bar 2 mm. c) Photos by C. Do Nascimiento." figureDoi="http://doi.org/10.5281/zenodo.8142278" httpUri="https://zenodo.org/record/8142278/files/figure.png" pageId="6" pageNumber="321">Fig. 2b</figureCitation>
). Lower lip wide, its margin with unbranched barblets and its surface covered with elongated, cylindrical, unbranched fleshy barblets, which resemble filaments; some of these barblets surpasses lower lip margin (
<figureCitation id="19DE2A6EF65D9004FC4D422D16BEFCE8" box="[355,439,799,825]" captionStart="FIGURE 2" captionStartId="5.[151,250,1944,1968]" captionTargetBox="[263,1324,180,1919]" captionTargetId="figure-15@5.[263,1324,180,1919]" captionTargetPageId="5" captionText="FIGURE 2. Cheilonimata minuta, MBUCV-V-32954, 54.3 mm SL, holotype. a) Detail of ventral view to show: head contour, mouth and abdomen cover. b) Close-up of mouth, detail of lower lip cover and translucent membrane on upper lip margin (red arrows). Scale bar 2 mm. c) Photos by C. Do Nascimiento." figureDoi="http://doi.org/10.5281/zenodo.8142278" httpUri="https://zenodo.org/record/8142278/files/figure.png" pageId="6" pageNumber="321">Fig. 2b</figureCitation>
). Palate with one elongate, cylindrical, unbranched, fleshy barblet behind premaxillaries at midline and one barblet with the same shape, lateral and external to each premaxilla (
<figureCitation id="19DE2A6EF65D9004F941427113CBFC8C" box="[1135,1218,835,861]" captionStart="FIGURE 2" captionStartId="5.[151,250,1944,1968]" captionTargetBox="[263,1324,180,1919]" captionTargetId="figure-15@5.[263,1324,180,1919]" captionTargetPageId="5" captionText="FIGURE 2. Cheilonimata minuta, MBUCV-V-32954, 54.3 mm SL, holotype. a) Detail of ventral view to show: head contour, mouth and abdomen cover. b) Close-up of mouth, detail of lower lip cover and translucent membrane on upper lip margin (red arrows). Scale bar 2 mm. c) Photos by C. Do Nascimiento." figureDoi="http://doi.org/10.5281/zenodo.8142278" httpUri="https://zenodo.org/record/8142278/files/figure.png" pageId="6" pageNumber="321">Fig. 2b</figureCitation>
,
<figureCitation id="19DE2A6EF65D9004F9E3427113D4FC8C" box="[1229,1245,835,861]" captionStart="FIGURE 3" captionStartId="6.[151,250,1795,1819]" captionTargetBox="[176,1411,884,1771]" captionTargetId="figure-335@6.[176,1411,884,1771]" captionTargetPageId="6" captionText="FIGURE 3. Cheilonimata minuta, ANSP 162156, 39.5 mm SL, paratype, detail of buccal ornamentation on mouth roof and behind premaxillaries (red arrow). Scale bar 1 mm. Photo by F. Provenzano." figureDoi="http://doi.org/10.5281/zenodo.8142280" httpUri="https://zenodo.org/record/8142280/files/figure.png" pageId="6" pageNumber="321">3</figureCitation>
).
</paragraph>
<caption id="D59A6663F65D9004FDB94631149FF8EE" ID-DOI="http://doi.org/10.5281/zenodo.8142280" ID-Zenodo-Dep="8142280" httpUri="https://zenodo.org/record/8142280/files/figure.png" pageId="6" pageNumber="321" startId="6.[151,250,1795,1819]" targetBox="[176,1411,884,1771]" targetPageId="6" targetType="figure">
<paragraph id="815A36EBF65D9004FDB94631149FF8EE" blockId="6.[151,1436,1795,1856]" pageId="6" pageNumber="321">
<emphasis id="B391EAF9F65D9004FDB94631161CF8CA" bold="true" box="[151,277,1795,1819]" pageId="6" pageNumber="321">FIGURE 3.</emphasis>
<taxonomicName id="46E54D68F65D9004FC304631154DF8CD" authority=", ANSP" authorityName="ANSP" box="[286,580,1795,1820]" family="Loricariidae" genus="Cheilonimata" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="321" phylum="Chordata" rank="species" species="minuta">
<emphasis id="B391EAF9F65D9004FC30463116FFF8CA" box="[286,502,1795,1819]" italics="true" pageId="6" pageNumber="321">Cheilonimata minuta</emphasis>
, ANSP
</taxonomicName>
162156, 39.5 mm SL, paratype, detail of buccal ornamentation on mouth roof and behind premaxillaries (red arrow). Scale bar 1 mm. Photo by F. Provenzano.
</paragraph>
</caption>
<paragraph id="815A36EBF65D9004FDE946A013FAF825" blockId="6.[151,1437,1938,2036]" pageId="6" pageNumber="321">
Both jaws with very small, but visible teeth, with similar size in each jaw. Premaxillary with two to three teeth and dentary with four to five teeth (
<figureCitation id="19DE2A6EF65D9004FF3346841568F801" box="[541,609,1974,2000]" captionStart="FIGURE 3" captionStartId="6.[151,250,1795,1819]" captionTargetBox="[176,1411,884,1771]" captionTargetId="figure-335@6.[176,1411,884,1771]" captionTargetPageId="6" captionText="FIGURE 3. Cheilonimata minuta, ANSP 162156, 39.5 mm SL, paratype, detail of buccal ornamentation on mouth roof and behind premaxillaries (red arrow). Scale bar 1 mm. Photo by F. Provenzano." figureDoi="http://doi.org/10.5281/zenodo.8142280" httpUri="https://zenodo.org/record/8142280/files/figure.png" pageId="6" pageNumber="321">Fig. 3</figureCitation>
). Teeth asymmetrically with outer cusp very small and pointed, sometimes not visible and inner cusp more developed, rounded; crown yellow or golden, stalk whitish (
<figureCitation id="19DE2A6EF65D9004F98E46E813ECF825" box="[1184,1253,2010,2036]" captionStart="FIGURE 3" captionStartId="6.[151,250,1795,1819]" captionTargetBox="[176,1411,884,1771]" captionTargetId="figure-335@6.[176,1411,884,1771]" captionTargetPageId="6" captionText="FIGURE 3. Cheilonimata minuta, ANSP 162156, 39.5 mm SL, paratype, detail of buccal ornamentation on mouth roof and behind premaxillaries (red arrow). Scale bar 1 mm. Photo by F. Provenzano." figureDoi="http://doi.org/10.5281/zenodo.8142280" httpUri="https://zenodo.org/record/8142280/files/figure.png" pageId="6" pageNumber="321">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="815A36EBF65C9005FDE941A51407FE58" blockId="7.[151,1437,151,861]" pageId="7" pageNumber="322">
Body sides with 2932 (mode 32) plates in mid-lateral series, 1720 (mode 18) coalescent plates and 914 (mode 14) posterior plates. Thoracic plates (between pectoral-fin to pelvic-fin, bases) 89 only observed in specimens with more than 65.0 mm SL. Post-dorsal plates 2223; post-anal plates 1920. Five plates along dorsal-fin base and three plates along anal-fin base. Dorsal-fin rays i,7, no spinelet nor vestigial bony plate observed; pectoral-fin rays i,6; pelvic-fin rays i,5; anal-fin rays i,5 and caudal-fin rays i,10,i. Pelvic-fin unbranched ray surpasses origin of anal-fin. Caudal fin with upper unbranched ray projected as filament (broken in
<typeStatus id="5E5E8849F65C9005FEEA40791322FEB4" box="[964,1067,331,357]" pageId="7" pageNumber="322">holotype</typeStatus>
). First unbranched ray of dorsal, pectoral, pelvic and anal fins not elongated as filaments.
</paragraph>
<paragraph id="815A36EBF65C9005FDE940A01497FD54" blockId="7.[151,1437,151,861]" pageId="7" pageNumber="322">
<emphasis id="B391EAF9F65C9005FDE940A01682FE7C" bold="true" box="[199,395,402,429]" pageId="7" pageNumber="322">Color in alcohol.</emphasis>
Dorsal surface of head and body light brown or yellowish, presence of six or seven transverse dark brown bands or stripes, sometimes faded, behind pectoral fin, first band at end of pectoral-fin base, second at dorsal-fin origin, third at end of dorsal-fin, remaining three or four, posterior to anal-fin base to caudal-fin origin (
<figureCitation id="19DE2A6EF65C9005FDB140CD17EBFDC8" box="[159,226,511,537]" captionStart="FIGURE 1" captionStartId="3.[151,250,1897,1921]" captionTargetBox="[183,1405,698,1872]" captionTargetId="figure-114@3.[183,1405,698,1872]" captionTargetPageId="3" captionText="FIGURE 1. Cheilonimata minuta, MBUCV-V-32954, 54.3 mm SL, holotype. Lateral, dorsal and ventral views. Scale bar 2 mm. Photos by C. Do Nascimiento." figureDoi="http://doi.org/10.5281/zenodo.8142276" httpUri="https://zenodo.org/record/8142276/files/figure.png" pageId="7" pageNumber="322">Fig. 1</figureCitation>
). Ventral surface of head and body yellowish or creamy, uniform (
<figureCitation id="19DE2A6EF65C9005FEF440CD1347FDC8" box="[986,1102,511,537]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[151,250,1897,1921]" captionStartId-1="5.[151,250,1944,1968]" captionTargetBox-0="[183,1405,698,1872]" captionTargetBox-1="[263,1324,180,1919]" captionTargetId-0="figure-114@3.[183,1405,698,1872]" captionTargetId-1="figure-15@5.[263,1324,180,1919]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionText-0="FIGURE 1. Cheilonimata minuta, MBUCV-V-32954, 54.3 mm SL, holotype. Lateral, dorsal and ventral views. Scale bar 2 mm. Photos by C. Do Nascimiento." captionText-1="FIGURE 2. Cheilonimata minuta, MBUCV-V-32954, 54.3 mm SL, holotype. a) Detail of ventral view to show: head contour, mouth and abdomen cover. b) Close-up of mouth, detail of lower lip cover and translucent membrane on upper lip margin (red arrows). Scale bar 2 mm. c) Photos by C. Do Nascimiento." figureDoi-0="http://doi.org/10.5281/zenodo.8142276" figureDoi-1="http://doi.org/10.5281/zenodo.8142278" httpUri-0="https://zenodo.org/record/8142276/files/figure.png" httpUri-1="https://zenodo.org/record/8142278/files/figure.png" pageId="7" pageNumber="322">Figs 12a</figureCitation>
). Dorsal, pectoral, pelvic and caudal-fins rays with dark bands; two longitudinal on dorsal fin, two or three transversal on pectoral fin, one on pelvic fin, sometime faded, and one on caudal fin, oblique; interradial membranes hyaline (
<figureCitation id="19DE2A6EF65C9005F99D437513F9FDB0" box="[1203,1264,583,609]" captionStart="FIGURE 1" captionStartId="3.[151,250,1897,1921]" captionTargetBox="[183,1405,698,1872]" captionTargetId="figure-114@3.[183,1405,698,1872]" captionTargetPageId="3" captionText="FIGURE 1. Cheilonimata minuta, MBUCV-V-32954, 54.3 mm SL, holotype. Lateral, dorsal and ventral views. Scale bar 2 mm. Photos by C. Do Nascimiento." figureDoi="http://doi.org/10.5281/zenodo.8142276" httpUri="https://zenodo.org/record/8142276/files/figure.png" pageId="7" pageNumber="322">Fig.1</figureCitation>
). Anal-fin rays uniform, whitish or yellowish, interradial membrane hyaline (
<figureCitation id="19DE2A6EF65C9005FE6543591485FD54" box="[843,908,619,645]" captionStart="FIGURE 1" captionStartId="3.[151,250,1897,1921]" captionTargetBox="[183,1405,698,1872]" captionTargetId="figure-114@3.[183,1405,698,1872]" captionTargetPageId="3" captionText="FIGURE 1. Cheilonimata minuta, MBUCV-V-32954, 54.3 mm SL, holotype. Lateral, dorsal and ventral views. Scale bar 2 mm. Photos by C. Do Nascimiento." figureDoi="http://doi.org/10.5281/zenodo.8142276" httpUri="https://zenodo.org/record/8142276/files/figure.png" pageId="7" pageNumber="322">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="815A36EBF65C9005FDE943BC157CFCC4" blockId="7.[151,1437,151,861]" pageId="7" pageNumber="322">
<emphasis id="B391EAF9F65C9005FDE943BC16FDFD79" bold="true" box="[199,500,654,680]" pageId="7" pageNumber="322">Geographic distribution.</emphasis>
Available records indicate
<taxonomicName id="46E54D68F65C9005FE1643BD14A2FD79" box="[824,939,655,681]" family="Loricariidae" genus="Cheilonimata" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="322" phylum="Chordata" rank="species" species="minuta">
<emphasis id="B391EAF9F65C9005FE1643BD14A2FD79" box="[824,939,655,681]" italics="true" pageId="7" pageNumber="322">C. minuta</emphasis>
</taxonomicName>
inhabits the upper Orinoco River and the Casiquiare River,
<collectingRegion id="4321F809F65C9005FC4F4381151CFD1C" box="[353,533,691,717]" country="Venezuela" name="Amazonas" pageId="7" pageNumber="322">Amazonas state</collectingRegion>
,
<collectingCountry id="F9F2767BF65C9005FF0F4381159DFD1C" box="[545,660,691,717]" name="Venezuela" pageId="7" pageNumber="322">Venezuela</collectingCountry>
(
<figureCitation id="19DE2A6EF65C9005FF8A438115E3FD1C" box="[676,746,691,717]" captionStart="FIGURE 4" captionStartId="7.[151,250,1874,1898]" captionTargetBox="[179,1409,885,1850]" captionTargetId="figure-316@7.[178,1409,884,1850]" captionTargetPageId="7" captionText="FIGURE 4. Map of Venezuela showing the geographic distribution of Cheilonimata minuta, red star type locality.Some symbols may represent more than one lot." figureDoi="http://doi.org/10.5281/zenodo.8142282" httpUri="https://zenodo.org/record/8142282/files/figure.png" pageId="7" pageNumber="322">Fig. 4</figureCitation>
). The information about capture sites indicates clear to black waters environments with sandy, rocky, gravel and pebbles bottoms. The water has low pH and conductivity levels, and little quantity of suspended sediments.
</paragraph>
</subSubSection>
<subSubSection id="C9FF6560F65C9005FDE9422D161DFC8C" pageId="7" pageNumber="322" type="etymology">
<paragraph id="815A36EBF65C9005FDE9422D161DFC8C" blockId="7.[151,1437,151,861]" pageId="7" pageNumber="322">
<emphasis id="B391EAF9F65C9005FDE9422D1647FCE8" bold="true" box="[199,334,799,825]" pageId="7" pageNumber="322">Etymology.</emphasis>
<taxonomicName id="46E54D68F65C9005FC72422D16A3FCE9" box="[348,426,799,824]" family="Loricariidae" genus="Cheilonimata" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="322" phylum="Chordata" rank="species" species="minuta">
<emphasis id="B391EAF9F65C9005FC72422D16A3FCE9" box="[348,426,799,824]" italics="true" pageId="7" pageNumber="322">minuta</emphasis>
</taxonomicName>
is taken from the Latin meaning small, which is a trait exhibited by the analyzed specimens.
</paragraph>
</subSubSection>
</treatment>
</document>